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Comitato Scientifico per la Fauna d’Italia CHECKLIST AND DISTRIBUTION OF THE ITALIAN FAUNA FAUNA THE ITALIAN AND DISTRIBUTION OF CHECKLIST 10,000 terrestrial and inland water species and inland water 10,000 terrestrial CHECKLIST AND DISTRIBUTION OF THE ITALIAN FAUNA 10,000 terrestrial and inland water species ISBNISBN 88-89230-09-688-89230- 09- 6 Ministero dell’Ambiente 9 778888988889 230091230091 e della Tutela del Territorio e del Mare CH © Copyright 2006 - Comune di Verona ISSN 0392-0097 ISBN 88-89230-09-6 All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, without the prior permission in writing of the publishers and of the Authors. Direttore Responsabile Alessandra Aspes CHECKLIST AND DISTRIBUTION OF THE ITALIAN FAUNA 10,000 terrestrial and inland water species Memorie del Museo Civico di Storia Naturale di Verona - 2. Serie Sezione Scienze della Vita 17 - 2006 PROMOTING AGENCIES Italian Ministry for Environment and Territory and Sea, Nature Protection Directorate Civic Museum of Natural History of Verona Scientifi c Committee for the Fauna of Italy Calabria University, Department of Ecology EDITORIAL BOARD Aldo Cosentino Alessandro La Posta Augusto Vigna Taglianti Alessandra Aspes Leonardo Latella SCIENTIFIC BOARD Marco Bologna Pietro Brandmayr Eugenio Dupré Alessandro La Posta Leonardo Latella Alessandro Minelli Sandro Ruffo Fabio Stoch Augusto Vigna Taglianti Marzio Zapparoli EDITORS Sandro Ruffo Fabio Stoch DESIGN Riccardo Ricci LAYOUT Riccardo Ricci Zeno Guarienti EDITORIAL ASSISTANT Elisa Giacometti TRANSLATORS Maria Cristina Bruno (1-72, 239-307) Daniel Whitmore (73-238) VOLUME CITATION: Ruffo S., Stoch F. (eds.), 2006. Checklist and distribution of the Italian fauna. Memorie del Museo Civico di Storia Naturale di Verona, 2.Serie, Sezione Scienze della Vita 17, with CD-ROM. CHAPTER CITATION: Minelli A., 2006. Annelida Hirudinea. In: Ruffo S., Stoch F. (eds.). Checklist and distribution of the Italian fauna. Memorie del Museo Civico di Storia Naturale di Verona, 2.Serie, Sezione Scienze della Vita 17: 77-78, with data on CD-ROM. This volume was published with the fi nancial support of the Italian Ministry for Environment and Territory and Sea, Nature Protection Directorate. CHECKLIST AND DISTRIBUTION OF THE ITALIAN FAUNA 10,000 terrestrial and inland water species INDEX Introduction 13 Crustacea Malacostraca Bathynellacea 101 The project CKmap (Checklist and distribution of the Italian fauna): methods Crustacea Malacostraca and informatical techniques 15 Thermosbaenacea 103 The checklist of the Italian fauna 21 Crustacea Malacostraca Mysidacea from groundwaters 105 The chorotypes of the Italian fauna 25 Crustacea Malacostraca Isopoda 107 Endemism in Italy 29 Crustacea Malacostraca Amphipoda 109 Analysis of the distribution in Italy of species richness and rarity 33 Crustacea Malacostraca Decapoda 113 The Italian faunal Provinces 37 Arachnida Pseudoscorpionida 115 Chronogeonemy analysis: some examples Arachnida Araneae Salticidae 119 regarding species of the Italian fauna 41 Arachnida Acari Actinedida Distribution of Italian fauna species versus Hydrachnidia 121 Eunis, Corine and Natura 2000 habitat types 47 Chilopoda 123 The exotic species of the Italian fauna 57 Insecta Ephemeroptera 127 Criteria for the compilation of the fi rst Red List of species of national and regional Insecta Odonata 131 interest 63 Insecta Blattaria 133 Strategies for the conservation of Italian fauna 67 Insecta Mantodea 135 Insecta Orthoptera 137 TAXONOMIC SECTION 71 Annelida Oligochaeta 73 Insecta Dermaptera 141 Annelida Hirudinea 77 Insecta Plecoptera 143 Mollusca Gastropoda Prosobranchia Insecta Heteroptera Nepomorpha and Heterobranchia Heterostropha 79 and Gerromorpha 147 Mollusca Bivalvia 83 Insecta Heteroptera Leptopodidae, Saldidae, Miridae (partim), Crustacea Branchiopoda Anostraca, Tingidae 151 Notostraca, Conchostraca 85 Insecta Homoptera Auchenorrhyncha Crustacea Branchiopoda Cladocera 87 (partim) 155 Crustacea Copepoda Calanoida 91 Insecta Coleoptera Carabidae (Carabini, Cychrini, Trechini, Abacetini, Crustacea Copepoda Cyclopoida 93 Stomini, Pterostichini) 159 Crustacea Copepoda Harpacticoida 97 Insecta Coleoptera Hydroadephaga 165 6 Insecta Coleoptera Hydrophiloidea 167 Insecta Mecoptera 235 Insecta Coleoptera Hydraenidae 169 Insecta Diptera Tipulidae 237 Insecta Coleoptera Georissidae 171 Insecta Diptera Simuliidae 239 Insecta Coleoptera Histeridae 173 Insecta Diptera Stratiomyidae 241 Insecta Coleoptera Cholevidae and Insecta Diptera Syrphidae Platypsyllidae 177 (Syrphinae, Syrphini) 243 Insecta Coleoptera Staphylinidae 181 Insecta Diptera Conopidae 245 Insecta Coleoptera Staphylinidae Pselaphinae 183 Insecta Diptera Sciomyzidae 247 Insecta Coleoptera Staphylinidae Insecta Trichoptera 249 Omaliinae 185 Insecta Lepidoptera Hepialidae 253 Insecta Coleoptera Staphylinidae Staphylininae 187 Insecta Lepidoptera Zygaenoidea 255 Insecta Coleoptera Staphylinidae Insecta Lepidoptera Papilionoidea Aleocharinae (Rhopalocera) 257 (genus Leptusa Kraatz, 1859) 189 Insecta Lepidoptera Noctuidae Insecta Coleoptera Lucanidae 191 (Plusiinae, Noctuinae) 263 Insecta Coleoptera Scarabaeoidea 193 Insecta Hymenoptera Chrysididae 267 Insecta Coleoptera Dryopoidea 197 Insecta Hymenoptera Dryinidae Embolemidae Sclerogibbidae 269 Insecta Coleoptera Elateridae 199 Insecta Hymenoptera Scolioidea Insecta Coleoptera Buprestidae 203 (Tiphiidae excluded) 271 Insecta Coleoptera Nitidulidae 205 Insecta Hymenoptera Aculeata Apoidea (partim) 273 Insecta Coleoptera Cucujidae 209 Agnatha Osteichthyes 277 Insecta Coleoptera Cryptophagidae 213 Amphibia and Reptilia 281 Insecta Coleoptera Tenebrionidae 215 Mammalia Insectivora 285 Insecta Coleoptera Cerambycidae 217 Mammalia Chiroptera 289 Insecta Coleoptera Chrysomelidae Cryptocephalinae 221 Mammalia Rodentia 293 Insecta Coleoptera Chrysomelidae Alticinae 225 AUTHORS LIST 297 Insecta Coleoptera Curculionoidea 229 ACKNOWLEDGEMENTS 303 Insecta Neuroptera, Megaloptera and Raphidioptera 233 7 Italy has a very rich fauna, with more than 57,000 species recorded so far. This high biodiversity is a privilege, but also a responsibility: its protection is one of our challen- ges and is a goal that everybody should take aim at. In fact, Italy together with another 180 countries ratifi ed the Rio de Janeiro Convention on Biodiversity, which in 1992 established at worldwide level the intrinsic value of biodiversity. Ten years later, in 2002, the World Conference for Sustainable Development in Johannesburg defi ned a precise target: to reduce the biodiversity loss by 2010, thus the countdown has already started. Conservation of biodiversity in general, and of animal biodiversity in particular, can not be obtained without a good knowledge of what we want to protect. For this purpose the Italian Ministry for the Environment and Territory and the Nature Protec- tion Division supported in the last years several projects aimed at the improvement of the scientifi c knowledge of the various components of biodiversity of Italy. Among them, the project “Checklist and distribution of the Italian fauna”, the results of which are presented and discussed in this volume, represents the most complete knowledge available on the fauna of a given country. The volume and the CD report more than 500,000 distribution data of about 10,000 terrestrial and freshwater species of the Ita- lian fauna. This huge amount of data is pride for Italy, but it is also an essential operati- ve tool to compile Red Lists, to create environmental quality models, and to contribute to implement the Habitat Directive and thus reduce biodiversity loss in Italy by the target year 2010. The Checklist is a knowledge tool of great relevance, which places Italy in fi rst place worldwide for biodiversity conservation. Aldo Cosentino General Director Italian Ministry for the Environment and Territory and Sea Nature Protection Directorate 9 It is with great pleasure that I present this volume, which represents the end point of a long and com- plex journey in the study of Italian fauna, to which I was fortunate to be involved at different levels and with different roles, and it is the starting point of a new phase, in which the knowledge we obtained so far will be applied to the management and protection of the fauna. This volume represents the most detailed study available on the fauna of a given country; it was the result of a long, painstaking work which involved all the zoology specialists of our country for more than 10 years. It was the result of the synergy between the Scientific Committee for Italian Fauna and the Italian Ministry for the Envi- ronment and Territory. The origin of this project can be found in the series “Fauna d’Italia”, created in 1952 under the auspi- ces of the Italian National Academy of Entomology and the Italian Zoological Union, programmed and realised by the Scientific Committee for Italian Fauna (formally established in 1989, with the two promoting agencies represented with the same status), and published by the Calderini publishing house. The first volume was issued in 1956, the 40th is in press in this moment, and several more are in progress. This is a prestigious and long-term work, of great importance, which suffered several inter- ruptions, and therefore it is still in progress. The productive relationship developed during the last 15 years between the Scientific Committee for Italian Fauna and the Italian Ministry for the Environment (now Italian Ministry