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A couple of partially-feathered creatures about the The Outside Story size of a turkey pop out of a stand of ferns. By the water you spot a flock of bigger animals, lean and predatory, catching fish. And then an even bigger pair of animals, each longer than a car, with ostentatious crests on their heads, stalk out of the heat haze. The fish-catchers dart aside, but the new pair have just come to drink. We can only speculate what a walk through Jurassic New England would be like, but the fossil record leaves many hints. According to Matthew Inabinett, one of the Beneski Museum of Natural History’s senior docents and a student of vertebrate paleontology, dinosaur footprints found in the sedimentary rock of the Connecticut Valley reveal much about these animals and their environment. At the time, the land that we know as New England was further south, close to where Cuba is now. A system of rift basins that cradled lakes ran right through our region, from North Carolina to Nova Scotia. As reliable sources of water, with plants for the herbivores and fish for the carnivores, the lakes would have been havens of life. While most of the fossil footprints found in New England so far are in the lower Connecticut Valley, Dinosaur Tracks they provide a window into a world that extended throughout the region. According to Inabinett, the By: Rachel Marie Sargent tracks generally fall into four groupings. He explained that these names are for the tracks, not Imagine taking a walk through a part of New the dinosaurs that made them, since, “it’s very England you’ve never seen—how it was 190 million difficult, if not impossible, to match a footprint to a years ago. -
Los Restos Directos De Dinosaurios Terópodos (Excluyendo Aves) En España
Canudo, J. I. y Ruiz-Omeñaca, J. I. 2003. Ciencias de la Tierra. Dinosaurios y otros reptiles mesozoicos de España, 26, 347-373. LOS RESTOS DIRECTOS DE DINOSAURIOS TERÓPODOS (EXCLUYENDO AVES) EN ESPAÑA CANUDO1, J. I. y RUIZ-OMEÑACA1,2 J. I. 1 Departamento de Ciencias de la Tierra (Área de Paleontología) y Museo Paleontológico. Universidad de Zaragoza. 50009 Zaragoza. [email protected] 2 Paleoymás, S. L. L. Nuestra Señora del Salz, 4, local, 50017 Zaragoza. [email protected] RESUMEN La mayoría de los restos fósiles de dinosaurios terópodos de España son dientes aislados y escasos restos postcraneales. La única excepción es el ornitomimosaurio Pelecanimimus polyodon, del Barremiense de Las Hoyas (Cuenca). Hay registro de terópodos en el Jurásico superior (Oxfordiense superior-Tithónico inferior), en el tránsito Jurásico-Cretácico (Tithónico superior- Berriasiense inferior) y en todos los pisos del Cretácico inferior, con excepción del Valanginiense. En el Cretácico superior únicamente hay restos en el Campaniense y Maastrichtiense. La mayor parte de las determinaciones son demasiado generales, lo que impide conocer algunas de las familias que posiblemente estén representadas. Se han reconocido: Neoceratosauria, Baryonychidae, Ornithomimosauria, Dromaeosauridae, además de terópodos indeterminados, y celurosaurios indeterminados (dientes pequeños sin dentículos). La mayoría de los restos son de Maniraptoriformes, siendo especialmente abundantes los dromeosáuridos. Las únicas excepciones son por el momento, el posible Ceratosauria del Jurásico superior de Asturias, los barionícidos del Hauteriviense-Barremiense de Burgos, Teruel y La Rioja, el posible carcharodontosáurido del Aptiense inferior de Morella y el posible abelisáurido del Campaniense de Laño. Además hay algunos terópodos incertae sedis, como los "paronicodóntidos" (entre los que se incluye Euronychodon), y Richardoestesia. -
Paleoherpetofauna Portuguesa
Rev. Esp. Herp. (2002): 17-35 17 Paleoherpetofauna Portuguesa E.G. CRESPO Centro de Biologia Ambiental – Fac. Ciências Univ. Lisboa Resumo: Nos últimos anos a importância da paleoherpetofauna portuguesa tem sido posta em evidência sobre- tudo através do seu grupo mais mediático, os dinossauros. As recentes descobertas em Portugal de vestígios de vários dinossauros, incluindo ossos, ovos, embriões, gastrólitos e pegadas, têm merecido ampla cobertura jorna- lística e têm sido oportunamente acompanhadas por intensas campanhas de divulgação, levadas a cabo pelo Mu- seu Nacional de História Natural de Lisboa, encabeçadas pelo geólogo, Professor Galopim de Carvalho. As pro- longadas e por vezes polémicas acções de sensibilização pública e política que foi necessário empreender para se preservarem muitos dos locais onde esses vestígios foram encontrados, contribuiram também para sustentar e até aumentar o interesse por este grupo de grandes répteis. A importância da paleoherpetofauna portuguesa está porém longe de se limitar apenas aos dinossauros! Em Portugal viveram muitos outros répteis e anfíbios de que existem vestígios desde o começo do Mesozói- co –Quelónios, Crocodilos, Ictiossauros, Plesiossauros, Pterossauros, Lepidossauros, “Estegossauros” e Lis- samphia– que, embora geralmente muito menos conhecidos, têm um significado evolutivo, paleogeográfico e paleoclimático extremamente importante. Na sua descoberta e estudo estiveram envolvidos, já desde o século passado, numerosos investigadores por- tugueses e estrangeiros, dos quais se destacam, entre outros, Georges Zbyszewski, Miguel Telles Antunes, Vei- ga Ferreira, H. Sauvage, A.F. Lapparent, L. Ginsburg, R.Thulborn, P. Galton. Muitos destes estudos encontram- se todavia dispersos por uma vasta gama de publicações em que, frequentemente, as referências aos répteis e aos anfíbios ou são laterais ou são apresentadas em contextos zoológicos mais abrangentes, pelo que, como parece que tem acontecido, têm passado praticamente despercebidos à maioria daqueles que se dedicam aos estudo da nossa herpetofauna actual. -
Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia. -
Coossified Tarsometatarsi in Theropod Dinosaurs and Their Bearing on the Problem of Bird Origins
HALSZKA OSM6LSKA COOSSIFIED TARSOMETATARSI IN THEROPOD DINOSAURS AND THEIR BEARING ON THE PROBLEM OF BIRD ORIGINS OSM6LSKA, H. : Coossified tarsometatarsi in theropod dinosaurs and their bearing on the problem of bird origins, Palaeontologia Polonica, 42, 79-95, 1981. Limb remains of two small theropod dinosaurs from the Upper Cretaceous deposits of Mongolia display fused tarsometatarsi. Presence of fusion in the tarsometatarsus in some theropods is consi dered as additional evidence for the theropod origin of birds. E/misaurus rarus gen. et sp. n. is described based upon a fragmentary skeleton represented by limbs. Family Elmisauridae novo is erected to include Elmisaurus, Chirostenotes GlLMORE and Ma crophalangia STERNBERG. Key words: Dinosauria, Theropoda, bird origins, Upper Cretaceous, Mongolia. Halszka Osmolska , ZakladPaleobiologii, Polska Akademia Nauk, Al. Zw irki i Wigury 93,02-089 War szawa, Po/and. Received: June 1979. Streszczenie. - W pracy opisano szczatki malych dinozaur6w drapieznych z osad6w gornokredo wych Mongolii . Stopa tych dinozaur6w wykazuje obecnosc zrosnietego tarsomet atarsusa. Zrosniecie to stanowi dodatkowy dow6d na pochodzenie ptak6w od dinozaur6w drapieznych, Opisano nowy rodzaj i gatunek dinozaura drapieznego E/misaurus rarus, kt6ry zaliczono do nowej rodziny Elmisau ridae . Do rodziny tej, opr6cz Elmisaurus, naleza: Chirostenotes GILMORE i Macr opha/angia STERNBERG. Praca byla finansowana przez Polska Akademie Nauk w ramach problemu rniedzyresorto wego MR 11-6. INTRODUCTION During the Polish-Mongolian -
The Distinctive Theropod Assemblage of the Ellisdale Site of New Jersey and Its Implications for North American Dinosaur Ecology and Evolution During the Cretaceous
Journal of Paleontology, 92(6), 2018, p. 1115–1129 Copyright © 2018, The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/15/0088-0906 doi: 10.1017/jpa.2018.42 The distinctive theropod assemblage of the Ellisdale site of New Jersey and its implications for North American dinosaur ecology and evolution during the Cretaceous Chase D. Brownstein Stamford Museum and Nature Center, Stamford CT 〈[email protected]〉 Abstract.—The Cretaceous landmass of Appalachia has preserved an understudied but nevertheless important record of dinosaurs that has recently come under some attention. In the past few years, the vertebrate faunas of several Appalachian sites have been described. One such locality, the Ellisdale site of the Cretaceous Marshalltown Forma- tion of New Jersey, has produced hundreds of remains assignable to dinosaurs, including those of hadrosauroids of several size classes, indeterminate ornithopods, indeterminate theropods, the teeth, cranial, and appendicular elements of dromaeosaurids, ornithomimosaurians, and tyrannosauroids, and an extensive microvertebrate assemblage. The theropod dinosaur record of the Ellisdale site is currently the most extensive and diverse known from the Campanian of Appalachia. Study of the Ellisdale theropod specimens suggests that at -
104Ornithodiraphyl
Millions of Years Ago 252.3 247.2 235.0 201.5 175.6 161.2 145.5 99.6 65.5 Triassic Jurassic Cretaceous Early Middle Late Early Middle Late Early Late Euparkeria Crurotarsi ? Scleromochlus ? Archosauria Pterosauria Lagerpetidae Ornithodira Marasuchus Genasauria Dinosauromorpha Silesauridae Neornithsichia Thyreophora Ornithischia Eocursor (esp. Dinosauria) et al. (2011), Yates (2007) Yates et al. (2011), Nesbitt etal.(2009), Sues (2007), Martinezet al.(2011), Irmis etal. Ezcurra (2006), EzcurraandBrusatte (2011), Phylogeny after Brusatteetal.(2010), Butleretal.(2007), Heterodontosauridae Pisanosaurus Dinosauria Ornithodira Sauropodomorpha Herrerasauria Saurischia Eodromeus Theropoda Daemonosaurus Tawa Neotheropoda Millions of Years Ago 253.0 247.2 235.0 201.5 175.6 161.2 145.5 99.6 65.5 Triassic Jurassic Cretaceous Early Middle Late Early Middle Late Early Late (2009), Norman et al. (2004), Thompson etal. (2011) (2009), Norman etal. (2004), Phylogeny afterButler etal. (2007a,b), Carpenter (2001),Galton &Upchurch (2004), Maidment etal.(2008), Mateus etal. Cerapoda Ornithopoda Eocursor Marginocephalia Neornithischia Othnielosaurus Genasauria (esp. Thyreophora) Genasauria (esp. Hexinlusaurus Stormbergia Genasauria Lesothosaurus Scutellosaurus Thyreophora Scelidosaurus Stegosauridae Stegosaurinae Dacentrurinae Stegosauria Kentrosaurus Tuojiangosaurus Huayangosauridae Gigantspinosaurus Eurypoda Tianchiasaurus Ankylosauria Nodosauridae Ankylosauridae Millions of Years Ago 253.0 247.2 235.0 201.5 175.6 161.2 145.5 99.6 65.5 Triassic Jurassic Cretaceous -
) Baieiicanjizseum
)SovitatesbAieiicanJizseum PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK 24, N.Y. NUMBER 1901 JULY 22, 1958 Coelurosaur Bone Casts from the Con- necticut Valley Triassic BY EDWIN HARRIS COLBERT1 AND DONALD BAIRD2 INTRODUCTION An additional record of a coelurosaurian dinosaur in the uppermost Triassic of the Connecticut River Valley is provided by a block of sandstone bearing the natural casts of a pubis, tibia, and ribs. This specimen, collected nearly a century ago but hitherto unstudied, was brought to light by the junior author among the collections (at present in dead storage) of the Boston Society of Natural History. We are much indebted to Mr. Bradford Washburn and Mr. Chan W. Wald- ron, Jr., of the Boston Museum of Science for their assistance in mak- ing this material available for study. The source and history of this block of stone are revealed in brief notices published at the time of its discovery. The Proceedings of the Boston Society of Natural History (vol. 10, p. 42) record that on June 1, 1864, Prof. William B. Rogers "presented an original cast in sand- stone of bones from the Mesozoic rocks of Middlebury, Ct. The stone was probably the same as that used in the construction of the Society's Museum; it was found at Newport among the stones used in the erec- tion of Fort Adams, and he owed his possession of it to the kindness of Capt. Cullum." S. H. Scudder, custodian of the museum, listed the 1 Curator of Fossil Reptiles and Amphibians, the American Museum of Natural History. -
Back Matter (PDF)
Index Note: Page numbers in italic denote figures. Page numbers in bold denote tables. Abel, Othenio (1875–1946) Ashmolean Museum, Oxford, Robert Plot 7 arboreal theory 244 Astrodon 363, 365 Geschichte und Methode der Rekonstruktion... Atlantosaurus 365, 366 (1925) 328–329, 330 Augusta, Josef (1903–1968) 222–223, 331 Action comic 343 Aulocetus sammarinensis 80 Actualism, work of Capellini 82, 87 Azara, Don Felix de (1746–1821) 34, 40–41 Aepisaurus 363 Azhdarchidae 318, 319 Agassiz, Louis (1807–1873) 80, 81 Azhdarcho 319 Agustinia 380 Alexander, Annie Montague (1867–1950) 142–143, 143, Bakker, Robert. T. 145, 146 ‘dinosaur renaissance’ 375–376, 377 Alf, Karen (1954–2000), illustrator 139–140 Dinosaurian monophyly 93, 246 Algoasaurus 365 influence on graphic art 335, 343, 350 Allosaurus, digits 267, 271, 273 Bara Simla, dinosaur discoveries 164, 166–169 Allosaurus fragilis 85 Baryonyx walkeri Altispinax, pneumaticity 230–231 relation to Spinosaurus 175, 177–178, 178, 181, 183 Alum Shale Member, Parapsicephalus purdoni 195 work of Charig 94, 95, 102, 103 Amargasaurus 380 Beasley, Henry Charles (1836–1919) Amphicoelias 365, 366, 368, 370 Chirotherium 214–215, 219 amphisbaenians, work of Charig 95 environment 219–220 anatomy, comparative 23 Beaux, E. Cecilia (1855–1942), illustrator 138, 139, 146 Andrews, Roy Chapman (1884–1960) 69, 122 Becklespinax altispinax, pneumaticity 230–231, Andrews, Yvette 122 232, 363 Anning, Joseph (1796–1849) 14 belemnites, Oxford Clay Formation, Peterborough Anning, Mary (1799–1847) 24, 25, 113–116, 114, brick pits 53 145, 146, 147, 288 Benett, Etheldred (1776–1845) 117, 146 Dimorphodon macronyx 14, 115, 294 Bhattacharji, Durgansankar 166 Hawker’s ‘Crocodile’ 14 Birch, Lt. -
Limb Design, Function and Running Performance in Ostrich-Mimics and Tyrannosaurs
GAIA Nº 15, LISBOA/LISBON, DEZEMBRO/DECEMBER 1998, pp. 257-270 (ISSN: 0871-5424) LIMB DESIGN, FUNCTION AND RUNNING PERFORMANCE IN OSTRICH-MIMICS AND TYRANNOSAURS Gregory S. PAUL 3109 N Calvert St. Side Apt., BALTIMORE MD 21218. USA ABSTRACT: Examination of the limb morphology of small ornithomimids and large tyranno- saurids shows that they remained remarkably constant in design regardless of size. The changes that were present were consistent with maintaining limb strength and function constant with size. It is concluded that ornithomimid and tyrannosaurid legs functioned in a similar manner, and always exhibit the features normally associated with a fast running gait. This is in contrast to modern animals, in which elephants as gigantic as large tyranno- saurids have limbs that are modified for a slow walking gait. INTRODUCTION to run observed in elephants. The hypothesis can be challenged if it can be shown that at least some ex- Because they had long, gracile, bird-like legs, it tinct giants retained the skeletal adaptations for run- has long been accepted that the smaller predatory ning observed in smaller species, and in their own dinosaurs were swift runners (O [& GREG- SBORN offspring. In turn, the hypothesis that giants can run ], 1916; COLBERT, 1961; RUSSELL, 1972; ORY fast if they retain limbs similar to smaller runners can C , 1978; THULBORN, 1982; PAUL, 1988; OOMBS be challenged if it is shown that the skeleton is too H , 1994). Much more controversial has been OLTZ vulnerable to structural failure. the locomotory abilities of their giant relatives, which have been restored as no faster than elephants BODY MASSES (LAMBE, 1917; HALSTEAD &HALSTEAD, 1981; THUL- BORN, 1982; BARSBOLD, 1983), able to run at only Sources for mass data for extinct and extant ani- modest speeds (COOMBS, 1978; MOLNAR &FAR- mals include PAUL (1988, 1997), NOWAK (1991) and LOW, 1990; HORNER &LESSEM, 1993; FARLOW, MATTHEWS (1994). -
New Tyrannosaur from the Mid-Cretaceous of Uzbekistan Clarifies Evolution of Giant Body Sizes and Advanced Senses in Tyrant Dinosaurs
New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs Stephen L. Brusattea,1, Alexander Averianovb,c, Hans-Dieter Suesd, Amy Muira, and Ian B. Butlera aSchool of GeoSciences, University of Edinburgh, Edinburgh EH9 3FE, United Kingdom; bZoological Institute, Russian Academy of Sciences, St. Petersburg 199034, Russia; cDepartment of Sedimentary Geology, Saint Petersburg State University, St. Petersburg 199178, Russia; and dDepartment of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560 Edited by Neil H. Shubin, The University of Chicago, Chicago, IL, and approved January 29, 2016 (received for review January 5, 2016) Tyrannosaurids—the familiar group of carnivorous dinosaurs in- We here report the first diagnostic tyrannosauroid from the mid- cluding Tyrannosaurus and Albertosaurus—were the apex predators Cretaceous, a new species from the Turonian (ca. 90–92 million in continental ecosystems in Asia and North America during the years ago) Bissekty Formation of Uzbekistan. This formation has latest Cretaceous (ca. 80–66 million years ago). Their colossal sizes recently emerged as one of the most important records of mid- and keen senses are considered key to their evolutionary and eco- Cretaceous dinosaurs globally (9–11). Possible tyrannosauroid logical success, but little is known about how these features devel- specimens from the Bissekty Formation were reported more than oped as tyrannosaurids evolved from smaller basal tyrannosauroids a half century ago (12), and, more recently, several isolated fossils that first appeared in the fossil record in the Middle Jurassic (ca. 170 were assigned to the group (9, 13), but none of these has been million years ago). -
Body-Size Evolution in the Dinosauria
8 Body-Size Evolution in the Dinosauria Matthew T. Carrano Introduction The evolution of body size and its influence on organismal biology have received scientific attention since the earliest decades of evolutionary study (e.g., Cope, 1887, 1896; Thompson, 1917). Both paleontologists and neontologists have attempted to determine correlations between body size and numerous aspects of life history, with the ultimate goal of docu- menting both the predictive and causal connections involved (LaBarbera, 1986, 1989). These studies have generated an appreciation for the thor- oughgoing interrelationships between body size and nearly every sig- nificant facet of organismal biology, including metabolism (Lindstedt & Calder, 1981; Schmidt-Nielsen, 1984; McNab, 1989), population ecology (Damuth, 1981; Juanes, 1986; Gittleman & Purvis, 1998), locomotion (Mc- Mahon, 1975; Biewener, 1989; Alexander, 1996), and reproduction (Alex- ander, 1996). An enduring focus of these studies has been Cope’s Rule, the notion that body size tends to increase over time within lineages (Kurtén, 1953; Stanley, 1973; Polly, 1998). Such an observation has been made regarding many different clades but has been examined specifically in only a few (MacFadden, 1986; Arnold et al., 1995; Jablonski, 1996, 1997; Trammer & Kaim, 1997, 1999; Alroy, 1998). The discordant results of such analyses have underscored two points: (1) Cope’s Rule does not apply universally to all groups; and (2) even when present, size increases in different clades may reflect very different underlying processes. Thus, the question, “does Cope’s Rule exist?” is better parsed into two questions: “to which groups does Cope’s Rule apply?” and “what process is responsible for it in each?” Several recent works (McShea, 1994, 2000; Jablonski, 1997; Alroy, 1998, 2000a, 2000b) have begun to address these more specific questions, attempting to quantify patterns of body-size evolution in a phylogenetic (rather than strictly temporal) context, as well as developing methods for interpreting the resultant patterns.