JOURNAL OF PALEONTOLOGY

A publication of THE SOCIETYOF ECONOMICPALEONTOLOGISTS AND MINERALOGISTS and THE PALEONTOLOGICALSOCIETY with the generous support and cooperation of THE AMERICAN ASSOCIATIONOF PETROLEUMGEOLOGISTS and THE GEOLOGICALSOCIETY OF AMERICA

VOLUME 24 JANUARY, 1950 NUMBER 1

TERTIARY NAUTILOIDS OF THE AMERICAS: SUPPLEMENT A. K. MILLER AND H. R. DOWNS State University of Iowa, Iowa City, Iowa

ABSTRAcT-Representativesof Eutrephocerasare describedfrom the Paleoceneand the Eocene of California; Cimomiafrom the Paleocene of Colombiaand Trinidad and the Eocene of Washington;Hercoglossa from the Paleocene of Baja California and the Eocene of Peru; Deltoidonautilusfrom the Eocene of Peru; and Aturia from the Eocene of Texas, California, Washington, and Peru, and Oligocene of Oregon and Washington, and the Miocene of California, Florida, and Cuba. All of these specimens that merit illustration are figured. A representative of Aturia from the Miocene of Sinai is discussedin an appendix,and illustrationsof it are included.

ATE in 1947 The Geological Society of ral History and Columbia University; Pro- America published a monograph of fessor W. P. Popenoe of the University of the "Tertiary nautiloids of the Americas" California (Los Angeles); Dr. H. H. Renz of by the senior author of the present report. the Mene Grande (Gulf) Oil Company Since that time, various paleontologists (Caracas), and Sr. Pedro Verastegui M. of have sent in additional specimens from Stanford University. Special acknowledg- widely separated localities. We propose to ment is also due to Professor Don L. Friz- treat all of this material in a single publica- zell of the Missouri School of Mines and tion, which can therefore be regarded as a Metallurgy for advice in regard to certain supplement to the recently published mono- Peruvian strata, and especially to Mr. graph. Howard Webster of Iowa City who re- The specimens on which this report is touched the photographs and made the based became available to us through the drawings which accompany this study. The courtesy of the following persons: Dr. P. J. completion of the report was made financial- Bermudez of the Creole Petroleum Corpo- ly possible by the Graduate College of the ration (Caracas); Professor Bernardo Boit State University of Iowa. of the Universidad de San Marcos; Professor Much of the material discussed in this J. Wyatt Durham of the University of supplement is from the collections of the California (Berkeley); Dr. Julia Gardner of University of California at Berkeley, and the United States Geological Survey; Dr. it came from various horizons and localities A. Myra Keen of Stanford University; Dr. in our Pacific coast Tertiary. It should H. G. Kugler of the North Venezuelan perhaps be mentioned in this connection Petroleum Company, Limited (Caracas); that Professor J. Wyatt Durham has in- Mr. Chas. R. Locklin of Union Lake (near formed us that it is now generally believed Pontiac), Michigan; Professor Norman D. that all of the Paleocene Martinez formation Newell of The American Museum of Natu- of California is as old as the Midway group 1 2 A. K. MILLER AND H. R. DOWNS

of the Gulf Coastal Plain, and that some of the Eocene Claiborne group of east Texas. it may be older. If this belief is correct Two others are delicate testiferous repre- Aturia, and possibly also Aturoidea, were sentatives of Aturia curvilineata Miller and almost certainly represented on our west Thompson from the Miocene Chipola for- coast long before they appeared on our mation of the Florida panhandle. A fifth Atlantic-Gulf coast. Professor Ewart M. is a representative of the rare species Aturia Baldwin of the University of Oregon has cubanensis (Lea) of the Miocene of Cuba; written us that the formation at Dallas, and along with the discussion of it is in- Oregon, which yielded the holotype of cluded some data on the occurrence in Cuba Eutrephoceras oregonense Miller, is not Oligo- of other congeneric forms. A sixth is a small cene in age (as stated in the original descrip- fragment of a conch from the Paleocene at tion of the species) but is Eocene, "probably a locality in northern Colombia from which equivalent to the Domengine stage of no nautiloids had previously been recorded California. Similar limy beds nearby, which in the literature. A seventh is an essentially also contain what I [Baldwin] believe to be complete internal mold from the Paleocene these same nautiloids, contain large Forami- in the Marac quarry of Trinidad, and it is nifera assigned to Pseudophragmina psila probably referable to Cimomia subrecta Mil- (Woodring), which tends to support the ler and Thompson, which was originally older Eocene age." Both Professors Durham described from the Midway group of our and Baldwin have assured us that no Plio- Gulf Coastal Plain. An eighth is an excep- cene strata are known in the coastal area tionally fine large internal mold of Herco- west of Eugene from which Dall (1909, p. 21) glossa peruviana Berry from the Eocene of stated that Thomas Condon had collected Peru. A ninth is a small representative of a "fine" nautiloid. Professor Baldwin adds the genus Aturia from the Miocene of Sinai. that the "rock that would come the closest to The other material described in this re- being called Pliocene is that at Newport, port is a collection made by Sr. Pedro where Dr. Condon splent many of his vaca- Verastegui M. from the Salina formation tions. This is now considered to be a part and the Lomitos sandstone (which was for- of the Astoria formation of approximately merly regarded as a member of the lower middle Miocene age. The only marine Plio- Talara shale) in northwestern Peru. It con- cene that we know of in Oregon is the Em- sists of an immature representative of Deltoi- pire formation at Coos Bay and the Empire donautilus from the Salina, and D. haughti and Port Orford formations at Cape Blanco." (Olsson) and Aturia peruviana Olsson from A discussion of a few sporadic specimens the Lomitos. The occurrence of the genus is also included in this report. One of these, Deltoidonautilus in South America is being which represents a new species of Hercoglos- recognized for the first time. sa, is the first Tertiary nautiloid to be re- Altogether, we are describing (and in corded from the Pacific coast region of most cases illustrating) representatives of Mexico-it came from the Paleocene of the following genera: Eutrephoceras from the Baja California. Another is an exceptionally Paleocene (Martinez) and the Eocene well preserved internal mold of Aturia (Capay) of California; Cimomia from the brazoensis Stenzel from the lower part of Paleocene of Colombia and Trinidad (Sol-

EXPLANATION OF PLATE 1 FIGS.1-3-Eutrephoceras sp. Three views of a small specimen from the Eocene Capay shale about 4 miles west of Winters, California,X2. (p. 3) 4, 5-Eutrephoceras aff. E. stephensoni(Dickerson). Two views of a portion of a phragmacone from the Paleocene Martinez formation near Selby station, California, X 1. (p. 3) 6-8-Aturia angustata (Conrad)? Two moderately small specimens from the Oligocene Twin Rivers formation an eighth of a mile east of the city of Twin Rivers (figs. 6, 7), and in a sea cliff east of the mouth of East Twin River (fig. 8), Washington, X2 and X1, respec- tively. (p. 9) 9-Aturia sp. Lateral view of a rather poorly preserved testiferous specimen from the Eocene Crescentformation in a sea cliff near Port Crescent,Washington, X . (p. 15) JOURNAL OF PALEONTOLOGY, VOL. 24 PLATE11

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Miller and Downs, Tertiarynautiloids TERTIARY NAUTILOIDS OF THE AMERICAS 3 dado) and the Eocene (Cowlitz) of Washihg- 6 mm. from the dorsum and 14 mm. from ton; Hercoglossa from the Paleocene of Baja the venter. California and the Eocene of Peru; Deltoi- Remarks.-The large size and particularly donautilus from the Eocene of Peru; and the reticulate surface ornamentation of the Aturia from the Eocene of Texas, Cali- test of this specimen indicate that it is fornia, Washington, and Peru, the Oligo- almost certainly not referable to E. stephen- cene of Washington and Oregon, and the soni; and as Vokes stated, its conch is Miocene of California, Florida, Cuba, and relatively wider than that of E. hallidayi Sinai. (Waring), the only other congeneric form SYSTEMATICPALEONTOLOGY known from the Martinez formation. It EUTREPHOCERASaff. E. STEPHENSONI probably represents an unnamed species, not make a holo- (Dickerson) but it would satisfactory Plate 1, figures 4, 5 type. Occurrence.-Martinez formation about Eutrephocerasstephensoni? VOKES, 1937, Jour. 200 ft. northwest of the south of Paleontology, vol. 11, p. 5; MILLER,1947, portal Geol. Soc. America, Mem. 23, p. 37. the Southern Pacific railroad tunnel near the smelter at Selby station, Contra Costa When Vokes discussed Eutrephoceras County, California. stephensoni in 1937, he stated: "There is in of California, the collections at the University of California Repository.-University Berkeley. a fragment of a very large specimen from the Martinez formation near Selby Station, EUTREPHOCERASSp. which this California, may represent species. Plate 1, figures 1-3 The greatest width of the conch was 85 mm., and the distance from the venter to In June of 1938 Miss Herdis Bentson the base of the impressed area over 30 mm. found the only nautiloid known from the It is too globose to be referable to E. hal- Eocene Capay shale of California. It is an lidayi, but is too imperfect to permit definite internal mold that is septate throughout specific determination." and is about 26 mm. in diameter-its maxi- This specimen represents parts of two mum width of conch measures about 19 (or more) volutions of a phragmacone, the mm. The umbilicus is small and inconspicu- preserved portion of which is estimated to ous, and in one of the umbilical depressions have been a little less than 100 mm. in (pl. 1, fig. 2) there is a "plug" of shell diameter. The shape of the cross section material which indicates that the conch, of the conch is elucidated by figure 4 on when complete, consisted of at least one plate 1. The surface of the test of at least more full volution. The external sutures are the penultimate of the volutions represented essentially straight and directly transverse, is reticulate as it is marked by moderately and the camerae are of about average length. coarse longitudinal and transverse lines, A structure that appears to represent the which are of about equal size and promi- siphuncle is small and is subcentral in nence. The transverse ones, growth lines, position. indicate that the conch bore a broad rounded Remarks.-Clearly this specimen belongs hyponomic sinus. The siphuncle is small in the genus Eutrephoceras. However, its and is located moderately close to the dorsal preservation leaves much to be desired, and wall of the conch. Where the distance be- its small size and general physiognomy indi- tween the dorsum and the venter measures cate that it probably represents only the about 20 mm., the siphuncle is about 2 immature portion of the conch. Therefore, mm. in diameter, and its center is about we are reluctant to express an opinion in

EXPLANATION OF PLATE 2 FIGS.1, 2-Cimomia subrectaMiller and Thompson.Lateral and aperturalviews of a specimen, from the Paleocene Soldado formation in Marac quarry, southern Trinidad, British West Indies, very slightly less than X . (p. 4) 4 A. K. MILLER AND H. R. DOWNS regard to its specific affinities, but it may sentative of the genus Cimomia to be re- well be referable to E. hannai Vokes. corded from the Pacific coast region of Occurrence.-Capay shale (65-90 feet North America. Its most distinctive char- stratigraphically below the Pliocene con- acters seem to be the shape of its cross glomerate contact) at the Fenn Ranch section and its sutures. It is not particularly locality, just above the stream bottom, close to any of the congeneric species that about 4 miles west of Winters, Yolo County, have been described from our Atlantic-Gulf California-it was found in association with Coastal Plain or from South America, brachiopods and Venericardia-[loc. A3073]. probably because most of them are from Repository. - University of California, older (that is, Paleocene) strata. Berkeley. The collections of the University of Cali- fornia at Berkeley contain a very frag- CIMOMIAHESPERIA M. & n. D., sp. mentary specimen which is probably con- Plate 4 3, figures 3, specific with the holotype just described. The holotype of this species is an internal It came from the same general horizon and mold which represents much of four camerae locality as the holotype, and it seems to of a phragmacone. Its whorls are helmet- represent an individual of about the same shaped in cross section as they are broadly size. However, it is so distorted and incom- rounded ventrally, flattened laterally, and plete that its affinities will always be open impressed dorsally. Clearly, the conch, when to question. Both of these specimens were complete, was subglobular, The over-all collected by Professor J. Wyatt Durham. length of the holotype measures about 66, Occurrence.-Type section of the upper mm.; and at the uncrushed septum which Eocene Cowlitz formation at the big bend forms its adapical end its height and width in the Cowlitz River (a well-known locality), of conch measure approximately 38 mm. Lewis County, Washington. and 45 mm., respectively, and the depth Holotype.-University of California, of the impressed zone is about 12? mm. The Berkeley. umbilical shoulders are rounded, and appar- CIMOMIASUBRECTA Miller and Thompson ently the umbilicus is small. Plate 2, figures 1, 2 Fragments of the test which adhere to this are thin. Traces of the Cimomia subrecta MILLERand THOMPSON,1933, specimen growth Jour. Paleontology, vol. 7, pp. 308-311, pl. 35, lines that are preserved on them show that figs. 1, 2; STENZEL,1942, [Illustrated card there was a broad shallow rounded hypo- catalogue of North American early Tertiary nomic sinus. fossils of Atlantic-Gulf Coastal Plain], Ceph- The camerae are Each alopoda, cards 9a, 9b; SHIMERand SHROCK, moderately long. 1944, Index fossils of North America, p. 550, suture forms a low broad rounded ventral pi. 225, fig. 6; MILLER,1947, Geol. Soc. Amer- saddle and on either side of it a similar but ica, Mem. 23, pp. 4, 6, 42, 46-47, 48, 49, pl. 31, smaller lateral lobe, a small but rather figs. 1, 2. prominent saddle on the umbilical shoulder, Dr. H. G. Kugler recently sent us for a slight lobe on the umbilical wall, and a study a rather large specimen from the low broadly rounded internal lateral saddle Paleocene of Trinidad that should probably which extends to the similar but medianly be referred to Miller and Thompson's flattened dorsal lobe. species. However, as explained below, it The siphuncle is small, and is subcentral may well be specifically distinct. in position, but is located considerably This specimen is an essentially complete closer to the dorsum than the venter. At internal mold which is only slightly distorted. the adapical end of the holotype, the siph- Its maximum diameter is about 140 mm., uncle is about 2 mm. in diameter and its and near its adoral end the maximum height center is about 10? mm. from the dorsum and width of its conch measure about 83 and 15 mm. from the venter. It appears to mm. and 88 mm., respectively. The umbili- be composed of cylindrical segments and cus, though deep, is small and inconspicu- presumably is orthochoanitic in structure ous, and the umbilical shoulders are rounded as in other congeneric species. and indefinite. No trace of the surface mark- Remarks.-This form is the first repre- ings of the test is preserved. The nature of TERTIARY NA UTILOIDS OF THE AMERICAS 5 the sutures and the general physiognomy of mia, and none of its characters that we are the specimen are elucidated by the accom- able to ascertain militates against this panying illustrations. generic reference. However, it may well Remarks.-It is clear from a direct com- belong in such genera as Eutrephoceras, parison of this specimen with the holotype Hercoglossa, or Deltoidonautilus; and with of C. subrecta that the saddles on the um- no knowledge as to the shape of its sutures bilical shoulders are much less prominent we can, of course, not determine its generic on the Trinidad individual. However, in the affinities. extreme adoral portion of the phragmacone Occurrence.-A buff-colored limestone bed of the holotype, those saddles become rela- in a Paleocene brown shale which outcrops tively low. In both specimens the adoral "in a small quebrada about 450 meters camera is considerably shorter than the west of the village of Manantial, along the preceding one, indicating that we are dealing Hato Nuevo-Manantial trail, about 12 with fully mature individuals in each case; kilometers northeast of Hato Nuevo, De- but the Trinidad specimen is about 20 per partamento de Magdalena, Rancheria Val- cent larger than the holotype. Nevertheless, ley, Colombia." most of the other characters of the two Repository.-University of California, do not seem to differ materially, and there- Berkeley. fore we are tentatively regarding them as HERCOGLOSSAPERUVIANA Berry conspecific. Plate Plate 2 Occurrence.-Soldado formation (Paleo- 6, figures 1, 2; 7, figure cene) in the Marac quarry in the south- An exceptionally fine large representative central part of Trinidad, British West of this species was sent to us for study Indies. All of the specimens that have through the courtesy of Professor Norman previously been referred to this species came D. Newell, who borrowed it from the Engi- from the Clayton formation (the basal neering School in Lima, Peru. This specimen part of the Paleocene Midway group) of is an internal mold to which a considerable Mississippi. portion of the test (or a replacement of it) Repository.-Museum of Natural History adheres. It is preserved in mottled light at Basel, Switzerland. brown limestone and is only slightly dis- torted. Inasmuch as it is septate throughout, CIMOMIA? sp. it represents only phragmacone. When com- Plate 7 3, figure plete, the conch was at least another half- Professor J. Wyatt Durham loaned us a volution in extent. fragment of a nautiloid which he secured The maximum diameter of this individual, from the Paleocene of northern Colombia at measured from the adoral end of the venter a locality where no representatives of this across the umbilicus to the opposite side of order had been found previously. Most of the specimen, is about 225 mm. The maxi- the beds exposed there are essentially non- mum height and width of the preserved fossiliferous; but one of them contains an part of the conch measure about 140 mm. abundance of small gastropods, pelecypods, and 150 mm., respectively; and near the and other fossils, and the nautiloid was adoral end of the specimen the impressed found in it. zone is almost 50 mm. deep. The diameter This specimen consists of umbilical and of the complete shell is estimated to have adjacent portions of a moderately small been more than 400 mm. individual-its maximum over-all measure- The test of the adapical portion of the ment is only about 30 mm. The umbilicus outer volution is about 1 mm. thick. On is small and closed, and the umbilical the surface of the test there are numerous shoulders are rounded and indefinite. The fine growth lines. Each of these forms a deep surface of the preserved portion of the test rather narrowly rounded ventral or hypo- is marked by fine transverse lines (presum- nomic sinus and on either side of it a broad ably growth lines) that are slightly sinuous. rounded asymmetrical lateral salient that The general physiognomy of this speci- extends clear to the umbilicus. No trace of men seems to us to be reminiscent of Cimo- the growth lines is discernible on the internal 6 A. K. MILLER AND H. R. DOWNS mold, but along its venter there is a rather The holotype of the new species appears prominent raised line. to be fairly complete, and it attains a maxi- There are some 20 camerae in the outer mum diameter of about 270 mm. Its adoral volution of this specimen. The sutures and half-volution is non-septate and presum- the siphuncle appear to be typical of the ably therefore represents living chamber. species. At its passage through the adoral Near the junction of the phragmacone and septum, the siphuncle is about 8 mm. in the living chamber, the conch is essentially diameter. free from distortion and it is about 108 mm. Remarks.-Recently, Miller (1947, pp. wide and 100 mm. high. The whorls are 58-59, pl. 23, fig. 2; pl. 24, figs. 1, 2; pl. 25, rather narrowly rounded ventrally, strongly figs. 1, 2; pl. 89, figs. 1, 2; pl. 90, fig. 4) flattened (but very broadly rounded) lateral- described this species in detail, listed its ly, and deeply impressed dorsally. The complete synonymy, and indicated how it lateral zones converge ventrally, and there- can be distinguished from the very similar fore the maximum width of conch is attained form, H. ulrichi (White). There is of course at the umbilical shoulders. These shoulders no need to duplicate that work here. are rounded, and the umbilicus is small, Occurrence.-Near Lobitos, Peru pre- closed, and inconspicuous. sumably from the Talara formation. Olsson The test is fairly thick, and at the junc- (1928, p. 96) has stated that this species is tion of the phragmacone and the living "strictly limited" to one "horizon" and is chamber its thickness along the ventrolateral "quite common" in it; and Frizzell has zone measures almost 3 mm. For the most recently written us that it occurs "in the part, only the inner layers of the test are lower part of the Talara formation in beds retained on the holotype. However, from that .. .[are] near the top of the middle preserved portions of the outer layer, it can Eocene." Furthermore the lithology and be ascertained that the surface bears small preservation of the specimen, which are low inconspicuous ribs and fine growth lines very distinctive, are precisely like those of which are parallel to them. Each of these conspecific individuals near Lobitos. forms a broad deep rather narrowly rounded Repository.-School of Engineering, Lima, ventral sinus and on either side of it a low Peru. broad rounded lateral salient which extends to the umbilicus. POPENOEI M. & n. HERCOGLOSSA D., sp. The length of the adoral camerae of the Plate 2 5, figures 1, holotype, measured along the venter, varies On February 14, 1948, Professor W. P. from about 19 to 23 mm. The adoral camera Popenoe (and party) found a large well of this specimen is distinctly longer than the preserved representative of Hercoglossa in preceding one. As shown by text figure 1, the northern half of Baja California and each external suture forms a broad high kindly loaned it to us for study. This form rounded ventral saddle, and on either side is not referable to any described species, and of it a deep rather narrowly rounded lateral we propose to name it in honor of its dis- lobe and a high rounded lateral saddle which coverer. extends to the umbilical shoulder.

EXPLANATION OF PLATE 3 FIGS. 1, 2-Aturia sp. Two views of an internal mold, from the Vacaville shale memberof the Eocene Capay formation about half a mile north of "Black Rocks" in Putah Creek, Yolo County California, X1. (p. 15) 3, 4-Cimomia hesperia M. & D., n. sp. Lateral and apical views of the holotype, from the Eocene Cowlitz formationat the big bend in the Cowlitz River, Lewis County, Washington, X1. (p.4) 5, 6-Aturia cf. A. myrlaeHanna. A small specimen, from the Eocene Arroyo Hondo formation north of Coalinga, California, X3. (p. 15) 7-Cimomia? sp. A fragmentof a moderatelysmall conch of uncertainaffinities, from a limestone bed in a brown Paleocene shale about 12 km. northeast of Hato Nuevo, Departamento de Magdalena, Colombia, X2. (p. 5) 8-Aturia angustata(Conrad)? An incomplete internal mold, from the Miocene Temblorforma- tion in Kern County, California,probably near Mt. Poso oil field, X1. (p. 9) JOURNAL OF PALEONTOLOGY, VOL. 24 PLATE 3

1 .3

Miller and Downs, Tertiarynautiloids JOURNAL OF PALEONTOLOGY, VOL. 24 PLATE 4

Miller and Downs, Tertiarynautiloids TERTIARY NA UTILOIDS OF THE AMERICAS 7

Remarks.-This specimen is the first nau- stream divides, in sea cliff on south side of tiloid to be recorded from the Paleocene of valley of Arroyo Santa Catarina, about 200 Btja California. The matrix in which it is feet above sea level and perhaps an eighth preserved is a dark-gray fine-grained cal- of a mile inland at unnamed point about careous sandstone. In the filling of the living 2? miles northwest of Punta Canoas, Santa

FIG. --Diagrammatic representationof an external suture of Hercoglossapopenoei M. & D., n. sp., based on the adoral sutures of the holotype, X . chamber there are several types of small Catarina Landing, Baja California (U. C. gastropods, one of which, according to W. L. A. loc. 2372). P. Popenoe, appears to be Retipirula cras- Holotype.-University of California, Los sitesta Gabb. Angeles, 10655. The new species is a typical representative of the genus Hercoglossa. It is close to the DELTOIDONAUTILUSHAUGHTI (Olsson) genotype, H. orbiculata (Tuomey), and to Plate 4, figures 1-3, 7, 8 H. mcglameryae Miller and Thompson, both Eutrephocerashaughti OLSSON, 1928, Bull. Am. of the Midway group of the Gulf Coastal Paleontology, vol. 14, no. 52, p. 101, pl. 23, Plain-in the former the conch is fig. 2; MAURY,1930, Serv. geol. e min. Brasil, relatively no. 250. wide and Mon., 8, pp. 249, and in the latter relatively small Cimomia haughti MILLERand THOMPSON,1933, narrow. Other similar species are H. merri- Jour. Paleontology, vol. 7, p. 305; MILLER, ami Dickerson of the Martinez formation of 1947, Geol. Soc. America, Mem. 23, pp. 6, 10, California in which the lateral lobes of the 42-43, 45, 46, pl. 16, figs. 1, 2; pl. 64, fig. 3. sutures are deeper and more narrowly Professor Bernardo Boit of the Univer- rounded, H. gardneri Stenzel of the Kincaid sidad de San Marcos kindly sent in for study formation of Texas in which the ventral 14 nautiloids that Sr. Pedro Verastegui \M. lobe of the sutures is more nearly straight- had collected from the Lomitos sandstone sided, and H. peruviana Berry of the Talara at a single locality near Cabo Blanco in formation of Peru which has a wider conch. northwestern Peru. Eleven of these appear Occurrence.-From concretion in fine- to represent one species, though they vary grained gray sandstone of Paleocene age in diameter from about 39 mm. (pl. 4, cropping out a few feet below the base of figs. 1-3) to approximately 172 mm. (pl. 4, the Tertiary lava beds capping the inter- figs. 7, 8). The smaller ones seem to resemble

EXPLANATION OF PLATE 4 FIGS.1-3-Deltoidonautilus haughti (Olsson). A small well preservedinternal mold (representingthe adolescent portion of the conch), from the Eocene Lomitos sandstone at Cerro Organos,6 miles northeast of Cabo Blanco, Provincia de Paita, Departamento de Piura, Peru, X 1i. (p. 7) 4-Aturia sp. A small specimen,from the lower Miocene on the south side of the mouth of Wadi Gharandelsome 2 or 3 miles north of Hammam Faraun, Sinai, X2. (p. 17) 5, 6-Aturia peruvianaOlsson. A small internal mold from the same horizon and locality as figures 1-3, X1. (p. 14) 7, 8-Deltoidonautilus haughti (Olsson). An internal mold of a large mature phragmacone, from the same horizonand locality as figures1-3, X 1 . (p. 7) 8 A. K. MILLER AND H. R. DOWNS

in all significant particulars the holotype saddle which centers well outside the um- of Eutrephoceras [Cimomia] haughti Olsson, bilical shoulder, apparently a lobe on the which came from the Talara formation at a umbilical wall, and a broad rounded inter- nearby locality in northwestern Peru and nal lateral saddle, which extends to a rather which is therefore of about the same age. deep narrowly rounded dorsal lobe. The However, the larger individuals possess all sinuosity of the sutures seems to increase of the characteristics of the genus Deltoido- progressively throughout ontogenetic de- nautilus, and the intermediate ones are velopment. The fact that the shape of the gradational between these two extremes. It sutures changes rapidly in the outer volution therefore seems to us that almost certainly of the small specimen represented by figures Olsson's species was based on an adolescent 1-3 on plate 4, indicates that it represents specimen, the fully mature portion of which only the immature portion of the conch. would have revealed that its generic affinities The siphuncle is moderately small and are with Deltoidonautilus. In view of the is located fairly close to the dorsum. Where fact that this genus is believed to have the conch is about 53 mm. high and is evolved from Eutrephoceras through Cimo- impressed to a depth of about 23 mm., the mia, it is not surprising that the inner siphuncle is about 4 mm. in diameter and volutions of its conch resemble those of its center is about 8 mm. from the dorsum mature representatives of these genera. and 22 mm. from the venter. The material now available for study Remarks.-In both the holotype and the shows that the phragmacone of this species small specimen which Miller (1947, p. 43, pl. attained a diameter of at least 172 mm., and 16, figs. 1, 2) doubtfully associated with it, a corresponding height and width of conch the siphuncle is also dorsad in position. In of some 115 mm. and 120 mm., respectively. the genotype of Deltoidonautilus it is simi- At full maturity, the ventral zone of the larly located, but its position in the genotype conch becomes so narrowly rounded as to of Cimomia is unfortunately not known. be almost subangular, the lateral zones are Nevertheless it seems probable that the greatly flattened and converged ventrally, dorsad position of the siphuncle is one of and therefore the volutions are subtriangu- the best ways to distinguish immature lar in cross section, dorsal impressed zone equal-sized specimens of Cimomia and disregarded. Only the adoral half volution Hercoglossa. of our largest specimens (for example, pl. It should perhaps be mentioned that the 4, figs. 7, 8) have this more or less deltoid genus Deltoidonautilus has not been recog- cross section, for the rest of the conch is nized previously in South America. Only one not as narrowly rounded ventrally or as representative of it is so far known from strongly flattened laterally. The living North America, D. elliotti Stenzel of the chamber attains a length of more than half middle Eocene Reklaw formation of Texas. a volution. That species seems to be very close to the The umbilicus is small but moderately one under consideration, and, like it, is a prominent, and the umbilical shoulders are rather primitive form of the genus. The fairly distinct. The test seems to be of about specimen described below, which is also average thickness except in the region of the from northwestern Peru seems to be an umbilicus where it is greatly thickened to immature representative of a more advanced form a callus. Its surface is marked by form. numerous very fine faint transverse lines Occurrence.-All 11 of the specimens in which form broad rounded lateral salients the collection under consideration came and a moderately deep ventral sinus. On from the Lomitos sandstone (which until well preserved internal molds, there is a recently was regarded as a member of the rather prominent raised line along the Talara shale) at Cerro Organos, about 6 venter (pl. 4, figs. 1, 2). miles northeast of Cabo Blanco, Provincia At full maturity, each suture forms a high de Paita, Departamento de Piura, Peru. very narrowly rounded ventral saddle, and The holotype is from the Talara formation on either side of it a rather deep broadly at a nearby locality in northwestern Peru, rounded lateral lobe, a smaller prominent that is, near Lagunitas, Piura; and the TERTIARY NA UTILOIDS OF THE AMERICAS 9

specimen, mentioned above, which Miller either side of it a shallow broad rounded questionably associated with it, came from lateral lobe, a small but rather prominent the same formation at a nearby locality. very narrowly rounded umbilical saddle, a Repository.-State University of Iowa, small rounded lobe on the umbilical wall, 9537 (pl. 4, figs. 1-3, 7, 8) and 9538 (9 and apparently a slight internal lateral unfigured specimens). saddle which extends to a similar dorsal lobe. DELTOIDONAUTILUS sp. The siphuncle is small and is dorsal but Plate 1-3 10, figures not quite marginal in position. At the adoral Through the courtesy of Dr. A. Myra end of the specimen, it is almost 1 mm. in Keen and Sr. Pedro Verastegui M., we were diameter and is about half a millimeter from loaned for study a small representative of the dorsum. Deltoidonautilus from the lower Eocene Remarks.-As a result of its small size Salina formation of northwestern Peru-its and particularly because the form of the generic affinities had already been recog- conch and the shape of the sutures change nized by Sr. Verastegui. This specimen is a rapidly throughout the length of the outer well preserved completely septate internal volution, we believe that this specimen mold that is only about 23 mm. in diameter. represents only the immature inner portion Near the adapical end of its outer volution, of a conch. However, its generic affinities the conch is rather narrowly rounded are almost certainly with Deltoidonautilus, ventrally and is much wider than high. and it appears to be a rather well developed However, adorally its ventral zone is progres- representative of that genus. That is, in sively narrowed and gradually becomes sub- spite of its small size and presumed im- angular, and the conch increases in height maturity, its ventral zone is subangular, more rapidly than in width. At the adoral and in that respect it is farther advanced end of the specimen, the width and height than is D. haughti (Olsson) at full maturity. of conch measure about 154 and 14? mm., Nevertheless, the beds which carry that respectively. The maximum width is at- species are considerably younger than are tained at or just outside the umbilical those which yielded the specimen under shoulders, which are fairly distinct. The consideration. umbilicus is small and apparently open but Occurrence.-This specimen was secured not perforate. by Sr. Pedro Verastegui M. from the lower A small portion of the test (or rather a Eocene Salina formation south of the village replacement of it) is preserved in one of the of Yasila, Provincia de Paita, Departamento umbilical depressions. It shows that the de Piura, Peru (approximately at latitude test is quite thin. No trace of growth lines 5? 08' S. and longitude 81? 10' E.). can be discerned on the specimen. Repository.-Stanford University. The camerae are rather long, but they ATURIA ANGUSTATA become progressively shorter orad. In (Conrad)? Plate Plate 3 the adapical half of the outer volution of 1, figures 6-8; figure 8; Plate 2 this specimen there are only about five 9, figures 1, camerae, whereas there are eight in the ador- The collections of the University of al half of the same volution. California at Berkeley contain four speci- On the adapical portion of the outer mens which are probably referable to this volution, the external sutures are essentially species and which have not been mentioned straight and directly transverse, though in the literature. These were made available they seem to form an incipient ventral to us for study by Professor J. Wyatt Dur- saddle. Throughout the length of that volu- ham. tion this saddle becomes higher. Also, The one illustrated by figures 6 and 7 on umbilical saddles are developed in the outer plate 1 represents only the inner volutions volution, and they increase in prominence of the phragmacone. Its maximum diameter relatively rapidly. As a result, the adoral is about 24 mm., and the maximum width suture of this specimen consists of a low of the preserved portion of its conch meas- broad subangular ventral saddle and on ures about 12 mm. This specimen shows 10 A. K. MILLER AND H. R. DOWNS that during rather early ontogenetic de- some unrecorded locality in Kern County, velopment, the conch is relatively wide, the California, probably near the Mt. Poso oil sutures are moderately far apart, and the field. The adoral portion of this specimen is lateral lobes of the sutures are bluntly essentially free from distortion and repre- pointed and are not in contact with the sents the complete circumference of the preceding suture. It was found by Professor conch. Near its adoral end, the maximum Durham in the upper Oligocene Twin Rivers height and width of conch measure about formation about an eighth of a mile east 57 mm. and 37 mm., respectively. As shown of the city of Twin Rivers on the coast of by the accompanying figure, the shape of the Stra,its of Juan de Fuca, Clallam County, the sutures is also elucidated by this portion Washington (locality A3678 of Durham, of the specimen. 1944, p. 131). Remarks.-Miller (1947, pp. 85-88) has The individual illustrated by figure 8 on recently published the extensive synonymy the same plate is from the same formation at of this species and a detailed discussion of a nearby locality, that is, in a sea cliff it. Inasmuch as his report is readily avail- inshore from conglomerate and shale on the able, there is no need for us to duplicate beach just east of the first Geodetic Survey his work. mark east of the mouth of East Twin River, Occurrence.--This species is known to Clallam County, Washington. This speci- range from the middle (lower?) Oligocene to men is very incomplete, and it is not at all the middle Miocene in the Coast Ranges of well preserved. It is septate throughout, Washington, Oregon, and California, where and its maximum diameter measures about it has been found at many localities. The 62 mm. In the adapical portion of the outer precise horizon and locality of each of the volution, the shape of the lateral lobes of specimens discussed above is given in the the sutures can be seen quite clearly. preceding paragraphs. Figures 1 and 2 on plate 9 represent a Repository.-University of California, well preserved early mature portion of a Berkeley. that Professor Durham se- phragmacone ATURIA BRAZOENSISStenzel cured from a nodule in the upper Oligocene Plate 9, figures 5, 6 Blakeley formation at Bean Point on Bain- bridge Island, Washington (Univ. Calif. Miss Julia Gardner recently sent us for loc. C-X-3). The maximum diameter of this study a fine representative of the genus specimen is about 36 mm., and near its Aturia that was found April 16, 1941, in adoral end its conch is about 17 mm. wide the lower portion of the Claiborne group in and 22 mm. high. Attention should perhaps East Texas. It is a well preserved internal be called to the fact that the conch is rather mold that is completely septate. Its maxi- broadly rounded ventrally, and the lateral mum diameter measures about 104 mm., lobes of the sutures, which are pointed, are and the portion of its conch that is repre- in contact with the preceding sutures. Por- sented attained a maximum height of about tions of the test which are preserved show 67 mm. and a corresponding width of a that its surface is marked by numerous fine little more than 50 mm. (estimated). Its but quite distinct growth lines which form general physiognomy and the nature of its broad rounded lateral salients that exltend umbilicus, septa, etc. are elucidated by the from a presumed hyponomic sinus clear to accompanying photographs. the umbilicus. This specimen is of almost exactly the The other specimen that we are illustrat- same size, shape, and proportions as the ing (pl. 3, fig. 8) and referring to this species holotype of A. brazoensis Stenzel (a testifer- is from the Miocene Temblor formation at ous specimen), and there is no doubt in our

EXPLANATION OF PLATE 5 FIGS.1, 2-Hercoglossa popenoei M. & D., n. sp. Ventral and lateral views of the holotype, from a dark-gray calcareous Paleocene sandstone about 24 miles northwest of Punta Canoas, Baja California,very slightly less than X 1. (p. 6) JOURNAL OF PALEONTOLOGY, VOL. 24 PLATE 5

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110 TERTIARY NA UTILOIDS OF THE A MERICA S 11 mind that the two are conspecific. That Miocene), El Abrasandstone and conglomer- species has previously been found to occur ate (upper Miocene), and Matanzas marly at several horizons and localities in the limestone (Pliocene). "In this section the Eocene Claiborne group, that is, in the only beds which really are of limestone be- Stone City beds of central Texas, the Tyus long to the lower Miocene, Giiines formation and Viesca members of the Weches forma- This cavernous limestone is characterized tion of central and east Texas, and possibly by the abundanceof macrofossilsbut shows the Cook Mountain formation of south very poor conditions of preservation. Below Texas. Miller (1947, pp. 88-90, pl. 69, figs. this limestone there is a good marl forma- 1-4; pl. 70, figs. 1, 2; pl. 71, figs. 1, 2; pl. tion (upper Oligocene, Cojimar formation) 72, figs. 6, 7; pI. 73, fig. 7) has recently dis- with abundant microfauna. It is possible cussed the species in detail and listed its that the type locality of Aturia cubaensis complete synonymy. is in the Guines formation rather than in Occurrence.-"Probablyrather low in the the Cojimar formation." Claiborne group" on the east side of the Dr. Bermuidezalso kindly loaned us for Chireno road directly south of the San study a portion of an internal mold of a Augustine-Nacogdoches highway, Nacog- phragmaconeof an Aturia which many years doches County, Texas. ago was given to him by a friend who lives Repository.-U. S. National Museum. in Matanzas and who informed him that it came from "the Giiines ATURIA CUBAENSIS limestone, exposed (Lea) in the Yumuri Gorge of Matanzas." He Plate 9, 4 figures 3, adds that its presevation is "very similar Nautilius CubaensisLEA, 1841, Am. Phil. Soc., to the fossils of the Giiines formation."This Tr., n. ser., vol. 7, p. 259, pl. 10, fig. 15. (?) Aturia zig-zag SALTERAIN,1880, Bol. Com. specimen, like the holotype of A. cubaensis, Mapa geol. Espafia, t. 7, p. 198. is preserved in "hard fine-grained white Aturia cubaensis MILLER,1947, Geol. Soc. Amer- limestone," and the two most Mem. 4-6. probably ica, 23, pp. 7, 8, 10, 92, pl. 83, figs. came from the same formationand represent All that Miller was able to ascertain in only one species. However, Dr. Bermudez's regard to the derivation of the holotype of specimen is somewhat crushed, and its this species is that it was obtained long ago proportionscan thereforenot be determined by Louis Vanuxem from a "white limestone accurately. Its general physiognomy and of Matanzas," Cuba. Dr. Pedro J. Bermfidez the shape of its sutures are elucidated by who is thoroughly familiar with the geology the accompanyingfigures. It should perhaps of Cuba, and particularly that of the Matan- be stated that the maximum height of the zas area, has recently written us that in portion of the conch that is represented "the vicinity of Matanzas city there is a measures about 41 mm., and no trace of good exposition of Tertiary sediments with any markings other than the sutures can beds of lower Eocene, upper Oligocene, be discerned on the specimen. Miocene, and Pliocene. The locality most Furthermore, Dr. Bermudez called our visited by geologists and fossil collectors is attention to the fact that in 1880 Salterain the Yumuri Gorge very close to the city. had listed "Aturia zig-zag (Sow.)" from In this place the river cuts the sediments Cantera (quarry) La Criolla,Habana, Cuba. very deeply. The Tertiary is well exposed in According to Dr. Berm(udez,the age of the this cut" and consists of the following for- beds exposed at "this locality is upper mations: Universidad radiolarian earth Oligocene and lower Miocene." This form (lower Eocene), Cojimar marl (upper may therefore well.be referable to A. cuba- Oligocene), Giiines limestone (lower Mi- ensis. ocene), Canimar marly limestone (middle It should also be mentioned in this con-

EXPLANATIONOF PLATE6 FIGS. 1, 2-Hercoglossa peruviana Berry. Lateral and ventral views of a large completely septate individual, from the Eocene Talara formation near Lobitos, Peru, X . Same specimen as figure 2 on plate 7. (p. 5) 12 A. K. MILLER AND H. R. DOWNS nection that Mrs. Katherine V. W. Palmer cal shoulders are low, rounded, and in- has written us that in one of the notebooks definite. of the late Dorothy K. Palmer, the genus The test, which in both of the specimens Aturia is stated to be represented in the under consideration is iridescent, is thin type section of the Jicotea, that is, in "gray except in the umbilical region where it lime ss., side of Carretera Central in Jico- forms a callus. Its surface is marked by tea," Provincia Santa Clara, Cuba (Palmer, numerous fine but very distinct growth R. H., 1948, p. 12-locality 114). According lines, each of which forms a deep narrowly to Bermudez (1938) the age of the Jicotea rounded ventral sinus and on either side of is middle Eocene, and its type section has it a broad rounded lateral salient which yielded Globorotalia centralis Cushman and extends clear to the umbilicus. Bermudez, which Mrs. Dorothy K. Palmer The sutures are not well exposed, but considered characteristic of that age. It is they do not seem to differ in any available therefore rather doubtful that this form is particular from those of other representatives very closely related to A. cubaensis, which of this species. The spacing of the septa also presumably is typical of much younger is typical of the species. beds. The specimen represented by figures 6 and Occurrence.-Giiines limestone (Lower Mi- 7 on plate 10 (the preserved portion of which ocene) in the Yumuri Gorge near Matanzas, is about 55 mm. in diameter) reveals the Cuba. siphuncle particularly well. It is located Repository.-Private collection of Dr. close to the dorsum in adapical infundibular Pedro J. Bermudez of Caracas, Venezuela. inflections of the septa which are telescoped to a considerable extent. these two ATURIACURVILINEATA Miller and Remarks.-Clearly specimens are specifically identical with the similar but Thompson somewhat smaller one from the same general Plate 10, figures 4-7 horizon and locality described and illus- (?) Nautilus (fragments)MAURY, 1902, Bull. Am. trated by Miller in 1947 as "Aturia sp." Paleontology, vol. 3, no. 15, p. 60 (370). as noted that it is Aturia curvilineataMILLER and THOMPSON,1937, Also, by author, quite Eclogae geol. Helvetiae, vol. 30, pp. 61, 69-70, likely that the specimens which in 1902 pl. 9, figs. 1-4; pl. 10, figs. 1, 2; MILLERand Maury listed as "Nautilus (fragments)" are FURNISH,1938, Jour. Paleontology, vol. 12, conspecific, but they seem to have been lost. Geol. Soc. pp. 150, 151, fig. 1G; MILLER,1947, However, the of the Floridian America, Mem. 23, pp. 7, 10, 11, 40, 86, 93-94, relationship 106, 110, 111, pl. 73, figs. 3, 4; pl. 84, figs. 1, 2, specimens to those from South America that 5-8; pi. 85, figs. 4-6. have been referred to this species is not Aturia sp. MILLER, 1947, Geol. Soc. America, quite as certain. The holotype, which came Mem. 23, pp. 7, 8, 110-111, pl. 88, figs. 2, 3. from the Miocene of northwestern Vene- Two exquisite but slightly broken aturias zuela, is relatively large and is somewhat were recently collected from the Miocene distorted and laterally compressed, but of Florida by Mr. Chas. R. Locklin and sent other Miocene South American specimens to Miss Julia Gardner, who kindly forwarded (from the same and different localities) have them to us for study. Both are testiferous slightly but distinctly narrower conchs. and are septate throughout. Opinions of various authors in regard to the The maximum diameter of the more taxonomic value of such variations differ, nearly complete individual (pl. 10, figs. 4, 5) but we are inclined to believe that for the measures about 54 mm., and the correspond- present, at least, they should not be regarded ing height and width of its conch are approx- as of sufficient importance to justify specific imately 37 mm. and 21? mm., respectively. separation. The maximum width of conch is attained Occurrence.-Both of the specimens we just outside the umbilical shoulders, the are studying came from the lower part of lateral zones are almost flat and are distinct- the Miocene Chipola formation at Ten Mile ly converged ventrad, and the ventral zone Creek on the Chipola River, about 18 miles is rather narrowly rounded. The umbilicus south of Marian and 2 miles west of the is small and inconspicuous, and the umbili- McCelland farm, Calhoun County, Florida. TERTIARY NA UTILOIDS OF THE AMERICAS 13

The specimen which Miller described and umbilical and lateral portions of a little more illustrated in 1947 came from the same for- than half of one side of a volution of a mation at or near the same locality, where- phragmacone (and the crushed inner whorls) as those listed by Maury in 1902 were from and it is so distorted and fragmentary that Bailey's Ferry, also in Calhoun County, its specific affinities are very uncertain. The Florida. In South America, specimens that preserved portion of this incomplete speci- are tentatively being regarded as con- men is estimated to have attained a diameter specific are known from the Miocene of of some 150 mm. The lateral lobes of the Venezuela, Trinidad, and Ecuador. sutures are asymmetrically attenuate, and Figured Specimens.-U. S. National Mu- at least these lobes and the lateral saddles seum and The Academy of Natural Sciences are very similar to those of the holotype. of Philadelphia. The collections under consideration also contain three ATURIA cf. A. GRANDIORSchenck large fragmentary specimens from the equivalent of the Keasey in south- Plate 7, figure 1; Plate 8, figures 1, 2 western Washington. None of these is The collections of the University of Cali- sufficiently well preserved or nearly enough fornia at Berkeley contain seven specimens complete to be identified specifically, but that may belong in this species, and Profes- all three appear to represent portions of sor J. Wyatt Durham kindly loaned them phragmacones of the same general size and for study. Two of them merit illustration proportions as typical A. grandior. We are and all seven at least brief description. therefore comparing them with that species. The largest (pl. 7, fig.l) is unfortunately The specimen illustrated by figures 1 and somewhat but it to be crushed, appears 2 on plate 8 is from the Oligocene Quimper essentially complete. It is testiferous and, sandstone of northwestern Washington. It in seems to resemble the holo- except size, is primarily an internal mold of the outer rather so that the two well type closely may volution of a moderately large specimen, be this one came from conspecific, though but considerable portions of the test adhere the Eocene Cowlitz formation whereas the to it. The adoral part of this specimen repre- is from an tuff-both were type Oligocene sents living chamber. The phragmacone found in western The maxi- Washington. attained a maximum height of conch of mum diameter of this in its specimen, about 95 mm., and the corresponding width present crushed state, is about 235 mm.- of conch and over all diameter are estimated that of the is about 160 mm. Be- holotype to have been 65 mm. and 150 mm., cause of the distortion which respec- the conch The lateral zones of the whorl are has other tively. undergone measurements would flattened and are con- have little As in the strongly distinctly significance. holotype, ventrad, and the maximum width the chamber is a little more than half verged living of conch is therefore attained ven- a volution in and the surface of the slightly length, trad of the low rounded indefinite umbilical test is marked fine but distinct by very shoulders. The ventral zone of the conch lines each of which forms a growth deep is rather and rounded. The ventral sinus and on either side of it a broadly evenly umbilicus is small and and broad rounded lateral salient that extends inconspicuous seems to be filled with a callus-elsewhere to the umbilicus. In the adoral portion of the test is less than 1 mm. thick. The shape this specimen the test is approximately 1 of the sutures is elucidated by the accom- mm. thick, except in the umbilical region, illustrations, and they seem to be where it forms a callus. The shape of the panying the same as those of the holo- adoral sutures is revealed, and they do not essentially we believe that this seem to differ materially from those of the type. All in all, specimen holotype. can be referred to A. grandior with a reason- The lower Oligocene Keasey shale (or its able degree of assurance. that we are stratigraphic equivalent) in northwestern The remaining specimen Oregon has also yielded a specimen which associating with this species is from the may be related to the holotype of this Oligocene Lincoln "formation" on Marrow- species. However, it represents only the stone Island of northwestern Washington. 14 A. K. MILLER AND H. R. DOWNS

It represents about a quarter of each of two large fragmentary specimens); (4) Keasey successive volutions of a phragmacone which equivalent in high bluff along Willapa River is estimated to have attained a diameter at Big Bend about 14 miles due north of of more than 200 mm. The test (or rather a Hanan foot bridge across river north of replacement of it) of the outer volution of Holcomb, Pacific County, Washington (loc. this specimen is crushed and is broken into A64 of Durham, 1944, p. 130); (5) con- a number of fragments, but it appears to cretion in Molopophorus gabbi zone of Quim- be very thick. The preserved part of the per sandstone (lower Oligocene) along beach penultimate volution (unlike that of the close to Willamette Meridian line, center outer volution) is largely free from distor- NE- SE' sec. 1, T. 29 N., R. 1 W., Jefferson tion, and it shows that the cross section of County, Washington (loc. A3702 of Durham the conch and the size and position of the 1944, p. 132); and (6) Turritella porterensis siphuncle are essentially the same as in zone of Lincoln "formation" (middle Olig- typical representatives of this species. ocene), 100 feet stratigraphically below Nevertheless, this specimen is so crushed and sandy shales of Marrowstone shale in sea fragmentary that its specific affinities are cliff, center NE' sec. 32, T. 30 N., R. 1 E., uncertain. Marrowstone Island, Jefferson County, Remarks.-Miller (1947, pp. 97-98, pl. 93, Washington (loc. A3696 of Durham, 1944, fig. 5; pl. 94, figs. 1, 2) has recently published p. 132). Durham (1944, p. 121) indicates a detailed description and illustrations of that this species also occurs in the Turritella the type specimens of this species, together porterensis zone of the Marrowstoneshale with a synonymy, and since his work is (middle Oligocene) in the NW4 NE- sec. readily available there is no need to duplicate 32, T. 30 N., R. 1 E., Marrowstone Island it. However, attention should be called to (loc. A3695 of Durham, 1944 p. 132). Both the fact that he did not note that Durham the holotype and the paratype came from (1944, pp. 120, 121 ,192) had referred to the the Oligocene of western Washington. species three (or more) specimens from the Repository.-University of California, Oligocene of northwestern Washington- Berkeley. two of these are described above. ATURIA PERUVIANA Olsson Occurrence.-The seven specimens under Plate 6 consideration are from the following horizons 4, figures 5, and localities: (1) Cowlitz formation (upper A collection of cephalopods made by Sr. Eocene) on Olequah Creek, just below the Pedro Verastegui M. at Cerro Organos, near falls and above the grits, Lewis County, Cabo Blanco in northwestern Peru, and sent Washington (presumably equivalent to to us by Professor Bernardo Boit of the Univ. Washington loc. 301); (2) Keasey Universidad de San Marcos contains three shale, or its equivalent, (lower Oligocene) in small completely septate internal molds south bank of Rock Creek just west of that are most probably referable to this highway and railroad bridges (which are species. All of these are of about the same side-by-side), 1.9 miles east of Keasey post general size. The best one is represented by office and 0.2 mile west of Tara station, figures 5 and 6 on plate 4. It is about 32?mm. Colombia County, Oregon; (3) Keasey equi- in diameter and near its adoral end, its valent in bluff on north side of bend in conch is about 20 mm. high and 18 mm. wide. Willapa River about half a mile east of The maximum diameter of the other two Holcomb, Pacific County, Washington (two specimens measures about 33 mm. and 35

EXPLANATION OF PLATE 7 FIG. 1-Aturia cf. A. grandiorSchenck. An essentially complete crushed testiferous individual, from the Eocene Cowlitz formation just below the falls and above the grits on Olequah Creek, Lewis County, Washington, X#. (p. 13) 2-Hercoglossa peruvianaBerry. Apertural view of a large completely septate individual, from the Eocene Talara formation near Lobitos, Peru, X 3. Same specimen as figures 1 and 2 on plate 6. (p. 5) 0 / L? r.. ?i ..

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*4 *-~~~t-i.jt.6 "???\ utL?".? L TERTIARY NA UTILOIDS OF THE AMERICAS 15 mm. The general physiognomy of the conch width measures about 20 mm. It is rather and the nature of the sutures are elucidated narrowly rounded ventrally, and its flat- by the accompanying illustrations. tened lateral zones converge ventrad rather Remarks.-These three specimens are of strongly. The surface of its test is marked about the same size and shape as one from by numerous fine growth lines, each of which the same general horizon and locality that forms a deep narrowly rounded ventral was figured by Miller (1947, pl. 83, figs. 1-3). sinus and on either side of it a broad rounded A direct comparison of it with those now lateral salient which extends clear to the under consideration leaves no doubt in closed umbilicus. Only small portions of the our mind that all four are conspecific. sutures are visible, but they do not seem to Miller's report, which is readily available, differ materially from those of similar con- contains a detailed description and a com- generic forms. This specimen came from a plete synonymy of this species. black fine-grained sandstone layer (with Occurrence.-All three of the specimens abundant Loxotrema) about 16 to 20 feet we are studying were found in association stratigraphically below the contact with the with Deltoidonautilus haughti (Olsson) in the basalt, in the Eocene Crescent formation Lomitos sandstone at Cerro Organos, 6 miles near the base of the sea cliff on a small northeast of Cabo Blanco, Provincia de point about 250 yards west of the beacon Paita, Departamento de Piura, Peru. This shown on the west side of Crescent Bay on species, as now interpreted, also occurs in the Port Crescent quadrangle, Clallam the upper Eocene Saman formation in the County, Washington. same general region and in the upper Eocene At one locality, the Vacaville shale of of southwestern Ecuador?, Panama, north- central California has yielded three aturias ern Colombia, and northwestern Venezuela. that are crushed and rather poorly pre- Furthermore, some of the paratypes of it served. Presumably all are conspecific, but came from the Oligocene of northwestern their preservation leaves so much to be Peru, and the middle Eocene of that area desired that we are uncertain in regard to has yielded a single small immature speci- their precise affinities. All are internal molds men that is probably conspecific. that are largely septate. The best one is Repository.-State University of Iowa, represented by figures 1 and 2 on plate 3. 9539 (pl. 4, figs. 5, 6) and 9540 (2 unfigured One of the others seems to represent a slight- specimens). ly larger individual, whereas the third is ATURIAspp. only a very small segment of a phragmacone Plate 1, figure 9; Plate 3, of uncertain size. Because of the distortion figures 1, 2, 5, 6 which all three of these have undergone, The collections under consideration con- precise measurements of them would have tain a few Pacific Coast Eocene aturias of little significance. The spacing and the shape which we are unable to determine the of their sutures, and particularly the form specific affinities. These came from two of the lateral lobes of those sutures (see pl. localities in California and one in Washing- 3, fig. 1) indicate that this form is a rather ton, and they were loaned to us for study primitive representative of the genus. All by Professor J. Wyatt Durham. three of these specimens are from the lower One of them (pl. 1, fig. 9) is from the Cres- Eocene Vacaville shale member of the cent formation of northwestern Washington. Capay formation about half a mile north It is a rather poorly preserved testiferous of "Black Rocks" in Putah Creek, 1280 feet specimen which is about 54 mm. in diameter. east and 1302 feet north of the southeast It appears to have been slightly crushed and corner of sec. 23, T. 8 N., R. 2 W. (Mount flattened laterally, but its present maximum Diablo Meridian), Yolo County, California.

EXPLANATION OF PLATE 8 FIGs. 1, 2-Aturia grandiorSchenck. Ventral and lateral views of a specimen from a concretion in the OligoceneQuimper sandstone along the beach close to the Willamette Meridianline, center NE4 SEI sec. 1, T. 29 N., R. 1 W., JeffersonCounty, Washington, X . (p 13) 16 A. K. MILLER AND H. R. DOWNS

The specimen represented by figures 5 tion may well be correct, but it is difficult and 6 on plate 3 is the one from the Arroyo to substantiate. This specimen came from Hondo formation of west-central California the Eocene Arroyo Hondo formation in the that was illustrated and briefly described by SW' NW' sec. 15, T. 18 S., R. 14 E., 100 Vokes (1939, p. 107, pl. 16, fig. 36). It is a feet up the fourth small draw from west completely septate internal mold which has end of ridge, immediately opposite the a maximum diameter of only about 12' mm. place where Urruttia Canyon enters Salt Its small size, and particularly the wide Creek, north of Coalinga, Fresno County, spacing of its septa indicate that it repre- California (Univ. Cal. loc. 1817).

I

FIG. 2-Diagrammatic cross section and representation of an adoral suture of a small specimen belonging in the genus Aturia, from the lower Miocene of the west-central portion of the Sinai Peninsula, both X4. sents only the immature portion of a phrag- Occurrence.-One of the forms described macone. This belief is strengthened by the above is from the Crescent formation of fact that the lateral lobes of the sutures are northwestern Washington, another is from rather bluntly pointed. Vokes states that the Vacaville shale (Capay) of central this specimen was found associated "with California, and the third is from the Arroyo abundant fragments of the outer shell of Hondo formation of west-central California. much larger individuals," and he referred The precise horizon and locality is given it to Aturia myrlae Hanna; that identifica- along with the discussion of each.

EXPLANATION OF PLATE 9 FIGS. 1, 2-Aturia angustata(Conrad)? A small well preserved internal mold, from a nodule in the Oligocene Blakeley formation at Bean Point on Bainbridge Island, Washington, X2. (p. 9) 3, 4-Aturia cubaensis(Lea)? A crushed fragment of an internal mold, from the lower Miocene Giines limestone in the Yumuri Gorge, near Matanzas, Cuba, X1. (p. 11) 5, 6-Aturia brazoensisStenzel. A well preservedinternal mold, from the lower part of the Eo- cene Claiborne group on the east side of the Chireno road directly south of the San Au- gustine-Nacogdoches highway, Texas, X1. (p. 10) JOURNAL OF PALEONTOLOGY, VOL. 24 PLATE 9

3

Miller and Downs, Tertiarynautiloids JOURNAL OF PALEONTOLOGY, VOL. 24 PLATE 1 0

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Miller and Downs, Tertiary nautiloids TERTIAR Y NA UTILOIDS OF THE AMERICAS 17

Repository.-All are in the paleontological this genus, which strengthens our conviction collections of the University of California that we are not dealing with a fully mature at Berkeley. The one from the Arroyo phragmacone. Hondo formation is numbered 15877. Although this specimen may well belong in Aturia aturi, which was orginally described APPENDIX from the Miocene of France, we hesitate to refer it to It is the AN ATURIA FROMTHE MIOCENEOF SINAI definitely any species. property of the Paleontological Museum Through the courtesy of Dr. A. Myra of Stanford University, where its catalogue Keen of Stanford University, we were kindly number is 7993 (type collection). loaned for study a small representative of the genus Aturia, which seems to merit REFERENCES special consideration because of the locality BERMfDEZ,P. J., 1938, Foraminiferosde la fauna from which it came. This specimen was de Jicotea (Eoceno Medio), Provincia Santa secured in March of 1921 by Mr. Robert Clara, Cuba: Soc. Cubana Hist. Nat. Felipe van Vleck Anderson from the "lower beds Poey, Mem. 12, nfim. 2, pp. 91-96. DALL, W. H., 1909, Contrioutions to the Ter- of [the] Miocene series" in a prominent tiary paleontology of the Pacific coast; I, The bluff on the south side of the mouth of Miocene of Astoria and Coos Bay, Oregon: Wadi Gharandel on the Gulf of Suez coast U. S. Geol. Surv. Prof. Pap. 59, pp. 1-278, pls. 1-23. of Sinai, some two or three miles north DURHAM,J. WYATT, 1944, Megafaunal zones of of Hammam Faraun. The label which ac- the Oligocene of northwestern Washington: companies it shows that it was identified California Univ. Publ., Dept. Geol. Sci., Bull., by H. G. Schenck as Aturia aturi (Basterot), vol. 27, pp. 101-212, pls. 13-18. Notice of the and it may well be the basis for his indi- LEA, ISAAC,1841, Oolitic formation in America, with descriptions of some of its cation (Schenck, 1931, map on p. 437) that organic remains: Am. Phil. Soc., Tr., n. ser., the genus occurs on this peninsula. vol. 7, pp. 251-260, pls. 8-10. This specimen is partially imbedded in a MAURY, C. J., 1902, A comparison of the Oligo- block of mottled with cene of western Europe and the southern light-colored (white United States: Bull. Am. vol. limestone which Paleontology, 3, brown) crystalline contains no. 15, pp. 1-118, pls. 20-29. some rather poorly preserved gastropods MILLER,A. K., 1947, Tertiary nautiloids of the and other small fossils. Its general physiog- Americas: Geol. Soc. America, Mem. 23, pp. is elucidated the 1-234, pls. 1-100. nomy by accompanying A. Contributions to the Ter- illustrations and text OLSSON, A., 1928, (pl. 4, fig. 4; fig. 2); tiary paleontology of northern Peru; Part 1, but attention should perhaps be called to Eocene Mollusca and Brachiopoda:Bull. Am. the fact that its maximum diameter is only Paleontology, vol. 14, no. 52, pp. 1-154, pls. about 23 and therefore we are of the 1-26. mm., -- Contributionsto the that it the , 1930, Tertiary paleon- opinion represents adolescent tology of northern Peru; Part 3, Eocene Mol- or early mature portion of a conch that was lusca: Bull. Am. Paleontology, vol. 17, no. 62, much larger at full maturity. At least its pp. 1-96, pls. 1-12. adoral sutures are so that the PALMER, R. H., 1948, List of Palmer Cuban fossil closely spaced localities: Bull. Am. vol. no. of the lateral Paleontology, 31, adapical portions ("tips") 128, pp. 1-178, 1 map. lobes are in contact with the adoral portions SALTERAIN,PEDRO, 1880, Apuntes para una de- (ventrolateral shoulders) of the ventral sad- scripcion fisico-geologica de las jurisdicciones dles of the sutures. The lateral de la Habana y Guanabacoa (Isla de Cuba): preceding Com. t. 161- lobes of the sutures seem to be rather Mapa geol. Expafia, Bol., 7, pp. 225, 1 map. primitive for a Miocene representative of SCHENCK,H. G., 1931, Cephalopods of the genus

EXPLANATION OF PLATE 10 FIGS. 1-3-Deltoidonautilus sp. A small immature specimen from the Eocene Salina formation south of the village of Yasila, Departamento de Piura, Peru, X 1 . (p. 9) 4-7-Aturia curvilineata Miller and Thompson. Two exceptionally well preserved testiferous specimens, from the Miocene Chipola formation at Ten Mile Creek on the Chipola River, Calhoun County, Florida, X1. 4, 5, USNM specimen. (p. 12) 18 A. K. MILLER AND H. R. DOWNS

Aturia from western North America: Cali- and Arroyo Hondo formations of the California fornia Univ. Publ., Dept. Geol. Sci., Bull., vol. Eocene: New York Acad. Sci., Ann., vol. 38, 19, pp. 435-491, pls. 66-78. pp. 1-246, pls. 1-22. VOKES, H. E., 1937, Nautiloid cephalopods from WIEDEY, L. W., and FRIZZELL, D. L., 1940, Revi- the Eocene of California: Jour. Paleontology, sion of the Eocene stratigraphy of northwestern vol. 11, pp. 3-9, pls. 1, 2. Peru: Pacific Sci. Cong., Sixth, California, - , 1939, Molluscan faunas of the Domengine 1939, Pr., vol. 2, pp. 527-528.

MANUSCRIPT RECEIVED JUNE 7, 1949.