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University Micrdnlms International 300 N INFORMATION TO USERS This reproduction was made from a copy of a document sent to us for microfilming. While the most advanced technology has been used to photograph and reproduce this document, the quality of the reproduction is heavily dependent upon the quality of the material submitted. The following explanation of techniques is provided to help clarify markings or notations which may appear on this reproduction. 1.The sign or “target” for pages apparently lacking from the document photographed is “Missing Page(s)”. If it was possible to obtain the missing page(s) or section, they are spliced into the film along with adjacent pages. This may have necessitated cutting through an image and duplicating adjacent pages to assure complete continuity. 2. 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These prints are available upon request from the Dissertations Customer Services Department. 5. Some pages in any document may have indistinct print. In all cases the best available copy has been filmed. University Micrdnlms International 300 N. Zeeb Road Ann Arbor, Ml 48106 8400300 Smoot, Edith L. PHLOEM ANATOMY AND PHYLOGENY OF SELECTED CARBONIFEROUS FERNS AND PTERIDOSPERMS The Ohio State University Ph.D. 1983 University Microfilms International 300 N. Zeeb Road, Ann Arbor, Ml 48106 PLEASE NOTE: In all cases this material has been filmed in the best possible way from the available copy. Problems encountered with this document have been identified here with a check mark V . 1. Glossy photographs or pages _ 2. Colored illustrations, paper or print_____ 3. Photographs with dark background 4. Illustrations are poor copy______ 5. Pages with black marks, not original copy ______ 6. Print shows through as there is text on both sides of page______ 7. Indistinct, broken or small print on several pages _____ 8. Print exceeds margin requirements ______ 9. Tightly bound copy with print lost in spine_____ 10. Computer printout pages with indistinct print ______ 11. Page(s)___________lacking when material received, and not available from school or author. 12. Page(s)___________seem to be missing in numbering only as text follows. 13. Two pages numbered ____________. Text follows. 14. Curling and wrinkled pages ______ 15. O t h e r _______________________________________________________________ University Microfilms International PHLOEM ANATOMY AND PHYLOGENY OP SELECTED CARBONIFEROUS FERNS AND PTERIDOSPEPMS DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University by Edith L. Smoot, B.S., M.Sc. * * * The Ohio State University 1983 Reading Cctrrnittee: Approved By Thanas N. Taylor Daniel J. Crawford Gary L. Floyd Robert D. Warmbrodt Copyright by Edith L. Smoot 1983 ACKNOWLEDGEMENTS I would like to thank Drs. Robert W. Baxter, Charles W. Good, Gilbert M. Leisman, Gar W. Rothwell, Jeffry T. Schabilion and Benton M. Stidd for their generosity in loaning sane of the specimens used in this study. I would also like to thank my reading connittee for their suggestions, especially Dr. Gary L. Floyd, who was always available for advice. Numerous discussions on all aspects of paleobotany and evolution with Michael A. Cichan were invaluable. This study was carried out under the direction of Dr. Thanas N. Taylor. He has shared his considerable expertise in botany with me over the years, and I would like to ejqpress my appreciation for his continual and generous support, encouragement, advice and friendship. Special thanks to my parents for their encouragement and to my husband, Dick and my daughter, Sarah, for their patience and understanding, especially in the last year. ii VITA August 21, 1 9 5 1 ........... B o m - Ft. Worth, Texas 1969....................... Undergraduate, Washington University St. Louis, Missouri 1971-1972, 1974 ........... Undergraduate, Ohio Wesleyan University Delaware, Ohio 1976....................... B.Sc., The Ohio State University Columbus, Ohio 1977-1978 ................. Research Assistant, Department of Botany The Ohio State University 1977-1978 ................. Teaching Assistant, Department of Botany The Ohio State University 1978....................... M.Sc., The Ohio State University Columbus, Ohio 1978-1979 ................. Research Assistant, Department of Botany The University of Texas Austin, Texas 1979-1982 ................. Teaching Assistant, Department of Botany The Ohio State University 1979-1982 ................. Research Assistant, Department of Botany The Ohio State University 1982-1983 ................. Graduate Fellow American Association of University Wcmen PROFESSIONAL AFFILIATIONS Botanical Society of America Paleabotanical Section Pteridological Section American Institute of Biological Sciences American Association for the Advancement of Science Palaeontological Association iii International Organization of Paleobotany Ohio Academy of Science The Society of the Sigma Xi Sigma Delta Epsilon, Graduate Women in Science International Association of Wood Anatomists Phi Kappa Phi Fellow, Linnean Society of London RESEARCH INTERESTS Fossil phloem— anatomy and phylogenetic trends Anatomy and morphology of Carboniferous vascular plants Investigations of Permian and Triassic plants from Antarctica Paleoecology of Permian petrified peat from Antarctica AWARDS AND HONORS 1975 Summa Award University College, The Ohio State University Recognition of superior scholarship 1976 The Arts and Sciences Award Colleges of the Arts and Sciences The Ohio State University Recognition of scholarship 1978 Isabel Cookson Award Paledbotanical Section Botanical Society of America Best student paper at annual meeting 1981 Elected a fellow Linnean Society of London 1982 Fellow American Association of University Women 1983 Best student paper at annual meeting Plant Sciences Section Ohio Academy of Science 1983 Graduate School Leadership Award The Ohio State University RESEARCH GRANTS 1979 Sigma Xi Grant-in-Aid of Research Project Title: Phloem Anatomy of Pennsylvanian Age Plants $250 iv 1979 National Science Foundation Doctoral Dissertation Research in the Field Sciences Project Title: Phloem Anatomy of Pennsylvanian Age Plants $3789 1982 Sigma Xi Grant-in-Aid of Research Project Title: Phloem Anatomy and Phylogenetic Trends in Selected Pennsylvanian Age Plants $400 1982 Dissertation Fellowship Educational Foundation American Association of University Wfcmen July, 1982 - June, 1983 $7000 1983 Graduate Student Alumni Research Award Graduate School, The Ohio State University Project Title: Phloem Anatomy and Evolution of Pennsylvanian Age Plants $400 1983 National Science Foundation Two-Year and Four-Year College Research Instrumentation Program Project Title: Purchase of Optical and Photographic Equipment for Research in Paleobotany $34,053.50 (Pending) UNIVERSITY SERVICE Department: Graduate Committee (Student Representative) 1979-1980 Faculty Meetings (Student Representative) 1979-1980 Chairman, Waller Lecture Committee 1977-1978 Member, Waller Lecture Committee 1979-1980 Colloquium Committee (Student Representative) 1980-1981 Space and Facilities Committee (Student Representative) 1981-1982 Research Committee (Student Representative) 1982-1983 College: Student Selection Committee for Position of Dean of Biological Sciences 1977-1978 University: Graduate Council, Student Representative for Biological Sciences College Research Committee 1980-1981 PUBLICATIONS BOOKS 1983a Paleobotany: Volume I— Precambrian through Permian. Benchmark Papers in Systematic and Evolutionary Biology. Hutchinson Ross Publishing Co. (in press). Eds. T.N. Taylor and E.L. Shoot. v BOOKS (Cont.) 1983b Paleobotany: Volume II— Triassic through Pliocene. Benchmark Papers in Systematic and Evolutionary Biology. Hutchinson Ross Publishing Co. (in press). Eds. T.N. Taylor and E.L. Smoot. PAPERS 1978a Phloem in the Lower Pennsylvanian fern Etapteris. ( M o J. Sci. 78: 21 (Abstract). 1978b A new species of coenopterid fern from the Pennsylvanian of North America. Ohio J. Sci. 78: 22 (Abstract) (E.L. Smoot and T.N. Taylor). 1978c Etapteris leclercqii sp. n., a zygopterid fern frcm the Pennsylvanian of North America. Amer. J. Bot. 65: 571-576 (E.L. Smoot and T.N. Taylor). 1978d Sieve areas in fossil phloem. Science 202: 1081-1083 (E.L. Smoot and T.N. Taylor). 1978e Phloem in the Lower Pennsylvanian fern Etapteris. Bot. Soc. Amer. Misc. Ser. Publ. 156: 31 (Abstract). 1979a The phloem of Etapteris leclercqii and Botryopteris tridentata. Amer. J. Bot. 66: 511-521. 1979b Phloem in two Pennsylvanian ferns. Bot. Soc. Amer. Misc. Ser. Publ. 157:
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