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Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications Sherwin Carlquist Santa Barbara Botanic Garden
Aliso: A Journal of Systematic and Evolutionary Botany Volume 14 | Issue 1 Article 2 1995 Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications Sherwin Carlquist Santa Barbara Botanic Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Carlquist, Sherwin (1995) "Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 14: Iss. 1, Article 2. Available at: http://scholarship.claremont.edu/aliso/vol14/iss1/2 Aliso, 14(1), pp. 1-17 © 1995, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 WOOD ANATOMY OF CARYOPHYLLACEAE: ECOLOGICAL, HABITAL, SYSTEMATIC, AND PHYLOGENETIC IMPLICATIONS SHERWIN CARLQUIST1 Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara, CA 93105 ABSTRACT Wood of Caryophyllaceae is more diverse than has been appreciated. Imperforate tracheary elements may be tracheids, fiber-tracheids, or libriform fibers. Rays may be uniseriate only, multiseriate only, or absent. Roots of some species (and sterns of a few of those same genera) have vascular tissue produced by successive cambia. The diversity in wood anatomy character states shows a range from primitive to specialized so great that origin close to one of the more specialized families of Cheno podiales, such as Chenopodiaceae or Amaranthaceae, is unlikely. Caryophyllaceae probably branched from the ordinal clade near the clade's base, as cladistic evidence suggests. Raylessness and abrupt onset of multiseriate rays may indicate woodiness in the family is secondary. Successive cambia might also be a subsidiary indicator of secondary woodiness in Caryophyllaceae (although not necessarily dicotyledons at large). -
Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications
Aliso, 14(1), pp. 1—17 © 1995, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 WOOD ANATOMY OF CARYOPHYLLACEAE: ECOLOGICAL, HABITAL, SYSTEMATIC, AND PHYLOGENETIC IMPLICATIONS SHERwIN CARLQuIsT’ Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara, CA 93105 ABSTRACT Wood of Caryophyllaceae is more diverse than has been appreciated. Imperforate tracheary elements may be tracheids, fiber-tracheids, or libriform fibers. Rays may be unisei-iate only, multiseriate only, or absent. Roots of some species (and stems of a few of those same genera) have vascular tissue produced by successive cambia. The diversity in wood anatomy character states shows a range from primitive to specialized so great that origin close to one of the more specialized families of Cheno podiales, such as Chenopodiaceae or Amaranthaceae, is unlikely. Caryophyllaceae probably branched from the ordinal dade near the dade’s base, as cladistic evidence suggests. Raylessness and abrupt onset of multiseriate rays may indicate woodiness in the family is secondary. Successive cambia might also be a subsidiary indicator of secondary woodiness in Caryophyllaceae (although not necessarily dicotyledons at large). Although a small shrub, Gyinnocarpos may be primitively woody, and because that genus appears primitive in many wood features the family as a whole may derive from woody ancestors, despite apparent secondary woodiness in many phylads. Systematic distribution of wood character states in the family corresponds closely to the Pax and Hoffmann system of three tribes and their progressive degrees of specialization (Paronychieae, Alsineae, Sileneae). Wood data support the idea that Caryophyllaceae represent a branch from near the base of the order Caryophyllales. -
Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument
Schmidt, Drost, Halvorson In Cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument Plant and Vertebrate Vascular U.S. Geological Survey Southwest Biological Science Center 2255 N. Gemini Drive Flagstaff, AZ 86001 Open-File Report 2006-1163 Southwest Biological Science Center Open-File Report 2006-1163 November 2006 U.S. Department of the Interior U.S. Geological Survey National Park Service In cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument By Cecilia A. Schmidt, Charles A. Drost, and William L. Halvorson Open-File Report 2006-1163 November, 2006 USGS Southwest Biological Science Center Sonoran Desert Research Station University of Arizona U.S. Department of the Interior School of Natural Resources U.S. Geological Survey 125 Biological Sciences East National Park Service Tucson, Arizona 85721 U.S. Department of the Interior Dirk Kempthorne, Secretary U.S. Geological Survey Mark Myers, Director U.S. Geological Survey, Reston, Virginia: 2006 Note: This document contains information of a preliminary nature and was prepared primarily for internal use in the U.S. Geological Survey. This information is NOT intended for use in open literature prior to publication by the investigators named unless permission is obtained in writing from the investigators named and from the Station Leader. Suggested Citation Schmidt, C. A., C. A. Drost, and W. L. Halvorson 2006. Vascular Plant and Vertebrate Inventory of Montezuma Castle National Monument. USGS Open-File Report 2006-1163. -
December 2012 Number 1
Calochortiana December 2012 Number 1 December 2012 Number 1 CONTENTS Proceedings of the Fifth South- western Rare and Endangered Plant Conference Calochortiana, a new publication of the Utah Native Plant Society . 3 The Fifth Southwestern Rare and En- dangered Plant Conference, Salt Lake City, Utah, March 2009 . 3 Abstracts of presentations and posters not submitted for the proceedings . 4 Southwestern cienegas: Rare habitats for endangered wetland plants. Robert Sivinski . 17 A new look at ranking plant rarity for conservation purposes, with an em- phasis on the flora of the American Southwest. John R. Spence . 25 The contribution of Cedar Breaks Na- tional Monument to the conservation of vascular plant diversity in Utah. Walter Fertig and Douglas N. Rey- nolds . 35 Studying the seed bank dynamics of rare plants. Susan Meyer . 46 East meets west: Rare desert Alliums in Arizona. John L. Anderson . 56 Calochortus nuttallii (Sego lily), Spatial patterns of endemic plant spe- state flower of Utah. By Kaye cies of the Colorado Plateau. Crystal Thorne. Krause . 63 Continued on page 2 Copyright 2012 Utah Native Plant Society. All Rights Reserved. Utah Native Plant Society Utah Native Plant Society, PO Box 520041, Salt Lake Copyright 2012 Utah Native Plant Society. All Rights City, Utah, 84152-0041. www.unps.org Reserved. Calochortiana is a publication of the Utah Native Plant Society, a 501(c)(3) not-for-profit organi- Editor: Walter Fertig ([email protected]), zation dedicated to conserving and promoting steward- Editorial Committee: Walter Fertig, Mindy Wheeler, ship of our native plants. Leila Shultz, and Susan Meyer CONTENTS, continued Biogeography of rare plants of the Ash Meadows National Wildlife Refuge, Nevada. -
Fort Ord Natural Reserve Plant List
UCSC Fort Ord Natural Reserve Plants Below is the most recently updated plant list for UCSC Fort Ord Natural Reserve. * non-native taxon ? presence in question Listed Species Information: CNPS Listed - as designated by the California Rare Plant Ranks (formerly known as CNPS Lists). More information at http://www.cnps.org/cnps/rareplants/ranking.php Cal IPC Listed - an inventory that categorizes exotic and invasive plants as High, Moderate, or Limited, reflecting the level of each species' negative ecological impact in California. More information at http://www.cal-ipc.org More information about Federal and State threatened and endangered species listings can be found at https://www.fws.gov/endangered/ (US) and http://www.dfg.ca.gov/wildlife/nongame/ t_e_spp/ (CA). FAMILY NAME SCIENTIFIC NAME COMMON NAME LISTED Ferns AZOLLACEAE - Mosquito Fern American water fern, mosquito fern, Family Azolla filiculoides ? Mosquito fern, Pacific mosquitofern DENNSTAEDTIACEAE - Bracken Hairy brackenfern, Western bracken Family Pteridium aquilinum var. pubescens fern DRYOPTERIDACEAE - Shield or California wood fern, Coastal wood wood fern family Dryopteris arguta fern, Shield fern Common horsetail rush, Common horsetail, field horsetail, Field EQUISETACEAE - Horsetail Family Equisetum arvense horsetail Equisetum telmateia ssp. braunii Giant horse tail, Giant horsetail Pentagramma triangularis ssp. PTERIDACEAE - Brake Family triangularis Gold back fern Gymnosperms CUPRESSACEAE - Cypress Family Hesperocyparis macrocarpa Monterey cypress CNPS - 1B.2, Cal IPC -
Literaturverzeichnis Acken, H. Von Und P. Müllner (1972): Zur
Literaturverzeichnis Anmerkung: Die Literaturliste wurde automatisch aus einer Datenbank erzeugt. Wir haben uns bemüht, grobe Fehler & Doppeleinträge nachträglich „auszumerzen“, doch der Teufel steckt ja bekanntlich im Detail, so werden sich wahrscheinlich noch einige „Unzulänglichkeiten“ finden lassen. Unterschiedliche Zitierweisen und uneinheitliche Abkürzungen bitten wir zu entschuldigen! Acken, H. von und P. Müllner (1972): Zur Unterscheidung von Digitaria sanguinalis (Setaria sanguinalis Panicum sanguinale, Bluthirse) und Digitaria ischaemum (Setaria ischaemum, Panicum lineare, Fingerhirse). Kieler Not. Pflanzenk. Schleswig-Holstein 4 (2/3): 37-38 Ade, A. (1901): Flora des bayerischen Bodenseegebietes - Übersicht über die im bayerischen Bodenseegebiet bis jetzt beobachteten wildwachsenden Phanerogamen und Gefässkryptogamen. Ber. Bayer. Bot. Ges. 8: 3-127 Ade, A. (1957): Die Gattung Rubus in Südwestdeutschland. Versuch einer Bearbeitung der Brombeerflora Hessens, des nördlichen Bayerns, Badens, und Württembergs, einschließlich Rheinhessens, der Pfalz und des Nahegebietes, sowie der gesamten Rhön. Schriftenreihe NatSchutzstelle Darmstadt 7: 1-217 Ade, M. (1989, 1990): Flora von Oberndorf am Neckar. Veröff. Landesstelle Naturschutz Landschaftspflege Bad.-Württ. 64/65: 509-583 Ade, U. (1990): Seltene und geschützte Pflanzen im Landkreis Böblingen. In: Ade, U. et al.: Naturnahe Lebensräume und Flora im Schönbuch und Gäu. : 52-76 Ade, U., B. Baumann, H. Baumann u. W. Wahrenburg (1990): Naturnahe Lebensräume und Flora in Schönbuch und Gäu. : 244 Adema, F. (1968): En "oude" voor Nederland nieuwe Salvia-bastaard. Gorteria 4: 72-74 Adema, F. u. M.T. Jansen (1979): Xanthium L. in Nederland. Gorteria 9(9): 300-303 Adler, W., K. Oswald u. R. Fischer (1994): Exkursionsflora von Österreich (Hrsg.: Fischer, M.). E. Ulmer : 1180 Adolphi, K. (1981): Muscari armeniacum, eine verwilderte Zierpflanze. -
Annotated Checklist of Vascular Flora, Bryce
National Park Service U.S. Department of the Interior Natural Resource Program Center Annotated Checklist of Vascular Flora Bryce Canyon National Park Natural Resource Technical Report NPS/NCPN/NRTR–2009/153 ON THE COVER Matted prickly-phlox (Leptodactylon caespitosum), Bryce Canyon National Park, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Bryce Canyon National Park Natural Resource Technical Report NPS/NCPN/NRTR–2009/153 Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741 Sarah Topp Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 January 2009 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner. The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. -
Vascular Flora of the Liebre Mountains, Western Transverse Ranges, California Steve Boyd Rancho Santa Ana Botanic Garden
Aliso: A Journal of Systematic and Evolutionary Botany Volume 18 | Issue 2 Article 15 1999 Vascular flora of the Liebre Mountains, western Transverse Ranges, California Steve Boyd Rancho Santa Ana Botanic Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Boyd, Steve (1999) "Vascular flora of the Liebre Mountains, western Transverse Ranges, California," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 18: Iss. 2, Article 15. Available at: http://scholarship.claremont.edu/aliso/vol18/iss2/15 Aliso, 18(2), pp. 93-139 © 1999, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 VASCULAR FLORA OF THE LIEBRE MOUNTAINS, WESTERN TRANSVERSE RANGES, CALIFORNIA STEVE BOYD Rancho Santa Ana Botanic Garden 1500 N. College Avenue Claremont, Calif. 91711 ABSTRACT The Liebre Mountains form a discrete unit of the Transverse Ranges of southern California. Geo graphically, the range is transitional to the San Gabriel Mountains, Inner Coast Ranges, Tehachapi Mountains, and Mojave Desert. A total of 1010 vascular plant taxa was recorded from the range, representing 104 families and 400 genera. The ratio of native vs. nonnative elements of the flora is 4:1, similar to that documented in other areas of cismontane southern California. The range is note worthy for the diversity of Quercus and oak-dominated vegetation. A total of 32 sensitive plant taxa (rare, threatened or endangered) was recorded from the range. Key words: Liebre Mountains, Transverse Ranges, southern California, flora, sensitive plants. INTRODUCTION belt and Peirson's (1935) handbook of trees and shrubs. Published documentation of the San Bernar The Transverse Ranges are one of southern Califor dino Mountains is little better, limited to Parish's nia's most prominent physiographic features. -
Vascular Flora of West Clear Creek Wilderness, Coconino and Yavapai
VASCULAR FLORA OF WEST CLEAR CREEK WILDERNESS, COCONINO AND YAVAPAI COUNTIES, ARIZONA By Wendy C. McBride A Thesis Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Science in Biology Northern Arizona University May 2016 Approved: Tina J. Ayers, Ph.D., Chair Randall W. Scott, Ph.D. Liza M. Holeski, Ph.D. ABSTRACT VASCULAR FLORA OF WEST CLEAR CREEK WILDERNESS, COCONINO AND YAVAPAI COUNTIES, ARIZONA WENDY C. MCBRIDE West Clear Creek Wilderness bisects the Mogollon Rim in Arizona, and is nested between the Colorado Plateau and Basin and Range physiographic provinces. Between 2013 and 2016, a floristic inventory vouchered 542 taxa and reviewed 428 previous collections to produce a total plant inventory of 594 taxa from 93 families and 332 genera. The most species rich families Were Asteraceae, Poaceae, Fabaceae, Brassicaceae, Rosaceae, Plantaginaceae, Cyperaceae, and Polygonaceae. Carex, Erigeron, Bromus, Muhlenbergia, and Oenothera Were the most represented genera. Nonnative taxa accounted for seven percent of the total flora. Stachys albens was vouchered as a new state record for Arizona. New county records include Graptopetalum rusbyi (Coconino), Pseudognaphalium pringlei (Coconino), Phaseolus pedicellatus var. grayanus (Coconino), and Quercus rugosa (Coconino and Yavapai). This study quantified and contrasted native species diversity in canyon versus non- canyon floras across the Southwest. Analyses based on eighteen floras indicate that those centered about a major canyon feature shoW greater diversity than non-canyon floras. Regression models revealed that presence of a canyon Was a better predictor of similarity between floras than was the distance betWeen them. This study documents the remarkable diversity found Within canyon systems and the critical, yet varied, habitat they provide in the southwestern U.S. -
Cryptantha of Southern California
Crossosoma 35(1), Spring-Summer 2009 1 CRYPTANTHA OF SOUTHERN CALIFORNIA Michael G. Simpson and Kristen E. Hasenstab Department of Biology San Diego State University San Diego, California 92182 USA [email protected]; [email protected] (Current address for K. Hasenstab: Rancho Santa Ana Botanic Garden, Claremont, 1500 N. College Ave., California 91711) ABSTRACT: The genus Cryptantha (Boraginaceae) contains 202 species, with 49 species and 56 taxa (including varieties) occurring in Southern California, defined here as including the entire Southwestern California region and Tehachapi Mountain region of the California Floristic province, the entire Desert province, and most of the White and Inyo Mountain subregion of the Great Basin province. The purposes of this article are 1) to summarize the current taxonomy of Cryptantha species and infraspecies in Southern California; 2) to provide taxonomic keys and images illustrating the diagnostic features for identification; and 3) to review the distribution, environmental factors, and current conservation status of these taxa. KEYWORDS: Cryptantha, Boraginaceae, taxonomy, identification. INTRODUCTION Taxonomic History and Nomenclature Cryptantha Lehmann ex G. Don, commonly known as “popcorn flower” or “cat’s eye,” is a genus within the family Boraginaceae. The circumscription of this family has changed repeatedly over the last twenty years [Engler and Prantl 1897, Heywood et al. 2007, Gottschling et al. 2001, Angiosperm Phylogeny Group (APG II) 2003], with various authors recognizing either a broad or narrow family concept. Here we accept the APG II (2003) system of classification, which recognizes a broad Boraginaceae. As treated in this manner, the family may be divided into subfamilies Boraginoideae, Cordioideae, Eretioideae, Heliotropoideae, Hydrophylloideae, and (possibly) Lennoideae (see Stevens 2001 onwards). -
Climate Change Vulnerability Assessment for Rare Plants of the San Juan Region of Colorado
Climate Change Vulnerability Assessment for Rare Plants of the San Juan Region of Colorado May 2014 CNHP’s mission is to preserve the natural diversity of life by contributing the essential scientific foundation that leads to lasting conservation of Colorado's biological wealth. Colorado Natural Heritage Program Warner College of Natural Resources Colorado State University 1475 Campus Delivery Fort Collins, CO 80523 (970) 491-7331 Report Prepared for: Tres Rios Bureau of Land Management & San Juan U.S. Forest Service Recommended Citation: J. Handwerk, B. Kuhn, R. Rondeau, and L. Grunau. 2014. Climate Change Vulnerability Assessment for Rare Plants of the San Juan Region of Colorado. Colorado Natural Heritage Program, Colorado State University, Fort Collins, Colorado. Front Cover from the top: Triteleia grandiflora, © Bob Dorn; Amsonia jonesii, © Joe Leahy; Eriophorum chamissonis, © Delia Malone; Drosera anglica, © Sara Brinton; Mimulus eastwoodiae, © Lori Brummer Climate Change Vulnerability Assessment for Rare Plants of the San Juan Region of Colorado Jill Handwerk, Bernadette Kuhn, Renée Rondeau, and Lee Grunau Colorado Natural Heritage Program Warner College of Natural Resources Colorado State University Fort Collins, Colorado 80523 May 2014 ABSTRACT The 5 million-acre San Juan Region lies in southwest Colorado and is considered part of the Four Corners area. The Colorado Natural Heritage Program tracks 122 plant species within this region. Recently published climate models for the Southern San Juan Region project a median decrease in May precipitation across southwestern Colorado and a 4.3 F increase in average annual temperatures, suggesting drought may become problematic for these species in the future (Lukas et al. 2014). -
Bulletin of the Natural History Museum
Bulletin of _ The Natural History Bfit-RSH MU8&M PRIteifTBD QENERAl LIBRARY Botany Series VOLUME 23 NUMBER 2 25 NOVEMBER 1993 The Bulletin of The Natural History Museum (formerly: Bulletin of the British Museum (Natural History)), instituted in 1949, is issued in four scientific series, Botany, Entomology, Geology (incorporating Mineralogy) and Zoology. The Botany Series is edited in the Museum's Department of Botany Keeper of Botany: Dr S. Blackmore Editor of Bulletin: Dr R. Huxley Assistant Editor: Mrs M.J. West Papers in the Bulletin are primarily the results of research carried out on the unique and ever- growing collections of the Museum, both by the scientific staff and by specialists from elsewhere who make use of the Museum's resources. Many of the papers are works of reference that will remain indispensable for years to come. All papers submitted for publication are subjected to external peer review for acceptance. A volume contains about 160 pages, made up by two numbers, published in the Spring and Autumn. Subscriptions may be placed for one or more of the series on an annual basis. Individual numbers and back numbers can be purchased and a Bulletin catalogue, by series, is available. Orders and enquiries should be sent to: Intercept Ltd. P.O. Box 716 Andover Hampshire SPIO lYG Telephone: (0264) 334748 Fax: (0264) 334058 WorW Lwr abbreviation: Bull. nat. Hist. Mus. Lond. (Bot.) © The Natural History Museum, 1993 Botany Series ISSN 0968-0446 Vol. 23, No. 2, pp. 55-177 The Natural History Museum Cromwell Road London SW7 5BD Issued 25 November 1993 Typeset by Ann Buchan (Typesetters), Middlesex Printed in Great Britain at The Alden Press.