A Revised Bibliography of the Psocoptera (Arthropoda: Insecta)
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Historical Biogeography of Thyrsophorini Psocids and Description of a New Neotropical Species of Thyrsopsocopsis (Psocodea: Psocomorpha: Psocidae)
European Journal of Taxonomy 194: 1–16 ISSN 2118-9773 http://dx.doi.org/10.5852/ejt.2016.194 www.europeanjournaloftaxonomy.eu 2016 · Román-Palacios C. et al. This work is licensed under a Creative Commons Attribution 3.0 License. Research article urn:lsid:zoobank.org:pub:96E9EA43-F6FE-492E-97BE-60DFB8EDE935 Historical biogeography of Thyrsophorini psocids and description of a new neotropical species of Thyrsopsocopsis (Psocodea: Psocomorpha: Psocidae) Cristian ROMÁN-PALACIOS 1,*, Alfonso N. GARCÍA ALDRETE 2 & Ranulfo GONZÁLEZ OBANDO 3 1,3 Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, Colombia. 2 Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apartado Postal 70-153, 04510 Mexico City, Mexico. * Corresponding author: [email protected] 1 urn:lsid:zoobank.org:author:E88D0518-B6CB-4FE7-9EFC-F789EA6F05AD 2 urn:lsid:zoobank.org:author:9E03B921-78AE-4ED6-B1EA-9DCA01BE20BC 3 urn:lsid:zoobank.org:author:16C7AD76-F035-4C8B-8C00-A228CCCD39B0 Abstract. When based on phylogenetic proposals, biogeographic historic narratives have a great interest for hypothesizing paths of origin of the current biodiversity. Among the many questions that remain unsolved about psocids, the distribution of Thyrsophorini represents still a remarkable enigma. This tribe had been considered as exclusively Neotropical, until the description of Thyrsopsocopsis thorntoni Mockford, 2004, from Vietnam. Three hypotheses have been proposed to explain this atypical distribution, recurring to dispersal, vicariance and morphological parallelism between lineages, but the lack of evidence has not allowed a unique support. Here, we describe a new Neotropical species of Thyrsopsocopsis, and also attempt to test the three biogeographical hypotheses in a phylogenetic context. -
Psocopteran Species Associated with Eastern Hemlock in the Southern Appalachians
224 Florida Entomologist (95)1 March 2012 PSOCOPTERAN SPECIES ASSOCIATED WITH EASTERN HEMLOCK IN THE SOUTHERN APPALACHIANS CARLA COOTS1,2, PARIS LAMBDIN1, JEROME GRANT1, RUSTY RHEA3, AND EDWARD MOCKFORD4 1Department of Entomology and Plant Pathology, The University of Tennessee, Knoxville, TN 37996 2Corresponding author, E-mail: [email protected] 3USDA Forest Service, Forest Health Protection, 200 Weaver Boulevard, Asheville, NC 28804. 4School of Biological Sciences, Illinois State University, Normal, IL 61790-4120. Eastern hemlock, Tsuga canadensis (L.) Car- collected using direct sampling (beat sheet, hand- rière, comprises a vital component of biological picking, and trunk vacuuming) with collected diversity (Jordan & Sharp 1967; Lapin 1994; Tin- specimens placed into labeled (date, site and tree gley et al. 2002; Buck 2004; Buck et al. 2005; Dill- location, sampling type) alcohol-filled vials and ing et al. 2007; Dilling et al. 2009) and economi- taken to the laboratory for identification. Beat- cal (Travel Industry Association 2006; Woodsen sheet samples (4 per tree) were taken at each 2001) and environmental stability (Evans 2002; direction by striking each branch 5 times with Snyder et al. 2004) within its geographical range. a one-meter rod, while visual observations were Hemlock woolly adelgid, Adelges tsugae Annand conducted on each tree for 5 min per stratum, and (Hemiptera: Adelgidae), is an exotic insect species direct trunk vacuuming occurred on 61 cm of the capable of rapidly reducing populations of eastern circumference of the tree’s trunk. hemlock throughout the eastern United States During this study, 3,740 adult and nymph pso- (McClure & Fergione 1977; Buck et al. 2005; copteran specimens were collected and identified, Ellison et al. -
BÖCEKLERİN SINIFLANDIRILMASI (Takım Düzeyinde)
BÖCEKLERİN SINIFLANDIRILMASI (TAKIM DÜZEYİNDE) GÖKHAN AYDIN 2016 Editör : Gökhan AYDIN Dizgi : Ziya ÖNCÜ ISBN : 978-605-87432-3-6 Böceklerin Sınıflandırılması isimli eğitim amaçlı hazırlanan bilgisayar programı için lütfen aşağıda verilen linki tıklayarak programı ücretsiz olarak bilgisayarınıza yükleyin. http://atabeymyo.sdu.edu.tr/assets/uploads/sites/76/files/siniflama-05102016.exe Eğitim Amaçlı Bilgisayar Programı ISBN: 978-605-87432-2-9 İçindekiler İçindekiler i Önsöz vi 1. Protura - Coneheads 1 1.1 Özellikleri 1 1.2 Ekonomik Önemi 2 1.3 Bunları Biliyor musunuz? 2 2. Collembola - Springtails 3 2.1 Özellikleri 3 2.2 Ekonomik Önemi 4 2.3 Bunları Biliyor musunuz? 4 3. Thysanura - Silverfish 6 3.1 Özellikleri 6 3.2 Ekonomik Önemi 7 3.3 Bunları Biliyor musunuz? 7 4. Microcoryphia - Bristletails 8 4.1 Özellikleri 8 4.2 Ekonomik Önemi 9 5. Diplura 10 5.1 Özellikleri 10 5.2 Ekonomik Önemi 10 5.3 Bunları Biliyor musunuz? 11 6. Plocoptera – Stoneflies 12 6.1 Özellikleri 12 6.2 Ekonomik Önemi 12 6.3 Bunları Biliyor musunuz? 13 7. Embioptera - webspinners 14 7.1 Özellikleri 15 7.2 Ekonomik Önemi 15 7.3 Bunları Biliyor musunuz? 15 8. Orthoptera–Grasshoppers, Crickets 16 8.1 Özellikleri 16 8.2 Ekonomik Önemi 16 8.3 Bunları Biliyor musunuz? 17 i 9. Phasmida - Walkingsticks 20 9.1 Özellikleri 20 9.2 Ekonomik Önemi 21 9.3 Bunları Biliyor musunuz? 21 10. Dermaptera - Earwigs 23 10.1 Özellikleri 23 10.2 Ekonomik Önemi 24 10.3 Bunları Biliyor musunuz? 24 11. Zoraptera 25 11.1 Özellikleri 25 11.2 Ekonomik Önemi 25 11.3 Bunları Biliyor musunuz? 26 12. -
Acari: Prostigmata: Cunaxidae
360 North-Western Journal of Zoology 13(2) / 2017 Kaczmarek, Ł., Diduszko, D., Michalczyk, Ł. (2011): New records of small arthropods (Skvarla et al. 2014). Addition- Mexican Tardigrada. Revista Mexicana de Biodiversidad 82: ally, some species can also feed on honeydew pro- 1324-1327. Kaczmarek, Ł., Jakubowska, N., Michalczyk, L. (2012): Current duced by their host plant (Walter & Proctor 1999). knowledge on Turkish Tardigrades with a description of The genus Cunaxa was defined by Von Hey- Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: den in 1826 with type species Scirus setirostris Milnesiidae, the granulatum group). Zootaxa 3589: 49-64. Kaczmarek, Ł., Michalczyk, Ł., McInnes, S.J. (2014): Annotated Hermann 1804 (Von Heyden 1826). It is the largest zoogeography of non-marine Tardigrada. Part I: Central in sub-family Cunaxinae Oudemans with ap- America. Zootaxa 3763(1): 1-107. proximately 50 valid species (Sergeyenko 2009, Maucci, W. (1978): Tardigradi muscicoli della Turchia (terzo contributo). Bollettino Museo civico Storia naturale 5: 111-140. Skvarla et al. 2014). And can be separated from McInnes, S. (1994): Zoogeographic distribution of other Cunaxinae genera by the following charac- terrestrial/freshwater tardigrades from current literature. ters: dorsal shields not reticulated, prodorsal Journal of Natural History 28: 257-352. Michalczyk, Ł., Kaczmarek, Ł. (2003): A description of the new shield smooth or striated, five segmented pedi- tardigrade Macrobiotus reinhardti (Eutardigrada, Macrobiotidae, palps, elongate apophyses or spine-like setae on harmsworthi group) with some remarks on the oral cavity inner margin of telofemur, genu, tibiotarsus, setal armature within the genus Macrobiotus Schultze. Zootaxa 331: 1- 24. formula of coxae II-IV 1-3-2 and long, slender, at- Michalczyk, L., Kaczmarek, L., Weglarska, B. -
Survs; “‘1’ 03‘ the of MECHEGAN
SURVs; “‘1’ 03‘ THE PSEUQOSCORPIONS OF MECHEGAN 1959 WI“NH“\llUllNHlH“llllH“|H21|1|NH||NL|W _ 3 1293 10402 LIB R A R Y Mi chig nState Uni! rsity SURVEY OF THE PSEUDOSCOFPIONS OF MICHIGAN By JOSEPH D. FFNSTWRMACHWR AN ABSTRACT Submitted to the School of Graduate Studies of Michigan State University in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE Deoartment of Zoology 1959 Approvedjflw_ fizz ABSTRACT This study was conducted as a preliminary survey of the species of pseudoscornions found in Michigan end their habi- tat preferences. Apnroximately 1450 specimens were collected by the author. Around 500 additional snecimens were made available for this survey from institutional and nersonal collections. Records published prior to this study list four species of pseudoscorpions in the state. With the nresent survey the list has been increased to include twelve species, ten genera, five families, and three suborders. Three of the four previously recorded species were taken at new locali- ties. An accurate species determination could not be made for six specimens. They appear to demonstrate the presence of three or four additional genera and species. Pseudoscorpions have varied habitat preferences. Four species were found in the leaf litter and ground cover of deciduous forests: ghfhonius tetraghelgtus, pactvlocheljfer cooiosus, Microbisium confusum, and Fselanhochernes oarvus. The latter two were taken also in rotten loss and wood de- bris. Microbisium brunneum_was collected from Sphagnum moss ”no -—-—.—_ and bog debris. Several species were taken near the habits- tions of man: Ehelifer cancroides in an abandoned shack; Lamnrocherngs minor and Eaispghelifer callus in a grain bin. -
In Alpinen Wald-Ökosystemen Südtirols Auf Den Dauerbeobachtungsflächen IT01 Ritten Und IT02 Montiggl Im Jahre 2006
forest observer vol.4 2008 249 - 292 Biomonitoring der Zikadenfauna (Auchenorrhyncha) in alpinen Wald-Ökosystemen Südtirols auf den Dauerbeobachtungsflächen IT01 Ritten und IT02 Montiggl im Jahre 2006 Michael Carl Abstract Ecological assessment in alpine forest ecosystems: Biomonitoring of and bioindication by the leafhopper-fauna (Auchenorrhyncha) at monitoring sites in northern Italy Concerning leafhopper communities the Ritten (IT01) and Montiggl (IT02) have become one of the most intensively explored mountain massifs of South Tyrol. From 1996 to 2006 more than 4900 individuals out of 81 species were collected. The use of leafhoppers as a management tool for monitoring status and change in forest ecosystems is critically evaluated. The fauna of both sites is dominated by a characteristic set of partially strongly specialized species. Species turnover as well as evaluation of precipitation and temperature of the monitoring sites show a strong reaction of the leafhopper communities on effects probably caused by climate change. Keywords: Leafhoppers, Auchenorrhyncha, species communities, mountaineous forests, climate change, Alps, Italy 1 Einleitung 1.1 Veranlassung und Fragestellung Seit einigen Jahren haben durch verschiedenste Individuenzahl in Grünland‑ und Waldhabitaten Forschungsprojekte die Zikaden (Insecta, Auchenor‑ vor und besiedeln deren gesamte dreidimensionale rhyncha) eine wachsende Bedeutung für die Bioin‑ Struktur von der Wurzel bis zur Baumspitze. Zikaden dikation erlangt. Diese Insektenordnung von Pflan‑ sind daher als -
Psocoptera Em Cavernas Do Brasil: Riqueza, Composição E Distribuição
PSOCOPTERA EM CAVERNAS DO BRASIL: RIQUEZA, COMPOSIÇÃO E DISTRIBUIÇÃO THAÍS OLIVEIRA DO CARMO 2009 THAÍS OLIVEIRA DO CARMO PSOCOPTERA EM CAVERNAS DO BRASIL: RIQUEZA, COMPOSIÇÃO E DISTRIBUIÇÃO Dissertação apresentada à Universidade Federal de Lavras, como parte das exigências do programa de Pós-Graduação em Ecologia Aplicada, área de concentração em Ecologia e Conservação de Paisagens Fragmentadas e Agroecossistemas, para obtenção do título de “Mestre”. Orientador Prof. Dr. Rodrigo Lopes Ferreira LAVRAS MINAS GERAIS – BRASIL 2009 Ficha Catalográfica Preparada pela Divisão de Processos Técnicos da Biblioteca Central da UFLA Carmo, Thaís Oliveira do. Psocoptera em cavernas do Brasil: riqueza, composição e distribuição / Thaís Oliveira do Carmo. – Lavras : UFLA, 2009. 98 p. : il. Dissertação (mestrado) – Universidade Federal de Lavras, 2009. Orientador: Rodrigo Lopes Ferreira. Bibliografia. 1. Insetos cavernícolas. 2. Ecologia. 3. Diversidade. 4. Fauna cavernícola. I. Universidade Federal de Lavras. II. Título. CDD – 574.5264 THAÍS OLIVEIRA DO CARMO PSOCOPTERA EM CAVERNAS DO BRASIL: RIQUEZA, COMPOSIÇÃO E DISTRIBUIÇÃO Dissertação apresentada à Universidade Federal de Lavras, como parte das exigências do programa de Pós-Graduação em Ecologia Aplicada, área de concentração em Ecologia e Conservação de Paisagens Fragmentadas e Agroecossistemas, para obtenção do título de “Mestre”. APROVADA em 04 de dezembro de 2009 Prof. Dr. Marconi Souza Silva UNILAVRAS Prof. Dr. Luís Cláudio Paterno Silveira UFLA Prof. Dr. Rodrigo Lopes Ferreira UFLA (Orientador) LAVRAS MINAS GERAIS – BRASIL ...Então não vá embora Agora que eu posso dizer Eu já era o que sou agora Mas agora gosto de ser (Poema Quebrado - Oswaldo Montenegro) AGRADECIMENTOS A Deus, pois com Ele nada nessa vida é impossível! Agradeço aos meus pais, Joaquim e Madalena, pela oportunidade e apoio. -
ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000). -
Morphology of Psocomorpha (Psocodea: 'Psocoptera')
Title MORPHOLOGY OF PSOCOMORPHA (PSOCODEA: 'PSOCOPTERA') Author(s) Yoshizawa, Kazunori Insecta matsumurana. New series : journal of the Faculty of Agriculture Hokkaido University, series entomology, 62, 1- Citation 44 Issue Date 2005-12 Doc URL http://hdl.handle.net/2115/10524 Type bulletin (article) File Information Yoshizawa-62.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP INSECTA MATSUMURANA NEW SERIES 62: 1–44 DECEMBER 2005 MORPHOLOGY OF PSOCOMORPHA (PSOCODEA: 'PSOCOPTERA') By KAZUNORI YOSHIZAWA Abstract YOSHIZAWA, K. 2005. Morphology of Psocomorpha (Psocodea: 'Psocoptera'). Ins. matsum. n. s. 62: 1–44, 24 figs. Adult integumental morphology of the suborder Psocomorpha (Psocodea: 'Psocoptera') was examined, and homologies and transformation series of characters throughout the suborder and Psocoptera were discussed. These examinations formed the basis of the recent morphology-based cladistic analysis of the Psocomorpha (Yoshizawa, 2002, Zool. J. Linn. Soc. 136: 371–400). Author's address. Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060-8589 Japan. E-mail. [email protected]. 1 INTRODUCTION Psocoptera (psocids, booklice or barklice) are a paraphyletic assemblage of non-parasitic members of the order Psocodea (Lyal, 1985; Yoshizawa & Johnson, 2003, 2005; Johnson et al., 2004), containing about 5500 described species (Lienhard, 2003). They are about 1 to 10 mm in length and characterized by well-developed postclypeus, long antennae, pick-like lacinia, reduced prothorax, well-developed pterothorax, etc. Phylogenetically, Psocoptera compose a monophyletic group (the order Psocodea) with parasitic lice ('Phtiraptera': biting lice and sucking lice) (Lyal, 1985; Yoshizawa & Johnson, 2003, in press; Johnson et al., 2004). The order is related to Thysanoptera (thrips) and Hemiptera (bugs, cicadas, etc.) (Yoshizawa & Saigusa, 2001, 2003, but see also Yoshizawa & Johnson, 2005). -
Reserva De La Biosfera Montes Azules, Selva Lacandona; Investigacion Para Su Conservacion
RESERVA DE LA BIOSFERA MONTES AZULES, SELVA LACANDONA; INVESTIGACION PARA SU CONSERVACION Editado por Miguel Angel Vásquez Sánchez y Mario A. Ramos Olmos PUBUCACIONES ESPECIALES ECOSFERA No. 1 Centro de Estudios para la Conservación de los Recursos Naturales, A. C. Centro de Estudios para la Conservación de los Recursos Naturales, A.C. -ECOSFERA- Este Centro fue fundado en 1989, con los objetivos de promover y realizar acciones orientadas al aprovechamiento sostenido y restauración de los recursos naturales, a la investigación sobre la diversidad biológica, el impacto de las actividades humanas en las áreas silvestres y al manejo de aquellas de importancia biológica. Los miembros del Centro trabajan jjermanentemente en el forta lecimiento de un grupo multidisciplinario, con capacidad de generar la información necesaria para resolver problemas locales y regionales desde una perspectiva integral. Adicionalmente tiene como objetivos, la for mación y capacitación de recursos humanos, así como la difusión de la información gene rada en sus investigaciones. Sus programas de investigación abarcan: Estudios del Me dio Físico, Conservación de Especies Ame nazadas y en Peligro de Extinción, Manejo y Aprovechamiento de Fauna Silvestre, Pla nificación y Manejo de Areas Silvestres, De sarrollo Comunitario y Conservación. Fotos de portada: Foto superior izquierda: Ilach Winik (H om bre verdadero). Bonampak (Foto: M. A. Vás quez) Foto superior derecha: Rana arborícola Hyla ebraccata (Foto; R.C. Vogt) Foto inferior derecha: Jaguar {Panthera onca). Foto; J.L. Patjane Foto inferior izquierda; Niños lacandones (Foto; L J. M arch) RESERVA DE LA BIOSFERA MONTES AZULES, SELVA LACANDONA: INVESTIGACION PARA SU CONSERVACION EC/333.711/R4/EJ. -
Pseudoscorpions
Colorado Arachnids of Interest Pseudoscorpions Class: Arachnida Order: Pseudoscorpiones Identification and Descriptive Features: Pseudoscorpions are tiny arachnids (typically Figure 1. Pseudoscorpion ranging from 1.25-4.5 mm body length). They possess pedipalps modified into pincers in a manner similar to scorpions. However, they differ in other features, notably possessing a broad, flattened abdomen that lacks the well developed tail and stinger. Approximately 200 species of pseudoscorpions have been described from North America. A 1961 review of pseudoscorpions within Colorado listed 30 species; however, these arachnids have only rarely been subjects for collection so their occurrence and distribution within Colorado is poorly known. The pseudoscorpion most often found within buildings is Chelifer cancroides, sometimes known as the “house pseudoscorpion”. It is mahogany brown color with a body length of about 3-4 mm and long pedipalps that may spread 8 mm across. Distribution in Colorado: Almost all pseudoscorpions that occur in Colorado are associated with forested areas although a few prairie species do occur. Conifer forests, including scrublands of pinyon and juniper, support several species. Others occur in association with Gambel oak and aspen. The house pseudoscorpion has an unusually broad distribution and is found associated with human dwellings over wide areas of North America and Europe. Life History and Habits: Pseudoscorpions usually occur under rocks, among fallen leaves or needles, under bark or similar moist sites where they hunt mites, springtails and small insects. Typically they wait in ambush within small crevices and grab passing prey with the pincers. In most species, connected to the movable “finger” of the pincer is a venom gland. -
Vertical Stratification and Co-Occurrence Patterns of The
Forests 2012, 3, 127-136; doi:10.3390/f3010127 OPEN ACCESS forests ISSN 1999–4907 www.mdpi.com/journal/forests Article Vertical Stratification and Co-Occurrence Patterns of the Psocoptera Community Associated with Eastern Hemlock, Tsuga canadensis (L.) Carrière, in the Southern Appalachians Carla Coots 1,*, Paris Lambdin 1, Jerome Grant 1, Rusty Rhea 2 and Edward Mockford 3 1 Department of Entomology and Plant Pathology, The University of Tennessee, Knoxville, TN 37996, USA; E-Mails: [email protected] (P.L.); [email protected] (J.G.) 2 USDA Forest Service, Forest Health Protection, 200 Weaver Boulevard, Asheville, NC 28804, USA; E-Mail: [email protected] 3 School of Biological Sciences, Illinois State University, Normal, IL 61790, USA; E-Mail: [email protected] * Author to whom correspondence should be addressed; E-Mail: [email protected]; Tel.: +1-865-974-4979; Fax: +1-865-974-4744. Received: 10 February 2012; in revised form: 1 March 2012 / Accepted: 7 March 2012 / Published: 21 March 2012 Abstract: Of the more than 300 species of Psocoptera described in North America, 44 species have been documented on eastern hemlock, Tsuga canadensis (L.) Carrière, in the southern Appalachians. However, the distribution and co-occurrence patterns of these species throughout the tree canopy are unknown. This study was initiated to evaluate specimen abundance, species richness and species composition among three designated strata in the canopy of eastern hemlock, assess species for vertical stratification patterns, and determine if co-occurrence patterns of Psocoptera species are random or non-random. During this study, 27 species representing 18 genera and 10 families were evaluated.