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Home , Adoretus, Anomalini, Aphodiinae, Ateuchus, Coprini, Dichotomius

and Stag of Hawaii and the Pacific* Checklist: of Hawaii**

Family Subfamily Tribe Author Status Apterocyclus 1 Lucanidae Lucaninae Waterhouse, 1871 native honoluluensis Paulson & Hawks, 2 Lucanidae Lucaninae Apterocyclus kawaii native 2014 Apterocyclus native, possibly 3 Lucanidae Lucaninae Sharp, 1908 munroi extirpated Apterocyclus native, possibly 4 Lucanidae Lucaninae Van Dyke, 1921 palmatus extirpated Apterocyclus 5 Lucanidae Lucaninae Sharp, 1908 native waterhousei 6 (Linnaeus, 1758) intentionally imported fimetarius

7 Scarabaeidae Aphodiinae Aphodiini Aphodius lividus (Olivier, 1789) adventive

8 Scarabaeidae Aphodiinae cognatus (Le Conte, 1859) adventive

9 Scarabaeidae Aphodiinae Eupariini Ataenius pacificus (Sharp, 1879) adventive

Ataenius 10 Scarabaeidae Aphodiinae Eupariini Harold, 1877 adventive peregrinator Ataenius 11 Scarabaeidae Aphodiinae Eupariini (Le Conte, 1859) adventive puncticollis Saprosites 12 Scarabaeidae Aphodiinae Eupariini Harold, 1877 adventive pygmaeus 13 Scarabaeidae Aphodiinae (Rakovic 1981) adventive hawaiiensis 14 Scarabaeidae Aphodiinae Psammodiini (Bates, 1887) adventive micros Pleurophorus 15 Scarabaeidae Aphodiinae Psammodiini (Chevrolat, 1864) adventive parvulus

16 Scarabaeidae Cetoniinae fusca (Herbst, 1790) adventive

(Gory & Percheron, 17 Scarabaeidae Cetoniinae Cetoniini Protaetia orientalis adventive 1833)

18 Scarabaeidae Cetoniinae Cetoniini Protaetia pryeri (Janson, 1888) adventive

(Gory and 19 Scarabaeidae Cetoniinae mutabilis quarantine Percheron, 1833) 20 Scarabaeidae Casey, 1915 adventive pasadenae Trypoxylus 21 Scarabaeidae Dynastinae Dynastini (Linnaeus, 1767) intentionally imported? dichotomus

22 Scarabaeidae Dynastinae Dynastini gideon? (Linnaeus, 1771) quarantine

23 Scarabaeidae Dynastinae Oryctini rhinoceros (Linnaeus, 1758) adventive

24 Scarabaeidae Dynastinae Ligyrus gibbosus (De Geer, 1774) quarantine

Temnorhynchus 25 Scarabaeidae Dynastinae Pentodontini Fabricius, 1781 adventive retusus 26 Scarabaeidae (Say, 1825) quarantine ephilida adventive (not 27 Scarabaeidae Melolonthinae castanea (Arrow, 1913) established)

28 Scarabaeidae Melolonthinae Sericini (Motschulsky, 1860) quarantine

Adoretus 29 Scarabaeidae Adoretini (Weber, 1801) adventive compressus

30 Scarabaeidae Rutelinae Adoretini sinicus Burmeister, 1855 adventive

31 Scarabaeidae Rutelinae albopilosa (Hope, 1839) quarantine

32 Scarabaeidae Rutelinae Anomalini Anomala cuprea (Hope, 1839) quarantine

33 Scarabaeidae Rutelinae Anomalini Anomala orientalis (Waterhouse, 1875) adventive

34 Scarabaeidae Rutelinae Anomalini Anomala sulcatula Burmeister, 1844 adventive, extirpated?

35 Scarabaeidae Rutelinae Anomalini Anomala viridana (Kolbe, 1886) quarantine

36 Scarabaeidae Rutelinae Anomalini japonica Newman, 1838 quarantine

37 Scarabaeidae Rutelinae Anomalini Popillia lewisi Arrow, 1913 quarantine

38 Scarabaeidae lecontei (Harold, 1874) intentionally imported

39 Scarabaeidae Scarabaeinae Canthonini humectus (Say, 1832) intentionally imported

Canthon intentionally imported, not 40 Scarabaeidae Scarabaeinae Canthonini Harold, 1868 indigaceus established

41 Scarabaeidae Scarabaeinae Canthonini (Linnaeus, 1758) intentionally imported

(Palisot de intentionally imported, not 42 Scarabaeidae Scarabaeinae Canthonini Beauvois, 1805) established intentionally imported, not 43 Scarabaeidae Scarabaeinae (Linnaeus, 1758) molossus established

44 Scarabaeidae Scarabaeinae Coprini incertus Say, 1835 intentionally imported

intentionally imported, not 45 Scarabaeidae Scarabaeinae Coprini Copris remotus Le Conte, 1866 established intentionally imported, not 46 Scarabaeidae Scarabaeinae Coprini (Linnaeus, 1767) carolinus established 47 Scarabaeidae Scarabaeinae Coprini (Harold, 1873) intentionally imported africanus Euoniticellus 48 Scarabaeidae Scarabaeinae Coprini (Reiche, 1847) intentionally imported intermedius

49 Scarabaeidae Scarabaeinae cinctus (Fabricius, 1775) intentionally imported?

(Laporte Comte de 50 Scarabaeidae Scarabaeinae Oniticellini Oniticellus militaris intentionally imported Castelau, 1840)

51 Scarabaeidae Scarabaeinae Onitini alexis Klug, 1835 intentionally imported

intentionally imported, not 52 Scarabaeidae Scarabaeinae Onitini Onitis phartopus Lansberge, 1875 established

53 Scarabaeidae Scarabaeinae Onitini Onitis vanderkelleni Lansberge, 1886 intentionally imported

Digitonthophagus 54 Scarabaeidae Scarabaeinae (Fabricius, 1787) intentionally imported gazella 55 Scarabaeidae Scarabaeinae Onthophagini (Thunberg, 1818) intentionally imported binodis Onthophagus intentionally imported, not 56 Scarabaeidae Scarabaeinae Onthophagini Blackburn, 1903 comperei established

Onthophagus intentionally imported, not 57 Scarabaeidae Scarabaeinae Onthophagini MacLeay, 1866 cuniculus established Onthophagus intentionally imported, not 58 Scarabaeidae Scarabaeinae Onthophagini Lansberge, 1886 foliaceus established Onthophagus intentionally imported, not 59 Scarabaeidae Scarabaeinae Onthophagini Boheman, 1858 granulatus established Onthophagus 60 Scarabaeidae Scarabaeinae Onthophagini Say, 1835 intentionally imported incensus Onthophagus intentionally imported, not 61 Scarabaeidae Scarabaeinae Onthophagini MacLeay, 1866 laminatus established Onthophagus 62 Scarabaeidae Scarabaeinae Onthophagini D'Orbigny, 1905 intentionally imported nigriventris Onthophagus intentionally imported, not 63 Scarabaeidae Scarabaeinae Onthophagini (Linnaeus, 1758) nuchicornis established Onthophagus 64 Scarabaeidae Scarabaeinae Onthophagini Brown, 1927 intentionally imported oklahomesis Onthophagus 65 Scarabaeidae Scarabaeinae Onthophagini (Fabricius, 1775) intentionally imported sagittarius Onthophagus 66 Scarabaeidae Scarabaeinae Onthophagini Harold, 1871 intentionally imported tuberculifrons intentionally imported, not 67 Scarabaeidae Scarabaeinae Phanaeini daphnis Harold, 1863 established Neosisyphus intentionally imported, not 68 Scarabaeidae Scarabaeinae Sisyphini Thunberg, 1818 spinipes established

69 Omorgus procerus (Harold, 1872) adventive

Omorgus intentionally imported, not 70 Trogidae (Fabricius, 1775) suberosus established

71 Trogidae (Linnaeus, 1767) adventive

*Dunlap, Joshua B., Mary Liz Jameson, Paul E. Skelley, Emmy L. Engasser, Amanda J. Redford. 2015. Scarab and Stag Beetles of Hawaii and the Pacific. USDA APHIS Identification Technology Program (ITP). Fort Collins, CO. Checklist last updated February, 2016

**Checklist last updated February, 2016