Turk J Bot 26 (2002) 453-475 © TÜB‹TAK Research Article The Flora of E¤erli Mountain (Amasya-Turkey)
Arzu CANSARAN Ondokuz May›s University, Faculty of Education Dept. of Sci., Amasya - TURKEY
Received: 30.10.2000 Accepted: 26.04.2002
Abstract: The floristical characteristics of E¤erli Mountain (E¤erli Da¤›) are given and the vascular plant species growing there are documented. The research was carried out between 1996 and 1998. Approximately 2000 plant specimens were collected from the area. At the end of the study, 650 taxa (at specific and infraspecific ranks) belonging to 70 families and 298 genera were identified. Fifty-five taxa were defined as new records for the A5 square. The largest family was found to be Asteraceae (78 species) and the second largest one was Fabaceae (77 species). The largest genera were Astragalus L. (23 species), Silene L. and Lathyrus L. (10 species each). The phytogeographic elements are represented in the study area as follows: Irano-Turanian 102 (15.69%), Euro- Siberian 97 (14.92%), Mediterranean 46 (7.07%), and multi-regional or of unknown phytogeographic origin 375 (57.69%). Eighty (12.30%) species collected in the area are endemics. Key Words: Vascular plants, Flora, E¤erli Mountain, Amasya
E¤erli Da¤›’n›n (Amasya-Türkiye) Floras›
Özet: Bu araflt›rmada, E¤erli Da¤› ve çevresinin floristik özellikleri ve burada yetiflti¤i saptanan damarl› bitki taksonlar›n›n listes i verilmifltir. Araflt›rma 1996-1998 y›llar› aras›nda yap›lm›flt›r. Yöreden toplanan 2000 bitki örne¤inin de¤erlendirilmesi ile 70 familyadan 298 cins ve 650 tür ve türalt› seviyede takson saptanm›flt›r. Bu taksonlardan 55’i A5 karesi için yeni kay›t olarak belirlenmifltir. En büyük familya Asteraceae (78 tür), ikinci Fabaceae (77 tür)’dir. En büyük cinsler ise 23 türle Astragalus L. ve 10’ar türle Silene L. ve Lathyrus L.’dur. Araflt›rma alan›nda fitoco¤rafik elementlerin da¤›l›m› ise flöyledir; ‹ran-Turan 102 (%15.69), Avrupa-Sibirya 97 (%14.92), Akdeniz 46 (%7.07), genifl yay›l›fll› veya bilinmeyen 375 (%57.69). Araflt›rma alan›ndan toplanan türlerin 80’i (%12.30) endemiktir. Anahtar Sözcükler: Vasküler bitkiler, Flora, E¤erli Da¤›, Amasya
Introduction Karacakaya, Afla¤› and Yukar› fiamba villages, in the north The research area is located between Central Anatolia by Keçiköy and Çetmi villages, and in the south by and the Middle Black Sea region. The area contains two Karahisar, Büyükdüvenci and K›flladeresi villages (Figure 1). The surface measurement of the study area is 566.10 phytogeographic regions: the Euro-Siberian and Irano- 2 Turanian. The mountain was selected as a research area km and this measure was found by using a planometer as it is little-studied and has some interesting on the map. In the study area one of the most important streams flowing continuously is Köseler Stream characteristics in terms of both flora and phytogeography 6 3 as it is a transition region. In addition there are different (Büyükçay), the flow speed of which is 10.10 m /year vegetation types such as steppe, degraded forest and (Anonymous, 1973). forest on E¤erli Mountain. The climate of the research area was examined using E¤erli Mountain is located in the west of Amasya data from the meteorology station in Amasya. The province and within the provincial boundary of Amasya climate indicates transition region characteristics in the and Çorum. According to the grid square system used in study area (between the continental climate of the Central the Flora of Turkey (Davis, 1965-1985), E¤erli Mountain Anatolia and the ocean climate of the Black Sea region). It is located in square A5. The research area is bordered in is possible to see the local climatic characteristics of both the west by Göçeri village, in the east by Al›c›k, areas in the research area (Anonymous, 1984).
453 The Flora of E¤erli Mountain (Amasya-Turkey)
A Figure 1. Location of the study area.
B 1500 Keçiköy D. C Gümüflsuyu Hanköy 12345678910 Çetmi Keçiköy
Koçköy Al›c›k Eymir 1000 Bulak
750 Çitli Kandil D. Yemiflen Karacakaya 1250
E⁄ERL‹ DA⁄I Afl. fiamba Diphac› Göçeri 1000 Köseler Köseler C.Ovabaafl› (‹spari) Karacakaya 1250 1500 Yuk. fiamba Kayabafl› E¤erli Da¤ E⁄ERL‹ DA⁄I Küçükçay Büyücay Güllüce Akp›nar (Kü.Elbiz) 1500 Pekmezci
E⁄ERL‹ DA⁄I
Güvenli KARAH‹SAR Kü Düvenci Çaydere 1000 1500 De¤irmendere K›flladeresi Boundary of Study area N Flowing water Büyük Düvenci 024 — — —
The most detailed geological study carried out in the habitats, altitudes, collection dates, collectors’ names, area was by Alp (Alp, 1972). The geological structure of numbers, their phytogeographical regions, and the research area mostly contains rocks belonging to the endemisms. The authors of the plant names were checked Quaternary, Neogene, Eocene volcanic and Cretaceous (Brummitt & Powell, 1992). An asterisk in front of a periods (Anonymous, 1973; Anonymous, 1980). There plant name indicates a new record for the A5 grid square. are three large soil groups in the study area, namely, All the specimens are kept at the Herbarium of the brown forest soil, brown soil and alluvial soil Education Faculty in Amasya. (Anonymous, 1975) because of climate, topography and Abbreviations used in the text and the appendix areas differences in the main substance. follows: E: East Ir.-Tur.: Irano-Turanian Materials and Methods El.: Element Medit.: Mediterranean The research material consists of about 2000 plant End.: Endemic N: North specimens collected from the area between 1996 and Euro-Sib.: Euro-Siberian S: South 1998. Efforts were made to collect both flowering and Eux.: Euxine W: West fruiting specimens. The specimens were prepared H.Eux.: Hyrcano-Euxin Approx.: Approximately according to established herbarium techniques. The Flora of Turkey (Davis, 1965-1985; Davis et al., 1988) and Discussion and Conclusion other related floras (Evan & Townsend, 1968; Heywood & Tutin, 1964-1981; Heywood, 1978) were utilized in At the end of the study on the 2000 vascular plant the identification of the specimens. Experts were specimens collected from E¤erli Mountain and its vicinity, 620 species (650 taxa) and 298 genera, belonging to 70 consulted in some controversial cases. families, were established. One species is a nonflowering The order of plant taxa follows that in the Flora of vascular plant (of Pteridophyta). Four species belong to Turkey (Davis, 1965-1985; Davis et al., 1988). The Gymnospermae, while the other 615 were floristic list is in the appendix. The list gives the localities Angiospermae. The dispersion of the plant taxa in the of the taxa first (the area lies entirely within “A5 study area into the large taxonomical groups is shown in Amasya”, which is omitted from the citation), followed by Table 1.
454 A. CANSARAN
Table 1. The dispersion of taxa into the large taxonomical groups. The ten largest families according to number of species are shown in Table 3. Number Number Number of of of taxa in total Table 3. Species totals of the largest families found in the area. Families Genera (sp., subsp., var.) Familia Genera Species % Species Pteridophyta 11 1 Spermatophyta 69 297 649 Asteraceae 36 78 12.58 Gymnospermae 22 4Fabaceae 22 77 12.41 Dicotyledones 61 258 576 Poaceae 23 42 6.77 Monocotyledones 63769Lamiaceae 19 40 6.45 Brassicaceae 20 35 5.64 Total 70 298 650 Caryophyllaceae 13 32 5.16 Rosaceae 16 31 5.00 The species of the study area, categorized according Boraginaceae 13 26 4.19 to phytogeographic region, can be listed as follows: Scrophulariaceae 9 25 4.03 Irano-Turanian elements 102 (15.69%), Euro-Siberian Apiaceae 20 23 3.70 elements 97 (16 Eux., 3 H.Eux.) (14.92%), and Other families 107 211 34.03 Mediterranean elements 46 (7.07%); the remaining 375 (57.69%) species are multi-regional or of unknown phytogeographic origin (Table 2). The species belonging to first ten families form about 65% of the total species number.
Table 2. Distribution of species in research area within The genera with the largest number of species in the phytogeographic regions. study area are shown in Table 4. The richest genera are Astragalus L. (23), Silene L. and Lathyrus L. (10). Phytogeographic elements Number of species Rates(%) Astragalus has a total of c. 2500 species in the world; it is one of the largest genera of flowering plants. The Irano-Turanian elements 102 15.69 estimated number of species in Turkey is c. 400. Euro-Siberian elements 97 14.92 Furthermore, the steppic areas are increasing day by day Mediterranean elements 46 7.07 because of excessive destruction. Therefore, the area Multi-regional or of unknown 375 57.69 contains by far the greatest number of these species. The phytogeographic region genera Silene and Lathyrus rank second. Silene is the fifth largest genus in Turkey. The species belonging to the According to these data the research area is located in genus Lathyrus are spread especially in the shrubbery and the transition zone between the Euro-Siberian and Irano- forest areas in the study area. Turanian floristic regions. However, the effect of the Irano-Turanian phytogeographic region is greater. Table 4. Species totals of the largest genera in the study area. When the number of taxa is taken into consideration, the family Asteraceae ranks first with 78 species. This is Genus Number of Genus Number of related to its being the largest family in the flora of Species Species Turkey, having many family members and greater ecological toleration and breaking up seeds easily. Astragalus 23 Ranunculus 7 Following Asteraceae, the family Fabaceae is represented Silene 10 Alyssum 7 by 77 species. The reason why Fabaceae is second is that Lathyrus 10 Rumex 7 it is the second largest family in the flora of Turkey and Trifolium 8 Vicia 7 involves large genera containing many species. The Galium 8 Centaurea 7 Poaceae family comes next, because the family Poaceae is Onosma 8 Verbascum 7 the fifth largest family in the flora of Turkey. Salvia 8 Veronica 7
455 The Flora of E¤erli Mountain (Amasya-Turkey)
Moreover, the six species of Erysimum L., Medicago between Central Anatolia and the Black Sea regions or its L., Pilosella Hill, Convolvulus L. and Orobanche L. and the adjacent areas. The study “Flora of the Area between five species of Crataegus L., Sedum L., Inula L. and Amasya Castle and the villages of Vermifl and Yuvac›k” Quercus L. have been found in the study area. (Cansaran & Aydo¤du, 1998) was performed in the same The number of endemic taxa in the study area is 80 transition zone but it contains the Yeflil›rmak River. The and the endemism rate is 12.30%. Forty of these taxa Mediterranean elements, which are mostly from the belong to the Irano-Turanian phytogeographic region. tertiary period, grow in the Mediterranean enclaves Furthermore, five of the endemic taxa belong to the which occur along the Yeflil›rmak Valley. Thus, this valley Mediterranean phytogeographic region and five of these represents the pecularities of the Mediterranean region. taxa the Euro-Siberian. The pluriregional endemics Additionally, according to endemism ratios the nearby number 30. The proportion of endemism in the area is areas are not rich enough like the research area. low (12.30%), below the average for Turkey (33.5%) The comparison of the first three families in the (Ekim, 1990). The main reason for this is that the research area (E¤erli Mountain) and other nearby areas in edaphic, climatic and topographic properties of the region the region in respect to species richness is shown in Table do not vary enough. 6. The families Asteraceae, Fabaceae, Lamiaceae, The distribution of species in the research area (E¤erli Brassicaceae and Poaceae share the first three places in Mountain) and of other studies carried out nearby these studies. The family Asteraceae ranks first in terms according to the phytogeographic regions and the of the number of species. In addition, these five families comparison of endemism ratios are shown in Table 5. In are also the first five families in the Flora of Turkey studies performed in adjacent areas the Irano-Turanian (Davis, 1988). Astragalus is first according to number of elements rank first and the Euro-Siberian elements species. second except in “Flora of the Area between Amasya Some of the taxa in the study area have different Castle and the villages of Vermifl and Yuvac›k” (Cansaran taxonomic properties from those in the Flora of Turkey. & Aydo¤du, 1998). This condition is related to the fact These properties are given in Table 7. that these regions are located in the transition zone
Table 5. The distribution of species in the research area (E¤erli Mountain) and others according to the phytogeographic regions and the comparison of endemism ratios.
The Areas Irano-Turanian Euro-Siberian Mediterranean Endemism Ratio
E¤erli Mountain 102 (15.69%) 97 (14.92%) 46 (7.07%) 80 (12.30%) Vermifl-Yuvac›k vill.-Amasya Castle (Cansaran & Aydo¤du, 1998) 70 (15.71%) 31 (7.38%) 40 (9.52%) 46 (11%) Amasya-Yozgat-Çorum (Kurt et al., 1998) 100 (18.65%) 71 (13.25%) 41 (7.65%) 38 (7.09%) Deveci Mountains (Yozgat-Tokat) (‹larslan, 1983) 238 (37.8%) 100 (15.9%) 63 (10.1%) 107 (17.1%) Çaml›bel Pass and Y›ld›z Mountain (Civelek, 1992) 245 (25.1%) 159 (16.2%) 32 (3.2%) 138 (14.1%)
Table 6. The comparison of the first three families in the research area (E¤erli Mountain) and other areas in the region in respect to species richness.
The Areas The largest 3 families The largest 3 genera
E¤erli Mountain Asteraceae, Fabaceae, Poaceae Astragalus, Silene -Lathyrus, Trifolium Galium-Onosma-Salvia Vermifl-Yuvac›k vill.-Amasya Castle Asteraceae, Lamiaceae, Fabaceae Astragalus, Salvia, Convolvulus-Euphorbia- (Cansaran & Aydo¤du, 1998) Silene-Verbascum Amasya-Yozgat Çorum (Kurt et al., 1998) Asteraceae, Fabaceae, Poaceae Astragalus, Trifolium, Silene Deveci mountains (Yozgat-Tokat (‹larslan, 1983) Asteraceae, Fabaceae, Lamiaceae Astragalus, Silene, Centaurea Akda¤ (Amasya) (Baytop & Alp›nar, 1980) Asteraceae, Fabaceae, Brassicaceae Astragalus, Trifolium-Silene-Alyssum, Centaurea
456 A. CANSARAN
Table 7. Some of the taxa in the E¤erli Mountain have different taxonomic properties from those in the Flora of Turkey.
Taxa Flora of Turkey E¤erli Mountain
Convolvulus cantabrica L. Leaves 7-15 mm broad Also there are leaves 3-4 mm broad Dorycnium pentaphyllum Scop. subsp. herbaceum (Vill.)Rouy Leaflets 1-6 mm broad Leaflets 1-9 mm broad Inula helenium L. subsp. orgyalis Capitulum 3.5-5 cm in length Capitulum 2.5-5 cm in length Onosma sericeum Willd. Corolla 15-20 mm in length Corolla 15-22 mm in length Rumex caucasicus Rech. It is known from 3100 m altitude It is collected from 1050 m altitude
Fifty-five taxa have been defined as new records for Acknowledgements the A5 square (Davis, 1965-1985; Davis et al., 1988; I thank Prof. Dr. Mustafa Aydo¤du from the Faculty Donner, 1985; Donner, 1987; Donner, 1990; Erik, of Education in Gazi University; Prof. Dr. Mecit Vural, 1995; Korkmaz et al., 1996; Y›ld›r›ml›, 1994; Y›ld›r›ml›, Prof. Dr. Hayri Duman and Assoc. Prof. Dr. Zeki Aytaç, 1996; Y›ld›r›ml›, 1997; Y›ld›r›ml›, 1999) throughout this from the Faculty of Arts and Science in Gazi University; study. As a result, various floristic data which contribute Prof. Dr. Ekrem Sezik from the Faculty of Pharmacy in to the flora of Turkey were provided. Gazi University; and Prof. Dr. Mehmet Koyuncu from the Faculty of Pharmacy in Ankara University for their kind help in the determination of some plant specimens.
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457 The Flora of E¤erli Mountain (Amasya-Turkey)
Appendix Ranunculus brutius Ten. Roemeria hybrida (L.) DC. subsp. hybrida Approx 5 km NW of K›flladeresi village, under E places of Köseler village, steppe; 1200 m, PTERIDOPHYTA forest; 1400 m, 19.VII.1997, Cansaran 16.VI.1996, Cansaran 1452. 2860. Euro-Sib. El. SPHENOPSIDA Papaver lacerum Popov R. constantinopolitanus (DC.) d’Urv. EQUISETACEAE Approx 1 km E of Güllüce village, steppe; Güllüce village surroundings, N places, stream 1400 m, 24.V.1997, Cansaran 2373. Equisetum ramosissimum Desf. bank; 1300 m, 05.V.1996, Cansaran 1100. Hypecoum imberbe Sibth. & Sm. Güllüce village surroundings, N places, stream R. oxyspermus Willd. bank; 1300 m, 05.V.1996, Cansaran 1100- Approx 2 km W of Güvenli village, roadsides; B. N places of E¤erli Mountain, around Yukar› 1100 m, 19.V.1996, Cansaran 1216. Kayabafl› village, open areas; 1650 m, SPERMATOPHYTA Corydalis solida (L.) Sw. subsp. solida 04.V.1996, Cansaran 1042. GYMNOSPERMAE R. isthmicus Boiss. subsp. stepporum N places of E¤erli Mountain, Yukar› Kayabafl› PINACEAE P.H.Davis village surroundings, open areas; 1600 m, 04.V.1996, Cansaran 1066. Pinus sylvestris L. 300 m W of Ovabafl› village, open forests; Fumaria officinalis L. Approx 4 km S of Yemiflen village, forest; 1100 m, 05.V.1996, Cansaran 1114. 1500 m, 02.VII.1997, Cansaran 2725. R.arvensis L. Between Ovabafl› and Köseler village, roadsides; 1000 m, 05.V.1996, C.1119 P.nigra Arn. subsp. pallasiana (Lamb.) 1 km E of Köseler village, steppe; 1200 m, Holmboe 16.VI.1996, Cansaran 1453. F. asepala Boiss. Around Köseler village, N slopes of Saray R. trichophyllus Chaix W places of Köseler village, open areas; 1300 Mountain, forest; 1320 m, 14. IV.1996, m, 26.V.1996, Cansaran 1319. Ir.-Tur. El. Cansaran1040. Approx 5 km E of Büyükçay village, in the water; 860 m, 19.V.1996, C .1160. BRASSICACEAE/CRUCIFERAE CUPRESSACEAE R. ficaria L. subsp. calthifolius (Reiche) Sinapis arvensis L. Juniperus communis L. subsp. alpina (Sm.) Archer Celak. Approx 1 km NE of Küçükçay village, degraded forest; 1000 m, 20.VII.1996, 1 km S of Yemiflen village, forest; 1530 m, S places of Yukar› Kayabafl› village, stream 04.V.1996, Cansaran 1051. bank, 1530 m, 04.V.1996, Cansaran 1071. Cansaran 1835. J. oxycedrus L. subsp. oxycedrus Ceratocephalus falcatus (L.) Pers. Cardaria draba (L.) Desv. subsp. draba Between Ovabafl› and Akp›nar village, Vicinity of Köseler village, E slopes of Saray Between E¤mir and Al›c›k, roadsides; 1100 degraded forest; 1200 m, 02.VIII.1996, Mountain, open areas; 1300 m, 14.IV.1996, m, 05.V.1996, C.1132. Cansaran 1023. Cansaran 1902. Aethionema arabicum (L.) Andrz. ex DC. ANGIOSPERMAE BERBERIDACEAE Approx 2 km W of Büyükçay village, steppe; DICOTYLEDONES Berberis vulgaris L. 1100 m, 21.VII.1996, Cansaran 1727. RANUNCULACEAE Approx 5 km E of Karahisar village, Thlaspi arvense L. roadsides; 1100 m, 19.V.1996, Cansaran Nigella arvensis L. var. glauca Boiss. Approx 4 km SW of Köseler village, open 1175. Approx 1 km N of Karahisar village, forest; humid places; 1400 m, 25.V.1996, roadsides; 1100 m, 10.VII.1996, Cansaran B. crataegina DC. Cansaran 1246. 1566. 100 m S of Ovabafl› village, forest; 1200 m, T. perfoliatum L. Consolida orientalis (Gay) Schröd. 03.VIII.1996, Cansaran 2225. App 2 km W of Güvenli village, roadsides; PAPAVERACEAE Approx 1 km NE of Küçükçay village, 1100 m, 19.V.1996, Cansaran 1229. degraded forest; 1500 m, 10.VII.1996, Chelidonium majus L. Capsella bursa-pastoris (L.) Medik. Cansaran 1545. Approx 7 km SE of Göçeri village, under E places of Büyükçay village, roadsides; 1050 Clematis vitalba L. forest; 1200 m, 01.VII.1997, Cansaran m, 19.V.1996, Cansaran 1158. Vicinity of Köseler village, E places, roadsides; 2703. Euro-Sib. El. *C. rubella Reut. 1200 m, 10.VII.1996, Cansaran 1592. *Glaucium grandiflorum Boiss. & E.Huet Adonis flammea Jacq. var. grandiflorum S places of Yukar› Kayabafl› village, open areas; 1500 m, 04.V.1996, Cansaran 1081. Between Ovabafl› and Köseler village, 1 km NE of Küçükçay village, open degraded Medit. El. roadsides; 950 m; 05.V.1996, Cansaran forest; 1100 m, 08.VI.1997, Cansaran 2412. 1118. Ir.-Tur. El.
458 A. CANSARAN
Boreava orientalis Jaub. & Spach Turritis laxa (Sibth. & Sm.) Hayek D. kochii (Petri) Schulz Between E¤mir and Al›c›k, roadsides; 1100 Approx 4 km S of Yemiflen village, under S places of Küçükçay village; 1080 m, m, 05.V.1996, Cansaran 1133. forest; 1470 m, 02.VII.1997, Cansaran 19.V.1996, Cansaran 1184. 2770. Neslia apiculata Fisch. Camelina rumelica Vell. Approx 4 km SW of Köseler village, open Matthiola longipetala (Vent.) DC. subsp. 1 km E of Köseler village, steppe; 1200 m, forest; 1400 m, 25.V.1996, Cansaran 1238. bicornis (Sibth. & Sm.) P.W.Ball 16.VI.1996, Cansaran 1425. Fibigia eriocarpa (DC.) Boiss. W places of Köseler village, 1300 m, RESEDACEAE 26.V.1996, Cansaran 1328. Approx 3 km NW of Akp›nar village, Reseda lutea L. var. lutea roadsides; 1200 m, 05.V.1996, Cansaran Malcolmia africana (L.) R. 1103. 1 km E of Al›c›k, roadsides; 700 m, 1 km NE of Küçükçay village, degraded 19.V.1996, Cansaran 1140. *Alyssum meniocoides Boiss. forest, 1100 m, 08.VI.1997, Cansaran 2424. R. lutea L. var. nutans Boiss. 300 m W of Ovabafl› village, steppe, 1100 m, Erysimum cuspidatum (M.Bieb.) DC. 05.V.1996, Cansaran 1111. Ir.-Tur. El. 1 km E of Köseler village, steppe; 1250 m, 3 km SW of Çetmi village, shrubbery, 950 m, 16.VI.1996, Cansaran 1423. *A. blepharocarpum Dudley & Hub.-Mor. 14.VI.1998, Cansaran 3011. R. luteola L. Approx 4 km SW of Köseler village, under E. pulchellum (Willd.) Gay forest; 1450 m, 25.V.1996, Cansaran 1271- 1 km W of Çitli (Ba¤l›ca) village, roadsides; A. End. Ir.-Tur. El. Approx 2 km S of Köseler village, under 1000 m, 08.VI.1997, Cansaran 2432. forest, 1300 m, 20.VII.1997, Cansaran A. desertorum Stapf var. desertorum CISTACEAE 2931. (det. F. Karaer) S places of Yukar› Kayabafl› village, open Cistus laurifolius L. E. leptophyllum (M.Bieb.) Andrz. areas; 1500 m, 04.V.1996, Cansaran 1089. Approx 3 km W of Çitli (Ba¤l›ca) village, Approx 9 km SE of Göçeri village, under A. minus (L.) Rothm. var. minus degraded forest; 1100 m, 08.VI.1997, forest; 1240 m, 01.VII.1997, Cansaran 300 m W of Ovabafl› village, steppe; 950 m, Cansaran 2451. 2676. 03.VIII.1996, Cansaran 2043. Helianthemum nummularium (L.) Mill. E. thyrsoideum Boiss. subsp. thyrsoideum A. strigosum Banks & Sol. subsp. strigosum subsp. nummularium Approx 5 km N of Karahisar village, 1100 m, Approx 1 km NE of Küçükçay village, steppe; 1 km NE of Köseler village, steppe; 1150 m, 19.V.1996, Cansaran 1188. 1100 m, 20.VII.1996, Cansaran 1653. 10.VII.1996, Cansaran 1546. E. smyrnaeum Boiss. & Ball A. pateri Nyár. subp. pateri H. nummularium (L.) Mill. subsp. tomentosum (Scop.) Schinz & Thell. 500 m N of Küçükçay village, open degraded Approx 1 km NE of Küçükçay village, forest; 1150 m 15.VI.1996, Cansaran 1370. shrubbery; 1100 m, 08.VI.1997, Cansaran Approx 4 km S of Ovabafl› village, open End. Ir.-Tur. El. 2380. shrubbery; 1250 m, 31.V.1998, Cansaran A. pateri Nyár. subsp. prostratum (Nyár.) E. graecum Boiss. & Heldr. 2969. Dudley Approx 3 km S of Yemiflen village, under H. canum (L.) Baumg. (Group D) Approx 1 km NE of Küçükçay village, steppe; forest; 1400 m, 19.VII.1997, Cansaran W places of Köseler village, Ekinala¤›, open 1150 m, 10.VII.1996, Cansaran 1481. End. 2795. areas; 1300 m, 26.V.1996, Cansaran 1320. Ir.-Tur. El. Alliaria petiolata (M.Bieb.) Cav. & Grande H. ledifolium (L.) Mill. var. lasiocarpum A. murale Waldst. & Kit. var. murale Between E¤mir and Al›c›k, roadsides; 1100 (Willd.) Bornm. 100 m S of Ovabafl› village, under forest, m, 05.V.1996, Cansaran 1134. Approx 1 km E of Güllüce village, steppe; 1200 m, 03.VIII.1996, Cansaran 2244. Sisymbrium altissimum L. 1400 m, 24.V.1997, Cansaran 2353. Clypeola jonthlaspi L. Between Ovabafl› and Köseler village, Fumana procumbens (Dun) Gren. & Godr. 300 m, W of Ovabafl› village, steppe; 1100 m, roadsides; 950 m, 05.V.199, Cansaran 1116. App 1 km NE of Küçükçay village, steppe; 05.V.1996, Cansaran 1112. S. orientale L. 1100 m, 08.VI.1997, Cansaran 2386. Erophila verna (L.) Chevall. subsp. verna E places of Köseler village, 1270 m, VIOLACEAE Around Köseler village, Gökdere, roadsides; 10.VII.1996, Cansaran 1600-B. Viola odorata L. 1100 m, 31.III.1996, Cansaran 1000. Descurainia sophia (L.) Webb ex Prantl Around Köseler village, S slopes of Saray Arabis sagittata (Bert) DC. Köseler village surroundings; 1100 m, Mountain, open areas; 1200 m, 14.IV.1996, Approx 7 km SE of Göçeri village, open Cansaran 1025. forest; 1200 m, 01.VII.1997, Cansaran 25.V.1996, Cansaran 1297. 2704.
459 The Flora of E¤erli Mountain (Amasya-Turkey)
V. sieheana Becker Cerastium dichotomum L. subsp. Saponaria glutinosa M.Bieb. dichotomum S places of Yukar› Kayabafl› village, open 1 km NE of Küçükçay village, degraded areas; 1700 m, 04.V.1996, Cansaran 1088. Approx 2 km W of Güvenli village, roadsides; forest; 1100 m. 20.;VII.1996, Cansaran 1100 m, 19.V.1996, Cansaran 1217. 1663-A. V. kitaibeliana Roem. & Schult. C. brachypetalum Pers. subsp. roeseri E of Köseler castle, Aliba¤›, open areas; 1180 S. prostrata Willd. subsp. prostrata (Boiss. & Heldr.) Nyman m, 14.IV.1996, Cansaran 1017. App 4 km SW of Köseler village, open forests; 500 m NE of Ovabafl› village, open forests; POLYGALACEAE 1500 m, 25.V.1996, Cansaran 1289. End. 1000 m, 03.VIII.1996, Cansaran 2267. Ir.-Tur. El. Polygala supina Schreb. Holosteum umbellatum L. var. glutinosum Vaccaria pyramidata Medik. var. Approx 5 km NW of K›flladeresi village, under (M.Bieb.) Gay grandiflora (Fisch. ex DC.) Cullen forest; 1400 m, 19.VII.1997, Cansaran E places of Köseler castle, open areas; 1250 500 m W of fiamba village, roadsides; 1050 2806. m, 14.IV.1996, Cansaran 1018. m, 15.VI.1996, Cansaran 1355. P. pruinosa Boiss. subsp. pruinosa H. umbellatum L. var. tenerrimum (Boiss.) Silene italica (L.) Pers. Approx 3 km SW of Çetmi village, under Gay Approx 4 km NW of K›flladeresi village, under degraded forest; 950 m, 14.VI.1998, 1 km E of Güllüce village, steppe; 1400 m, Cansaran 3003. 24.V.1997, Cansaran 2372. forest; 1400 m, 19.VII.1997, Cansaran 2849. CARYOPHYLLACEAE Dianthus anatolicus Boiss. S. chlorifolia Sm. Arenaria serpyllifolia L. Between Akp›nar and Güllüce, village 1 km E of Köseler village, steppe; 1260, Approx 3 km S of Köseler village, under roadsides; 1300 m, 10.VII.1996, Cansaran 16.VI.1996, Cansaran 1427. Ir.-Tur. El. forest; 1200 m, 20.VII.1997, Cansaran 1456. End. 2938. D. carmelitarum Reut. ex Boiss S. otites (L.) Wibel A. ledebouriana Fenzl var. ledebouriana Approx 4 km S of Yemiflen village, under 500 m W of Güllüce village, steppe; 1470 m, 500 m W of Güllüce village, steppe; 1470 m, forest, 1500 m, 02.VII.1997, Cansaran 10.VII.1996, Cansaran 1472. 2752. End. Eux. El. 10.VII.1996, Cansaran 1479. End. S. spergulifolia (Desf.) M.Bieb. D. calocephalus Boiss. M. multinervis (Boiss.) Bornm. Approx 4 km W of Küçükçay village, steppe; Approx 2 km NE of Çitli (Ba¤l›ca) village, Approx 3 km NE of Çitli (Ba¤l›ca) village, open 1060 m, 19.V.1996, Cansaran 1146. Ir.- under shrubbery; 1020 m, 20.VI.1997, degraded forest; 1060 m, 20.VI.1997, Tur. El. Cansaran 2588. Cansaran 2621. S. supina M.Bieb. subsp. pruinosa (Boiss.) Petrorhagia alpina (Hablitz) Ball & Heywood M. corymbulosa (Boiss. & Ball) McNeil var. Chowdhuri subsp. alpina corymbulosa 1 km NE of Küçükçay village, steppe; 1150 Approx 5 km NW of K›flladeresi village, under 300 m W of Ovabafl› village, open shrubbery; m, 10.VII.1996, Cansaran 1480. forest; 1400 m, 19.VII.1997, Cansaran 1100 m, 05.V.1996, Cansaran 1121. End. 2846. S. vulgaris (Moench) Garcke var. vulgaris Ir.-Tur. El. P.alpina (Hablitz) Ball & Heywood subsp. 1 km E of Al›c›k, roadsides; 700 m, M. hybrida (Vill.) Schischk. subsp. hybrida olympica (Boiss.) Ball & Heywood 19.V.1996, Cansaran 1141. Approx 2 km NE of Çitli (Ba¤l›ca) village, Approx 3 km S of Yemiflen village, under S. compacta Fisch. degraded forest; 1020 m, 20.VI.1997, forest; 1500 m, 02.VII.1997, Cansaran 2711. Cansaran 2589. Approx 7 km SE of Göçeri village, open Velezia fasciculata Boiss. forests; 1200 m, 01.VII.1997, Cansaran Stellaria media (L.) Vill. subsp. media Approx 2 km NE of Çitli (Ba¤l›ca) village, 2708. Between Ovabafl› and Köseler village, degraded forest; 1030 m, 20.VI.1997, S. alba (Mill.) Krause subsp. divaricata roadsides; 950 m, 05.V.1996, Cansaran Cansaran 2609. (Reiche) Walters 1120. V. rigida L. Approx 3 km S of Köseler village, under S. holostea L. 1 km N of Karahisar village, roadsides, 1100 forest; 1350 m, 27.VI.1998, Cansaran 1300. Approx 8 km SE of Göçeri village, under m, 10.VII.1996, Cansaran 1570. forest, 1230 m, 01.VII.1997, Cansaran S. alba (Mill.) Krause subsp. eriocalycina *V. pseudorigida Hub.-Mor. 2647. Euro.-Sib. El. (Boiss.) Walters 1 km W of Çitli (Ba¤l›ca) village, under Myosoton aquaticum (L.) Moench Approx 2 km SW of Çetmi village, degraded shrubbery; 1100 m, 27.VI.1998, Cansaran forest; 950 m, 14.VI.1998, Cansaran 3015. S places of Küçükçay village; 1080 m, 3166. End. E. Medit. El. 19.V.1996, Cansaran 1182. Euro-Sib. El.
460 A. CANSARAN
S. dichotoma Ehrh. subsp. euxina (Rupr.) *R. caucasicus Rech. MALVACEAE Coode & Cullen 500 m W of fiamba village, roadsides; 1050 Malva neglecta Wallr. 1 km NE of Küçükçay village, steppe; 1100 m, 15.VI.1996, Cansaran 1356. Eux. El. Approx 4 km SW of Köseler village, open m, 20.VII.1996, Cansaran 1627. Eux.El. R. crispus L. forests, 1400 m, 25.V.1996, Cansaran S. conica L. 1241. Approx 3 km S of Yemiflen village, open Approx 2 km W of Çitli (Ba¤l›ca) village, forests; 1450 m, 02.VII.1997, Cansaran Alcea apterocarpa (Fenzl) Boiss. under shrubbery; 1100 m, 08.VI.1997, 2783. 1 km W of Köseler village, roadsides, steppe Cansaran 2455. R. obtusifolius L. subsp. subalpinus (Schur) areas; 1100 m, 10.VII.1996, Cansaran 1582. Agrostemma githago L. Celak. End. Ir.-Tur. El. 1 km S of Güvenli village, open degraded 1 km W of Çitli (Ba¤l›ca) village, open LINACEAE forest; 1160 m, 15.VI.1996, Cansaran 1376. shrubbery; 1100 m, 27.VI.1998, Cansaran Linum nodiflorum L. ILLECEBRACEAE 3167. 1 km E of Köseler village, steppe; 1000 m, Herniaria incana Lam. *R. nepalensis Spreng. 24.V.1997, Cansaran 2327. Medit. El. 300 m SE of Karahisar village, roadsides; Approx 4 km S of Yemiflen village, open L. olympicum Boiss. 1050 m, 15.VI.1996, Cansaran 1409. forest; 1450 m, 02.VII.1997, Cansaran SE places of Karahisar village, roadsides; Paronychia kurdica Boiss. subsp. kurdica 2780. 1050 m, 15.VI.1996, Cansaran 1408, End. var. kurdica CHENOPODIACEAE L. hirsutum L. subsp. pseudoanatolicum W places of Köseler village, Ekinala¤›, steppe; Chenopodium botrys L. P.H.Davis 1300 m, 26.V.1996, Cansaran 1308. Between Ovabafl› and Akp›nar village, 1 km NE of Küçükçay village, open shrubbery; *Scleranthus annuus L. subsp. annuus degraded forest; 1180 m, 02.VIII.1996, 1100 m, 08.VI.1997, Cansaran 2420. End. Approx 4 km SW of Köseler village, open Cansaran 1987. Ir.-Tur. El. forest; 1400 m, 25.V.1996, Cansaran 1274. C. foliosum (Moench) Archer L. tenuifolium L. POLYGONACEAE Vicinity of Köseler village, roadsides; 1200 m, 1 km W of Köseler village, Gökdere, open Polygonum cognatum Meisn. 10.VII.1996, Cansaran 1595. areas; 1100 m, 15.VI.1997, Cansaran 2502. Approx 4 km S of Yemiflen village, open TAMARICACEAE GERANIACEAE forests; 1450 m, 02.VII.1997, Cansaran 2782. Myricaria germanica (L.) Desv. Geranium robertianum L. P. aviculare L. 1 km W of Köseler village, open degraded Approx 4 km SW of Köseler village, under forest; 1400 m, 25.V.1996, Cansaran 1273. Between Akp›nar and Güllüce village, forest, humid places; 1200 m, 04.VIII.1996, roadsides; 1470 m, 10.VII.1996, Cansaran Cansaran 2172. (det. H. Duman) G. pusillum Burm. 1557. HYPERICACEAE / GUTTIFERAE Between Akp›nar and Güllüce village, degraded forest; 1400 m, 10.VII.1996, *P. equisetiforme Sibth. & Sm. Hypericum perfoliatum L. Cansaran 1550. Between Ovabafl› and Akp›nar village, Approx 7 km SE of Göçeri village, open shrubbery; 1250 m, 02.VIII.1996, Cansaran G. tuberosum L. subsp. tuberosum forests; 1200 m, 01.VII.1997, Cansaran 2002. 2699. Medit. El. (det. F.Karaer) Between E¤mir and Al›c›k, roadsides; 1100 P. convolvulus L. m, 05.V.1996, Cansaran 1122. H. origanifolium Willd. 1 km E of Köseler village, steppe; 1250 m, G. macrostylum Boiss. Approx 3 km SW of Çetmi village, shrubbery, 16.VI.1996, Cansaran 1434. 970 m, 14.VI.1998, Cansaran 3045. 1 km E of Güllüce village, steppe; 1400 m, Rumex acetosella L. 24.V.1997, Cansaran 2304. E. Medit. El. H. aviculariifolium Jaub. & Spach subsp. Approx 5 km N of Karahisar village, open aviculariifolium var. aviculariifolium Erodium ciconium (L.) L’Hér. areas; 1100 m, 19.V.1996, Cansaran1187. Approx 4 km SW of Köseler village, open Approx 3 km NW of Akp›nar village, R. scutatus L. forest; 1400 m, 25.V.1996, Cansaran 1291. roadsides; 1150 m, 05.V.1996, Cansaran 1 km W of Köseler village, roadsides, steppe End. E. Medit. El. 1107. areas; 1100 m, 10.VII.1996, Cansaran 1577. E. cicutarium (L.) L’Hér. subsp. cicutarium H. perforatum L. R. patientia L. Approx 2 km SW of Çetmi village, under 300 m SE of Karahisar village, steppe; 1050 shrubbery; 930 m, 14.VI.1998, Cansaran 1 km SW of Köseler village, forest; 1250 m, m, 15.VI.1996, Cansaran 1400. 04.VIII.1996, Cansaran 2127. 3020.
461 The Flora of E¤erli Mountain (Amasya-Turkey)
E. acaule (L.) Bech. & Thell. Argyrolobium biebersteinii Ball A. schottianus Boiss. Köseler village surroundings, S slopes of Approx 5 km NW of K›flladeresi village, under Approx 4 km NW of K›flladeresi village, Saray Mountain, open areas; 1400 m, forest; 1400 m, 19.VII.1997, Cansaran forest; 1450 m, 19.VII.1997, Cansaran 14.IV.1996, Cansaran 1029. Medit El. 2807. 2848. End. ACERACEAE Galega officinalis L. A. micropterus Fisch. Acer campestre L. subsp. campestre Between Akp›nar and Güllüce village, Approx 2 km W of Büyükçay village, steppe; Between Ovabafl› and Akp›nar village, roadsides; 1400 m, 10.VII.1996, Cansaran 1100 m, 21.VII.1996, Cansaran 1741. End. degraded forest; 1230 m, 02.VIII.1996, 1549. Euro.-Sib. El. Ir.-Tur. El. Cansaran 1894. Colutea cilicica Boiss. & Ball A. albifolius Freyn & Sint. RHAMNACEAE Approx 3 km SW of Çetmi village, degraded 1 km N of Karahisar village, roadsides; 1100 Rhamnus catharticus L. forest; 950 m, 14.VI.1998, Cansaran 3023. m, 10.VII.1996, Cansaran 1567. End. Ir.- Tur. El. Approx 3 km SW of Çetmi village, shrubbery, Astragalus leucothrix Freyn & Bornm. 950 m, 14.VI.1998, Cansaran 3007. Euro- A. andrachneifolius Fenzl Sib. El. 1 km W of Köseler village, Gökdere, steppe; 1100 m, 15.VI.1997, Cansaran 2503. End. Approx 2 km W of Çitli (Ba¤l›ca) village, R. petiolaris Boiss. Ir.-Tur. El. shrubbery; 1100 m, 08.VI.1997, Cansaran 1 km NE of Küçükçay village, degraded 2438. A. glycyphyllos L. subsp. glycyphyllos forest; 1100 m, 20.VII.1996, Cansaran *A. cephalotes Banks & Sol. var. brevicalyx 1667. End.? Approx 10 km SE of Göçeri village, under Eig ANACARDIACEAE forest; 1250 m, 01.VII.1997, Cansaran 2682. Euro-Sib. El. 1 km W of Köseler village, degraded forest; Cotinus coggyria Scop. 1200 m, 04.VIII.1996, Cansaran 2139 End. *A. melanophrurius Boiss. Approx 2 km SW of Çetmi village, degraded *A. strictispinis Boiss. forest; 930 m, 14.VI.1998, Cansaran 3024. Approx 3 km N of Karahisar village, roadsides; 1050 m, 19.V.1996, Cansaran 300 m SW of Ovabafl› village, steppe; 960 m, Rhus coriaria L. 1197. End. Ir.-Tur. El. 03.VIII.1996, Cansaran 2072. End. Between Ovabafl› and Akp›nar village, shrubbery; 1100m, 02.VIII.1996, Cansaran A. pinetorum Boiss. *A. podperae Sirj. 1898. Approx 4 km NW of Büyükçay village, steppe; 500 m W of Güllüce village, stony places; CELASTRACEAE 1200 m, 12.VII.1998, Cansaran 3170. End. 1470 m, 10.VII.1996, Cansaran 1470. End. Ir.-Tur. El. Ir.-Tur. El. Euonymus verrucosus Scop. A. pseudocaspius Fisch. A. brachypterus Fisch. 1 km NE of Küçükçay village, degraded forest; 1200 m, 20.VII.1996, Cansaran 300 m W of Büyükçay village, steppe; 1050 1 km NE of Küçükçay village, steppe; 1150 1682. m, 21.VII.1996, Cansaran 1837. End. m, 10.VII.1996, Cansaran 1515. End. Ir.- Tur. El. FABACEAE / LEGUMINOSAE A. microcephalus Willd. A ponticus Pall. Cytisus procumben s (Waldst. & Kit. ex Approx 3 km N of Büyükçay village, steppe; Willd.) Spreng. 1200 m, 15.VI.1996, Cansaran 1395. Ir.- 300 m SE of Karahisar village, roadsides; Approx 5 km E of Karahisar village; 1100 m, Tur. El. 1050 m, 15.VI.1996, Cansaran 1413. 19.V.1996, Cansaran 1177. *A. tokatensis Fisch. A. karamasicus Boiss. & Ball Chamaecytisus pygmaeus (Willd.) Rothm. 1 km NE of Küçükçay village, steppe; 1060 Approx 4 km SW of Köseler village, open 500 m W of Güllüce village, shrubbery; 1470 m, 20.VII.1996, Cansaran 1769. End. Ir.- forests; 1500 m, 25.V.1996, Cansaran m, 10.VIII.1996, Cansaran 1475. Euro-Sib. Tur. El. 1295. End. Ir.-Tur. El. El. A. barba-jovis DC. var. barba-jovis A. Iycius Boiss. *Gonocytisus pterocladus (Boiss.) Spach Between Ovabafl› and Akp›nar village, 1 km E of Güllüce village, steppe; 1400 m, Approx 8 km SE of Göçeri village, under degraded forest; 1250 m, 02.VIII.1996, 24.V.1997, Cansaran 2348. End. forest; 1230 m, 01.VII.1997, Cansaran Cansaran 2013. End. Ir.-Tur. El. 2643. E. Medit. El. *A. acmonotrichus Fenzl A. plumosus Willd. var. plumosus Genista tinctoria L. Approx 4 km SW of Köseler village, under 100 m S of Ovabafl› village, open forest; 1200 1 km W of Küçükçay village, steppe; 1050 m, forest; 1400 m, 25.V.1996, Cansaran 1259. m, 03.VIII.1996, Cansaran 2220. Euro-Sib. 20.VII.1996, Cansaran 1880. End. E. Medit. El. El.
462 A. CANSARAN
A. elongatus Willd. subsp. elongatus Lathyrus aureus (Stev.) D.Brândzâ T. campestre Schreb. 1 km NE of Küçükçay village, steppe; 1100 Approx 8 km SE of Göçeri village, under 1 km W of Köseler village, degraded forest; m, 08.VI.1997, Cansaran 2390. forest; 1230 m, 01.VII.1997, Cansaran 1200 m, 04.VIII.1996, Cansaran 2143. 2650. Eux. El. A. angustifolius Lam. subsp. pungens T. physodes Stev. ex M.Bieb. var. physodes L. tukhtensis Czeczott (Willd.) Hayek Approx 3 km W of Çitli (Ba¤l›ca) village, Approx 3 km S of Yemiflen village, open Approx 3 km S of Yemiflen village, under shrubbery; 1100 m, 08.VI.1997, Cansaran forests; 1450 m, 02.VII.1997, Cansaran forest; 1500 m, 02.VII.1997, Cansaran 2449. Medit. El. 2742. End. 2784. T. pratense L. var. pratense *L. elongatus (Bornm.) Sirj. Psoralea bituminosa L. Approx 5 km NW of K›flladeresi village, under 500 m N of Küçükçay village, open degraded 1 km E of Güllüce village, steppe; 1400 m, forest; 1400 m, 19.VII.1997, Cansaran forests, 1130, 15.VI.1996, Cansaran 1368. 24.V.1997, Cansaran 2306. End. 2804. Vicia cracca L. subsp. cracca L. pratensis L. T. medium L. var. medium 1 km E of Köseler village, steppe; 1000 m, 1 km NE of Küçükçay village, shrubbery; Approx 8 km SE of Göçeri village, under 24.V.1997, Cansaran 2332. Euro.-Sib. El. 1100 m, 08.VI.1997, Cansaran 2406. Euro- forest; 1230 m, 01.VII.1997, Cansaran Sib. El. 2644. V. cracca L. subsp. tenuifolia (Roth) Gaudin L. laxiflorus (Desf.) O.Kuntze subsp. T. pannonicum Jacq. subsp. elongatum 500 m N of Küçükçay village, open degraded laxiflorus (Willd.) Zohary forest; 1130 m, 15.VI.1996, Cansaran 1367. Approx 4 km S of Yemiflen village, forest; Approx 4 km S of Yemiflen village, under V. cracca L. subsp. stenophylla Vell. 1450 m, 02.VII.1997, Cansaran 2777. forest; 1500 m, 02.VI.1997, Cansaran 2728. Approx 2 km W of Güvenli village; 1100 m, L. czeczottianus Bässler End. 19.V.1996, Cansaran1236. Between Ovabafl› and Akp›nar village, T. hirtum All. V. hirsuta (L.) S.F.Gray degraded forest; 1300 m, 24.V.1996, Approx 2 km W of Çitli (Ba¤l›ca) village, 1 km NE of Çitli (Ba¤l›ca) village, shrubbery, Cansaran 2320. End. under shrubbery; 1100 m, 08.VI.1997, 1030 m. 20.VI.1997, Cansaran 2577. *L. sphaericus Retz. Cansaran 2448. Medit. El. V. anatolica Turrill Approx 2 km W of Çitli (Ba¤l›ca) village, T. arvense L. var. arvense shrubbery; 1100 m, 08.VI.1997, Cansaran 1 km E of Köseler village, roadsides; 1000 m, Between E¤mir and Al›c›k, roadsides; 1050 2445. 19.V.1996, Cansaran 1198-B. m, 05.V.1996, Cansaran 1130. Ir.-Tur. El. L. cicera L. Melilotus officinalis (L.) Desr. V. grandiflora Scop. var. grandiflora 1 km E of Köseler village, steppe; 1000 m, Approx 2 km E of Küçükçay village, roadsides; 1 km NE of Çitli (Ba¤l›ca) village, degraded 24.V.1997, Cansaran 2328. 1000 m, 19.V.1996, Cansaran 1167. forest, 1030 m, 20.VI.1997, Cansaran 2571. E. Medit-Euro-Sib. El. L. sativus L. M. alba Desr. V. sativa L. subsp. sativa Between E¤mir and Al›c›k, roadsides; 1000 Köseler village surroundings, E places; 1200 m, 05.V.1996, Cansaran 1124. m, 10.VII.1996, Cansaran 1586-B. Approx 2 km E of Küçükçay village, steppe; 1050 m, 19.V.1996, Cansaran 1165. L. nissolia L. Trigonella fischeriana Ser. *V. sativa L. subsp. incisa (M.Bieb.) Archibald Between Ovabafl› and Akp›nar village, Around Al›c›k, roadsides; 1100 m, var. cordata (Wulfen ex Hoppe) Archibald degraded forest; 1300 m, 24.V.1997, 24.V.1997, Cansaran 2299. Ir.-Tur. El. Cansaran 2314. Approx 2 km W of Çitli (Ba¤l›ca) village, *T. foenum-graecum L. Ononis pusilla L. shrubbery; 1100 m, 08.VI.1997, Cansaran 1 km NE of Küçükçay village, cultivated land; 2439. 300 m W of Büyükçay village, steppe; 1060 1000 m, 20.VII.1996, Cansaran 1834. m, 21.VII.1996, Cansaran 1754. Medit. El. cultivated plant. V. bithynica L. O. spinosa L. subsp. leiosperma (Boiss.) _irj. Medicago lupulina L. Between E¤mir and Al›c›k, roadsides; 1000 m, 05.V.1996, Cansaran 1131. 100 m S of Ovabafl› village, open forests; Approx 2 km W of Büyükçay village, steppe; 1000 m, 03.VIII.1996, Cansaran 2213. 1100 m, 21.VII.1996, Cansaran 1720. V. narbonensis L. var. narbonensis Trifolium repens L. var. repens M. sativa L. subsp. sativa 500 m W of fiamba village, roadsides; 1050 m, 15.VI.1996, Cansaran 1354. W places of Köseler village, open forest; 1300 1 km E of Köseler village, steppe; 1200 m, m, 26.V.1996, Cansaran 1317. 16.VI.1996, Cansaran 1433.
463 The Flora of E¤erli Mountain (Amasya-Turkey)
M. x varia Martyn C. varia L. subsp. varia R. hirtus Waldst. & Kit. 1 km NE of Küçükçay village, degraded Approx 4 km S of Yemiflen village, under Vicinity of Köseler village, E places; 1200 m, forest; 1200 m, 10.VII.1996, Cansaran forest; 1500 m, 02.VII.1997, Cansaran 10.VII.1996, Cansaran 1594. Euro-Sib. El. 1541. 2735. Potentilla recta L. M. coronata (L.) Bartal. Hedysarum varium Willd. Approx 2 km SW of Çetmi village, shrubbery; 1 km NE of Çitli (Ba¤l›ca) village, under 1 km NE of Küçükçay village, steppe; 1150 950 m, 14.VI.1998, Cansaran 2999. (Group shrubbery; 1030 m, 20.VI.1997, Cansaran m, 10.VII.1996, Cansaran 1505. Ir.-Tur. El. A) 2578. Medit. El. Onobrychis armena Boiss. & E.Huet Between Ovabafl› and Akp›nar village, M. minima (L.) Bartal. var. minima degraded forest; 1200 m, 02.VIII.1996, 1 km NE of Küçükçay village, steppe, 1150 Cansaran 1968. (Group B) 300 m SE of Karahisar village, steppe; 1050 m, 10.VII.1996, Cansaran 1507. End. m, 15.VI.1996, Cansaran 1403. S places of Köseler village; 1300 m, ROSACEAE 27.VI.1998, Cansaran 3143. (Group C) M. rigidula (L.) All. var. rigidula Prunus spinosa L. subsp. dasyphylla (Schur) P. micrantha Ramond ex DC. 1 km E of Köseler village, steppe; 1000 m, Domin 24.V.1997, Cansaran 2323. S places of Yukar› Kayabafl› village, open 1 km NE of Küçükçay village, degraded areas; 1600 m, 04.V.1996, Cansaran 1084. M. rigidula (L.) All. var. cinerascens (Jord.) forest; 1200 m, 20.VII.1996, Cansaran Rouy 1658. Euro-Sib. El. Geum urbanum L. 1 km E of Güllüce village, steppe; 1400 m, P. x domestica L. Approx 3 km W of Küçükçay village, 24.V.1997, Cansaran 2345. roadsides; 1020 m, 19.V.1996, Cansaran Approx 10 km SE of Göçeri village, forest; 1142. Euro-Sib. El. Dorycnium graecum (L.) Ser. 1250 m, 01.VII.1997, Cansaran 2689. Agrimonia eupatoria L. Between Ovabafl› and Akp›nar village, P. divaricata Ledeb. subsp. ursina (Kotschy) degraded forest; 1250 m, 02.VIII.1996, Browicz Köseler village surroundings, E places; 1200 Cansaran 1938. Eux. El. m, 10.VII.1996, Cansaran 1590. Approx 5 km NW of K›flladeresi village, forest; D. pentaphyllum Scop. subsp. herbaceum 1400 m, 19.VII.1997, Cansaran 2831. A. repens L. (Vill.) Rouy *Cerasus microcarpa (C.A.Mey.) Boiss. Köseler village surroundings, NE places; 1300 Approx 3 km SW of Çetmi village, under subsp. tortuosa (Boiss. & Hausskn.) Browicz m, 10.VII.1996, Cansaran 1591. degraded forest; 950 m, 14.VI.1998, Approx 4 km SW of Köseler village, Sanguisorba minor Scop. subsp. muricata Cansaran 3028. shrubbery; 1500 m, 25.V.1996, Cansaran (Spach) Briq. D. pentaphyllum Scop. subsp. anatolicum 1280. Ir.-Tur. El. Approx 2 km W of Büyükçay village, steppe; (Boiss.) Gams Filipendula vulgaris Moench 1100 m, 21.VII.1996, Cansaran 1713. Approx 5 km S of Köseler village, under Approx 3 km SW of Çetmi village, degraded Alchemilla mollis (Buser) Rothm. forest; 1330 m, 27.VI.1998, Cansaran 3069. forest; 950 m, 14.VI.1998, Cansaran 2988. Approx 2 km S of Köseler village, stream Lotus corniculatus L. var. corniculatus Euro-Sib. El. bank; 1100 m, 20.VII.1997, Cansaran 2959. Approx 4 km NW of K›flladeresi village, Rubus idaeus L. Rosa foetida Herrm. forest; 1400 m, 19.VII.1997, Cansaran Approx 2 km S of Köseler village, forest; 2855. Approx 5 km E of Büyükçay village, 1160 m, 20.VII.1997, Cansaran 2899. roadsides; 900 m, 19.V.1996, Cansaran Anthyllis vulneraria L. subsp. boissieri R. sanctus Schreb. 1164. Ir.-Tur. El. (Sagorski) Bornm. Approx 4 km NW of Büyükçay village, damp R. canina L. Approx 10 km SE of Göçeri village, under places; 1200 m, 12.VII.1998, Cansaran forest; 1250 m, 01.VII.1997, Cansaran Approx 4 km NW of K›flladeresi village, forest; 3173. 2685. 1400 m, 19.VII.1997, Cansaran 2800. R. discolor Weihe & Nees Coronilla scorpiodies (L.) Koch Cotoneaster nummularia Fisch. & C.A.Mey. 1 km W of Çitli (Ba¤l›ca) village, open 1 km E of Köseler village, steppe; 1200 m, 100 m S of Ovabafl› village, forest; 1200 m, degraded forest; 1100 m, 27.VI.1998, 16.VI.1996, Cansaran 1454. 03.VIII.1996, Cansaran 2216. Cansaran 3168. C. orientalis Mill. var. orientalis Pyracantha coccinea Roem. R. canescens DC. var. canescens Approx 3 km N of Büyükçay village, steppe; 1 km NE of Küçükçay village, shrubbery; Approx 3 km S of Yemiflen village, under 1200 m, 15.VI.1996, Cansaran 1391. 1000 m, 20.VII.1996, Cansaran 1817. forest; 1470 m, 02.VII.1997, Cansaran 2747.
464 A. CANSARAN
*Crataegus x bornmuelleri Zabel E. hirsutum L. Chaerophyllum aureum L. 500 m W of Güllüce village, roadsides; 1470 1 km W of Köseler village, roadsides, humid Approx 4 km S of Yemiflen village, under m, 10.VII.1996, Cansaran 1468. End. places; 1100 m, 10.VII.1996, Cansaran forest; 1480 m, 02.VII.1997, Cansaran 1580. 2767. C. orientalis Pall. ex M.Bieb. var. orientalis E. montanum L. Scandix stellata Banks & Sol. 1 km NE of Küçükçay village, degraded forest; 1200 m, 10.VIII.1996, Cansaran Approx 3 km N of Büyükçay village, damp Güllüce village surroundings, N places, open 1525. places; 1200 m, 15.VI.1996, Cansaran 1388. areas; 1300 m, 05.V.1996, Cansaran 1097. Euro-Sib. El. *C. szovitsii Pojark. S. iberica M.Bieb. CUCURBITACEAE Approx 3 km W of Çitli (Ba¤l›ca) village, 1 km E of Güllüce village, steppe; 1400 m, shrubbery; 1100 m, 08.VI.1997, Cansaran Bryonia alba L. 24.V.1997, Cansaran 2344. 2444. Ir.-Tur. El.? Vicinity of Köseler village, humid places; 1170 S. pecten – veneris L. m, 26.V.1996, Cansaran 1305. Euro-Sib. El. C. monogyna Jacq. subsp. monogyna Between E¤mir and Al›c›k, roadsides; 1100 DATISCACEAE Approx 8 km SE of Göçeri village, forest; m, 05.V.1996, Cansaran 1128. 1250 m, 01.VII.1997, Cansaran 2668. Datisca cannabina L. Bifora radians M.Bieb. C. microphylla C.Koch 1 km W of Köseler village, degraded forest; 1 km E of Köseler village, steppe; 1200 m, 1200 m, 04.VIII.1996, Cansaran 2170. Approx 3 km NW of Akp›nar village, 16.VI.1996, Cansaran 1415. roadsides; 1150 m, 05.V.1996, Cansaran CRASSULACEAE Seseli tortuosum L. 1109. H. Eux. El. Sedum obtusifolium C.A.Mey. 1 km NE of Güllüce village, rocky areas; 1400 Sorbus umbellata (Desf.) Fritsch var. Between Akp›nar and Güllüce village, m, 02.VIII.1996, Cansaran 1981. umbellata shrubbery; 1470 m, 10.VII.1996, Cansaran Conium maculatum L. Approx 3 km SW of Köseler village, degraded 1551. forest; 1500 m, 25.V.1996, Cansaran 1280. S. acre L. Between Akp›nar and Güllüce village, roadsides; 1300 m, 10.VII.1996, Cansaran S. torminalis (L.) Crantz var. torminalis Between Akp›nar and Güllüce village, 1461. roadsides, stony places; 1300 m, 100 m S of Ovabafl› village, forest; 1200 m, 10.VII.1996, Cansaran 1463. Bupleurum rotundifolium L. 03.VIII.1996, Cansaran 2224. Euro-Sib. El.? S. album L. 500 m N of Küçükçay village, open degraded Malus sylvestris Mill. subsp. orientalis forest; 1150 m, 15.VI.1996, Cansaran 1371. (Uglitzk.) Browicz var. orientalis 100 m S of Ovabafl› village, open forests; 1200 m, 03.VIII.1996, Cansaran 2222. B. gerardii All. Around Güllüce village, N places; 1300 m, 05.V.1996, Cansaran 1101-B. S. hispanicum L. var. hispanicum Approx 2 km W of Çitli (Ba¤l›ca) village, open shrubbery; 1100 m, 08.VI.1997, Cansaran Pyrus communis L. subsp. communis 1 km E of Köseler village, steppe; 1260 m, 16.VI.1996, Cansaran 1424. 2450. Approx 2 km S of Köseler village, open Falcaria vulgaris Bernh. forests; 1300 m, 20.VII.1997, Cansaran S. pallidum M.Bieb. var. pallidum 2945. Approx 3 km W of Çitli (Ba¤l›ca) village, Approx 3 km S of Yemiflen village, open degraded forest; 1030 m, 20.VI.1997, forest; 1500 m, 02.VII.1997, Cansaran P. elaeagnifolia Pall. subsp. elaeagnifolia Cansaran 2579. 2718. Between Ovabafl› and Akp›nar village, S. pallidum M.Bieb. var. bithynicum (Boiss.) Peucedanum palimbioides Boiss. shrubbery; 1200 m, 02.VIII.1996, Cansaran Chamb. 1916. 300 m W of Büyükçay village, steppe; 1100 500 m W of Güllüce village, rocky areas; m, 21.VI.1996, Cansaran 1733. End. Ir.-Tur. LYTHRACEAE 1470 m, 10.VII.1996, Cansaran 1469. Eux. El. Lythrum salicaria L. El. Pastinaca sativa L. subsp. urens (Req. ex 1 km E of Köseler village, stream bank; 1000 Sempervivum brevipilum Muirhead Godr.) Celak. m, 21.VII.1996, Cansaran 1863. Euro-Sib. 1 km N of Karahisar village, steppe; 1100 m, Between Ovabafl› and Akp›nar village, El. 10.VII.1996, Cansaran 1572. End. shrubbery; 130 m, 02.VIII.1996, Cansaran 1897. ONAGRACACEAE APIACEAE / UMBELLIFERAE Malabaila secacul Banks & Sol. (Group A) Epilobium angustifolium L. Eryngium campestre L. var. virens Link Approx 2 km SW of Çetmi village, degraded App 2 km S of Köseler village, under forest; 300 m SW of Ovabafl› village, steppe; 950 m, 1200 m, 20.VII.1997, Cansaran 2918. 03.VIII.1996, Cansaran 2036. forest; 970 m, 14.VI.1998, Cansaran 3052.
465 The Flora of E¤erli Mountain (Amasya-Turkey)
Heracleum platytaenium Boiss. S. nigra L. *G. consanguineum Boiss. 500 m W of fiamba village, steppe; 1050 m, 1 km S of Yemiflen village, open forests; 500 m W of Köseler village, steppe; 1160 m, 15.VI.1996, Cansaran 1358, End. Eux. El.? 1530 m, 04.V.1996, Cansaran 1052. Euro- 04.VIII.1996, Cansaran 2202. Ir.-Tur. El. Zosima absinthifolia (Vent.) Link Sib. El. (det. F. Karaer) 1 km W of Küçükçay village, open shrubbery; Viburnum lantana L. G. album Mill. subsp. prusense (C.Koch) 1000 m, 08.VI.1997, Cansaran 2465. W places of Köseler village, roadsides; 1220 Ehrend. & Krendl Laser trilobum (L.) Borkh. m, 26.V.1996, Cansaran 1346. Euro-Sib. El. 100 m S of Ovabafl› village, forest; 1200 m, 03.VIII.1996, Cansaran 2215. 1 km SW of Köseler village, under forest; Lonicera caucasica Pall. subsp. orientalis 1250 m, 04.VIII.1996, Cansaran 2101. (Lam.) Chamb. & Long G. incanum Sm. subsp. elatius (Boiss.) Laserpitium hispidum M.Bieb. Approx 2 km E of Büyükçay village, degraded Ehrend. forest; 1250 m, 10.VII.1996, Cansaran Between Ovabafl› and Akp›nar village, 1 km E of Köseler village, roadsides; 1100 m, 1604. End. degraded forest; 1180 m, 02.VIII.1996, 19.V.1996, Cansaran 1202. Ir.-Tur. El. Cansaran 1988. Euro-Sib. El. L. etrusca Santi var. etrusca G. spurium L. subsp. spurium Torilis leptophylla (L.) Reiche 1 km W of Köseler village, roadsides; 1100 1 km NE of Çitli (Ba¤l›ca) village, degraded Approx 2 km W of Büyükçay village, steppe; m, 10.VII.1996, Cansaran 1574. Medit. El. forest; 1030 m, 20.VI.1997, Cansaran 2562. 1100 m, 21.VII.1996, Cansaran 1721. RUBIACEAE Euro-Sib. El. Astrodaucus orientalis (L.) Drude Crucianella bithynica Boiss. G. tricornutum Dandy 1 km N of Karahisar village, roadsides, steppe Approx 2 km NE of Çitli (Ba¤l›ca) village, Approx 4 km SW of Köseler village, forest; areas; 1100 m, 10.VII.1996, Cansaran 1564. shrubbery; 1030 m, 20.VI.1997, Cansaran 1400 m, 25.V.1996, Cansaran 1240. Medit. Ir.-Tur. El.? 2604. E. Medit. El. El. Caucalis platycarpos L. *C. disticha Boiss. G. tenuissimum M.Bieb. subsp. trichophorum (Kar. & Kir.) Ehrend. Approx 4 km SW of Köseler village, forest; 1 km W of Çitli (Ba¤l›ca) village, shrubbery, 1400 m, 25.V.1996, Cansaran 1242. 1100 m, 27.VI.1998., Cansaran 3161. End. App 2 km NE of Çitli (Ba¤l›ca) village, Turgenia latifolia (L.) H.Hoffm. Ir.-Tur. El. shrubbery; 1060 m, 20.VI.1997, Cansaran 2614. Ir.-Tur. El. 1 km E of Al›c›k, roadsides; 670 m, *C. exasperata Fisch. & C.A.Mey. 19.V.1996, Cansaran 1139. Cruciata taurica (Pall. ex Willd.) Ehrend. Approx 5 km NW of K›flladeresi village, under Artedia squamata L. forest; 1400 m, 19.VII.1997, Cansaran E places of Köseler castle, open areas; 1280 1 km E of Güllüce village, steppe; 1400 m, 2838. Ir.-Tur. El. m, 14.IV.1996, Cansaran 1012. Ir.-Tur. El. 24.V.1997, Cansaran 2336. Asperula lilaciflora Boiss. subsp. phrygia VALERIANACEAE ARALICACEAE (Bornm.) Schönb.-Tem. Centranthus longiflorus Stev. subsp. Hedera helix L. 1 km W of Küçükçay village, steppe; 1050 m, longiflorus 20.VII.1996, Cansaran 1882. End. 1 km W of Köseler village, degraded forest; Approx 5 km N of Karahisar village, 1200 m, 04.VIII.1996, Cansaran 2166. A. involucrata Wahlenb. roadsides; 1100 m, 19.V.1996, Cansaran 1195. Ir.-Tur. El. CORNACEAE Approx 3 km S of Yemiflen village, under forest; 1500 m, 02.VII.1997, Cansaran Valerianella pumila (L.) DC. Cornus sanguinea L. subsp. australis 2710. Eux. El. (C.A.Mey.) Jáv. 1 km E of Güllüce village, steppe; 1400 m, A. arvensis L. 24.V.1997, Cansaran 2366. 1 km E of Köseler village, humid places; 1200 m, 16.VI.1996, Cansaran 1431. Euro-Sib. El. 1 km E of Güllüce village, steppe; 1400 m, V. vesicaria (L.) Moench 24.V.1997, Cansaran 2357. Medit. El. C. mas L. 300 m W of Ovabafl› village, open shrubbery; Galium verum L. subsp. verum. 1 km NE of Küçükçay village, shrubbery; 1100 m, 05.V.1996, Cansaran 1115. 500 m W of Küçükçay village, open 1200 m., 20.VII.1996, Cansaran 1677. MORINACEAE shrubbery; 1130 m, 15.VI.1996, Cansaran Euro-Sib. El.? 1366. Euro-Sib. El. Morina persica L. CAPRIFOLIACEAE G. margaceum Ehrend. & Schönb.-Tem. Approx 3 km N of Büyükçay village, steppe; Sambucus ebulus L. Between Ovabafl› and Akp›nar village, 1200 m, 15.VI.1996, Cansaran 1396. Ir.- 1 km E of Köseler village, steppe; 1200 m, degraded forest; 1250 m, 02.VIII.1996, Tur. El. 16.VI.1996, Cansaran 1448. Euro-Sib. El. Cansaran 1949. End. (det. F. Karaer)
466 A. CANSARAN
DIPSACACEAE F. pyramidata L. *A. teretifolia Willd. Dipsacus laciniatus L. Approx 4 km S of Köseler village, under Between Ovabafl› and Akp›nar village, roadsides; 1100 m, 02.VIII.1996, Cansaran 1 km E of Köseler village, roadsides, damp forest; 1330 m, 27.VI.1998, Cansaran 3067. 1935-A. End. Ir.-Tur. El. places; 1000 m, 21.VII.1996, Cansaran (det. F. Karaer) 1864. Bellis perennis L. A. millefolium L. subsp. pannonica (Scheele) Hayek Scabiosa columbaria L. subsp. ochroleuca E places of Köseler castle, Dereci village, (L.) Celak. var. ochroleuca (L.) Coult. stream bank; 1300 m, 14.IV.1996, Cansaran Approx 3 km S of Yemiflen village, open forests; 1450 m, 02.VII.1997, Cansaran 100 m S of Ovabafl› village, forest; 1200 m, 1009. Euro-Sib. El. 2785. Euro-Sib. El. 03.VIII.1996, Cansaran 2239. Doronicum orientale H.Hoffm. A. biebersteinii Afan. S. argentea L. Approx 8 km SE of Göçeri village, forest; 500 m W of Güllüce village, steppe; 1470 m, 1230 m, 01.VII.1997, Cansaran 2649. 300 m SE of Karahisar village, steppe; 1050 10.VII.1996, Cansaran 1473. m, 15.VI.1996, Cansaran 1398. Ir.-Tur. El. Senecio vernalis Waldst. & Kit. S. micrantha Desf. Tanacetum poteriifolium (Ledeb.) Grierson 1 km E of Köseler village, steppe; 1260 m, Approx 8 km SE of Göçeri village, under Approx 2 km NE of Çitli (Ba¤l›ca) village, 16.VI.1996, Cansaran 1442. degraded forest; 1030 m, 20.VI.1997, forest; 1230 m, 01.VII.1997, Cansaran Tussilago farfara L. Cansaran 2596. 2660. Eux. El. S. rotata M.Bieb. Köseler village surroundings, roadsides; 1170 T. parthenium (L.) Sch. Bip. m, 31.III.1996, Cansaran 1005. Euro-Sib. El. App 3 km N of Büyükçay village, steppe; Approx 8 km S of Göçeri village, forest; 1230 1200 m, 15.VI.1996, Cansaran 1389. Ir.- Petasites hybridus (L.) Gaertn. m, 01.VII.1997, Cansaran 2636. Tur. El. S of Yukar› Kayabafl› village, NE places of T. vulgare L. ASTERACEAE / COMPOSITAE E¤erci hill, humid places; 1600 m, Approx 2 km S of Köseler village, open forest; 04.V.1996, Cansaran 1091, Euro-Sib. El. *Inula helenium L. subsp. orgyalis (Boiss.) 1100 m, 20.VII.1997, Cansaran 2958. Grierson Anthemis cretica L. subsp. albida (Boiss.) Tripleurospermum callosum (Boiss. & Grierson 1 km NE of Güllüce village, stony areas; 1400 Heldr.) E.Hossain m, 02.VIII.1996, Cansaran 1979. End. Eux. Approx 2 km W of Güvenli village, steppe; 1 km E of Köseler village, roadsides; 1100 m, El. 1100 m, 19.V.1996, Cansaran 1237. 21.VII.1996, Cansaran 1868. End. I. salicina L. A. cretica L. subsp. tenuiloba (DC.) Grierson Artemisia absinthium L. Between Ovabafl› and Akp›nar village, Approx 2 km W of Çitli (Ba¤l›ca) village, Approx 2 km S of Köseler village, open degraded forest; 1100 m, 02.VIII.1996, degraded forest; 1100 m, 08.VI.1997, forests, damp places; 1200 m, 20.VII.1997, Cansaran 1918. Euro-Sib. El. Cansaran 2443. Cansaran 2947. I. ensifolia L. A. cretica L. subsp. anatolica (Boiss.) *A. caucasica Willd. Approx 2 km W of Büyükçay village, steppe; Grierson 300 m SE of Karahisar village, stony places; 1100 m, 21.VII.1996, Cansaran 1711. Euro- Approx 5 km N of Karahisar village, steppe; 1050 m, 15.VI.1996, Cansaran 1397. Sib. El. 1100 m, 19.V.1996, Cansaran 1192. *Cirsium osseticum (Adams) Petr. I. oculus-christi L. A. cretica L. subsp. umbilicata (Boiss. & Approx 5 km NW of K›flladeresi village, forest; 1 km NE of Küçükçay village, under E.Huet) Grierson 1450 m, 19.VII.1997, Cansaran 2842. shrubbery; 1100 m, 20.VII.1996, Cansaran 1 km E of Köseler village, steppe; 1280 m, C. hypoleucum DC. 1670. Euro-Sib. El.? 16.VI.1996, Cansaran 1455. Approx 7 km SE of Göçeri village, open I. montbretiana DC. *A. cretica L. subsp. argaea (Boiss. & Ball) forests; 1200 m, 01.VII.1997, Cansaran 500 m W of Güllüce village, rocky areas; Grierson 2701. Eux. El. 1470 m, 10.VII.1996, Cansaran 1474. Ir.- C. pseudopersonata Boiss. & Ball subsp. Tur. El. 1 km E of Güllüce village, steppe; 1400 m, 24.V.1996, Cansaran 2369. End. Ir.-Tur. El. pseudopersonata Helichrysum plicatum DC. subsp. plicatum A. tinctoria L. var. tinctoria Approx 4 km S of Yemiflen village, under 300 m W of Büyükçay village, steppe; 1050 forest; 1470 m, 02.VII.1997, Cansaran m, 21.VII.1996, Cansaran 1752. 100 m S of Ovabafl› village, forest; 1200 m, 2775. End. Eux. El. 03.VIII.1996, Cansaran 2249. Filago eriocephala Guss. C. arvense (L.) Scop. subsp. vestitum Approx 2 km NE of Çitli (Ba¤l›ca) village, open *Achillea phrygia Boiss. & Ball (Wimm. & Grab.) Petr. shrubbery; 1040 m, 20.VI.1997, Cansaran Around Al›c›k, roadsides; 700 m, 24.V.1997, 500 m W of fiamba village, roadsides, steppe; 2610. E. Medit. El. Cansaran 2292. End. Ir.-Tur. El. 1050 m, 15.VI.1996, Cansaran 1357.
467 The Flora of E¤erli Mountain (Amasya-Turkey)
Ptilostemon afer (Jacq.) Greuter subsp. Crupina vulgaris Cass. Tragopogon longirostris Bisch. ex Sch. Bip. eburneus Greuter var. abbreviatus Boiss. 1 km E of Köseler village, steppe; 1000 m, Approx 2 km E of Büyükçay village, steppe; 24.V.1997, Cansaran 2325. 1 km E of Güllüce village, steppe; 1400 m, 1350 m, 10.VII.1996, Cansaran 1611. End. 24.V.1997, Cansaran 2350. C. crupinastrum (Moris) Vis. Carduus nutans L. sensu lato T. coloratus C.A.Mey. Approx 3 km SW of Çetmi village, edge of 1 km E of Köseler village, roadsides; 1000 m, fields; 900 m, 14.VI.1998, Cansaran 3042. 1 km E of Köseler village, steppe; 1280 m, 19.V.1996, Cansaran 1211. 16.VI.1996, Cansaran 1421. Ir.-Tur. El. Cnicus benedictus L. var. benedictus *C. acanthoides L. subsp. acanthoides T. dubius Scop. 1 km N of Karahisar village, eroded places; 1 km N of Karahisar village, open areas; 1100 1100, 10.VII.1996, Cansaran 1565-A. 1 km E of Al›c›k, roadsides; 650 m, m, 10.VII.1996, Cansaran 1560. Euro-Sib. 19.V.1996, Cansaran 1137. Carthamus dentatus Vahl El. T. aureus Boiss. 300 m SW of Ovabafl› village, steppe; 950 m, C. pycnocephalus L. subsp. albidus (M.Bieb.) 03.VIII.1996, Cansaran 2056. 1 km N of Karahisar village, eroded places; Kazmi 1100 m, 10.VII.1996, Cansaran 1569. End. Carlina oligocephala Boiss. & Kotschy subsp. Approx 5 km E of Büyükçay village, oligocephala Leontodon hispidus L. var. hispidus roadsides; 860 m, 19.V.1996, Cansaran Approx 5 km NW of K›flladeresi village, under 1163. 500 m W of Köseler village, steppe; 1160 m, 04.VIII.1996, Cansaran 2179. forest; 1400 m, 19.VII.1997, Cansaran Centaurea virgata Lam. 2799. C. lanata L. 1 km NE of Küçükçay village, steppe; 1150 L. asperrimus (Willd.) Ball m, 10.VII.1996, Cansaran 1502. (Group A) Approx 5 km NW of K›flladeresi village, forest; App 2 km W of Çitli (Ba¤l›ca) village, 1450 m, 19.VII.1997, Cansaran 2845. 1 km NE of Küçükçay village, steppe; 1100 degraded forest; 1100 m, 08.VI.1997, Medit. El. m, 20.VII.1996, Cansaran 1624-A. (Group Cansaran 2443. Ir.-Tur. El. Xeranthemum annuum L. C) L. crispus Vill. subsp. asper (Waldst. & Kit.) C. solsitalis L. subsp. solsitialis 300 m SW of Ovabafl› village, steppe; 950 m, Röhl. var. asper 03.VIII.1996, Cansaran 2054-B. 1 km N of Karahisar village, eroded places; Approx 3 km S of Çetmi village, shrubbery; 1100 m, 10.VII.1996, Cansaran 1565-B Echinops ritro L. 950 m, 14.VI.1998, Cansaran 2995. C. urvillei DC. subsp. stepposa Wagenitz 1 km NE of Küçükçay village, steppe; 1150 Picris strigosa M.Bieb. m, 10.VII.1996, Cansaran 1501. Approx 2 km E of Büyükçay village, steppe; 500 m W of Köseler village, steppe; 1160 m, 1350 m, 10.VII.1996, Cansaran 1612. Ir.- E. galaticus Freyn 04.VIII.1996, Cansaran 2194. Ir.-Tur. El. Tur. El. Between Ovabafl› and Akp›nar village, Sonchus asper (L.) Hill subsp. glaucescens C. carduiformis DC. subsp. carduiformis shrubbery; 1100 m, 02.VIII.1996, Cansaran (Jord.) Ball var. carduiformis. 1966. Eux. El. 1 km NE of Küçükçay village, shrubbery; 1100 m, 08.VI.1997, Cansaran 2427. 1 km NE of Küçükçay village, steppe; 1150 Cichorium intybus L. m, 10.VII.1996, Cansaran 1490. Approx 2 km W of Büyükçay village, steppe; Reichardia glauca Matthews *C. pichleri Boiss. subsp. pichleri 1100 m, 21.VII.1996, Cansaran 1708. Approx 4 km NW of Büyükçay village, steppe; 1200 m, 12.VII.1998, Cansaran 3187, Ir.- Approx 2 km W of Güvenli village, steppe; Scorzonera cana (C.A.Mey.) H.Hoffm. var. Tur. El. 1100 m, 19.V.1996, Cansaran 1221. jacquiniana (W.Koch) Chamb. Hieracium medianiforme (Litv. & Zahn) C. pichleri Boiss. subsp. extrarosularis Approx 2 km W of Güvenli village, steppe; Juxip (Hayek & Siehe) Wagenitz 1100 m, 19.V.1996, Cansaran 1227. App 3 km S of Yemiflen village, forest; 1470 1 km E of Güllüce village, steppe; 1400 m, *S. eriophora DC. m, 02.VII.1997, Cansaran 2762. Eux. El. 24.V.1997, Cansaran 2355. 1 km NE of Güllüce village, rocky areas; 1400 H. labillardierei Arv.-Touv. C. triumfetti All. (Group B) m, 02.VIII.1996, Cansaran 1980. End. S places of Köseler village, under forest; 1300 S. tomentosa L. 1 km E of Al›c›k, roadsides; 650 m, m, 27.VI.1998, Cansaran 3111. 19.V.1996, Cansaran 1138. 1 km. SW of Köseler village, open degraded H. bornmuelleri Freyn C. depressa M.Bieb. forests; 1250 m, 04.VIII.1996, Cansaran 2097. End. Ir.-Tur. El. 1 km NE of Küçükçay village, shrubbery; 1 km E of Büyükçay village, roadsides; 1140 1100 m, 20.VII.1996, Cansaran 1669. End. m, 08.VI.1997, Cansaran 2474.
468 A. CANSARAN
Pilosella hoppeana (Schultes) C.H. & *L. communis L. subsp. grandiflora Legousia falcata (Ten.) Fritsch F.W.Schultz subsp. testimonialis (Nägeli ex (M.Bieb.) P.D.Sell Approx 3 km W of Köseler village, Gökdere, Peter) P.D.Sell & West Approx 3 km S of Ovabafl› village, open shrubbery; 1325 m, 15.VI.1997, Cansaran 500 m NE of Ovabafl› village, forest; 1000 m, degraded forest; 1250 m, 31.V.1998, 2532. Medit. El. 03.VIII.1996, Cansaran 2262. Euro-Sib. El. Cansaran 2965. Eux. El. L. speculum-veneris (L.) Chaix P. hoppeana (Schultes) C.H. & F.W.Schultz Chondrilla juncea L. var. juncea 1 km S of Güvenli village, roadsides; 1160 m, subsp. troica (Zahn) P.D.Sell & West App 2 km W of Büyükçay village, steppe; 15.VI.1996, Cansaran 1377. Medit. El. Approx 5 km NW of K›flladeresi village, under 1100 m, 21.VII.1996, Cansaran 1722. forest; 1400 m, 19.VII.1997, Cansaran L. pentagonia (L.) Thell. Crepis foetida L. subsp. rhoeadifolia 2794-B. Approx 2 km W of Çitli (Ba¤l›ca) village, open (M.Bieb.) Celak. P. x ruprechtii (Boiss.) P.D.Sell & West shrubbery; 1100 m, 08.VI.1997, Cansaran Approx 2 km E of Büyükçay village, steppe; 2436. E. Medit. El. 1 km NE of Küçükçay village, degraded 1350 m, 10.VII.1996, Cansaran 1615. forest; 1000 m, 20.VII.1996, Cansaran ERICACEAE C. sancta (L.) Babcock 1813. Orthilia secunda (L.) House Vicinity of Güllüce village, N slopes, steppe; P. piloselloides (Vill.) Soják subsp. 1300 m, 05.V.1996, Cansaran 1099. Approx 8 km SE of Göçeri village, under piloselloides forest; 1250 m, 01.VII.1997, Cansaran 2672 CAMPANULACEAE Approx 2 km S of Köseler village, forest; (det. H. Duman) 1200 m, 20.VII.1997, Cansaran 2908. Campanula rapunculoides L. subsp. PRIMULACEAE rapunculoides P. piloselloides (Vill.) Soják subsp. Primula vulgaris Huds. subsp. vulgaris megalomastix P.D.Sell & West Between Akp›nar and Güllüce village, degraded forest; 1300 m, 10.VII.1996, Around Yukar› Kayabafl› village, stream bank; Approx 8 km SE of Göçeri village, forest; Cansaran 1460. Euro-Sib. El. 1700 m, 04.V.1996, Cansaran 1061. Euro- 1230 m, 01.VII.1997, Cansaran 2657-A. Sib. El. C. rapunculoides L. subsp. cordifolia P. x fallax (Willd.) Arv.-Touv. (C.Koch) Damboldt Androsace maxima L. App 8 km SE of Göçeri village, forest; 1230 Approx 4 km S of Yemiflen village, under m, 01.VII.1997, Cansaran 2657-B. Vicinity of Köseler village, S places of Saray forest; 1470 m, 02.VII.1997, Cansaran Mountain, open areas; 1320 m, 14.IV.1996, P. echioides (Lumn.) C.H. & F.W.Schultz 2758. Cansaran 1027. subsp. procera (Friese) P.D.Sell & West C. glomerata L. subsp. hispida (Witasek) Cyclamen coum Mill. var. coum Approx 4 km SW of Köseler village, forest; Hayek 1500 m, 25.V.1996, Cansaran 1286. Köseler village surroundings, Gökdere, damp 1 km NE of Küçükçay village, shrub; 1150 m, places; 1170 m, 31.VIII.1996, Cansaran P. x auriculoides (F.A.Láng) P.D.Sell & West 10.VII.1996, Cansaran 1540. Euro-Sib. El. 1006. Approx 2 km W of Çitli (Ba¤l›ca) village, C. involucrata Aucher ex A.DC. Lysimachia verticillaris Spreng. shrubbery; 1100 m, 08.VI.1997, Cansaran 2443. 100 m S of Ovabafl› village, forest; 1200 m, Between Ovabafl› and Akp›nar village, 03.VIII.1996, Cansaran 2221. Ir.-Tur. El. degraded forest; humid places; 1200 m, Scariola viminea (L.) F.W.Schmidt *C. rapunculus L. var. rapunculus 02.VIII.1996, Cansaran 1952. H. Eux. El. Approx 2 km W of Büyükçay village, steppe; Anagallis arvensis L. var. caerulea (L.) 1100 m, 21.VII.1996, Cansaran 1714. Approx 3 km SW of Çetmi village, shrub; 950 m, 14.VI.1998, Cansaran 3005. Gouan S. orientalis (Boiss.) Soják C. rapunculus L. var. lambertiana (A.DC.) 500 m W of fiamba village, roadsides, eroded 300 m W of Büyükçay village, steppe; 1050 Boiss. places; 1050 m, 15.VI.1996, Cansaran 1353. m, 21.VII.1996, Cansaran 1751. Ir.-Tur. El. 1 km E of Köseler village, steppe; 1260 m, OLEACEAE Lapsana communis L. subsp. alpina (Boiss. 16.VI.1996, Cansaran 1438. Euro-Sib. El. Jasminum fruticans L. & Ball) P.D.Sell Asyneuma limonifolium (L.) Janch. subsp. Approx 5 km N of Karahisar village, Approx 4 km NW of K›flladeresi village, pestalozzae (Boiss.) Damboldt roadsides; 1100 m, 19.V.1996, Cansaran forest; 1400 m, 19.VII.1997, Cansaran 1190. Medit. El. 2801. Eux. El. App 3 km S of Köseler village, open forest; 1230 m, 20.VII.1997, Cansaran 2929. End. Ligustrum vulgare L. L. communis L. subsp. intermedia (M.Bieb.) Hayek A. rigudum (Willd.) Grossh. subsp. rigidum Approx 3 km N of Büyükçay village, open areas; 1200 m, 15.VI.1996, Cansaran 1392. Between Ovabafl› and Akp›nar village, 1 km NE of Küçükçay village, degraded Euro-Sib. El. shrubbery; 1230 m, 02.VIII.1996, Cansaran forest; 1150 m, 10.VII.1996, Cansaran 1890. 1519. Ir.-Tur. El.
469 The Flora of E¤erli Mountain (Amasya-Turkey)
CONVOLVULACEAE Rochelia disperma (L.f.) C.Koch var. O. isauricum Boiss. & Heldr. Convolvulus cantabrica L. disperma Approx 7 km SE of Göçeri village, open Approx 3 km N of Büyükçay village, steppe; Approx 3 km S of Ovabafl› village, open forests; 1200 m, 01.VII.1997, Cansaran 1200 m, 15.VI.1996, Cansaran 1394. shrubbery; 1250 m, 31.V.1998, Cansaran 2694. End. Ir.-Tur. El. 2966. C. holosericeus M.Bieb. subsp. holosericeus O. bracteosum Hausskn. & Bornm. Myosotis ramosissima Rochel ex Schultes Approx 2 km W of Büyükçay village, steppe; Approx 4 km W of Küçükçay village, stony subsp. ramosissima 1100 m, 21.VII.1996, Cansaran 1716. places; 1060 m, 19.V.1996, Cansaran 1147. S places of Yukar› Kayabafl› village, steppe End. Ir.-Tur. El. C. holosericeus M.Bieb. subsp. areas; 1600 m, 04.V.1996, Cansaran 1072. macrocalycinus Hausskn. & Bornm. ex O. bourgaei Boiss. Bornm. M. stricta Link ex Roem. & Schultes App 3 km S of Ovabafl› village, open degraded 1 km W of Küçükçay village, steppe; 1050 m, Approx 2 km W of Güvenli village, roadsides; forest; 1250 m, 31.V.1998, Cansaran 2977. 20.VII.1996, Cansaran 1885. End. Ir.-Tur. 1100 m, 19.V.1996, Cansaran 1231. Euro- Ir.-Tur. El. El. Sib. El. O. albo-roseum Fisch. & C.A.Mey. subsp. C. assyricus Griseb. M. arvensis (L.) Hill subsp. arvensis albo-roseum var. albo-roseum 1 km N of Karahisar village, open areas; 1060 Approx 3 km SW of Çetmi village, degraded 1 km E of Köseler village, roadsides, rocky m, 19.V.1996, Cansaran 1152. End. Ir.-Tur. forest; 1000 m, 14.VI.1998, Cansaran 2996. areas; 1000 m, 05.V.1996, Cansaran 1110. El. Euro-Sib. El. Ir.-Tur. El. C. arvensis L. M. lithospermifolia (Willd.) Hornem. O. angustissimum Hausskn. & Bornm. 1 km N of Karahisar village, roadsides; 1050 1 km E of Güllüce village, steppe; 1400 m, Approx 4 km S of Ovabafl› village, steppe m, 19.V.1996, Cansaran 1161. 24.V.1997, Cansaran 2303. areas; 1140 m, 31.V.1998, Cansaran 2979. End. Ir.-Tur. El. C. galaticus Rostan ex Choisy Paracaryum ancyritanum Boiss. O. aucheranum DC. 1 m S of Güvenli village, roadsides; 1160 m, Approx 3 km W of Çitli (Ba¤l›ca) village, 15.VI.1996, Cansaran 1380. End. Ir.-Tur. El. shrubbery; 1100 m, 08.VI.1997, Cansaran 300 m SE of Karahisar village, roadsides; C. betonicifolius Mill. subsp. betonicifolius 2459. End. Ir.-Tur. El. 1050 m, 15.VI.1996, Cansaran 1414. E. Medit. El. 500 m N of Küçükçay village, open degraded Cynoglossum creticum Mill. forest; 1130 m, 15.VI.1996, Cansaran 1362. O. roussaei DC. 1 km NE of Küçükçay village, open degraded CUSCUTACEAE forest; 1100 m, 08.VI.1997, Cansaran 2426. Approx 3 km S of Ovabafl› village, open degraded forests; 1150 m, 24.V.1997, *Cuscuta kotschyana Boiss. subsp. caudata C. montanum L. Bornm. & Schwarz Cansaran 2293. Ir.-Tur. El. App 4 km SW of Köseler village, shrubbery; Cerinthe minor L. subsp. auriculata (Ten.) Approx 5 km NW of K›flladeresi village, 1400 m, 25.V.1996, Cansaran 1284. Euro- Domac forest, on Fabaceae; 1400 m, 19.VII.1997, Sib. El. Cansaran 2809. Ir.-Tur. El Approx 2 km W of Güvenli village, roadsides; Lithospermum officinale L. *C. palaestina Boiss. subsp. balansae 1100 m, 19.V.1996, Cansaran 1222. S places of Köseler village, 1200 m, (Yunck.) Plitmann Anchusa leptophyla Roem. & Schultes 27.VI.1998, Cansaran 3141. Euro-Sib. El. 1 km W of Köseler village, roadsides, on subsp. leptophylla L. purpurocaeruleum L. Salvia; 1100 m, 10.VII.1996, Cansaran 1 km E of Köseler village, open forests; 1100 1583. Approx 2 km SW of Çetmi village, degraded m, 19.V.1996, Cansaran 1208. forest; 950 m, 14.VI.1998, Cansaran 3030. *C. planiflora Ten. A. strigosa Labill. Euro-Sib. El. Around Köseler village, E places, roadsides; Approx 2 km E of Küçükçay village, roadsides; Buglossoides arvensis (L.) I.M.Johnst. on Veronica; 1200 m, 10.VII.1996, Cansaran 1000 m, 19.V.1996, Cansaran 1168. 1598. E places of Köseler castle, open areas; 1200 Alkanna orientalis (L.) Boiss. var. orientalis m, 14.V.1996, Cansaran 1016. BORAGINACEAE E places of Çitli (Ba¤l›ca) village, roadsides; Neatostema apulum (L.) I.M.Johnst. Heliotropium suaveolens M.Bieb. 1000 m, 14.IV.1996, Cansaran 1019. Ir.- N slopes of E¤erli Mountain, Yukar› Kayabafl› 1 km E of Köseler village, roadsides; 1000 m, Tur. El. village surroundings, open areas; 1600 m, 21.VII.1996, Cansaran 1867. E. Medit. El.? SOLANACEAE 04.V.1996, Cansaran 1044. Medit. El. Lappula barbata (M.Bieb.) Gürke Hyoscyamus niger L. Onosma sericeum Willd. Approx 5 km N of Karahisar village, steppe; Approx 3 km W of Küçükçay village, 1 km NE of Küçükçay village steppe; 1150 m, 1100 m, 19.V.1996, Cansaran 1189. Ir.- roadsides; 1000 m, 08.VI.1997, Cansaran 10.VII.1996, Cansaran 1512. Ir.-Tur. El. Tur. El. 2479.
470 A. CANSARAN
SCROPHULARIACEAE S. canina L. subsp. bicolor (Sm.) Greuter Euphrasia pectinata Ten. Verbascum wiedemannianum Fisch. & W places of Köseler village; 1300 m, Approx 4 km S of Yemiflen village, open C.A.Mey. 26.V.1996, Cansaran 1323. E. Medit. El. forests; 1450 m, 02.VII.1997, Cansaran 2789. Euro-Sib. El. Approx 2 km SW of Çetmi village, shrubbery; Anarrhinum orientale Benth. 970 m, 14.VI.1998, Cansaran 3055. End. Pedicularis comosa L. var. sibthorpii 1 km NE of Küçükçay village, steppe; 1100 Ir.-Tur. El. (Boiss.) Boiss. m, 20.VII.1996, Cansaran 1625. Ir.-Tur. El. V. pyramidatum M.Bieb. 1 km E of Güllüce village, steppe; 1400 m, Linaria corifolia Desf. 24.V.1997, Cansaran 2309. Approx 7 km SE of Göçeri village, open 1 km NE of Küçükçay village, steppe; 1150 forest; 1200 m, 01.VII.1997, Cansaran OROBANCHACEAE m, 10.VII.1996, Cansaran 1509. End. Ir.- 2695. H. Eux. El. Tur. El. Orobanche nana Noë ex G.Becker V. caudatum Freyn & Bornm. Digitalis ferruginea L. subsp. ferruginea 1 km E of Köseler village, steppe; 1250 m, Between Akp›nar and Güllüce village, 16.VI.1996, Cansaran 1445. 1 km SW of Köseler village, forest; 1250 m, roadsides; 1300 m, 10.VII.1996, Cansaran 04.VIII.1996, Cansaran 2124. Euro-Sib. El. O. cernua Loefl. 1464. End. Ir.-Tur. El. D. lamarckii Ivan. 1 km SW of Köseler village, forest; 1250 m, *V. bellum Hub.–Mor. 04.VIII.1996, Cansaran 2134. Between Ovabafl› and Akp›nar village, 1 km NE of Küçükçay village, steppe; 1100 degraded forest; 1100 m, 02.VIII.1996, O. alba Stephan m, 20.VII.1996, Cansaran 1655-A. End. E. Cansaran 1911. End. Ir.-Tur. El. Medit. El. W places of Köseler village, Ekinala¤›; 1170 Veronica reuterana Boiss. m, 26.V.1996, Cansaran 1351. V. lasianthum Boiss. ex Benth. E places of Köseler castle, open areas; 1170 O. minor Sm. 1 km SW of Köseler village, under forest; m, 14.IV.1996, Cansaran 1007. Ir.-Tur. El. 1250 m, 04.VIII.1996, Cansaran 2090. App 3 km E of Küçükçay village, damp places; *V. hispidula Boiss. & E.Huet subsp. 1000 m, 21.VII.1996, Cansaran 1869. *V. songaricum Schrenk ex Fisch. & hispidula C.A.Mey. subsp. subdecurrens Hub.-Mor. O. caryophyllacea Sm. 1 km NE of Çitli (Ba¤l›ca) village, shrubbery; Approx 5 km NW of K›flladeresi village, Around Köseler village, E places, roadsides; 1030 m, 20.VI.1997, Cansaran 2566. Ir.- forest; 1400 m, 19.VII.1997, Cansaran 1200 m, 10.VII.1996, Cansaran 1585. Tur. El. 2816. End. Ir.-Tur. El. *O. anatolica Boiss. & Reuter V. arvensis L. V. cherianthifolium Boiss. var. App 2 km SW of Çetmi village, degraded Approx 4 km NW of K›flladeresi village, under cherianthifolium forest; 970 m, 14.VI.1998, Cansaran 3038. forest; 1400 m, 19.VII.1997, Cansaran 300 m W of Büyükçay village, steppe; 1060 2821. Euro-Sib. El. ACANTHACEAE m, 21.VII.1996, Cansaran 1765. V. anagallis- aquatica L. Acanthus hirsutus Boiss. V. cherianthifolium Boiss. var. asperulum Approx 5 km NW of Büyükçay village, steppe (Boiss.) Murb. Approx 4 km SW of Köseler village, stream bank; 1400 m, 25.V.1996, Cansaran 1244. areas; 1200 m, 12.VII.1998, Cansaran 3189. Approx 2 km W of Büyükçay village, steppe; End. 1100 m, 21.VII.1996, Cansaran 1717. End. V. jacquinii Baumg. GLOBULARIACEAE Scrophularia scopolii [Hoppe ex] Pers. var. N places of Güllüce village, steppe; 1300 m, Globularia trichosantha Fisch. & C.A.Mey. adenocalyx Sommer & Lev. 05.V.1996, Cansaran 1094. Euro-Sib. El. 1 km S of Yemiflen village, open forest; 1530 S places of E¤erli Mountain, vicinity of V. multifida L. m, 13.IV.1996, Cansaran 1048. Bo¤ac›k village; 1200 m, 15.VI.1996, 1 km NE of Küçükçay village, steppe; 1100 Cansaran 1373. Eux. El. m, 20.VII.1996, Cansaran 1631. End. Ir.- LAMIACEAE / LABIATAE S. xanthoglossa Boiss. var. decipiens (Boiss. Tur. El. Ajuga orientalis L. & Kotschy) Boiss. V. officinalis L. Vicinity of Güllüce village, open areas; 1300 Approx 2 km N of Karahisar village, humid Approx 8 km SE of Göçeri village, under m, 05.V.1996, Cansaran 1101-A. places; 1080 m, 19.V.1996, Cansaran 1149. forest; 1260 m, 01.VII.1997, Cansaran A chamaepitys (L.) Schreb. subsp. chia Ir.-Tur. El. 2663-A. Euro-Sib. El. (Schreber) Arcang. var. chia S. striata Boiss. Melampyrum arvense L. var. arvense Around Köseler village, S places of Saray 1 km W of Çitli (Ba¤l›ca) village, roadsides; W places of Köseler village, roadsides; 1250 Mountain, open areas; 1350 m, 14.IV.1996, 980 m, 08.VI.1997, Cansaran 2433. Ir.-Tur. m, 26.V.1996, Cansaran 1350. Euro-Sib. El. Cansaran 1032. El.
471 The Flora of E¤erli Mountain (Amasya-Turkey)
Teucrium orientale L. var. orientale Marrubium parviflorum Fisch. & C.A.Mey. *Origanum vulgare L. subsp. hirtum (Link) subsp. parviflorum 1 km W of Çitli (Ba¤l›ca), open shrubbery; Ietsw. 1100 m, 27.VI.1998, Cansaran 3162. Ir.- 1 km N of Karahisar village, steppe; 1100 m, Approx 4 km NW of Büyükçay village, open Tur. El. 10.VII.1996, Cansaran 1559. Ir.-Tur. El. areas; 1200 m, 12.VII.1998, Cansaran 3179. T. orientale L. var. puberulens Ekim M. globosum Montbret & Aucher ex Benth. E. Medit. El. subsp. globosum Approx 3 km S of Yemiflen village, open O. vulgare L. subsp. viride (Boiss.) Hayek forests; 1450 m, 02.VII.1997, Cansaran 1 km N of Karahisar village, roadsides; 1060 E places of Köseler village, open forests; 2781. Ir.-Tur. El. m, 19.V.1996, Cansaran 1154. End. Ir.-Tur. 1270 m, 10.VIII.1996, Cansaran 1600-A. El. T. chamaedrys L. subsp. chamaedrys Satureja wiedemanniana (Lallem.) Velen. Sideritis montana L. subsp. remota (d’Urv.) Approx 5 km NW of K›flladeresi village, forest; P.W.Ball ex Heywood 1 km SW of Köseler village, open forests; 1400 m, 19.VII.1997, Cansaran 2827. Euro- 1250 m, 04.VIII.1996, Cansaran 2096-A. Sib. El. 1 km E of Köseler village, steppe; 1000 m, End. 24.V.1997, Cansaran 2329. E. Medit. El. T. polium L. Clinopodium vulgare L. subsp. vulgare S. amasiaca Bornm. 1 km NE of Küçükçay village, steppe; 1150 500 m N of Küçükçay village, steppe; 1130 m, 10.VII.1996, Cansaran 1513-A. 1 km SW of Köseler village, under forest; 1250 m, 04.VIII.1996, Cansaran 2083. End. m, 15.VI.1996, Cansaran 1372. Scutellaria velenovskyi Rech. Stachys byzantina C.Koch Cansaran vulgare L. subsp. arundanum App 7 km SE of Göçeri village, open forest; (Boiss.) Nyman 1200 m, 01.VII.1997, Cansaran 2698. E. 1 km W of Çitli (Ba¤l›ca) village, roadsides; Medit. El. 980 m, 08.VI.1997, Cansaran 2429. Euro- Approx 2 km S of Köseler village, forest; Sib. El. 1130 m, 20.VII.1997, Cansaran 2891. S. salviifolia Benth. S. iberica M.Bieb. subsp. stenostachya Acinos rotundifolius Pers. App 3 km S of Ovabafl› village, roadsides; (Boiss.) Rech. 1250 m, 31.V.1998, Cansaran 2972. End. Approx 4 km S of Ovabafl› village, roadsides; 1 km NE of Küçükçay village, steppe; 1150 1250 m, 31.V.1998, Cansaran 2973. S. orientalis L. subsp. pinnatifida m, 10.VII.1996, Cansaran 1488. Ir.-Tur. El. Edmondtson Thymus sipyleus Boiss. subsp. rosulans S. annua (L.) L. subsp. annua var. annua (Borbás) Jalas 1 km NE of Küçükçay village, steppe; 1150 m, 10.VII.1996, Cansaran 1497. Approx 4 km S of Ovabafl› village, open 1 km NE of Küçükçay village, steppe; 1100 degraded forest; 1250 m, 31.V.1998, Phlomis pungens Willd. var. pungens m, 08.VI.1997, Cansaran 2394. Cansaran 2968. Between Akp›nar and Güllüce, roadsides; Mentha longifolia (L.) Huds. subsp. S. annua (L.) L. subsp. annua var. Iycaonica 1300 m, 10.VII.1996, Cansaran 1457. longifolia Bhattacharjee P. armeniaca Willd. 1 km E of Köseler village, stream bank; 1000 App 5 km N of Karahisar village, steppe; m, 21.VII.1996, Cansaran 1866. Approx 2 km E of Büyükçay village, steppe; 1100 m, 19.V.1996, Cansaran 1191. Ir.- Ziziphora capitata L. 1350 m, 10.VII.1996, Cansaran 1606. End. Tur. El. Ir.-Tur. El. 500 m N of Küçükçay village, steppe; 1130 Nepeta italica L. Lamium amplexicaule L. m, 15.VI.1996, Cansaran 1364. Ir.-Tur. El. 1 km W of Köseler village, roadsides; 1100 Salvia tomentosa Mill. Güllüce village surroundings, steppe; 1300 m, m, 10.VII.1996, Cansaran 1579. 05.V.1996, Cansaran 1098. Euro-Sib. El. Between Ovabafl› and Akp›nar, degraded N. nuda L. subsp. albiflora (Boiss.) Gams L. purpureum L. var. purpureum forest; 1100 m, 02.VIII.1996, Cansaran Approx 5 km NW of K›flladeresi village, under 1904. Medit. El. N places of E¤erli Mountain, vicinity of Yukar› forest; 1400 m, 19.VII.1997, Cansaran Kayabafl› village, steppe areas; 1500 m, 2825. S. bracteata Banks & Sol. 04.V.1996, Cansaran 1060. Euro-Sib. El. Prunella vulgaris L. 300 m SE of Karahisar village, steppe; 1050 L. album L. m, 15.VI.1996, Cansaran 1411. Ir.-Tur. El. Between Akp›nar and Güllüce, roadsides, Approx 4 km SW of Köseler village, open damp places; 1470 m, 10.VII.1996, Cansaran S. syriaca L. forest; 1500 m, 25.V.1996, Cansaran 1275. 1558. Euro-Sib. El. 1 km N of Karahisar village, roadsides; 1100 Euro-Sib. El. P. laciniata (L.) L. m, 10.VII.1996, Cansaran 1562. Ir.-Tur. El. Wiedemannia orientalis Fisch. & C.A.Mey. Approx 3 km SW of Çetmi village, degraded S. viridis L. Between Al›c›k-E¤mir village, roadsides; 1100 forest; 950 m, 14.VI.1998, Cansaran 3040. 1 km E of Köseler village, steppe; 1000 m, m, 05.V.1996, Cansaran 1126. End. Ir.-Tur. Euro-Sib. El. 24.V.1997, Cansaran 2334. Medit. El. El.
472 A. CANSARAN
S. sclarea L. T. billardieri Boiss. Q. macranthera Fisch. & C.A.Mey. ex Hohen. subsp. syspirensis (C.Koch) Menitsky Between Akp›nar and Güllüce roadsides; 1300 S places of Yukar› Kayabafl› village, open m, 10.VII.1996, Cansaran 1458. areas; 1550 m, 04.V.1996, Cansaran 1075. Approx 3 km W of Köseler village, Gökdere, S. aethiopis L. Ir.-Tur. El. shrubbery, 1300 m, 15.VI.1997, Cansaran 2481. End. App 2 km N of Güvenli village, steppe; 1170 LORANTHACEAE Q pubescens Willd. m, 15.VI.1996, Cansaran 1359. Viscum album L. subsp. album Approx 2 km SW of Çetmi village, degraded S. virgata Jacq. S places of Köseler village, in forest on trees; forest; 950 m, 14.VI.1998, Cansaran 2987. 100 m S of Ovabafl› village, open forests; 1300 m, 27.VI.1998, Cansaran 3149. Q. cerris L. var. cerris 1000 m, 03.VIII.1996, Cansaran 2211. Ir.- EUPHORBIACEAE Tur. El.? 1 km NE of Küçükçay village, shrubbery; Andrachne telephioides L. S. verticillata L. subsp. amasiaca (Freyn & 1250 m, 20.VII.1996, Cansaran 1689. Approx 3 km S of Ovabafl› village, open Bornm.) Bornm. CORYLACEAE shrubbery; 1140 m, 31.V.1998, Cansaran Approx 3 km S of Yemiflen village, open 2981. Carpinus betulus L. forests; 1450 m, 02.VII.1997, Cansaran W places of Köseler village, degraded forest; 2788. Ir.-Tur. El. Euphorbia cardiophylla Boiss. & Heldr. 1250 m, 26.V.1996, Cansaran 1348. Euro- PLUMBAGINACEAE 1 km NE of Küçükçay village, open shrubbery; Sib. El. 1100 m, 08.VI.1998, Cansaran 2404. End. Plumbago europaea L. Corylus avellana L. var. avellana E. szovitsii Fisch. & C.A.Mey. var. szovitsii 100 m S of Ovabafl› village, open forest; 1000 Approx 8 km SE of Göçeri village, forest; m, 03.VIII.1996, Cansaran 2212. Euro-Sib. App 2 km S of Köseler village, stony places; 1250 m, 01.VII.1997, Cansaran 2670. Euro- El. 1200 m, 20.VII.1997, Cansaran 2957. Ir.- Sib. El. Tur. El. Acantholimon acerosum (Willd.) Boiss. var. SALICACEAE acerosum E. falcata L. var. galilaea (Boiss.) Boiss. Populus tremula L. App 2 km E of Büyükçay village, steppe; 1300 1 km N of Karahisar village, open areas; 1100 Approx 4 km SW of Köseler village, forest; m, 10.VII.1996, Cansaran 1614. Ir.-Tur. El. m, 10.VII.1996, Cansaran 1571. 1400 m, 25.V.1996, Cansaran 1269. Euro- A. glumaceum (Jaub. & Spach) Boiss. E. myrsinites L. Sib. El. Approx 4 km NW of K›flladeresi village, forest; E places of Köseler castle, Aliba¤›, open areas; MONOCOTYLEDONES 1400 m, 19.VII.1997, Cansaran 2862. Ir.- 1200 m, 14.IV.1996, Cansaran 1015. LILIACEAE Tur. El. URTICACEAE Allium paniculatum L. subsp. paniculatum PLANTAGINACEAE Urtica dioica L. 1 km NE of Küçükçay village, steppe; 1060 Plantago lanceolata L. Köseler village surroundings, humid places; m, 20.VII.1996, Cansaran 1766. Medit. El. W places of Köseler village, roadsides; 1300 1170 m, 26.V.1996, Cansaran 1302. Euro- (det. M. Koyuncu) m, 26.V.1996, Cansaran 1339. Sib. El. A. atroviolaceum Boiss. ELAEAGNACEAE MORACEAE 1 km NE of Küçükçay village, open shrubbery; Hippophae rhamnoides L. subsp. caucasica Morus rubra L. 1100 m, 08.VI.1997, Cansaran 2421. Rousi 1 km W of Köseler village, degraded forest; A. scorodoprasum L. subsp. rotundum (L.) Approx 3 km W of Küçükçay village, roadside; 1200 m, 04.VIII.1996, Cansaran 2164. Stearn 1000 m, 08.VI.1997, Cansaran 2478. FAGACEAE 1 km W of Köseler village, steppe; 1100 m, Elaeagnus angustifolia L. 10.VII.1996, Cansaran 1578. Medit. El. Quercus robur L. subsp. robur 100 m S of Ovabafl› village, open forests; Ornithogalum oligophyllum E.D.Clarke Approx 3 km W of Köseler village, Gökdere, 1000 m, 03.VIII.1996, Cansaran 2214. N places of E¤erli Mountain, around Yukar› degraded forest; 1275 m, 15.VI.1997, Kayabafl› village, open areas; 1430 m, SANTALACEAE Cansaran 2541. Euro-Sib. El. 04.V.1996, Cansaran 1057. *Thesium bergeri Zucc. Q. hartwissiana Steven O. armeniacum Baker Approx 2 km NE of Çitli (Ba¤l›ca) village, Between Ovabafl› and Akp›nar village, degraded forest; 1030 m, 20.VI.1997, Approx 2 km W of Güvenli village, open areas; degraded forest; 1230 m, 02.VIII.1996, Cansaran 2587. E. Medit. El. 1100 m, 19.V.1996, Cansaran 1223. E. Cansaran 1893. Medit. El.
473 The Flora of E¤erli Mountain (Amasya-Turkey)
Muscari armeniacum Leichtlin ex Baker O. mascula (L.) L. subsp. pinetorum (Boiss. POACEAE / GRAMINEAE & Kotschy) E.G.Camus Between Ovabafl› and Akp›nar, shrubbery; Brachypodium sylvaticum (Huds.) P.Beauv. 1300 m, 24.V.1997, Cansaran 2317. Approx 4 km SW of Köseler village, under Approx 2 km W of Büyükçay village, steppe; M. bourgaei Baker forest; 1400 m, 25.V.1996, Cansaran 1268. 1100 m, 21.VII.1996, Cansaran 1719. Euro- E. Medit. El. (det. E. Sezik) N places of E¤erli Mountain, vicinity of Yukar› Sib. El. Kayabafl› village, open areas; 1700 m, Dactylorhiza romana (Seb.) Soó subsp. *Elymus repens (L.) Gould subsp. 04.V.1996, Cansaran 1041. End. Medit El. romana elongatiformis (Drobow) Melderis (det. M. Koyuncu) 1 km S places of Yemiflen village, forest; 1 km NE of Küçükçay village, steppe; 1150 Gagea villosa (M.Bieb.) Duby var. villosa 1500 m, 04.V.1996, Cansaran 1047. Medit. m, 10.VII.1996, Cansaran 1495. Ir.-Tur. El. El. (det. E.Sezik) E places of Köseler castle, open areas; 1280 *E. hispidus (Opiz) Melderis subsp. hispidus m, 14.IV.1996, Cansaran 1020. Medit. El.? D. romana (Seb.) Soó subsp. georgica (Klán) 1 km NE of Küçükçay village, steppe; 1100 (det. M. Koyuncu) Soó ex Renz & Taub. m, 20.VII.1996, Cansaran 1623. IRIDACEAE 1 km S places of Yemiflen village, forest; *E. hispidus (Opiz) Melderis subsp. 1530 m, 04.V.1996, Cansaran 1046. Eux. Iris kerneriana Asch. & Sint. ex Baker barbulatus (Schur) Melderis El. (det. E. Sezik) Between Ovabafl› and Akp›nar village, 500 m W of Köseler village, steppe; 1160 m, D. osmanica (Kl.) Soó var. osmanica degraded forest; 1200 m, 02.VIII.1996, 04.VIII.1996, Cansaran 2190. Cansaran 1965. End. Euro-Sib. El.? (det. M. 1 km S of Güvenli village; 1160 m, Aegilops triuncialis L. subsp. triuncialis Koyuncu) 15.VI.1996, Cansaran 1383. End. Ir.-Tur. El. Approx 2 km W of Büyükçay village, steppe; Crocus ancyrensis (Herbert) Maw (det. E. Sezik) 1100 m, 21.VII.1996, Cansaran 1704. Köseler village surroundings, roadsides; 1260 JUNCACEAE Ae. biuncialis Vis. m, 31.III.1996, Cansaran 1002. End. Ir.-Tur. Juncus heldreichianus T.Marsson ex Parl. 1 km W of Küçükçay village, steppe; 1050 m, El. subsp. heldreichianus 20.VII.1996, Cansaran 1884. ORCHIDACEAE 1 km NE of Küçükçay village, wet areas; Hordeum bulbosum L. Cephalanthera rubra (L.) Rich. 1100 m, 08.VI.1997, Cansaran 2403. E. Medit. El. 1 km E of Güllüce village, steppe; 1400 m, Approx 8 km SE of Göçeri village, forest; 24.V.1997, Cansaran 2361. 1230 m, 01.VII.1997, Cansaran 2634. (det. Luzula forsteri (Sm.) DC. Taeniatherum caput-medusae (L.) Nevski E. Sezik) App 4-5 km NW of K›flladeresi village, under subsp. crinitum (Schreb.) Melderis C. longifolia (L.) Fritsch forest; 1450 m, 19.VII.1997, Cansaran 2843. Euro-Sib. El. 1 km W of Çitli (Ba¤l›ca) village, open W places of Köseler village, Ekinala¤›, open shrubbery; 1100 m, 27.VI.1998, Cansaran areas; 1350 m, 26.V.1996, Cansaran 1341. CYPERACEAE 3159. Ir.-Tur. El.? Euro-Sib. El. (det. E.Sezik) Eleocharis uniglumis (Link) Schultes Bromus arvensis L. C. damasonium (Mill.) Druce App 5 km E of Büyükçay village, watery S places of Köseler village, under forest; 1290 Approx 4 km SW of Köseler village, under places, 900 m, 19.V.1996, Cansaran 1159. m, 27.VI.1998, Cansaran 3107. forest; 1500 m, 25.V.1996, Cansaran 1290. Carex otrubae Podp. Euro-Sib. El. (det. E. Sezik) B. japonicus Thunb. subsp. japonicus 1 km NE of Küçükçay village, shrubbery; Epipactis helleborine (L.) Crantz 1 km NE of Küçükçay village, steppe; 1150 1200 m, 20.VII.1996, Cansaran 1683. Euro- m, 10.VII.1996, Cansaran 1496. Approx 3 km S of Yemiflen village, forest; Sib. El.? (det. E.Hamzao¤lu) 1470 m, 02.VII.1997, Cansaran 2759. (det. B. tectorum L. C. ovalis Good E. Sezik) Approx 2 km W of Güvenli village, 1100 m, Orchis tridentata Scop. Approx 4 km NW of K›flladeresi village, under 19.V.1996, Cansaran 1214. forest; 1400 m, 19.VII.1997, Cansaran B. tomentellus Boiss. W places of Köseler village, Ekinala¤›, open 2859. Euro-Sib. El. areas; 1300 m, 26.V.1996, Cansaran 1324. 300 m, W of Büyükçay village, steppe; 1100 C. distans L. Medit. El. (det. E. Sezik) m, 21.VII.1996, Cansaran 1694. Ir.-Tur. El. Between Ovabafl› and Akp›nar village, O. purpurea Huds. Trisetum flavescens (L.) P.Beauv. degraded forest; 1200 m, 02.VIII.1996, W places of Köseler village, Ekinala¤› location; Cansaran 1961. Euro-Sib. El. App 5 km NW of K›flladeresi village, under open areas; 1300 m, 26.V.1996, Cansaran forest; 1450 m, 19.VII.1997, Cansaran 1340. Euro-Sib. El. (det. E.Sezik). 2841. Euro-Sib. El.
474 A. CANSARAN
Koeleria cristata (L.) Pers. F. valesiaca Schleid. ex Gaudin Dactylis glomerata L. subsp. hispanica (Roth) Nyman 1 km NE of Çitli (Ba¤l›ca) village, shrubbery; 300 m W of Büyükçay village, steppe; 1100 1030 m, 20.VI.1997, Cansaran 2590. m, 21.VII.1996, Cansaran 1748. Approx 8 km SE of Göçeri village, forest; 1260 m, 01.VII.1997, Cansaran 2655. *Milium vernale M.Bieb. subsp. montianum F. callieri (Hackel ex St.-Yves) Markgr. apud (Parl.) Al.Jahn & Maire Hayek subsp. callieri Briza media L. Approx 8 km SE of Göçeri village, forest; Approx 4 km SW of Köseler village, forest; App 3 km S of Yemiflen village, under forest; 1260 m, 01.VII.1997, Cansaran 2658. 1400 m, 25.V.1996, Cansaran 1252. 1460 m, 02.VII.1997, Cansaran 2737. Medit. El. Lolium perenne L. B. humilis L. Alopecurus arundinaceus Poir. Approx 5 km NW of K›flladeresi village, forest; Between Akp›nar and Güllüce village, E places of Büyükçay village, roadsides; 1060 1400 m, 19.VII.1997, C .2824. Euro-Sib El. roadsides; 1300 m, 10.VII.1996, Cansaran m, 19.V.1996, Cansaran 1155. Euro-Sib. El. 1467. L. rigudum Gaudin var. rigidum A. myosuroides Huds. var. myosuroides Echinaria capitata (L.) Desf. 1 km SW of Köseler village, forest; 1250 m, App 2 km W of Güvenli village, open areas; 500 m W of Köseler village, steppe; 1160 04.VIII.1996, Cansaran 2133. E. Medit. El. 1100 m, 19.V.1996, Cansaran 1234. Euro- mm, 04.VIII.1996, Cansaran 2198. Sib. El. Poa annua L. Melica uniflora Retz. *Phleum alpinum L. S places of Yukar› Kayabafl› village, open Approx 8 km SE of Göçeri village, under areas; 1500 m, 04.V.1996, Cansaran 1079. Between Ovabafl› and Akp›nar, shrubbery; forest; 1260 m, 01.VII.1997, Cansaran 1300 m, 02.VIII.1996, Cansaran 2000. P. trivialis L. 2662. Euro-Sib. El. Euro-Sib. El. Approx 2 km W of Güvenli village, 1100 m, M. ciliata L. subsp. ciliata P. bertolonii DC. 19.V.1996, Cansaran 1215. 1 km W of Köseler village, roadsides; 1100 1 km NE of Küçükçay village, steppe; 1100 P. pratensis L. m, 10.VII.1996, Cansaran 1581. m, 08.VI.1997, Cansaran 2381. Approx 4 km SW of Köseler village, open Stipa holosericea Trin. P. subulatum (Savi) Asch. & Graebn. subsp. forests; 1500 m, 25.V.1996, Cansaran Approx 2 km W of Güvenli village, open areas; subulatum 1279. 1100 m, 19.V.1996, Cansaran 1226. Ir.- Approx 3 km W of Köseler village, degraded P. bulbosa L. Tur. El. forest; 1275 m, 15.VI.1997, Cansaran 2506. Approx 2 km W of Güvenli village, roadsides; S. ehrenbergiana Trin. & Rupr. P. montanum C.Koch subsp. montanum 1100 m, 19.V.1996, Cansaran 1213. Approx 2 km W of Büyükçay village, steppe; 300 m E of Ovabafl› village, forest; 1200 m, 1100 m, 21.VII.1996, Cansaran 1730. Ir.- *Eremopoa persica (Trin.) Roshev. 03.VIII.1996, Cansaran 2252. Tur. El. 1 km NE of Çitli (Ba¤l›ca) village, shrubbery; P. exaratum Hochst. ex Griseb. subsp. Chrysopogon gryllus (L.) Trin. subsp. gryllus 1050 m, 20.VI.1997, Cansaran 2565. Ir.- exaratum Tur. El. 1 km W of Çitli (Ba¤l›ca) village, shrubbery; 300 m W of Büyükçay village, steppe; 1050 1100 m, 27.VI.1998, Cansaran 3160. Catabrosa aquatica (L.) P.Beauv. m, 21.VII.1996, Cansaran 1839. Bothriochloa ischaemum (L.) Keng Approx 4 km SW of Köseler village, watery Festuca jeanpertii (St.-Yves) Markgr. apud places; 1400 m, 25.V.1996, Cansaran 1245. 1 km W of Küçükçay village, steppe; 1060 m, Hayek subsp. jeanpertii 20.VII.1996, Cansaran 1844-B. 8 km SE of Göçeri village, under forest; 1230 m, 01.VII.1997, Cansaran 2635. E. Medit. El.
475