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Insect Fauna of Korea Insect Fauna of Korea Fauna Insect Insect Fauna of Korea Volume 12, Number 1 Arthropoda: Insecta: Coleoptera: Scarabaeoidea Pleurosticti Vol. 12, Vol. No. 1 Pleurosticti Flora and Fauna of Korea National Institute of Biological Resources Ministry of Environment National Institute of Biological Resources NIBR Ministry of Environment Russia CB Chungcheongbuk-do CN Chungcheongnam-do HB GB Gyeongsangbuk-do China GG Gyeonggi-do YG GN Gyeongsangnam-do GW Gangwon-do HB Hamgyeongbuk-do JG HN Hamgyeongnam-do HWB Hwanghaebuk-do HN HWN Hwanghaenam-do PB JB Jeollabuk-do JG Jagang-do JJ Jeju-do JN Jeollanam-do PN PB Pyeonganbuk-do PN Pyeongannam-do YG Yanggang-do HWB HWN GW East Sea GG GB (Ulleung-do) Yellow Sea CB CN GB JB GN JN JJ South Sea Insect Fauna of Korea Volume 12, Number 1 Arthropoda: Insecta: Coleoptera: Scarabaeoidea Pleurosticti 2011 National Institute of Biological Resources Ministry of Environment Insect Fauna of Korea Volume 12, Number 1 Arthropoda: Insecta: Coleoptera: Scarabaeoidea Pleurosticti Jin Ill Kim Sungshin Women’s University Copyright ⓒ 2011 by the National Institute of Biological Resources Published by the National Institute of Biological Resources Environmental Research Complex, Gyeongseo-dong, Seo-gu Incheon 404-708, Republic of Korea www.nibr.go.kr All rights reserved. No part of this book may be reproduced, stored in a retrieval system, or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior permission of the National Institute of Biological Resources. ISBN : 9788994555447-96470 Government Publications Registration Number 11-1480592-000145-01 Printed by Junghaengsa, Inc. in Korea on acid-free paper Publisher : Chong-chun Kim Project Staff : Hong-Yul Seo, Ye Eun, Joo-Lae Cho Published on February 28, 2011 The Flora and Fauna of Korea logo was designed to represent six major target groups of the project including vertebrates, invertebrates, insects, algae, fungi, and bacteria. The book cover and the logo were designed by Jee-Yeon Koo. Preface Biological resources are important elements encompassing organisms, genetic resources, and parts of organisms which provide potential values essential for human lives. The creation of high-valued products such as new varieties of organisms, new substances, and the development of new drugs by harnessing biological resources is now widely perceived to be one of the major indices of national competitiveness. In the wake of the “Convention of Biological Diversity”, which was adopted in 1992 in recognition of national sovereignty over indigenous biological and genetic resources, all the countries of the world are now concerting their efforts on the discovery of original materials for the bio-industry, initiating international competition in the 21st century. Competition among countries for biological resources is now entering an intense phase following the adoption of the ABS (Access to genetic resources and Benefit-Sharing) international regime in Nagoya in 2010. For this reason, the National Institute of Biological Resources of the Korean Ministry of Environment recognizes the preservation and management of the biological resources in Korea for the bio-industry as a first priority project for the future, and has begun publication of Flora and Fauna of Korea for the systematic preservation and efficient management of our biological resources. Korea has been acclaimed as a country with a high level of biological diversity, the total number of described species in Korea to date being about 37,000. Beginning in 2006, the National Institute of Biological Resources embarked on the publication of Flora and Fauna of Korea which, containing comprehensive and diverse information on our invaluable native species, has become the standard textbook of native species. The systematic survey of diverse taxa in all parts of Korea led by a group of professionals in the field of taxonomy over the past four years has finally come to fruition and culminated in the appearance of 16 monographs in the 2010 volumes of Flora and Fauna of Korea encompassing 1,037 species in 158 families belonging to 9 phyla, along with further volumes of Flora and Fauna of Korea encompassing 1,163 species in 112 families belonging to 7 phyla due to appear this year. This is the first volume of Flora and Fauna of Korea in which a taxon of organisms of the Korean Peninsula is extensively treated at the level of species. Flora and Fauna of Korea will contribute to raising the standard of Korean taxonomy and improve pride in the management of our biological resources through enhanced understanding of the true nature of our native species. In addition, I am confident that the ongoing publication of Flora and Fauna of Korea will significantly contribute to paving the way for sustainable, wise use of biological resources. I would like to express my sincerest gratitude to Professor Jin Ill Kim, Sungshin Women’s Univer- sity, who is responsible for writing this publication of Flora and Fauna of Korea. This series will play a pivotal role in the census of native Korean species, which are estimated to number 100,000. By promoting innovative and taxonomic research for the identification of the totality of native Korean species and by continuously publishing such results in Flora and Fauna of Korea, I sincerely hope that a valuable foundation will be laid for the sustainable use of our national biological resources through the extensive research, development and for their profitable use by a prosperous bio-industry in the creation of high-valued products such as natural products, medicines, cosmetics and essential supple- ments in our country. Chong-chun Kim, Ph. D. President NIBR 1 Contents List of Taxa 3 Introduction 8 Materials and Methods 10 Taxonomic Notes 25 Family Cetoniidae Leach 27 1. Valgus Scriba 29 2. Nipponovalgus Sawada 30 3. Osmoderma Le Peletier and Serville 32 4. Gnorimus Le Peletier and Serville 34 5. Trichius Fabricius 35 6. Dicronocephalus Hope 39 7. Rhomborhina Hope 41 8. Pseudotorrynorrhina Miksi´ˇ c 41 9. Cetonia Fabricius 44 10. Protaetia Burmeister 46 11. Anthracophora Burmeister 53 12. Glycyphana Burmeister 55 13. Gametis Burmeister 56 14. Clinterocera Motschulsky 59 Family Rutelidae MacLeay 61 15. Adoretus Dejean 62 16. Parastasia Westwood 66 17. Popillia Dejean 68 18. Proagopertha Reitter 76 19. Phyllopertha Stephens 77 20. Mimela Kirby 80 21. Callistethus Burmeister 88 22. Blitopertha Reitter 90 23. Anomala Samuelle 95 24. Bifurcanomala Kim 113 25. Chejuanomala Kim 114 Family Dynastidae MacLeay 115 26. Allomyrina Arrow 116 27. Eophileurus Arrow 118 28. Pentodon Hope 120 Family Melolonthidae Leach 121 29. Ectinohoplia Redtenbacher 123 30. Hoplia Illiger 125 2 Insect Fauna of Korea·Pleurosticti 31. Apogonia Kirby 130 32. Brahmina Blanchardt 133 33. Lasiopsis Erichson 136 34. Pseudosymmchia Dalla Torre 138 35. Miridiba Reitter 140 36. Sophrops Fairmaire 141 37. Holotrichia Hope 144 38. Heptophylla Motschulsky 156 39. Hilyotrogus Fairmaire 157 40. Polyphylla Harris 160 41. Melolontha Fabricius 161 42. Gastroserica Brenske 166 43. Paraserica Reitter 167 44. Maladera Mulsant 168 45. Sericania Motschulsky 188 46. Nipponoserica Nomura 193 47. Serica MacLeay 196 Literature Cited 202 Plates 214 Index to Korean Names 244 Index to Korean Names as Pronounced 247 Index to Scientific Names 250 3 List of Taxa Class Insecta Order Coleoptera Scarabaeoidea Latreille, 1802 Pleurosticti Family Cetoniidae Leach, 1815 Subfamily Valginae Mulsant, 1842 Genus Valgus Scriba, 1790 Valgus koreanus Sawada, 1944 Genus Nipponovalgus Sawada, 1941 Nipponovalgus angusticollis (Waterhouse), 1875 Subfamily Trichiinae Fleming, 1821 Genus Osmoderma Le Peletier de Saint-Fargeau and Serville, 1828 Osmoderma opicum Lewis, 1887 Genus Gnorimus Le Peletier de Saint-Fargeau and Serville, 1828 Gnorimus subopacus Motschulsky, 1860 Genus Lasiotrichius Reitter, 1899 Lasiotrichius succinctus (Pallas), 1781 Subfamily Cetoniinae Leach, 1815 Tribe Phaedimini Schoch, 1894 Genus Dicranocephalus Hope, 1831 Dicronocephalus adamsi Pascoe, 1863 Tribe Coryphocerina Burmeister, 1842 Genus Rhomborrhina Hope, 1837 Rhomborrhina polita Waterhouse, 1875 Genus Pseudotorynorrhina Miksi´ˇ c, 1974 Pseudotorynorrhina japonica (Hope), 1841 Tribe Cetoniini Leach, 1815 Genus Cetonia Fabricius, 1775 Cetonia pilifera (Motschulsky), 1860 Cetonia pilifera magnifica (Ballion) Genus Protaetia Burmeister, 1842 Protaetia lugubris (Herbst), 1786 Protaetia brevitarsis seulensis (Kolbe), 1886 Protaetia famelica scheini (Miksi´ˇ c), 1959 Protaetia mandschuriensis (Schürhoff), 1933 Protaetia orientalis submarmorea (Burmeister), 1842 Genus Anthracophora Burmeister, 1842 Anthracophora rusticola Burmister, 1842 Genus Glycyphana Burmeister, 1842 Glycyphana fulvistemma Motschulsky, 1860 Genus Gametis Burmeister, 1842 Gametis jucunda (Faldermann), 1935 4 Insect Fauna of Korea·Pleurosticti Tribe Cremastocheilini Burmeister and Schaum, 1841 Genus Clinterocera Motschulsky, 1858 Clinterocera obsoleta (Fairmaire), 1878 Family Rutelidae MacLeay, 1819 Tribe Adoretini Burmeister, 1844 Genus Adoretus Dejean, 1833 Adoretus tenuimaculatus Waterhouse, 1875 Adoretus hirsutus Ohaus, 1914 Tribe Rutelini MacLeay, 1819 Genus Parastasia Westwood, 1842 Parastasia ferrieri Nonfried, 1895 Tribe Anomalini Streubel, 1839 Subtribe Popillina Ohaus, 1918 Genus Popillia Dejean, 1821 Popillia flavosellata Fairmaire, 1886 Popillia mutans Newman, 1838 Popillia quadriguttata (Fabricius), 1787 Popillia quadriguttata japonica Newmann Popillia formosana
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