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Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications Sherwin Carlquist Santa Barbara Botanic Garden
Aliso: A Journal of Systematic and Evolutionary Botany Volume 14 | Issue 1 Article 2 1995 Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications Sherwin Carlquist Santa Barbara Botanic Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Carlquist, Sherwin (1995) "Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 14: Iss. 1, Article 2. Available at: http://scholarship.claremont.edu/aliso/vol14/iss1/2 Aliso, 14(1), pp. 1-17 © 1995, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 WOOD ANATOMY OF CARYOPHYLLACEAE: ECOLOGICAL, HABITAL, SYSTEMATIC, AND PHYLOGENETIC IMPLICATIONS SHERWIN CARLQUIST1 Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara, CA 93105 ABSTRACT Wood of Caryophyllaceae is more diverse than has been appreciated. Imperforate tracheary elements may be tracheids, fiber-tracheids, or libriform fibers. Rays may be uniseriate only, multiseriate only, or absent. Roots of some species (and sterns of a few of those same genera) have vascular tissue produced by successive cambia. The diversity in wood anatomy character states shows a range from primitive to specialized so great that origin close to one of the more specialized families of Cheno podiales, such as Chenopodiaceae or Amaranthaceae, is unlikely. Caryophyllaceae probably branched from the ordinal clade near the clade's base, as cladistic evidence suggests. Raylessness and abrupt onset of multiseriate rays may indicate woodiness in the family is secondary. Successive cambia might also be a subsidiary indicator of secondary woodiness in Caryophyllaceae (although not necessarily dicotyledons at large). -
Gymnosperms the MESOZOIC: ERA of GYMNOSPERM DOMINANCE
Chapter 24 Gymnosperms THE MESOZOIC: ERA OF GYMNOSPERM DOMINANCE THE VASCULAR SYSTEM OF GYMNOSPERMS CYCADS GINKGO CONIFERS Pinaceae Include the Pines, Firs, and Spruces Cupressaceae Include the Junipers, Cypresses, and Redwoods Taxaceae Include the Yews, but Plum Yews Belong to Cephalotaxaceae Podocarpaceae and Araucariaceae Are Largely Southern Hemisphere Conifers THE LIFE CYCLE OF PINUS, A REPRESENTATIVE GYMNOSPERM Pollen and Ovules Are Produced in Different Kinds of Structures Pollination Replaces the Need for Free Water Fertilization Leads to Seed Formation GNETOPHYTES GYMNOSPERMS: SEEDS, POLLEN, AND WOOD THE ECOLOGICAL AND ECONOMIC IMPORTANCE OF GYMNOSPERMS The Origin of Seeds, Pollen, and Wood Seeds and Pollen Are Key Reproductive SUMMARY Innovations for Life on Land Seed Plants Have Distinctive Vegetative PLANTS, PEOPLE, AND THE Features ENVIRONMENT: The California Coast Relationships among Gymnosperms Redwood Forest 1 KEY CONCEPTS 1. The evolution of seeds, pollen, and wood freed plants from the need for water during reproduction, allowed for more effective dispersal of sperm, increased parental investment in the next generation and allowed for greater size and strength. 2. Seed plants originated in the Devonian period from a group called the progymnosperms, which possessed wood and heterospory, but reproduced by releasing spores. Currently, five lineages of seed plants survive--the flowering plants plus four groups of gymnosperms: cycads, Ginkgo, conifers, and gnetophytes. Conifers are the best known and most economically important group, including pines, firs, spruces, hemlocks, redwoods, cedars, cypress, yews, and several Southern Hemisphere genera. 3. The pine life cycle is heterosporous. Pollen strobili are small and seasonal. Each sporophyll has two microsporangia, in which microspores are formed and divide into immature male gametophytes while still retained in the microsporangia. -
Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications
Aliso, 14(1), pp. 1—17 © 1995, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 WOOD ANATOMY OF CARYOPHYLLACEAE: ECOLOGICAL, HABITAL, SYSTEMATIC, AND PHYLOGENETIC IMPLICATIONS SHERwIN CARLQuIsT’ Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara, CA 93105 ABSTRACT Wood of Caryophyllaceae is more diverse than has been appreciated. Imperforate tracheary elements may be tracheids, fiber-tracheids, or libriform fibers. Rays may be unisei-iate only, multiseriate only, or absent. Roots of some species (and stems of a few of those same genera) have vascular tissue produced by successive cambia. The diversity in wood anatomy character states shows a range from primitive to specialized so great that origin close to one of the more specialized families of Cheno podiales, such as Chenopodiaceae or Amaranthaceae, is unlikely. Caryophyllaceae probably branched from the ordinal dade near the dade’s base, as cladistic evidence suggests. Raylessness and abrupt onset of multiseriate rays may indicate woodiness in the family is secondary. Successive cambia might also be a subsidiary indicator of secondary woodiness in Caryophyllaceae (although not necessarily dicotyledons at large). Although a small shrub, Gyinnocarpos may be primitively woody, and because that genus appears primitive in many wood features the family as a whole may derive from woody ancestors, despite apparent secondary woodiness in many phylads. Systematic distribution of wood character states in the family corresponds closely to the Pax and Hoffmann system of three tribes and their progressive degrees of specialization (Paronychieae, Alsineae, Sileneae). Wood data support the idea that Caryophyllaceae represent a branch from near the base of the order Caryophyllales. -
July 2008 Volume 31 Number 4
Sego Lily July 2008 31 (4) July 2008 Volume 31 Number 4 Salt Lake Chapter Trek to the Beaver Dam Mountains By Bill Gray In April, eight Salt Lake Chapter members took a break from their long winter and headed south as far as they could get, actually ending up just over the border in Arizona. Somehow the Virgin River gorge and that part of the Beaver Dam Moun- tains feel as though they really be- long in Utah. Most of you have probably driven through the gorge on Interstate 15 en route to/from Las Vegas or Southern California, and have ad- mired the steep rugged cliffs where the river has carved its way down. From a speeding car it is possible to spot a few of the obvious standout plants of the Mojave Desert – Joshua tree (Yucca brevifolia), Above: Crevice penstemon, Penstemon petiolatus, grows in Creosote bush (Larrea tridentata) mostly inaccessible crevices in rugged limestone cliffs in and rather modest specimens of Le- Utah’s Beaver Dam Mountains. Photo by Bill Gray conte's barrel cactus (Ferocactus acanthodes var. lecontei). But we wanted to take time to get In this issue: out and really experience first hand the other unusual plants from this Salt Lake Chapter Trek to the Beaver Dam Mountains . 1 area. In this we were very fortunate UNPS and Chapter News . 2 to have one of the best possible Bulletin Board . 3 guides in Dr. Larry Higgins. Larry Q and A: Growing Sego Lilies . 6 grew up in the area, getting to know Richard Joshua Shaw (1923-2008) . -
Annotated Checklist of Vascular Flora, Bryce
National Park Service U.S. Department of the Interior Natural Resource Program Center Annotated Checklist of Vascular Flora Bryce Canyon National Park Natural Resource Technical Report NPS/NCPN/NRTR–2009/153 ON THE COVER Matted prickly-phlox (Leptodactylon caespitosum), Bryce Canyon National Park, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Bryce Canyon National Park Natural Resource Technical Report NPS/NCPN/NRTR–2009/153 Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741 Sarah Topp Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 January 2009 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner. The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. -
Vascular Flora of the Liebre Mountains, Western Transverse Ranges, California Steve Boyd Rancho Santa Ana Botanic Garden
Aliso: A Journal of Systematic and Evolutionary Botany Volume 18 | Issue 2 Article 15 1999 Vascular flora of the Liebre Mountains, western Transverse Ranges, California Steve Boyd Rancho Santa Ana Botanic Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Boyd, Steve (1999) "Vascular flora of the Liebre Mountains, western Transverse Ranges, California," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 18: Iss. 2, Article 15. Available at: http://scholarship.claremont.edu/aliso/vol18/iss2/15 Aliso, 18(2), pp. 93-139 © 1999, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 VASCULAR FLORA OF THE LIEBRE MOUNTAINS, WESTERN TRANSVERSE RANGES, CALIFORNIA STEVE BOYD Rancho Santa Ana Botanic Garden 1500 N. College Avenue Claremont, Calif. 91711 ABSTRACT The Liebre Mountains form a discrete unit of the Transverse Ranges of southern California. Geo graphically, the range is transitional to the San Gabriel Mountains, Inner Coast Ranges, Tehachapi Mountains, and Mojave Desert. A total of 1010 vascular plant taxa was recorded from the range, representing 104 families and 400 genera. The ratio of native vs. nonnative elements of the flora is 4:1, similar to that documented in other areas of cismontane southern California. The range is note worthy for the diversity of Quercus and oak-dominated vegetation. A total of 32 sensitive plant taxa (rare, threatened or endangered) was recorded from the range. Key words: Liebre Mountains, Transverse Ranges, southern California, flora, sensitive plants. INTRODUCTION belt and Peirson's (1935) handbook of trees and shrubs. Published documentation of the San Bernar The Transverse Ranges are one of southern Califor dino Mountains is little better, limited to Parish's nia's most prominent physiographic features. -
List of Plants for Great Sand Dunes National Park and Preserve
Great Sand Dunes National Park and Preserve Plant Checklist DRAFT as of 29 November 2005 FERNS AND FERN ALLIES Equisetaceae (Horsetail Family) Vascular Plant Equisetales Equisetaceae Equisetum arvense Present in Park Rare Native Field horsetail Vascular Plant Equisetales Equisetaceae Equisetum laevigatum Present in Park Unknown Native Scouring-rush Polypodiaceae (Fern Family) Vascular Plant Polypodiales Dryopteridaceae Cystopteris fragilis Present in Park Uncommon Native Brittle bladderfern Vascular Plant Polypodiales Dryopteridaceae Woodsia oregana Present in Park Uncommon Native Oregon woodsia Pteridaceae (Maidenhair Fern Family) Vascular Plant Polypodiales Pteridaceae Argyrochosma fendleri Present in Park Unknown Native Zigzag fern Vascular Plant Polypodiales Pteridaceae Cheilanthes feei Present in Park Uncommon Native Slender lip fern Vascular Plant Polypodiales Pteridaceae Cryptogramma acrostichoides Present in Park Unknown Native American rockbrake Selaginellaceae (Spikemoss Family) Vascular Plant Selaginellales Selaginellaceae Selaginella densa Present in Park Rare Native Lesser spikemoss Vascular Plant Selaginellales Selaginellaceae Selaginella weatherbiana Present in Park Unknown Native Weatherby's clubmoss CONIFERS Cupressaceae (Cypress family) Vascular Plant Pinales Cupressaceae Juniperus scopulorum Present in Park Unknown Native Rocky Mountain juniper Pinaceae (Pine Family) Vascular Plant Pinales Pinaceae Abies concolor var. concolor Present in Park Rare Native White fir Vascular Plant Pinales Pinaceae Abies lasiocarpa Present -
Draft Programmatic EIS for Fuels Reduction and Rangeland
NATIONAL SYSTEM OF PUBLIC LANDS U.S. DEPARTMENT OF THE INTERIOR U.S. Department of the Interior March 2020 BUREAU OF LAND MANAGEMENT BUREAU OF LAND MANAGEMENT Draft Programmatic EIS for Fuels Reduction and Rangeland Restoration in the Great Basin Volume 3: Appendices B through N Estimated Lead Agency Total Costs Associated with Developing and Producing this EIS $2,000,000 The Bureau of Land Management’s multiple-use mission is to sustain the health and productivity of the public lands for the use and enjoyment of present and future generations. The Bureau accomplishes this by managing such activities as outdoor recreation, livestock grazing, mineral development, and energy production, and by conserving natural, historical, cultural, and other resources on public lands. Appendix B. Acronyms, Literature Cited, Glossary B.1 ACRONYMS ACRONYMS AND ABBREVIATIONS Full Phrase ACHP Advisory Council on Historic Preservation AML appropriate management level ARMPA Approved Resource Management Plan Amendment BCR bird conservation region BLM Bureau of Land Management BSU biologically significant unit CEQ Council on Environmental Quality EIS environmental impact statement EPA US Environmental Protection Agency ESA Endangered Species Act ESR emergency stabilization and rehabilitation FIAT Fire and Invasives Assessment Tool FLPMA Federal Land Policy and Management Act FY fiscal year GHMA general habitat management area HMA herd management area IBA important bird area IHMA important habitat management area MBTA Migratory Bird Treaty Act MOU memorandum of understanding MtCO2e metric tons of carbon dioxide equivalent NEPA National Environmental Policy Act NHPA National Historic Preservation Act NIFC National Interagency Fire Center NRCS National Resources Conservation Service NRHP National Register of Historic Places NWCG National Wildfire Coordination Group OHMA other habitat management area OHV off-highway vehicle Programmatic EIS for Fuels Reduction and Rangeland Restoration in the Great Basin B-1 B. -
United States Department of the Interior Bureau of Land Management
UNITED STATES DEPARTMENT OF THE INTERIOR BUREAU OF LAND MANAGEMENT FINDING OF NO SIGNFICANT IMPACT San Rafael Desert Travel Management Plan DOI-BLM-UT-G020-2018-0004-EA August 2020 Price Field Office 125 South 600 West Price, Utah 84501 435-636-3600 San Rafael Desert Travel Management Plan DOI-BLM-UT-G020-2018-0004-EA FINDING OF NO SIGNFICANT IMPACT I have reviewed the San Rafael Desert Travel Management Plan Environmental Assessment (EA) DOI-BLM-UT-G020-2018-0004-EA. After considering the environmental effects as described in the EA, and incorporated herein, I have determined that Alternative D, as identified in the EA and modified in the Decision Record (Modified Alternative D), will not significantly affect the quality of the human environment and that an Environmental Impact Statement is not required to be prepared. I have determined that the proposed action, which is to designate a comprehensive off-highway vehicle (OHV) travel management plan (TMP) for the San Rafael Desert Travel Management Area (TMA), is in conformance with the approved 2008 Price Field Office Record of Decision and Approved Resource Management Plan (2008 RMP) and is consistent with applicable plans and policies of county, state, Tribal and Federal agencies. This finding is based on my consideration of the Council on Environmental Quality’s (CEQ) criteria for significance (40 CFR 1508.27) regarding the context and the intensity of impacts described in the EA. Context The TMA that forms the basis of the San Rafael Desert TMP contains 377,609 acres of Bureau of Land Management (BLM)-managed lands, and an existing road inventory containing 1,180.8 miles of roads. -
Annotated Checklist of Vascular Flora, Cedar Breaks National
National Park Service U.S. Department of the Interior Natural Resource Program Center Annotated Checklist of Vascular Flora Cedar Breaks National Monument Natural Resource Technical Report NPS/NCPN/NRTR—2009/173 ON THE COVER Peterson’s campion (Silene petersonii), Cedar Breaks National Monument, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Cedar Breaks National Monument Natural Resource Technical Report NPS/NCPN/NRTR—2009/173 Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741 Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 February 2009 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner. The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientifi c literature reviews; and peer- reviewed proceedings of technical workshops, conferences, or symposia. -
Sensitive Species That Are Not Listed Or Proposed Under the ESA Sorted By: Major Group, Subgroup, NS Sci
Forest Service Sensitive Species that are not listed or proposed under the ESA Sorted by: Major Group, Subgroup, NS Sci. Name; Legend: Page 94 REGION 10 REGION 1 REGION 2 REGION 3 REGION 4 REGION 5 REGION 6 REGION 8 REGION 9 ALTERNATE NATURESERVE PRIMARY MAJOR SUB- U.S. N U.S. 2005 NATURESERVE SCIENTIFIC NAME SCIENTIFIC NAME(S) COMMON NAME GROUP GROUP G RANK RANK ESA C 9 Anahita punctulata Southeastern Wandering Spider Invertebrate Arachnid G4 NNR 9 Apochthonius indianensis A Pseudoscorpion Invertebrate Arachnid G1G2 N1N2 9 Apochthonius paucispinosus Dry Fork Valley Cave Invertebrate Arachnid G1 N1 Pseudoscorpion 9 Erebomaster flavescens A Cave Obligate Harvestman Invertebrate Arachnid G3G4 N3N4 9 Hesperochernes mirabilis Cave Psuedoscorpion Invertebrate Arachnid G5 N5 8 Hypochilus coylei A Cave Spider Invertebrate Arachnid G3? NNR 8 Hypochilus sheari A Lampshade Spider Invertebrate Arachnid G2G3 NNR 9 Kleptochthonius griseomanus An Indiana Cave Pseudoscorpion Invertebrate Arachnid G1 N1 8 Kleptochthonius orpheus Orpheus Cave Pseudoscorpion Invertebrate Arachnid G1 N1 9 Kleptochthonius packardi A Cave Obligate Pseudoscorpion Invertebrate Arachnid G2G3 N2N3 9 Nesticus carteri A Cave Spider Invertebrate Arachnid GNR NNR 8 Nesticus cooperi Lost Nantahala Cave Spider Invertebrate Arachnid G1 N1 8 Nesticus crosbyi A Cave Spider Invertebrate Arachnid G1? NNR 8 Nesticus mimus A Cave Spider Invertebrate Arachnid G2 NNR 8 Nesticus sheari A Cave Spider Invertebrate Arachnid G2? NNR 8 Nesticus silvanus A Cave Spider Invertebrate Arachnid G2? NNR -
Flora-Lab-Manual.Pdf
LabLab MManualanual ttoo tthehe Jane Mygatt Juliana Medeiros Flora of New Mexico Lab Manual to the Flora of New Mexico Jane Mygatt Juliana Medeiros University of New Mexico Herbarium Museum of Southwestern Biology MSC03 2020 1 University of New Mexico Albuquerque, NM, USA 87131-0001 October 2009 Contents page Introduction VI Acknowledgments VI Seed Plant Phylogeny 1 Timeline for the Evolution of Seed Plants 2 Non-fl owering Seed Plants 3 Order Gnetales Ephedraceae 4 Order (ungrouped) The Conifers Cupressaceae 5 Pinaceae 8 Field Trips 13 Sandia Crest 14 Las Huertas Canyon 20 Sevilleta 24 West Mesa 30 Rio Grande Bosque 34 Flowering Seed Plants- The Monocots 40 Order Alistmatales Lemnaceae 41 Order Asparagales Iridaceae 42 Orchidaceae 43 Order Commelinales Commelinaceae 45 Order Liliales Liliaceae 46 Order Poales Cyperaceae 47 Juncaceae 49 Poaceae 50 Typhaceae 53 Flowering Seed Plants- The Eudicots 54 Order (ungrouped) Nymphaeaceae 55 Order Proteales Platanaceae 56 Order Ranunculales Berberidaceae 57 Papaveraceae 58 Ranunculaceae 59 III page Core Eudicots 61 Saxifragales Crassulaceae 62 Saxifragaceae 63 Rosids Order Zygophyllales Zygophyllaceae 64 Rosid I Order Cucurbitales Cucurbitaceae 65 Order Fabales Fabaceae 66 Order Fagales Betulaceae 69 Fagaceae 70 Juglandaceae 71 Order Malpighiales Euphorbiaceae 72 Linaceae 73 Salicaceae 74 Violaceae 75 Order Rosales Elaeagnaceae 76 Rosaceae 77 Ulmaceae 81 Rosid II Order Brassicales Brassicaceae 82 Capparaceae 84 Order Geraniales Geraniaceae 85 Order Malvales Malvaceae 86 Order Myrtales Onagraceae