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Los Restos Directos De Dinosaurios Terópodos (Excluyendo Aves) En España
Canudo, J. I. y Ruiz-Omeñaca, J. I. 2003. Ciencias de la Tierra. Dinosaurios y otros reptiles mesozoicos de España, 26, 347-373. LOS RESTOS DIRECTOS DE DINOSAURIOS TERÓPODOS (EXCLUYENDO AVES) EN ESPAÑA CANUDO1, J. I. y RUIZ-OMEÑACA1,2 J. I. 1 Departamento de Ciencias de la Tierra (Área de Paleontología) y Museo Paleontológico. Universidad de Zaragoza. 50009 Zaragoza. [email protected] 2 Paleoymás, S. L. L. Nuestra Señora del Salz, 4, local, 50017 Zaragoza. [email protected] RESUMEN La mayoría de los restos fósiles de dinosaurios terópodos de España son dientes aislados y escasos restos postcraneales. La única excepción es el ornitomimosaurio Pelecanimimus polyodon, del Barremiense de Las Hoyas (Cuenca). Hay registro de terópodos en el Jurásico superior (Oxfordiense superior-Tithónico inferior), en el tránsito Jurásico-Cretácico (Tithónico superior- Berriasiense inferior) y en todos los pisos del Cretácico inferior, con excepción del Valanginiense. En el Cretácico superior únicamente hay restos en el Campaniense y Maastrichtiense. La mayor parte de las determinaciones son demasiado generales, lo que impide conocer algunas de las familias que posiblemente estén representadas. Se han reconocido: Neoceratosauria, Baryonychidae, Ornithomimosauria, Dromaeosauridae, además de terópodos indeterminados, y celurosaurios indeterminados (dientes pequeños sin dentículos). La mayoría de los restos son de Maniraptoriformes, siendo especialmente abundantes los dromeosáuridos. Las únicas excepciones son por el momento, el posible Ceratosauria del Jurásico superior de Asturias, los barionícidos del Hauteriviense-Barremiense de Burgos, Teruel y La Rioja, el posible carcharodontosáurido del Aptiense inferior de Morella y el posible abelisáurido del Campaniense de Laño. Además hay algunos terópodos incertae sedis, como los "paronicodóntidos" (entre los que se incluye Euronychodon), y Richardoestesia. -
Paleoherpetofauna Portuguesa
Rev. Esp. Herp. (2002): 17-35 17 Paleoherpetofauna Portuguesa E.G. CRESPO Centro de Biologia Ambiental – Fac. Ciências Univ. Lisboa Resumo: Nos últimos anos a importância da paleoherpetofauna portuguesa tem sido posta em evidência sobre- tudo através do seu grupo mais mediático, os dinossauros. As recentes descobertas em Portugal de vestígios de vários dinossauros, incluindo ossos, ovos, embriões, gastrólitos e pegadas, têm merecido ampla cobertura jorna- lística e têm sido oportunamente acompanhadas por intensas campanhas de divulgação, levadas a cabo pelo Mu- seu Nacional de História Natural de Lisboa, encabeçadas pelo geólogo, Professor Galopim de Carvalho. As pro- longadas e por vezes polémicas acções de sensibilização pública e política que foi necessário empreender para se preservarem muitos dos locais onde esses vestígios foram encontrados, contribuiram também para sustentar e até aumentar o interesse por este grupo de grandes répteis. A importância da paleoherpetofauna portuguesa está porém longe de se limitar apenas aos dinossauros! Em Portugal viveram muitos outros répteis e anfíbios de que existem vestígios desde o começo do Mesozói- co –Quelónios, Crocodilos, Ictiossauros, Plesiossauros, Pterossauros, Lepidossauros, “Estegossauros” e Lis- samphia– que, embora geralmente muito menos conhecidos, têm um significado evolutivo, paleogeográfico e paleoclimático extremamente importante. Na sua descoberta e estudo estiveram envolvidos, já desde o século passado, numerosos investigadores por- tugueses e estrangeiros, dos quais se destacam, entre outros, Georges Zbyszewski, Miguel Telles Antunes, Vei- ga Ferreira, H. Sauvage, A.F. Lapparent, L. Ginsburg, R.Thulborn, P. Galton. Muitos destes estudos encontram- se todavia dispersos por uma vasta gama de publicações em que, frequentemente, as referências aos répteis e aos anfíbios ou são laterais ou são apresentadas em contextos zoológicos mais abrangentes, pelo que, como parece que tem acontecido, têm passado praticamente despercebidos à maioria daqueles que se dedicam aos estudo da nossa herpetofauna actual. -
Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia. -
Coossified Tarsometatarsi in Theropod Dinosaurs and Their Bearing on the Problem of Bird Origins
HALSZKA OSM6LSKA COOSSIFIED TARSOMETATARSI IN THEROPOD DINOSAURS AND THEIR BEARING ON THE PROBLEM OF BIRD ORIGINS OSM6LSKA, H. : Coossified tarsometatarsi in theropod dinosaurs and their bearing on the problem of bird origins, Palaeontologia Polonica, 42, 79-95, 1981. Limb remains of two small theropod dinosaurs from the Upper Cretaceous deposits of Mongolia display fused tarsometatarsi. Presence of fusion in the tarsometatarsus in some theropods is consi dered as additional evidence for the theropod origin of birds. E/misaurus rarus gen. et sp. n. is described based upon a fragmentary skeleton represented by limbs. Family Elmisauridae novo is erected to include Elmisaurus, Chirostenotes GlLMORE and Ma crophalangia STERNBERG. Key words: Dinosauria, Theropoda, bird origins, Upper Cretaceous, Mongolia. Halszka Osmolska , ZakladPaleobiologii, Polska Akademia Nauk, Al. Zw irki i Wigury 93,02-089 War szawa, Po/and. Received: June 1979. Streszczenie. - W pracy opisano szczatki malych dinozaur6w drapieznych z osad6w gornokredo wych Mongolii . Stopa tych dinozaur6w wykazuje obecnosc zrosnietego tarsomet atarsusa. Zrosniecie to stanowi dodatkowy dow6d na pochodzenie ptak6w od dinozaur6w drapieznych, Opisano nowy rodzaj i gatunek dinozaura drapieznego E/misaurus rarus, kt6ry zaliczono do nowej rodziny Elmisau ridae . Do rodziny tej, opr6cz Elmisaurus, naleza: Chirostenotes GILMORE i Macr opha/angia STERNBERG. Praca byla finansowana przez Polska Akademie Nauk w ramach problemu rniedzyresorto wego MR 11-6. INTRODUCTION During the Polish-Mongolian -
104Ornithodiraphyl
Millions of Years Ago 252.3 247.2 235.0 201.5 175.6 161.2 145.5 99.6 65.5 Triassic Jurassic Cretaceous Early Middle Late Early Middle Late Early Late Euparkeria Crurotarsi ? Scleromochlus ? Archosauria Pterosauria Lagerpetidae Ornithodira Marasuchus Genasauria Dinosauromorpha Silesauridae Neornithsichia Thyreophora Ornithischia Eocursor (esp. Dinosauria) et al. (2011), Yates (2007) Yates et al. (2011), Nesbitt etal.(2009), Sues (2007), Martinezet al.(2011), Irmis etal. Ezcurra (2006), EzcurraandBrusatte (2011), Phylogeny after Brusatteetal.(2010), Butleretal.(2007), Heterodontosauridae Pisanosaurus Dinosauria Ornithodira Sauropodomorpha Herrerasauria Saurischia Eodromeus Theropoda Daemonosaurus Tawa Neotheropoda Millions of Years Ago 253.0 247.2 235.0 201.5 175.6 161.2 145.5 99.6 65.5 Triassic Jurassic Cretaceous Early Middle Late Early Middle Late Early Late (2009), Norman et al. (2004), Thompson etal. (2011) (2009), Norman etal. (2004), Phylogeny afterButler etal. (2007a,b), Carpenter (2001),Galton &Upchurch (2004), Maidment etal.(2008), Mateus etal. Cerapoda Ornithopoda Eocursor Marginocephalia Neornithischia Othnielosaurus Genasauria (esp. Thyreophora) Genasauria (esp. Hexinlusaurus Stormbergia Genasauria Lesothosaurus Scutellosaurus Thyreophora Scelidosaurus Stegosauridae Stegosaurinae Dacentrurinae Stegosauria Kentrosaurus Tuojiangosaurus Huayangosauridae Gigantspinosaurus Eurypoda Tianchiasaurus Ankylosauria Nodosauridae Ankylosauridae Millions of Years Ago 253.0 247.2 235.0 201.5 175.6 161.2 145.5 99.6 65.5 Triassic Jurassic Cretaceous -
Uncompahgre Dinosaur Fauna: a Preliminary Report
Great Basin Naturalist Volume 45 Number 4 Article 8 10-31-1985 Uncompahgre dinosaur fauna: a preliminary report James A. Jensen Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/gbn Recommended Citation Jensen, James A. (1985) "Uncompahgre dinosaur fauna: a preliminary report," Great Basin Naturalist: Vol. 45 : No. 4 , Article 8. Available at: https://scholarsarchive.byu.edu/gbn/vol45/iss4/8 This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. UNCOMPAHGRE DINOSAUR FAUNA: A PRELIMINARY REPORT James A. Jensen Abstract.—A diverse late Jurassic dinosaur fauna, discovered in western Colorado in 1963, contains many unde- scribed taxa that may represent evolutionary trends at the generic level not previously reported from the Morrison Formation. A preliminary faunal hst is given. Bones of the largest known dinosaur, Ultrasaurus , are present as are a variety of small animals, including Pterosaurs, in which one sacrum displays avianlike fused sacral neural spines. A new family, the Torvosauridae , erected, based on the genus Torvosaunis that is redescribed. One of the most diverse Jurassic dinosaur their field investigations "failed to find any faunas in North America was found on the convincing evidence of evolution at the Uncompahgre Upwarp in western Colorado generic level within the Morrison Forma- in 1963. This fauna contains more unde- tion." There are familiar forms in the Uncom- scribed taxa than has been encountered in any pahgre fauna, but there is also consistent evi- other North American Jurassic assemblage in dence of change, or "evolution at the generic this century. -
Caracterización De Una Relación Alométrica En Theropoda (Dinosauria) Con Énfasis En Su Extensión E Implicaciones Evolutivas
CARACTERIZACIÓN DE UNA RELACIÓN ALOMÉTRICA EN THEROPODA (DINOSAURIA) CON ÉNFASIS EN SU EXTENSIÓN E IMPLICACIONES EVOLUTIVAS Tesis Entregada A La Universidad De Chile En Cumplimiento Parcial De Los Requisitos Para Optar Al Grado De Magíster en Ciencias Biológicas Facultad De Ciencias Por José Antonio Palma Liberona Mayo, Año 2018 Director de Tesis Dr. Alexander O. Vargas Milne Co-Director de Tesis Dr. Marco A. Méndez Torres Proyecto Financiado mediante proyecto Anillo ACT172099 y Fondecyt 1150906 FACULTAD DE CIENCIAS UNIVERSIDAD DE CHILE INFORME DE APROBACIÓN TESIS DE MAGÍSTER Se informa a la Escuela de Postgrado de la Facultad de Ciencias que la Tesis de Magíster presentada por el candidato. José Antonio Palma Liberona Ha sido aprobada por la comisión de Evaluación de la tesis como requisito para optar al grado de Magíster en Ciencias Biológicas, en el examen de Defensa Privada de Tesis rendido el día 4 de Abril del 2018 Director de Tesis: D r . Alexander O. Vargas Milne ………………… Co-Director de Tesis D r . M a r c o A . Méndez Torres ………………… Comisión de Evaluación de la Tesis D r . Claudio P. Veloso Iriarte ………………… D r . Mauricio Canals Lambarri ………………… “He impaired his vision by holding the object too close. He might see, perhaps, one or two points with unusual clearness, but in doing so he, necessarily, lost sight of the matter as a whole. Thus there is such thing as being too profound.” ̶ Edgar Allan Poe, The Murders in the Rue Morgue iii BIOGRAFÍA José Antonio Palma Liberona nació el 1 de marzo de 1990, hijo de José Palma Keller y Flavia Liberona Céspedes. -
Avialan Status for Oviraptorosauria
Avialan status for Oviraptorosauria TERESA MARYAŃSKA, HALSZKA OSMÓLSKA, and MIECZYSŁAW WOLSAN Maryańska, T., Osmólska, H., and Wolsan, M. 2002. Avialan status for Oviraptorosauria. Acta Palaeontologica Polonica 47 (1): 97–116. Oviraptorosauria is a clade of Cretaceous theropod dinosaurs of uncertain affinities within Maniraptoriformes. All pre− vious phylogenetic analyses placed oviraptorosaurs outside a close relationship to birds (Avialae), recognizing Dromaeo− sauridae or Troodontidae, or a clade containing these two taxa (Deinonychosauria), as sister taxon to birds. Here we pres− ent the results of a phylogenetic analysis using 195 characters scored for four outgroup and 13 maniraptoriform (ingroup) terminal taxa, including new data on oviraptorids. This analysis places Oviraptorosauria within Avialae, in a sister−group relationship with Confuciusornis. Archaeopteryx, Therizinosauria, Dromaeosauridae, and Ornithomimosauria are suc− cessively more distant outgroups to the Confuciusornis−oviraptorosaur clade. Avimimus and Caudipteryx are succes− sively more closely related to Oviraptoroidea, which contains the sister taxa Caenagnathidae and Oviraptoridae. Within Oviraptoridae, “Oviraptor” mongoliensis and Oviraptor philoceratops are successively more closely related to the Conchoraptor−Ingenia clade. Oviraptorosaurs are hypothesized to be secondarily flightless. Emended phylogenetic defi− nitions are provided for Oviraptoridae, Caenagnathidae, Oviraptoroidea, Oviraptorosauria, Avialae, Eumaniraptora, Maniraptora, and Maniraptoriformes. -
The Family Deinodontidae, with Notice of a New Genus from the Cretaceous of Alberta
56.81,9(117:71.2) Article VI.—THE FAMILY DEINODONTIDAE, WITH NOTICE OF A NEW GENUS FROM THE CRETACEOUS OF ALBERTA By W. D. Matthew and Barnum Brown I. —Introductory note. II. —Distinctive Characters of the Megalosaurs, Ccelurids, Ornithomimids, and Deinodonts. III. —Chronological List of Described Species of Deinodontidse and Ornithomimidae. IV. —Provisional Systematic Arrangement. V. —Remarks upon the Nomenclature of the Deinodontidse. VI. —A New Genus of Carnivorous Dinosaurs from the Cretaceous of Alberta. I.—INTRODUCTORY NOTE This is the first of a series of preliminary notices to be published by Mr. Brown and myself to place upon record various contributions to a knowledge of the Cretaceous Dinosaurs resulting from the prepara tion of the collections secured in Alberta by the Museum parties of 1910 to 1915 under Mr. Brown’s leadership. It had been planned that these results should be studied and published by him, but owing to his absence from the Museum for some years past, mostly on field work abroad, his researches have been long delayed. Some obvious preliminary results we have thought advisable to publish now, postponing the more com plete research and publication of the material until Mr. Brown’s return to the Museum enables him to resume his more intensive studies upon the several groups of Cretaceous dinosaurs. As should appear from the situation above outlined, the junior author should be credited with the new evidence and data placed upon record, as the results of his splendidly successful series of expeditions in the Western Cretaceous formations. The senior author is chiefly re sponsible for the interpretation of the data, the revision of previous con clusions and taxonomy, and for various possible errors which will later be corrected in the extended researches planned by his absent friend and colleague, with whose approval and in the interest of the American Mu seum these contributions are placed upon record.—W. -
New Data on Small Theropod Dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA
Volumina Jurassica, 2014, Xii (2): 181–196 Doi: 10.5604/17313708 .1130142 New data on small theropod dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA Sebastian G. DALMAN1 Key words: dinosaurs, Theropoda, Upper Jurassic, Morrison Formation, Como Bluff, Wyoming, western USA. Abstract. In 1879, Othniel C. Marsh and Arthur Lakes collected in the Upper Jurassic Morrison Formation Quarry 12 at Como Bluff, Wyoming, USA, several isolated axial and appendicular skeletal elements of small theropod dinosaurs. Since the discovery the specimens remained unnoticed for over a century. The skeletal remains of small theropods are rare at Como Bluff and throughout the Morrison Forma- tion. Their bones are delicately constructed, so they are not as well-preserved as the bones of large-bodied theropods. The bones of small theropods described here were found mixed with isolated crocodile teeth and turtle shells. Comparison of the skeletal materials with other known theropods from the Morrison Formation reveals that some of the bones belong to a very small juvenile Allosaurus fragilis and Tor vosaurus tanneri and also to a new ceratosaur taxon, here named Fosterovenator churei, whereas the other bones represent previously unidentified juvenile taxa of basal tetanuran and coelurid theropods. The discovery and description of these fossil materials is significant because they provide important information about the Upper Jurassic terrestrial fauna of Quarry 12, Como Bluff, Wyoming. The presence of previously unidentified theropod taxa in the Morrison Formation indicates that the diversity of basal tetanuran and coelurid theropods may have been much greater than previously expected. Although the fossil material here described is largely fragmentary, it is tenable that theropods of different clades co-existed in the same ecosystems at the same time and most likely competed for the same food sources. -
T Riassic Jurassic Cretaceous
Millions of Years Ago 252.17 247.2 237.0 201.3 174.1 163.5 145.0 100.5 66.0 Avemetatarsalia Triassic Jurassic Cretaceous Early Middle Late Early Middle Late Early Late Scleromochlus Pterosauria Ornithodira Lagerpetidae Marasuchus Dinosauromorpha Genasauria Silesauridae Neornithsichia Nyasasaurus Dinosauriformes Thyreophora Ornithischia Eocursor ? Heterodontosauridae Pisanosaurus Dinosauria Sauropodomorpha Herrerasauria Saurischia Eodromaeus Theropoda Daemonosaurus Tawa Neotheropoda Millions of Years Ago 252.17 247.2 237.0 201.3 174.1 163.5 145.0 100.5 66.0 Triassic Jurassic Cretaceous Early Middle Late Early Middle Late Early Late Cerapoda Ornithopoda Marginocephalia Eocursor Parksosauridae Laquintasaura Kulindadromeus Othnielosaurus Jeholosauridae Neornithischia Leaellynasaura Hexinlusaurus Agilisaurus Stegosauridae Lesothosaurus Stegosaurinae Stegosauria Dacentrurinae Kentrosaurus Tuojiangosaurus Thyreophoroidea Struthiosaurinae Genasauria Huayangosauridae Eurypoda Gigantspinosaurus Panoplosaurinae Nodosauridae Sauropelta Polacanthinae Scelidosaurus Gargoyleosaurus Ankylosauridae Thyreophora Scutellosaurus Kunbarrasaurus Ankylosauria Hylaeosurus Shamosaurinae Ankylosaurinae Pinacosaurus Tianchiasaurus Ankylosaurini Millions of Years Ago 252.17 247.2 237.0 201.3 174.1 163.5 145.0 100.5 66.0 Triassic Jurassic Cretaceous Euhadrosauria Early Middle Late Early Middle Late Early Late Hadrosaurinae Hadrosauridae Hadrosauromorpha Lambeosaurinae Eotrachodon Tethyshadros Hadrosauriformes Hadrosauria Telmatosaurus Bactrosaurus Protohadros Styracosterna -
Zhao 1985 Trans.Rtf
Original Article- Zhao Xijin, 1985. The Jurassic Reptilia. In Wang Si-en, Cheng Zhengwu and Wang Neiwen (eds.). The Jurassic System of China. Stratigraphy of China. 11, 286-289, 347, plates 10 and 11. Translation by Leo W Sham, January 3, 2011. (6) Jurassic Reptilian Fauna The Jurassic period was a time of profuse development of reptiles, especially dinosaurs. For convenience of description, [we will] now separate our analysis into the following three stages. Early Jurassic The Early Jurassic was an early stage of dinosaur evolution. Dinosaurs and other Mesozoic reptiles then were largely primitive. Fossils are not only few but also relatively poorly preserved. These primitive bauplans dictate the paucity of difference between genera; it is more difficult to determine the identity of fossils compared to later times. The Early Jurassic was the starting stage of dinosaur evolution. Exposed strata were most developed [ sic ] in southwestern China, for example, Lower Lufeng Group (alas Fengjiahe Formation) of Yunnan, Lower Ziliujing Group of Sichuan, and Daye Group of Tibet. The large amount of dinosaurian and other reptilian fossils preserved in these strata provided strong evidence to determine the chronology these terrestrial strata. Here, we first analyze the fauna of the representative Lower Lufeng Group to demonstrate their primitiveness. Sauropods and prosauropods coexisted in the Lower Lufeng Group. In general, sauropods appeared at the beginning of the Jurassic; thus the existence of sauropod fossils in a stratum would point to its earliest Jurassic age because prosauropods, evolving since the Late Triassic, had much declined with lineages lingering into the Early Jurassic. This phenomenon is particular in Lufeng, Yunnan, where the prosauropod Lufengosaurus described by Professor CC Young coexisted with the primitive sauropod Dachongosaurus yunnanensis , which also showed certain advanced characteristics.