National Park Service U.S. Department of the Interior

Natural Resource Program Center

Annotated Checklist of Vascular Flora Cedar Breaks National Monument

Natural Resource Technical Report NPS/NCPN/NRTR—2009/173 ON THE COVER Peterson’s campion (Silene petersonii), Cedar Breaks National Monument, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Cedar Breaks National Monument

Natural Resource Technical Report NPS/NCPN/NRTR—2009/173

Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

February 2009

U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner.

The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientifi c literature reviews; and peer- reviewed proceedings of technical workshops, conferences, or symposia.

Views, statements, fi ndings, conclusions, recommendations, and data in this report are solely those of the author(s) and do not necessarily refl ect views and policies of the U.S. Department of the Interior, National Park Service. Mention of trade names or commercial products does not constitute endorsement or recommendation for use by the National Park Service.

Printed copies of reports in this series may be produced in limited quantity, and are only available as long as the supply lasts. You may send a request to:

Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

This report is also available electronically from the Northern Colorado Plateau Network website, http://science.nature.nps.gov/im/units/ncpn/index.cfm, or at http://www.nature.nps.gov/publications/NRPM/index.cfm.

Please cite this publication as: Fertig, W. 2009. Annotated checklist of vascular fl ora: Cedar Breaks National Monument. Natural Resource Technical Report NPS/NCPN/NRTR—2009/173. National Park Service, Fort Collins, Colorado.

NPS D-225, February 2009 ii Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justifi cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 2 1.3.3 Vegetation ...... 2 1.3.4 Previous botanical studies ...... 4

2 Methods ...... 9

3 Results ...... 11 3.1 Summary of the fl ora of Cedar Breaks National Monument ...... 11 3.2 Life form and biogeographic diversity ...... 12 3.3 Plants of special concern ...... 12 3.4 Non-native species and noxious weeds ...... 12 4 Discussion ...... 15

5 Literature Cited ...... 17

Appendices ...... 21 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument organized by family and scientifi c name with taxonomic and biogeographic notes...... 23 Appendix B. Potential (unconfi rmed) vascular plant taxa of Cedar Breaks National Monument...... 56 Appendix C. Rejected (falsely reported or questionable) vascular plant taxa of Cedar Breaks National Monument...... 69 Appendix D. Confi rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument organized by life form with ecological notes...... 73

Contents iii

Figures

Figure 1.3.1. Location of Cedar Breaks National Monument, Iron County, Utah...... 3 Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Cedar Breaks National Monument by decade, 1930–2007...... 7

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Cedar Breaks National Monument...... 5 Table 3.1. Statistical summary of the fl ora of Cedar Breaks National Monument...... 11 Table 3.3. Plant species of conservation concern from Cedar Breaks National Monument...... 13

Contents v

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Cedar Breaks National Monument. This entailed a thorough review of existing literature, re-examination of specimens from the Cedar Breaks National Monument herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports and locate new species. Based on this study, 345 vascu- lar plant taxa have been confi rmed or reliably reported as occuring in the monument (338 are known from validated herbarium specimens and 7 from unvouchered literature reports). Another 40 species formerly considered part of the monument fl ora have now been shown to be falsely reported. The cur- rent fl ora of Cedar Breaks National Monument represents 9.6% of the known fl ora of Utah and 21.5% of the fl ora of the Utah High Plateaus ecoregion. At least 18 of these species were formerly candidates for potential listing under the Endangered Species Act or are considered species of concern by the state’s natural heritage program. Introduced plants make up 5.2% of the monument fl ora (18 species) but none are considered noxious weeds by the State of Utah. Further surveys in Cedar Breaks National Monument are likely to fi nd additional new species (217 are known from similar habitats outside the monument boundary), especially among fall-fl owering, weedy, or wetland taxa. Further fl oristic stud- ies will be needed if the Ashdown Gorge Wilderness Area ultimately becomes part of Cedar Breaks National Monument.

Contents vii

Acknowledgements

I would like to thank the following individuals for their assistance with this project: Dr. Angela Even- den, botanist and former leader of the NCPN program in Moab, was an active proponent of this pro- ject and helped secure funding for fi eld work and herbarium studies in 2006. Dr. Douglas N. Reynolds assisted with fi eld work in 2007, helping me add 19 new species to the Cedar Breaks fl ora and keeping me from falling off of cliff s. Leslie Courtright, curator of museum collections for Zion National Park and Cedar Breaks National Monument, provided access to all mounted and unmounted herbarium specimens from the monument. Laura Fertig recorded specimen-label data from all mounted col- lections in the Cedar Breaks National Monument herbarium. Paul Roelandt, Cedar Breaks National Monument Superintendent, provided research permits for work in 2006–2007. Libby Nance, former database specialist with the NCPN, assisted with development and quality control of the monu- ment’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the monument. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep allowed access to collections at the Utah State University herbarium. Dr. Ron Hartman and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Sarah Topp and Amy Tendick, of the NCPN vegetation mapping team, shared collections and fi eld data from their 2007 fi eldwork. Rebecca Harms and Larry Stevens shared specimen data from their 2005 springs and seeps inventory. Thanks to technical reviewers Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance for their comments and suggestions. To any others whom I may have inadvertently forgotten, please accept my apologies (and thanks).

Research was conducted under permit # CEBR-2006-SCI-0002 for study number CEBR-00401. Fun- ding was provided by the National Park Service through the Rocky Mountains Cooperative Ecosystem Studies Unit and the University of Wyoming.

Contents ix

1 Introduction (Evenden et al. 2002). This eff ort has entailed re- viewing existing species checklists, corroborating the identifi cation of specimens in park museum In 2006, the National Park Service (NPS) estab- and herbarium collections, locating additional lished a cooperative agreement with the Univer- specimen records from other museums, mining sity of Wyoming, through the Rocky Mountains relevant literature and datasets, and conducting Cooperative Ecosystem Studies Unit, to produce fi eld surveys to locate new taxa or confi rm un- a series of technical reports summarizing the vas- substantiated reports.† All of these data have been cular plant fl ora of each of the 16 national park synthesized in NPSpecies, the park service’s stan- units of the Northern Colorado Plateau Network dardized database for documenting park status (NCPN). These reports are based on existing in- (present, probably present, unconfi rmed, falsely formation in the park service’s NPSpecies data- reported), abundance, residency, nativity, and base system and augmented by further data min- rarity. With the production of a technical series of ing, reviews of state and regional herbaria, and reports on the fl ora of each park, the NPSpecies new fi eld surveys to resolve the status of many data will now be available to a broader audience. reported and unconfi rmed species. The objective of these reports is to summarize the current state of knowledge of the fl ora of each park for the 1.2 Objectives benefi t of NPS biologists and interpretive staff , The objectives of this project were to: outside researchers, and the general public. The following report addresses the vascular fl ora of 1. Document the current status (present, Cedar Breaks National Monument, in southwest- reported, potential, or falsely reported) of ern Utah. Other NCPN park units are addressed all vascular plant species that have been in separate reports. cited as occurring in Cedar Breaks National Monument, based on a review of herbarium collections and past literature. 1.1 Background and justifi cation One of the primary missions of NPS is “to con- 2. Develop an annotated checklist of the fl ora serve unimpaired the natural and cultural resourc- of Cedar Breaks National Monument with es and values of the national park system for the supplemental information on taxonomic enjoyment of this and future generations” (NPS synonyms, distribution, within-park abun- 2000). Due to their high degree of protection and dance, fl owering period, growth form, nativ- distribution across a wide variety of habitat types, ity, habitat, and sources for each record. the nation’s national park units make a signifi - cant contribution to the protection of biological 1.3 Study area diversity (Noss and Cooperrider 1994). Unfortu- nately, the full contribution of the NPS system is 1.3.1 History and setting not known, as many park units have traditionally To Native Americans, the multi-colored badlands lacked comprehensive biological inventories or of what is now Cedar Breaks National Monu- monitoring programs to identify the composition ment (NM) were known as “the circle of painted and condition of their biota. Knowing the status cliff s,” or the “place where the rocks are sliding of park fl ora and fauna is critical for eff ective park down all the time.” Early Mormon settlers named management and for systematically identifying the area Cedar Breaks after the abundance of ju- and fi lling gaps in the nation’s protected area net- niper (locally known as “cedar”) and the abrupt work (Margules and Pressey 2000). cliff s, or breaks. In August 1933, President Frank- lin Roosevelt established Cedar Breaks National To rectify this shortcoming, Congress passed the Monument to preserve the area’s “spectacular National Parks Omnibus Management Act in 1998 cliff s, canyons, and features of scenic, scientifi c, to provide guidance and funding for an inventory and education interest contained therein. . . .” and monitoring program across the entire Na- (Evenden et al. 2002). tional Park System. Since 2000, the NCPN Inven- tory and Monitoring Program has been develop- Cedar Breaks NM is located in southeastern Iron ing and verifying species lists of vertebrates and County, Utah, approximately 18 miles east vascular plants for each park unit in the network

†Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 of Cedar City (Figure 1.3.1). The monument in- These layers have been undergoing extensive ero- cludes the colorful badlands of the Cedar Breaks sion for the past 10 million years as the Marka- Amphitheater, as well as the head of Ashdown gunt Plateau has been uplifted. The carving of the Canyon and the western rim of the Markagunt amphitheater rim continues to this day at a rate of Plateau. To the west, the monument is bordered 24–120 cm per century (Chronic 1988). by the Ashdown Gorge Wilderness Area, man- aged by Dixie National Forest. Recently, the Iron The area of Cedar Breaks NM to the east of the County commissioners have proposed expand- amphitheater rim is capped by lake and stream ing the monument’s boundary to incorporate the deposits of the Oligocene-age Brian Head For- Ashdown Wilderness and redesignate the area as mation. Some of the Brian Head sediments were Cedar Breaks National Park. derived from tuff deposited by extensive volcanic fi elds to the north and west of the Markagunt As presently confi gured, Cedar Breaks NM cov- Plateau. In places, the Brian Head Formation is ers 2,491 hectares (6,155 acres), and ranges in overtopped by the Markagunt Megabreccia, a elevation from 2,469 meters (8,100 feet) at the deposit of coarse, broken rock embedded in fi ne, bottom of Ashdown Creek Canyon to 3,250 me- volcanic ash. These deposits were originally laid ters (10,662 feet) above the amphitheater rim down east of the monument, but moved into their (Evenden et al. 2002). The Cedar Breaks area is current position by massive gravity sliding along characterized by cold winters (averaging –1.9° C weak shear planes in the poorly consolidated in January) with abundant snow (averaging 1.9–2 Brian Head layers (Hatfi eld et al. 2003). Also scat- meters), and short, cool summers (average July tered along the east rim of the plateau are Qua- temperature is 16.6° C). About 24% of annual ternary-age landslides (such as the area around precipitation comes from summer thunderstorms Alpine Pond) and eolian deposits. The extensive (Roberts and Jean 1989). Pleistocene basalt fl ows found over much of the Markagunt Plateau are not present within the 1.3.2 Geology monument. Cedar Breaks National Monument is located 1.3.3 Vegetation along the west rim of the Markagunt Plateau, an east-dipping sedimentary fault block uplifted Roberts and Jean (1989) recognized 17 habitat nearly 10,000 feet by the Hurricane Fault (Hat- types and 12 seral community types within Cedar fi eld et al. 2003). The lowermost strata exposed Breaks National Monument. These can be aggre- within the monument are Upper Cretaceous shal- gated into fi ve main vegetation types following low marine sandstones and shales of the Straight elevation, soil, and moisture gradients (more de- Cliff s Formation, and river deposits of the Grand tailed vegetation descriptions and maps are avail- Castle and Wahweap sandstones. Along Ash- able in NCPN in ed.). down Creek, these layers have been buried by Quaternary-age alluvial terraces and fan deposits. The bottom of Ashdown Canyon and the lower- The Cedar Breaks Amphitheater exposes over most slopes of the amphitheater are dominated 1,400 feet of reddish-orange and white Claron by forests of white fi r (Abies concolor) and Doug- limestone that has been eroded by water, freeze- las-fi r (Pseudotsuga menziesii). These forests are thaw action, and mass wasting into irregular strongly correlated with outcrops of Cretaceous spires, hoodoos, and steep cliff s. The Claron For- sandstones or Quaternary alluvium. Moister sites mation (sometimes also referred to as the Cedar have an understory of Oregon-grape (Mahonia City or Wasatch Formation) consists of alternat- repens) or common juniper (Juniperus communis), ing bands of ledge-forming sandy limestone and while drier areas contain greenleaf manzanita slope-forming calcareous sandstone that were (Arctostaphylos patula), Rocky Mountain maple deposited at the bottom of a large, freshwater lake (Acer glabrum), or curl-leaf mountain mahogany during the Paleocene and Eocene (Hatfi eld et al. (Cercocarpus ledifolius). Quaking aspen (Populus 2003). This lake extended from near the Nevada tremuloides) may be locally abundant within white state line east toward Bryce Canyon and Boulder fi r forests, especially on deeper, loamier soils. Mountain, and was slightly larger than modern Lake Erie (Chronic 1988). The lower to upper- A narrow band of riparian forest vegetation middle slopes of the amphitheater consist of the dominated by narrow-leaved cottonwood Red Member of the Claron, while the rim of the (Populus angustifolia), Engelmann spruce (Picea breaks is the White Member (Hatfi eld et al. 2003). engelmannii), and water birch (Betula occidental-

2 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Cedar Breaks NationalCedar MonumentCapito BreaksZion l Reef NationalNational National ParkMonument Park Utah Utah

143

143

Visitor Center

148

010.5 Map location Kilometers 010.5 Miles

Established August 22, 1933 2,469–3,250 m elevation Legend (8,100–10,662 ft elevation) 2,491 ha (6,154 ac) Intermittent streams Private

Paved roads U.S. Forest Service (USFS)

Ashdown Gorge Park boundary (NPS) (USFS wilderness area) NCPN boundary

MAP CREATED BY CYNTHIA WANSCHURA, ZION NATIONAL PARK

Figure 1.3.1. Location of Cedar Breaks National Monument, Iron County, Utah.

Chapter 1: Introduction 3 is) occurs on alluvial terraces bordering Ashdown Subalpine meadows occur above the rim and Creek and its main tributaries. A number of small on the summit fl ats of Cedar Breaks NM in springs are located above the terraces in the bot- openings interspersed among subalpine fi r/En- tom of Ashdown Canyon and are locally domi- gelmann spruce forests. Better-drained soils sup- nated by white fi r or blue spruce (Picea pungens), port the bear daisy (Erigeron ursinus)/Letterman horsetails, and various sedges (Roberts and Jean needlegrass (Stipa lettermanii) habitat type of 1989). Small areas of wetland vegetation associ- Roberts and Jean (1989). During early summer, ated with springs or small, perennial streams also these meadows support a rich array of blooming occur on the summit plateau above the amphithe- wildfl owers, including Markagunt penstemon ater rim, often on volcanic-derived soils or land- (Penstemon leiophyllus), little sunfl ower (Helian- slide deposits. Alpine Pond supports dense stands thella unifl ora), elkweed (Swertia radiata), and of water sedge (Carex aquatilis), small-wing sedge scarlet paintbrush (Castilleja miniata). This com- (C. microptera), and heartleaf bittercress (Car- munity grades into the wedge-leaf cinquefoil (Po- damine cordifolia), while marshy meadows near tentilla diversifolia)/tufted hairgrass habitat type Sunset Spring are dominated by beaked sedge on more permanently saturated soils (Roberts (Carex utriculata), tufted hairgrass (Deschampsia and Jean 1989). cespitosa), Baltic rush (Juncus arcticus), shrubby cinquefoil (Potentilla fruticosa), and a variety of 1.3.4 Previous botanical studies forbs (Springer et al. 2006). These wetlands grade into several subalpine fi r forest and meadow habi- The full extent of botanical studies done prior tat types described by Roberts and Jean (1989). to the establishment of Cedar Breaks National Monument remains poorly documented. Siebert Barren slopes and rim areas of the Red and White (1924) conducted one of the earliest ecological members of the Claron Formation are dominated studies of the Cedar Breaks area, focusing on al- by cushion plants and widely scattered western pine ecology (Table 1.3.4). The oldest confi rmed bristlecone pine (Pinus longaeva) or limber pine herbarium records from the monument area (P. fl exilis). Roberts and Jean (1989) named the were made in 1925–1926, by NPS collector An- Panguitch buckwheat (Eriogonum panguicense)/ gus Woodbury (who also documented much of interior bluegrass (Poa glauca) habitat type to de- the early fl ora of Zion National Park). Pioneer scribe the sparsely vegetated cushion plant stands Utah botanists Marcus E. Jones and Walter Cot- on calcareous, clayey soils along rim areas. Bris- tam made at least two collections in the area in tlecone/limber pine stands are more extensive, 1929. George Goodman and C. Leo Hitchcock covering most of the slopes of the Cedar Breaks collected at least a dozen specimens for the New Amphitheater. Claron rims and slopes provide York Botanical Garden in 1930, including the ho- habitat for most of the rare, endemic plant spe- lotypes of Breaks draba (Draba subalpina) and cies of the monument (Fertig and Reynolds 2007; Cedar Breaks wild buckwheat (Eriogonum pan- Roberts and Jean 1989). guicense var. alpestre) (Table 1.3.4, Figure 1.3.4).

Forests above the amphitheater rim and on the Records become more complete after the cre- summit of Cedar Breaks NM consist primarily of ation of Cedar Breaks NM in 1933. During the subalpine fi r (Abies lasiocarpa) and Engelmann late 1930s, 48 new species were discovered in spruce, though pockets of aspen, Douglas-fi r, the monument through the eff orts of G. Y. Croft, limber pine, or western bristlecone pine also oc- collecting for the Cedar Breaks NM herbarium casionally occur. These forests are associated with (CEBR), and Bassett Maguire, of Utah State Uni- cobbly, clay loams derived from volcanic rhyolite versity, who collected the type of Cedar Breaks and Claron limestone. Dominant understory spe- daisy (Erigeron sionis var. trilobatus) in or near cies include aspen bluebell (Mertensia arizonica) the monument. Many other prominent taxono- on wetter soils and alpine prickly currant (Ribes mists visited Cedar Breaks NM in the late 1930s, montigenum), Oregon-grape, common juniper, including Ralph Gierisch, Alice Eastwood, and and Ross’ sedge (Carex rossii) on drier sites. John T. Howell. While still common in the monument, 80–90% of mature Engelmann spruces have been killed by Collecting was more sporadic from 1940 to 1975. spruce bark beetles in an epidemic that began in Thirty-six new species were added to the monu- 1993 (Evenden et al. 2002). ment fl ora during the 1940s, by W. S. Boyle and J. L. Crawford (collecting for the CEBR herbarium)

4 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Table 1.3.4. Chronology of plant-collecting efforts in Cedar Breaks National Monument. # of new Year Collector(s) Comments taxa 1924 F. R. Siebert ? Florence Siebert (1924) studied the alpine ecology of Cedar Breaks area for her Master’s thesis at the University of Chicago, but her collections were not examined for this study. 1925– A. M. Woodbury 2 Mountain death camas (Zigadenus elegans) and common yarrow (Achillea 1926 millefolium) were the fi rst two plant species collected from the monument in the CEBR herbarium. 1929 M. E. Jones and W. 2 Marcus E. Jones and Walter Cottam made at least two collections from the Cottam monument (specimens at BRY and NY). 1930 G. Goodman & C. L. 12 George Goodman and C. Leo Hitchcock collected the type specimens of Draba Hitchcock subalpina and Eriogonum panguicense var. alpestre from Cedar Breaks. 1934– B. Maguire & G. Y. 48 Bassett Maguire collected the types of King’s plateau sandwort (Arenaria kingii 1939 Croft ssp. plateauensis, now a synonym of A. fendleri var. glabrescens) and Cedar Breaks daisy (Erigeron proselyticus, now E. sionis var. trilobatus) from Cedar Breaks in 1934 (specimens at NY). G. Y. Croft made more than 60 collections for the CEBR herbarium. Additional noteworthy collectors during this period included Ralph Gierisch, C. L. Hitchcock, Alice Eastwood, and John T. Howell. 1940– W. S. Boyle & B. F. 11 W. S. Boyle made more than 30 collections for the CEBR herbarium, including 1941 Harrison 10 new taxa. B. F. Harrison added one new species in 1940. 1942 O. Degener & L. 9 Otto Degener and Leroy Peiler made more than 20 collections (mostly at NY), Peiler including the fi rst record of Navajo Lake milkvetch (Astragalus limnocharis) from the monument. Agrostologist Frank Gould also collected in Cedar Breaks in 1942. 1946– J. L. Crawford and L. 16 J. L. Crawford collected 51 specimens for the CEBR herbarium, including fi rst 1947 Woodbury records for 16 taxa. L. Woodbury added one new species (BRY records). 1952 M. E. Lewis 1 Mont Lewis discovered kobresia sedge (Carex elynoides) in Cedar Breaks National Monument. 1954– A. F. Bruhn 44 A. F. Bruhn made nearly 100 collections for the CEBR herbarium, including 44 1955 fi rst records for the monument. 1966 R. Barneby 1 Rupert Barneby added Panguitch milkvetch (Astragalus argophyllus var. panguicensis) to the local fl ora. 1970 R. A. Nelson 15 Ruth Nelson collected more than 40 specimens (mostly deposited at RM), of which 15 represented new species for the monument (including the fi rst record of Payson’s daisy, Erigeron vagus). 1977– S. Dickman & H. 17 S. Dickman and Hayle Buchanan made more than 120 collections for the CEBR 1979 Buchanan herbarium, including fi rst reports for 17 new taxa. Noteworthy among these is the fi rst record of Zion draba (Draba asprella var. zionensis). 1981 B. C. Palmer & L. A. 76 Brent Palmer, of Southern Utah University, collected more than 220 specimens Palmer while conducting a systematic fl oristic inventory of Cedar Breaks NM. The Palmers compiled an unpublished checklist of 245 species in 47 families in 1981 (Roberts and Jean 1989). 1982 S. L. Welsh 1 Welsh’s collection of mountain kentrophyta (Astragalus kentrophyta var. reported tegetarius, deposited at NY) has not been corroborated. (All specimens I’ve examined from the monument have been var. elatus.) 1986– C. Johnson 3 C. Johnson documented three new exotic species for the monument. 1987 1988– C. Jean 44 Cathie Jean collected or reported (without vouchers) 49 new species while 1989 (+ 5 mapping major vegetation types and the distribution of selected rare plant reported) species of Cedar Breaks NM (Roberts and Jean 1989). Jean discovered several new rare species, including Reveal’s paintbrush (Castilleja parvula var. revealii), Cedar Breaks goldenbush (Haplopappus zionis), Zion jamesia (Jamesia americana var. zionis), Podunk groundsel (Senecio malmstenii), and Bryce Canyon townsendia (Townsendia montana var. minima).

Chapter 1: Introduction 5 Table 1.3.4. Chronology of plant-collecting efforts in Cedar Breaks National Monument. # of new Year Collector(s) Comments taxa 1994 N. D. Atwood 0 Duane Atwood documented the fi rst two populations of Arizona willow (Salix arizonica) in Cedar Breaks NM. In 2005, I found a misidentifi ed collection of this species by Brent Palmer in the CEBR collection, moving the earliest date for Arizona willow back to 1981. 1996 K. Mason 1 Karen Mason discovered lambsquarter (Chenopodium album var. album) in the monument. 2001 D. Louie, M. Malm, J. 1 Margaret Malm, Denise Louie, and Jason Alexander collected 24 specimens Alexander from the monument, including the fi rst record of King’s lupine (Lupinus kingii var. kingii). 2005 R. Harms and NPS 4 Rebecca Harms discovered 4 new riparian species for Cedar Breaks NM as part vegetation mapping (+ 1 of the NCPN springs and seeps inventory (Springer et al. 2006). The NCPN (in team reported) ed.) vegetation mapping team reported one new species for the monument (without a voucher). 2006– W. Fertig, D. N. 31 Fertig and Reynolds (2007) documented 30 new species while conducting rare 2007 Reynolds, & A. plant surveys across Cedar Breaks NM. Amy Tendick, of the NCPN vegetation Tendick mapping team, added one new species in 2007. BRY = Brigham Young University herbarium; CEBR = Cedar Breaks National Monument herbarium; NY = New York Botanical Garden; RM = University of Wyoming Rocky Mountain herbarium

and B. F. Harrison, Otto Degener, and LeRoy Peil- Palmer collected more than 220 specimens for the er (collecting for Brigham Young University and CEBR herbarium and documented 76 new taxa. the New York Botanical Garden). Mont Lewis Palmer’s work resulted in an unpublished monu- and A. F. Bruhn added 45 new species during the ment checklist containing 245 species (Jean and 1950s. New discoveries were especially slow in Palmer 1987). Cathie Jean added 49 species to the the 1960s and early 1970s, with Rupert Barneby Cedar Breaks fl ora in the course of her vegetation (of the New York Botanical Garden) and Ruth A. mapping project (Roberts and Jean 1989). Rebecca Nelson (working for the National Park Service) Harms collected four new riparian species in 2005, adding just 16 new taxa. during a survey of springs and seeps (Springer et al. 2006). In 2006–2007, vegetation mapping crews Since 1977, several signifi cant botanical surveys from the Northern Colorado Plateau Network have been conducted at Cedar Breaks National (in ed.) reported two new species, and Fertig and Monument. Hayle Buchanan and S. Dickman Reynolds (2007) located at least 30 new taxa dur- added 17 new species to the monument fl ora from ing a rare plant survey of the monument. 1977 to 1979 (Buchanan 1992). In 1981, Brent

6 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument 400

350

300

250

200

150

100 Number of vascular taxa

50

0 1930 1940 1950 1960 1970 1980 1990 2000 2007

Year

Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Cedar Breaks National Monument by decade, 1930–2007.

Chapter 1: Introduction 7

formation on life form, geographic range (with- 2 Methods in the context of Utah), population size in the monument (derived from Roberts and Jean 1989; In July 2005, I reviewed all mounted specimens NCPN in ed.; and fi eld observations), fl owering in the Cedar Breaks National Monument herbar- period, distribution across general vegetation ium (housed at Zion National Park in Springdale, types, nativity, data source (collector and collec- Utah) to determine whether each was correctly tion number from a sample specimen or literature identifi ed to species, subspecies, or variety fol- citation), year of earliest collection (from herbar- lowing the nomenclature of Welsh et al. (2003). ium records), and relevant synonymy and taxo- As necessary, I made corrections and annotations nomic problems (based on review of Barkworth to each specimen and to the voucher records in et al. 2003, 2007; Barneby 1989; Cronquist 1994; the monument’s Automated National Cataloging Cronquist et al. 1972, 1977, 1984, 1997; Dorn System (ANCS+) and NPSpecies database. 2001; Flora of North America Editorial Commit- tee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, Based on this herbarium work and a review of 2006a, 2006b, 2006c; Holmgren et al. 2005; and monument references (Buchanan 1992; Roberts Weber and Wittmann 2001). The annotations and Jean 1989), I developed four lists: to these lists and label data from the CEBR her- barium review were then incorporated into the 1. A revised checklist of species that were pres- NPSpecies database for the monument. ent in the park (confi rmed by an authenti- cated voucher; included in Appendices A Work in 2006–2007 focused on confi rmation of and D); reported and potential species for the monument 2. A list of species that were reported in the lit- fl ora through fi eld surveys and review of collec- erature but not corroborated with a voucher tions from state and regional herbaria. I made 14 (equivalent to “probably present” in NPSpe- collecting trips to Cedar Breaks NM in July and cies terminology; included in Appendices A September 2006 and June–August 2007 to tar- and D); get missing species, emphasizing under-sampled wetland, weedy, and late-fl owering taxa. Herbar- 3. A list of potential species (“unconfi rmed” in ium studies were undertaken at Brigham Young NPSpecies) known from the vicinity of Ce- University (BRY), Utah State University (USU), dar Breaks NM based on Albee et al. (1988) and the University of Wyoming (RM) in March, and an unpublished list developed for the June, and October 2006, and February and March NCPN by Dr. N. Duane Atwood of Brigham 2007. I also documented monument specimens Young University (included in Appendix B); from the Virtual Herbarium website of the New and York Botanical Garden (http://sciweb.nybg.org/ science2/hcol/ intf/index.asp). Finally, I found 4. A list of species that are now known to be additional new species reports by reviewing plot misidentifi ed or highly questionable (Ap- data from recent vegetation mapping, weed sur- pendix C). veys, and wetland inventories (Dewey and An- Each list was annotated with supplemental in- dersen 2005; NCPN in ed.; Springer et al. 2006).

Chapter 2: Methods 9

New York Botanical Garden Virtual Herbarium. 3 Results Field surveys during this time netted another 31 new species (Fertig and Reynolds 2007) and 18 3.1 Summary of the fl ora of Cedar additional species previously reported by Rob- erts and Jean (1989) without a voucher. Overall, Breaks National Monument the documented fl ora of Cedar Breaks NM has Of the 691 vascular plant vouchers in the CEBR increased by 58 species (17.2%), bringing the to- herbarium that I examined in 2005 (excluding 72 tal number of taxa to 338 (Table 3.1, Appendices specimens collected outside of the monument), A, D). 132 were misidentifi ed (19.1%), 95 were revised to a more current name (13.7%), 141 had variety Currently, seven additional taxa are reported for or subspecies names added (20.4%), and 323 were Cedar Breaks National Monument but have yet to found to be correctly determined (46.7%). Based be corroborated with a voucher specimen (Table solely on these collections, the confi rmed fl ora of 3.1, Appendix A). Of these, fi ve were reported by Cedar Breaks National Monument stood at 280 Roberts and Jean (1989) and one by the NCPN taxa (Fertig 2006). From 2006 to 2007, I located (in ed.) vegetation mapping crew. One other un- 10 new taxa from collections at Brigham Young confi rmed report (Astragalus kentrophyta var. University, the University of Wyoming, and the tegetarius) comes from the New York Botanical

Table 3.1. Statistical summary of the fl ora of Cedar Breaks National Monument. Number Number Category confi rmed additionally Total in park reported for park Taxonomic diversity Total taxa 338 7 345 (including varieties and subspecies) Full species 326 5 331 (excluding varieties and subspecies) Families 55 0 55 Life form diversity Tree taxa 14 1 15 Shrub taxa 35 1 36 Perennial forb taxa 196 3 199 Annual forb taxa 13 1 14 Perennial graminoid taxa 71 1 72 Annual graminoid taxa 3 0 3 Fern taxa 6 0 6 Biogeographic diversity Introduced taxa 17 1 18 Native taxa Locally endemic taxa 17 0 17 Regionally endemic taxa 19 0 19 Disjunct taxa 1 0 1 Peripheral taxa 0 0 0 Sparse taxa 6 0 6 Widespread taxa 278 6 284 Total native taxa 321 6 327 The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003). See Appendices, “Codes and Defi nitions”, for an explanation of the various categories.

Chapter 3: Results 11 Garden website (Welsh 21286), but may be mis- Of the monument’s 327 native taxa, 284 (86.8%) identifi ed. Taken together, the number of con- are widespread across Utah and western North fi rmed and reported vascular plant taxa in Cedar America. Only 17 species (5.2% of the monu- Breaks National Monument stands at 345 taxa ment’s native fl ora) have extremely limited global (Table 3.1, Appendix A). distributions (local endemics found within an area of less than 1° of latitude × 2° of longitude; Entering the 2006 fi eld season, I recognized 271 see Appendix A for complete defi nitions of bio- taxa as potentially occurring in Cedar Breaks geographic categories). Regional endemics (large- NM, based on the proximity of the monument ly restricted to the Colorado Plateau) comprise to known populations (Albee et al. 1988; Fertig an additional 5.8% of the monument fl ora (Table 2006). At least 40 of these species have since been 3.1). Species that are disjunct or occur sparsely recorded within the monument based on new across Utah account for just 2.1% of the monu- fi eld collections or relocated herbarium speci- ment fl ora. mens. The current number of potential species that I am recognizing for Cedar Breaks National Monument is 217 (Appendix B). 3.3 Plants of special concern Cedar Breaks National Monument has no plants Based on my review of specimens in the CEBR listed as threatened or endangered under the U.S. and BRY herbaria, at least 38 species previously Endangered Species Act. Arizona willow (Salix cited for the monument are now known to be arizonica) was proposed for listing in 1995, but falsely reported based on misidentifi ed vouch- the decision was postponed following implemen- ers. Two additional species reported by Buchanan tation of an interagency conservation agreement (1992) and the New York Botanical Garden web- and strategy (U.S. Forest Service Intermountain site are considered questionable because Cedar Region et al. 1995). At least 13 other monument Breaks NM is well outside their known or ex- taxa were formerly candidates for potential listing pected range (Appendix C). (Category 2 or 3C) under the act (Table 3.3) (Fer- tig and Reynolds 2007). Eight of these species are The confi rmed and reported fl ora of Cedar currently managed as sensitive by the U.S. Forest Breaks NM represents 9.6% of the 3,594 native Service and the U.S. Bureau of Land Manage- and naturalized taxa documented for the state of ment’s Utah State Offi ce. The Utah Conservation Utah by Welsh et al. (2003), and 36% of the state’s Data Center (UTCDC) lists 17 species from Ce- 151 plant families (Table 3.1). Within Iron Coun- dar Breaks NM on its rare, watch, or taxonomic ty, the monument fl ora captures 39.6% of the 871 problems lists (UDWR 1998). Although not cur- taxa reported by Welsh et al. (2003). At the ecore- rently cited on any agency list, Payson’s daisy (Er- gional scale, Cedar Breaks NM contains 21.5% igeron vagus) was included on the list of potential of the 1,608 species reported for the Utah High rare species of Cedar Breaks NM by Fertig and Plateaus ecoregion (Shultz et al. 2006). Reynolds (2007), based on information from the Brigham Young University Herbarium that south- 3.2 Life form and biogeographic ern Utah populations may represent an unde- diversity scribed (and locally endemic) taxon. Although ecologically dominant, tree and shrub species make up only 14.8% of the confi rmed 3.4 Non-native species and noxious and reported plant taxa of Cedar Breaks National weeds Monument (Table 3.1). Perennial forbs are the Introduced plant species (those not historically most species-rich life form in the monument fl o- native to Utah or North America) make up 5.2% of ra, with 199 known and reported taxa (57.7% of the confi rmed and reported fl ora of Cedar Breaks the total fl ora). Annual forbs and grass-like plants National Monument (Table 3.1). This fi gure is comprise less than 5% of the fl ora. Like trees and substantially lower than the statewide average of shrubs, perennial graminoids are ecologically im- 13.5% non-native species (Fertig 2007; Welsh et portant but relatively species-poor, representing al. 2003). Of the monument’s 18 introduced plant less than 21% of the total fl ora. The diversity of species, none are listed as noxious by the State of ferns and fern-allies in Cedar Breaks NM is espe- Utah (Dewey and Andersen 2005; State of Utah cially low (1.7%) (Table 3.1). Department of Agriculture and Food 2007).

12 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Table 3.3. Plant species of conservation concern from Cedar Breaks National Monument. TNC UTCDC Family Species Common name global Legal status status rank Caryophyllaceae Silene petersonii Peterson’s campion G2G3 USFS: Region 4 Sensitive; Watch, USFWS: former C2 candidate S2S3 Compositae Aster wasatchensis Markagunt aster G3? Watch, (Asteraceae) var. wasatchensis TNR S3? Compositae Aster welshii Welsh’s aster G2 Watch, S2 (Asteraceae) Compositae Erigeron sionis var. Cedar Breaks daisy G2 USFWS: former C2 candidate Watch, S2 (Asteraceae) trilobatus Compositae Erigeron vagus Payson’s daisy G4 SNR (Asteraceae) Compositae Haplopappus zionis Cedar Breaks G2 UT BLM: Sensitive; Rare, S2 (Asteraceae) goldenbush USFWS: former C2 candidate Compositae Senecio malmstenii Podunk groundsel G1 USFS: Region 4 Sensitive; Rare, S1? (Asteraceae) USFWS: former C2 candidate Compositae Townsendia Bryce Canyon G4T3 USFWS: former 3C Watch, S3 (Asteraceae) montana var. minima townsendia Cruciferae Draba asprella var. Zion draba G4T3? USFWS: former 3C Watch, (Brassicaceae) zionensis S3? Cruciferae Draba subalpina Breaks draba G3 USFWS: former 3C Watch, S3 (Brassicaceae) Leguminosae Astragalus Navajo Lake milkvetch G2T1 USFS: Region 4 Sensitive; Rare, S1 (Fabaceae) limnocharis var. USFWS: former C2 candidate limnocharis Polemoniaceae Ipomopsis tridactyla Tushar gilia G5T2 Rare, S2 Polygonaceae Eriogonum Cedar Breaks wild G3 USFWS: former 3C TaxProb, panguicense var. buckwheat T2T3Q SSYN alpestre Salicaceae Salix arizonica Arizona willow G2G3 USFS: Region 4 Sensitive; USFWS: Rare, S2 former C1 candidate (dropped in 1995; now equivalent of 3C) Saxifragaceae Jamesia americana Zion jamesia G5T2 USFS: Region 4 Sensitive; BLM Rare, S2 (Hydrangeaceae) var. zionis (UT): Sensitive; USFWS: former C2 candidate Scrophulariaceae Castilleja parvula var. Reveal’s paintbrush G2 USFS: Region 4 Sensitive; Rare, S2 revealii USFWS: former C2 candidate Umbelliferae Cymopterus minimus least spring-parsley G1G2Q USFS: Region 4 Sensitive; USFWS: Rare, S1S2 (Apiaceae) former C2 candidate Umbelliferae Lomatium minimum least lomatium G3 USFWS: former 3C Watch, S3 (Apiaceae) List derived from Fertig and Reynolds (2007).

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Utah under the column UTCDC status. Under legal status, USFWS = U.S. Fish and Wildlife Service. C2 = Category 2 candidate (a former category used for taxa that might warrant being proposed for Threatened or Endangered status following additional research). 3C = Category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. BLM (UT) = U.S. Bureau of Land Management (Utah state offi ce). UTCDC status includes conservation categories adopted by the state’s natural heritage program to prioritize endemic and rare plant taxa (UDWR 1998). These categories include: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western U.S.), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species appear rare, but more information needed before being assigned to a different category).

Chapter 3: Results 13

consist of cryptic species that fl ower late in the 4 Discussion growing season, or occur in undersampled wet- land or weedy habitats. Confi rmation of reported Following a review of herbarium records and re- species should remain a high priority, as some of cent fi eld work, the vascular fl ora of Cedar Breaks these taxa may ultimately prove to be erroneous. National Monument currently stands at 345 con- Additional studies would also be useful to better fi rmed and reported taxa (Table 3.1, Appendix refi ne the abundance and distribution of species of A). Based on available habitats and proximity, at high management interest (especially rare plants least 217 additional plant taxa may occur within and invasive non-natives). Further fl oristic stud- the monument (Appendix B). Should all of these ies are needed in the Ashdown Gorge Wilderness potential species be found, the documented fl ora Area, especially if this ultimately becomes part of of the monument would increase by nearly 41%. an expanded Cedar Breaks National Park. Many of the “missing” potential or reported taxa

Chapter 4: Discussion 15

5 Literature Cited York Botanical Garden. 584 pp. ——. 1984. Volume 4: Subclass Asteridae (except Asteraceae). Intermountain fl ora: Vascular Albee, B. J., L. M. Shultz, and S. Goodrich. 1988. plants of the Intermountain West, U.S.A. Atlas of the vascular plants of Utah. +Occa- Bronx, NY: New York Botanical Garden. sional Publ. 7. Salt Lake City: Utah Museum 573 pp. of Natural History. 670 pp. Cronquist, A., N. H. Holmgren, and P. K. Hol- Barkworth, M. E., K. M. Capels, S. Long, and M. mgren, eds. 1997. Volume 3, Part A, Subclass B. Piep, eds. 2003. Magnoliophyta: Com- Rosidae (except Fabales). Intermountain melinidae (in part): Poaceae, part 2. Flora of fl ora: Vascular plants of the Intermountain North America north of Mexico, Volume 25. West, U.S.A. Bronx, NY: New York Botanical New York: Oxford University Press. 783 pp. Garden. 446 pp. Barkworth, M. E., K. M. Capels, S. Long, L. K. Dewey, S., and K. Andersen. 2005. An inven- Anderton, and M. B. Piep, eds. 2007. Mag- tory of invasive non-native plants in Cedar noliophyta: Commelinidae (in part): Poace- Breaks National Monument (2004): Final ae, part 1. Flora of North America north report. Utah State University. Weed Science of Mexico, Volume 24. New York: Oxford Research Project # SD0515A. University Press. 911 pp. Dorn, R. D. 2001. Vascular plants of Wyoming. Barneby, R. C. 1989. Volume 3, Part B. Fabales. Third edition. Cheyenne, Wy.: Mountain In Cronquist, A., A. H. Holmgren, N. H. West Publishing. 412 pp. Holmgren, J. L. Reveal, and P. K. Holmgren, eds. Intermountain fl ora: Vascular plants of Evenden, A., M. Miller, M. Beer, E. Nance, S. the Intermountain West, U.S.A. Bronx, NY: Daw, A. Wight, M. Estenson, and L. Cudlip. New York Botanical Garden. 279 pp. 2002. Northern Colorado Plateau Vital Signs Network and Prototype Cluster, plan Buchanan, H. 1992. Wildfl owers of southwestern for natural resources monitoring: Phase Utah. Helena, Mt.: Bryce Canyon Natural I report, October 1, 2002 [two volumes]. History Association and Falcon Press Pub- National Park Service, Northern Colorado lishing. 119 pp. Plateau Network, Moab, Ut. 138 pp + app. Chronic, H. 1988. Pages of stone: Geology of Fertig, W. 2006. Herbarium and fi eld studies of western national parks and monuments. vascular plant fl ora of CEBR for National Grand Canyon and the plateau country. Park Service Inventory and Monitoring pro- Seattle, Wa.: The Mountaineers. 158 pp. gram. Invesigator’s Annual Report, National Cronquist, A. 1994. Volume 5, Asterales. In Park Service. 2 pp. Cronquist, A., A. H. Holmgren, N. H. Hol- ——. 2007. Introduced and naturalized plants of mgren, J. L. Reveal, and P. K. Holmgren, eds. Utah. Sego Lily 30(5):7–11. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, NY: New Fertig, W., and D. N. Reynolds. 2007. Survey York Botanical Garden. 496 pp. of rare plants of Cedar Breaks National Monument: Year One report. Prepared for Cronquist, A., A. H. Holmgren, N. H. Holmgren, Cedar Breaks National Monument, Colo- and J. L. Reveal, eds. 1972. Volume 1: Geo- rado Plateau Cooperative Ecosystem Studies logical and botanical history of the region, Unit, and Southern Utah University under its plant geography and a glossary. The CPCESU Cooperative Agreement # H1200- vascular cryptogams and the gymnosperms. 004-0002. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, NY: New Flora of North America Editorial Committee. York Botanical Garden. 270 pp. 1993. Pteridophytes and Gymnosperms. Flora of North America north of Mexico, Cronquist, A., A. H. Holmgren, N. H. Hol- Volume 2. New York: Oxford University mgren, J. L. Reveal, and P. K. Holmgren, Press. 475 pp. eds. 1977. Volume 6: The monocotyledons. Intermountain fl ora: Vascular plants of the ——. 1997. Magnoliophyta: Magnoliidae and Intermountain West, U.S.A. Bronx, NY: New Hamamelidae. Flora of North America

Chapter 5: Literature Cited 17 north of Mexico, Volume 3. New York: Ox- Jean, C., and B. Palmer. 1987. Vascular plant spe- ford University Press. 590 pp. cies list, Cedar Breaks National Monument. 8 pp. ——. 2000. Magnoliophyta: Alismatidae, Are- cidae, Commelinidae (in part), and Zin- Margules, C. R., and R. L. Pressey. 2000. Sys- giberidae. Flora of North America north tematic conservation planning. Nature of Mexico, Volume 22. New York: Oxford 405:243–253. University Press. 352 pp. National Park Service. 2000. Management poli- ——. 2002a. Magnoliophyta: Commelinidae (in cies. U.S. Department of Interior, National part): Cyperaceae. Flora of North America Park Service, Washington, D.C. Publication north of Mexico, Volume 23. New York: # NPS D1416. 137 pp. Oxford University Press. 608 pp. Northern Colorado Plateau Network (NCPN). ——. 2002b. Magnoliophyta: Liliidae: Liliales and in ed. Cedar Breaks National Monument Orchidales. Flora of North America north vegetation classifi cation and mapping of Mexico, Volume 26. New York: Oxford project. Natural Resource Technical Report. University Press. 723 pp. National Park Service, Denver, Co. ——. 2003. Magnoliophyta: Caryophyllidae, part Noss, R. F., and A. Y. Cooperrider. 1994. Saving 1. Flora of North America north of Mexico. nature’s legacy: Protecting and restoring Volume 4. New York: Oxford University biodiversity. Washington, D.C.: Island Press. Press. 559 pp. 416 pp. ——. 2005. Magnoliophyta: Caryophyllidae, part Roberts, D. W., and C. Jean. 1989. Plant com- 2. Flora of North America north of Mexico. munity and rare and exotic species dis- Volume 5. New York: Oxford University tribution and dynamics in Cedar Breaks Press. 656 pp. National Monument. Department of Forest Resources and Ecology Center, Utah State ——. 2006a. Magnoliophyta: Asteridae, part 6: University, Logan, Ut. 144 pp. Asteraceae, part 1. Flora of North America north of Mexico. Volume 19. New York: Shultz, L. M., R. D. Ramsey, and W. Lindquist. Oxford University Press. 579 pp. 2006. Revised atlas of Utah Plants. College of Natural Resources, Utah State University, ——. 2006b. Magnoliophyta: Asteridae, part 7: Logan, Ut. http://earth.gis.usu.edu/plants. Asteraceae, part 2. Flora of North America north of Mexico. Volume 20. New York: Siebert, F. R. 1924. An ecological study of the Oxford University Press. 666 pp. alpine zone of Cedar Breaks, southern Utah. Master’s thesis, University of Chicago, Chi- ——. 2006c. Magnoliophyta: Asteridae, part 8: cago, Il. 63 pp. Asteraceae, part 3. Flora of North America north of Mexico. Volume 21. New York: Springer, A. E., L. E. Stevens, and R. Harms. Oxford University Press. 616 pp. 2006. Inventory and classifi cation of selected National Park Service springs on the Colo- Hatfi eld, S. C., P. D. Rowley, E. G.Sable, D. J. rado Plateau. Northern Arizona University, Maxwell, B. V. Cox, M. D. McKell, and D. E. Flagstaff , Az. NPS Cooperative Agreement # Kiel. 2003. Geology of Cedar Breaks Nation- CA 1200-99-009; Task # NAU-118. al Monument. Pages 138–154 in Sprinkel, D. A., T. C. Chidsey, Jr., and P. B. Anderson, eds. State of Utah Department of Agriculture and Geology of Utah’s parks and monuments. Food. 2007. Utah noxious weed list. http:// Utah Geological Association Publication 28 ag.utah.gov/plantind/nox_utah.pdf. (second edition). U.S. Forest Service Intermountain Region and Holmgren, N. H., P. K. Holmgren, and A. Cron- Southwestern Region, National Park Service quist. 2005. Volume 2, part B: Subclass Dille- Rocky Mountain Region, U.S. Fish and niidae. Intermountain fl ora: Vascular plants Wildlife Service Mountain-Prairie Region of the Intermountain West, U.S.A. Bronx, and Southwest Region. 1995. Arizona willow NY: New York Botanical Garden. 488 pp. (Salix arizonica) draft conservation agree- ment and strategy. 61 pp. + appendices.

18 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Utah Division of Wildlife Resources (UDWR). Weber, W. A., and R. C. Wittmann. 2001. Colo- 1998. Inventory of sensitive species and rado fl ora: Western Slope. Third edition. ecosystems in Utah. Endemic and rare plants Boulder: University Press of Colorado. 488 of Utah: An overview of their distribution pp. and status. Report prepared for the Utah Welsh, S. L., N. D. Atwood, S. Goodrich, and L. Reclamation Mitigation and Conservation C. Higgins. 2003. A Utah fl ora. Third edition, Commission and U.S. Department of the revised. Provo, Ut.: Brigham Young Univer- Interior. 566 pp + app. sity. 912 pp.

Chapter 5: Literature Cited 19

Holmgren et al. 2005; Weber and Wittmann 2001). No ef- Appendices fort has been made to identify additional synonyms from monographs and fl oras preceding the early 1970s. The following appendices contain annotated checklists of the vascular fl ora of Cedar Breaks National Monument. Common name (Appendices A, B, C, and D) Appendix A lists all species that have been confi rmed for Common names are derived primarily from Welsh et al. the fl ora with an authenticated voucher specimen (includ- (2003) and the Intermountain Flora (Barneby 1989; Cron- ing extant and historical records) or that have been reliably quist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Hol- reported for the monument in the literature (Buchanan mgren et al. 2005). 1992; Dewey and Andersen 2005; Roberts and Jean 1989; NCPN in ed.; Springer et al. 2006). Vouchers were exam- Life form (Appendices A, B, C, and D) ined from the CEBR herbarium (housed in Zion National Life form is based on the stature and growth habit of typi- Park) and collections from Brigham Young University cal members of a species at reproductive maturity. Trees (BRY), Utah State University (USU), the University of Wy- are woody perennials with a single stem or trunk and are oming (RM), and the digital museum database of the New typically over 3.5 m tall. Shrubs are woody perennials York Botanical Garden (NY). Species entries in Appendix with one to many trunks arising from near the base of the A are organized alphabetically by family and genus. plant and are usually less than 3.5 m tall. Perennial Forbs Appendix B includes species that may potentially occur (PerF) are broad-leaved dicots or monocots with non- within Cedar Breaks NM based on the proximity of the woody stems (at least above ground level) that live for mul- monument to other populations or the presence of suit- tiple years. Annual Forbs (AnnF) are broad-leaved dicots able habitat. or monocots with non-woody stems that complete their life cycle (mature, fl ower, and die) in one year. Perennial Appendix C lists species that have been previously report- Graminoids (PerG) are linear-leaved, grass-like monocots ed for Cedar Breaks NM but are now considered errone- that live for multiple years. Annual Graminoids (AnnG) ous because they are based on misidentifi ed specimens or are linear-leaved, grass-like monocots that complete their questionable literature records. life cycle (mature, fl ower, and die) in one year. Ferns and fern-allies (Fern) are non-fl owering vascular plants that Appendix D contains the same confi rmed, historical, and reproduce by spores. reported taxa as in Appendix A, but organized by life form rather than by family in order to better meet the needs of Range (Appendices A, B, and C) NPS interpretive specialists. Range represents the distribution of a species within Utah and the state’s contribution to the plant’s overall global The codes and references for each fi eld in the appropriate range. Disjunct (Disj) taxa have their Utah distribution appendices are summarized below: separated from the main, contiguous portion of their range by a gap of more than 800 km (ca 500 miles). Intro- Family (Appendices A, B, C, and D) duced (Intro) taxa are those that are not native to Utah Family concepts and nomenclature follow A Utah Flora, or North America but have become naturalized (breeding third edition (Welsh et al. 2003). Alternative family names on their own without human assistance). Local Endemics and treatments from other regional fl oras (Dorn 2001; (LocEn) are taxa whose entire global range is restricted Flora of North America Editorial Committee 1993, 2002b; to an area of less than 16,500 km2 (ca 6,370 mi2, or 1° of Holmgren et al. 2005; Weber and Wittman 2001) are listed latitude × 2° of longitude). Peripheral (Periph) taxa have a in parentheses. widespread global distribution but occur at the margin of their contiguous range in Utah and occupy less than 5% Species name (Appendices A, B, C, and D) of the state’s area (usually only within a few miles of the Nomenclature for scientifi c names and taxonomic con- state border). Regional Endemics (RegEn) are taxa with cepts are derived from Welsh et al. (2003). Scientifi c names a global range of 16,500–250,000 km2 (an area about the are listed in alphabetical rather than phylogenetic order. size of the state of Wyoming). Sparse taxa occur widely across Utah or North America but their range within Utah Synonyms/Taxonomic notes (Appendices A, B, and C) is small and patchy, with populations restricted to special- Pertinent synonyms are included for species with alter- ized or uncommon habitats. Widespread (Wide) taxa have native scientifi c names in other recent state and regional global ranges exceeding 250,000 km2 and occur over at fl oras (Barkworth et al. 2003, 2007; Barneby 1989; Cron- least 10% of the state. quist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of North America Editorial Committee 1993, Park status (Appendices A, C, and D) 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Park status identifi es whether a species is Present (Pres),

Chapter 6: Appendices 21 Historical (Hist), or Reported (Rep) for Cedar Breaks Year doc. (Appendix A) National Monument. Species are considered Present if a Year documented indicates the fi rst year in which a spe- confi rmed voucher or documented observation has been cies was confi rmed or reported to occur in Cedar Breaks made within monument boundaries since 1970. Histori- National Monument. This year does not necessarily cor- cal species have been confi rmed with a voucher from the respond with the date of the voucher specimen cited un- monument but have not been relocated since before 1970. der Source. “NA” indicates that the fi rst year is not known Reported species are listed for the monument in the lit- (used mostly for ‘reported’ taxa). erature (Roberts and Jean 1989; NCPN in ed.; Springer et al. 2006), but have not been corroborated with a voucher Comments (Appendices A, B, C, and D) specimen. Species that may potentially occur in CEBR are This column includes additional comments on the status listed in Appendix B. Rejected taxa (listed in Appendix C) of a particular species in the monument, the place of ori- have either been falsely reported (FalsRep) in the litera- gin for introduced species, legal status, and other data. ture based on misidentifi ed herbarium specimens or are considered Questionable (Ques?) because park reports Nativity (Appendix D) are from well outside their known or expected range. Nativity indicates whether a species is native to Utah or Pop. size (Appendix A) Introduced (Intro) based on Fertig (2007). Population size refers to the abundance and frequency Flower time (Appendix D) with which a species occurs within Cedar Breaks National Flower time refers to the months in which a species is most Monument. Abundance data are derived from Roberts likely to be in fl ower. NA indicates a species that is a non- and Jean (1989), NCPN (in ed.), and personal observa- fl owering fern or gymnosperm. tions. Abundant (Abun) taxa have large populations, a broad ecological amplitude (often being dominant in one Habitat (Appendix D) to several vegetation types), or occur extensively across the monument. Common (Com) taxa have large local popula- The distribution of species across generalized habitats in tions but may be restricted to a single vegetation type or do Cedar Breaks National Monument is indicated by an “x” not occur across the entire monument. Uncommon (Unc) in the columns SubFir through Wet and is derived from taxa have low to medium-sized populations, a narrow eco- the Cedar Breaks NM vegetation classifi cation (NCPN in logical amplitude (or at least contribute relatively little to ed.), Roberts and Jean (1989), Buchanan (1992), and Welsh total vegetative cover), or a small range across the monu- et al. (2003). SubFir = Subalpine fi r/Engelmann spruce ment. Rare taxa usually have low population numbers and forest. White Fir = White Fir–Douglas-fi r–Aspen forests. are restricted to a single location within the monument. A SubMdw = Subalpine meadows above the amphitheater “?” indicates that population size is unknown. rim. Wet = Wetlands. Additional comments are included for species that are restricted to a particular soil/geologic Source (Appendix A) type (including the Claron endemics) or are found mostly in disturbed sites. Sources provide the basis for the assigned park status. In the case of Present and Historical taxa, sources are her- barium records (with collector, collection number, and repository) that were corroborated during this study. For reported taxa, the source is a published or unpublished reference citing the species within the monument. Speci- men repositories include the Cedar Breaks National Mon- ument herbarium (CEBR), Brigham Young University (BRY), Utah State University (USU), New York Botanical Garden (NY), and the University of Wyoming’s Rocky Mountain Herbarium (RM).

22 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument S. c name c var. var. ed . specimens are transitional to A. glabrum diffusum specimens of racemosa were based on misidentifi Comments 1988 Some CEBR 2007 1989 1981 1989 1981 1934 1946 1946 2007 Previous reports Year Year doc. (CEBR) Reynolds 23345 (CEBR) Jean (1989) Palmer 1048 (CEBR) 22981 (CEBR) Palmer 1048 (CEBR) Palmer 1122 (CEBR) F-56 (CEBR) Palmer 1074 (CEBR) Reynolds 23348 (CEBR) Source Pop. size Park status Range Shrub Wide Unc PresPerF Jean 170 Wide UncPerF Pres Wide Fertig & Unc Rep Roberts & PerF Wide Unc Pres Palmer & Shrub Wide Unc Pres Palmer & Life form Rocky Mountain maple spreading dogbane common dogbane Oregon-grape Shrub Widewater birch Com Pres Tree Palmer & Wide Unc Pres Fertig southwestern stoneseed aspen bluebell PerF Wide harebellParry’s PerF Abun Pres Wide Palmer & NA Hist Crawford black twinberry blue elderberry Shrub Wide Unc Pres Fertig & Common name

& parryi tripartitum caerulea, S. & var. ssp. Includes vars. neomexicanum, douglasii, angustifolium Includes vars. cannabinum, glaberrimum, Distegia involucrata cerulea Synonyms/ notes Taxonomic

var. orum Acer glabrum glabrum Apocynum androsaemifolium var. androsaemifolium Apocynum cannabinum Mahonia repens Berberis repens Betula occidentalis B. fontinalis Lithospermum multifl Mertensia arizonica Campanula parryi C. parryi Lonicera involucrata Sambucus caerulea S. nigra National Monument organized by family and scientifi vascular plant taxa of Cedar Breaks rmed, historical, and reported FamilyAceraceae Species name Apocynaceae Apocynaceae Berberidaceae Betulaceae Boraginaceae Boraginaceae Campanulaceae Caprifoliaceae Caprifoliaceae (Adoxaceae) Appendix A. Confi with taxonomic and biogeographic notes.

Chapter 6: Appendices 23 is Cedar plateauensis Breaks Rim, Cedar Breaks NM; Iron Co., UT (Maguire 19024 NY). Comments c name with taxonomic 1954 1981 1981 2006 1981 1930 locality of var. Type 1942 Year Year doc. s.n. (CEBR) Palmer 1113 (CEBR) Palmer 1120 (CEBR) 22749 (CEBR) Palmer 1214 (CEBR) (CEBR) Peiler 16431 (NY) Source Pop. size Park status Range Shrub Wide Unc Pres Palmer & PerF Wide Unc Pres Palmer & Life form PerF Wide Rare Pres Fertig PerF Wide Rare Pres Palmer & PerF Wide NA Hist Degener & y red elderberry Shrub Wide Unc Pres Buchanan mountain snowberry Drummond’s Drummond’s catchfl Common name Menzies’ campion Eastwood’s Eastwood’s sandwort basin sandwort PerF Wide Unc Pres Croft 44CB sprawling sandwort

& .

by some

ssp. included

ssp. menziesii

var. var. var. glabrescens, E. uintahensis ssp. includes vars var. var. racemosa drummondii, eastwoodiae, A. drummondii, S. microbotrys; in var. authors S. rotundifolius striata; striata, kruckebergii, & Gastrolychnis drummondii menziesii, Anotites menziesii Synonyms/ notes Taxonomic Eremogone eastwoodiae var. eastwoodiae A. saxosa, lanuginosa saxosa, Spergulastrum lanuginosum saxosum Silene drummondii ssp. S. drummondii kingii A. kingii uintahensis, compacta, plateauensis, Eremogone kingii

var. var. eastwoodiae glabrescens Sambucus racemosa microbotrys Symphoricarpos oreophilus utahensis Arenaria fendleri var. Silene menziesii S. menziesii Arenaria fendleri var. Arenaria lanuginosa Lychnis Lychnis drummondii rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCaprifoliaceae (Adoxaceae) Species name Caprifoliaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae

24 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument 2005 Flora of in based on collection cannot be reliably determined to variety but are presumed to be var. purpurea range. candidate, USFS Region 4: Sensitive. locality is 1 mile Type above Baldy Ranger Station, Sanpete Co., UT (Maguire 20000 NY) North America monantha Indian Creek, Tushar Indian Creek, Tushar Mtns., Beaver Co., UT (Martin & Welsh 17148 BRY) Comments c name with taxonomic 1981 Specimens in CEBR 1934 USFWS: former C2 1947 1981 1926 1954 Includes var. 1996 Native to Eurasia 1939 2006 locality is Type Year Year doc. Palmer 1134 (CEBR) 11217 (BRY) Palmer 1132 (CEBR) Palmer 1068 (CEBR) s.n. (CEBR) (CEBR) (CEBR) 1133 (CEBR) 22927 (CEBR) Source Pop. size Park status Range PerF Wide Unc Pres Palmer & PerF RegEn Unc Pres Higgins PerF Wide Unc Pres Palmer & Shrub Wide Unc Pres Palmer & Life form PerF Wide Rare Pres Bruhn s.n. AnnF Wide Rare Pres Palmer AnnF Wide Rare Pres Fertig PerF Wide Com Pres Woodbury purplish agoseris Peterson’s Peterson’s campion James’ chickweed mountain- lover Common name long-stalk starwort lambsquarter AnnF Intro Unc Pres Mason s.n. mountain goosefoot smallhead goosefoot common yarrow

& ssp.

S. clokeyi minor var. var. and petersonii Includes vars. Paxistima myrsinites, Pachistima myrsinites Synonyms/ notes Taxonomic Stellaria longipes stricta C. album C. fremontii atrovirens C. overi, foliosum A. lanulosa, millefolium occidentalis

var. var. var. album var. longipes purpurea Silene petersonii Stellaria jamesianajamesiana Pseudostellaria Stellaria longipes var. Pachystima myrsinites Chenopodium album Agoseris aurantiaca Chenopodium atrovirens parvicapitatum Chenopodium capitatum Achillea millefolium lanulosa rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCaryophyllaceae Species name Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Celastraceae Chenopodiaceae Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Compositae (Asteraceae)

Chapter 6: Appendices 25

A. rosea represent form. Comments c name with taxonomic 1979 1989 1979 1981 1970 Some specimens 1981 1981 1954 1954 2005 1934 Year Year doc. Palmer 1051 (CEBR) (CEBR) Mansor s.n. (CEBR) Palmer 1081 (CEBR) Palmer 1186 (CEBR) Palmer 1030 (BRY) Palmer 1277 (CEBR) (CEBR) s.n. (CEBR) (CEBR) (CEBR) Source Pop. size Park status Range Shrub Wide Unc Pres Palmer & PerF Wide UncPerF Wide Pres Palmer & Rare Pres Palmer & PerF Sparse RarePerF Pres Wide Palmer & Unc Pres Bruhn s.n. PerF Wide Unc Pres Buchanan Life form elds agoseris PerF Sparse Rare Pres Harms s.n. silver sagebrush tarragon PerF Wide Unc Pres Jean 1145 hairy arnica PerF Wide Unc Pres Buchanan & rayless arnica PerF Wide Unc Pres Palmer & small-leaf pussyoes common pussytoes showy pussytoes Breaks pussytoes heartleaf arnica Common name fi short agoseris PerF Wide Unc Pres Nelson 8981

ssp. . ssp by some parryi Utah Flora var. A. ovata A. rosea in viscidulum Oligosporus dracunculus. not recognized in Vars. Flora of North America (2006) Seriphidium canum A. pulcherrima pulcherrima (2003). Includes Synonyms/ notes Taxonomic authors Included in var. Included in var. dasycephala Includes Vars. not recognized in Vars. Flora of North America (2006). Misspelled var. cordata

var. ssp. agrestis cordifolia dasycephala Artemisia dracunculus glauca Arnica parryi A. parryi Agoseris glauca var. Artemisia cana viscidula Antennaria parvifolia Antennaria pulcherrima Antennaria rosulata Arnica cordifolia var. Arnica mollis Agoseris glauca var. Antennaria microphylla rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

26 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument based on collection cannot be reliably determined to variety but are presumably var. ludoviciana range. Tate Mine near Tate Marysvale, Piute Co., UT (Jones 5861 POM). Canyon, Zion NP, Co., UT Washington & Hays 24379 (Welsh BRY). Comments c name with taxonomic 1988 Specimens in CEBR 2006 1954 1934 1970 1977 locality is Type 2007 locality is Zion Type 2007 Year Year doc. (CEBR) 22987 (CEBR) (CEBR) 22907 (CEBR) Palmer 1296 (CEBR) s.n. (CEBR) Reynolds 23356 (CEBR) Reynolds 23342 (CEBR) Source Pop. size Park status Range PerF Wide Unc Pres Jean 107 PerF LocEn Com Pres Buchanan PerF Wide Unc Pres Fertig & Life form c aster c PerF Wide Unc Pres Fertig Louisiana wormwood Pacifi asterCanby’s PerF asterParry’s Wide PerF Unc Wide Pres Unc Bruhn s.n. Pres Fertig western aster PerF Wide Unc Pres Palmer & Markagunt aster asterWelsh’s PerF LocEn Rare Pres Fertig & hoary dusty- maiden Common name &

canbyi parryi douglasii ssp. var. var. adscendens, A. adscendens, Symphyotrichum ascendens, Virgulaster ascendens Symphyotrichum foliaceum Symphyotrichum foliaceum Symphyotrichum spathulatum Eurybia wasatchensis, Eucephalus wasatchensis, Herrickia wasatchensis Includes vars. montana Synonyms/ notes Taxonomic

var. var. var. wasatchensis ludoviciana Aster ascendens A. chilensis canbyi parryi Aster foliaceus Aster foliaceus Aster spathulatus A. occidentalis, Aster wasatchensis var. Artemisia ludoviciana Aster welshii Symphyotrichum welshii Chaenactis douglasii rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) and biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 27 is nitively C2 candidate. locality of Type E. proselyticus “canyon to Cedar Breaks”; Iron Co., UT (Maguire 14947 NY). Comments vegetative and cannot be defi determined to variety c name with taxonomic 1970 1954 1934 USFWS: former 1977 1954 Year Year doc. 1981 2006 1954 2007 Specimen is 1946 Palmer 1082 (CEBR) Palmer 1156 (CEBR) (CEBR) s.n. (CEBR) Louie, & Alexander s.n. (CEBR) Source Palmer 1293 (CEBR) 22984 (CEBR) (CEBR) Reynolds 23392 (CEBR) F-80 (CEBR) Pop. size Park status Range PerF Wide Unc Pres Palmer & PerF LocEn Unc Pres Jean 1103 Life form PerF Wide Unc Pres Malm, PerF Wide Unc Pres Palmer & Shrub Wide Unc Pres Fertig Shrub Wide Unc Pres Fertig & PerF Wide Com Pres Crawford eabane trailing daisy PerF Wide Unc Pres Palmer & wandering fl Cedar Breaks daisy aspen daisy PerF Wide Com Pres Buchanan Common name cut-leaved daisy hispid golden- aster Nevada rabbitbrush Arizona thistle PerF Wide Unc Pres Bruhn s.n. greenish rabbitbrush Wheeler’s Wheeler’s thistle

var. var. var. var. var. . var.

var.

var Included var. trilobatus, arizonicum var. hispida, by some var. var. oreophila. C. nauseosus nidulum agellaris E. peregrinus scaposus, E. glacialis glacialis E. proselyticus, fl Var. not recognized by Var. some authors Synonyms/ notes Taxonomic discoideus C. villosa Heterotheca villosa hispida, H. villosa minor Ericameria nauseosa consimilis, E. nauseosa var. Ericameria parryi nevadensis, C. parryi attenuatus Includes vars. & in authors oreophilus Vars. not recognized in Vars. Flora of North America (2006) E. compositus

var. var. . var. var. var

agellaris var. macranthus minor wheeleri Erigeron peregrinus callianthemus trilobatus Erigeron sionis Erigeron speciosus var. Chrysopsis villosa var. Erigeron fl Chrysothamnus nauseosus consimilis nevadensis Cirsium arizonicum Cirsium wheeleri var. Chrysothamnus parryi Erigeron compositus glabratus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

28 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Comments USFWS: former C2 candidate. Type locality is Cedar Canyon; Iron Co., UT (Anderson, Welsh, & Chatterley 5094 BRY). c name with taxonomic 1946 Year Year doc. 1946 1978 1934 1970 1989 1940 1934 1989 BLM (UT): Sensitive; 1977 F-53 (CEBR) Source F-26 (CEBR) Palmer 1152 (CEBR) Palmer 1210 (CEBR) (RM) 23337 (CEBR) (CEBR) (CEBR) (CEBR) Palmer 1149 (CEBR) Pop. size Park status Range Life form PerF Wide Unc Pres Crawford PerF Wide Unc Pres Palmer & Shrub Wide Unc Pres Fertig PerF WideShrub Unc Wide Unc Pres Boyle 759 Pres Croft 60CB Shrub LocEn Rare Pres Jean 1112 PerF Wide Unc Pres Palmer & ower PerF Wide Unc Pres Crawford Common name little sunfl hairy pretty daisy common hyalineherb bear daisy PerF Wide Com Pres Palmer & Payson’s daisyPayson’s PerF Wide Unc Pres Nelson 9025 broom snakeweed Clement’s Clement’s goldenweed narrowleaf goldenweed Cedar Breaks goldenbush orange sneezeweed

var.

& var. var. orus, linearis, E.

. discoidea, var. var canescens, Ericameria Synonyms/ notes Taxonomic E. subtrinervis subtrinervis tomentosus Includes vars. lugens, paucifl megacephalus, Vars. not recognized in Vars. Flora of North America (2006) Utah material may represent an undescribed variety Pyrrocoma clementis clementis discoidea Macronema discoideum H. macronema linearis, H. macronema var. discoidea Ericameria zionis Dugaldia hoopesii

var. ora mollis ursinus lifolius Erigeron speciosus var. Erigeron ursinus var. Hymenopappus fi cinereus Erigeron vagus Gutierrezia sarothrae Haplopappus clementis Haplopappus macronema Haplopappus zionis Helenium hoopesii Hymenoxys hoopesii, Helianthella unifl rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 29 eld Co., eld miles N of Orderville, Kane Co., UT (Maguire 18740 NY) Bromide Pass, Henry Mts., Garfi UT (Jones 5695y US) Sensitive, USFWS: former C2 candidate. locality is Little Type Podunk Creek, Kane Co., UT (Malmsten 131 US). Comments c name with taxonomic 1935 locality is 15 Type 1935 locality is Type 1946 1934 1981 1977 2007 1933 1989 USFS Region 4: 1981 1947 Year Year doc. (RM) 1263 (CEBR) F-77 (CEBR) 1244 (CEBR) 1172 (CEBR) s.n. (CEBR) Reynolds 23350 (CEBR) s.n. (CEBR) (CEBR) 1026 (CEBR) F-146 (CEBR) Source Pop. size Park status Range PerF RegEn NA Pres Nelson 9013 PerF RegEnPerF NA WidePerF Hist Unc Crawford WidePerF Pres Unc Wide Palmer Pres UncPerF Palmer RegEn Pres Rare Buchanan PerF Pres Fertig & Wide UncPerF LocEn Pres Rare Dickman Pres Jean 1110 PerF Wide Com Pres Palmer Life form PerF Wide NA Hist Crawford ower alpine hyalineherb Bigelow’s asterBigelow’s PerF RegEn Unc Pres Palmer montane conefl black groundsel woolly groundsel cut-leaved groundsel Hart’s groundsel western groundsel Podunk groundsel Common name basin groundsel broom groundsel

var. commixta var. var. M. bigelovii commixta, Dieteria bigelovii R. occidentalis montana Packera multilobata Synonyms/ notes Taxonomic Vars. not recognized in Vars. Flora of North America (2006)

var. var.

var. spartioides lifolius Hymenopappus fi nudipes Machaeranthera commixta Rudbeckia montana kingii Senecio atratus Senecio canusSenecio Packera cana eremophilus Senecio hartianus Packera hartiana Senecio integerrimus exaltatus Senecio malmstenii Packera malmstenii Senecio multilobatus Senecio spartioides var. rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

30 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument eld Type locality is rim of Type Bryce Canyon, Garfi Canyon NP, Co., Utah (Eastwood & Howell 727 CAS). Comments c name with taxonomic 1934 1934 1935 1981 1977 Native to Eurasia 2007 1989 USFWS: Category 3C. 1981 Native to Europe 2007 1934 Year Year doc. Palmer 1261 (CEBR) Palmer 1129 (CEBR) Palmer 1301 (CEBR) Palmer 1257 (CEBR) & Dickman s.n. (CEBR) Reynolds 23257 (CEBR) (CEBR) Palmer 1115 (CEBR) Reynolds 23393 (CEBR) (CEBR) Source Pop. size Park status Range PerF Wide Unc Pres Palmer & PerF Wide Com Pres Palmer & PerF Wide Unc Pres Palmer & PerF Wide Unc Pres Palmer & PerF Intro Unc Pres Buchanan PerF Wide Rare Pres Fertig & PerF LocEn Unc Pres Jean 117 Shrub Wide Unc Pres Fertig & PerF Wide Unc Pres Croft 24CB Life form showy goldeneye Rocky Mountain goldenrod Parry’s Parry’s goldenweed three-nerve goldenrod common dandelion common townsendia Bryce Canyon townsendia yellow salsify PerF Intro Unc Pres Palmer & Common name red-osier dogwood Drummond’s Drummond’s rockcress

var. ora minima, major var. ora var. var. ora, S. velutina ora sparsifl Oreochrysum parryi T. alpigena T. T. minima T. Heliomeris multifl T. dubius T. multifl C. stolonifera, Swida sericea drummondii, B. stricta Synonyms/ notes Taxonomic var. var.

ora var.

var. ora var. multifl cinale Cornus sericea Solidago multiradiata scopulorum Solidago parryi Haplopappus parryi, Solidago velutina S. sparsifl Taraxacum Taraxacum offi Townsendia Townsendia leptotes Townsendia montana minima var. Tragopogon Tragopogon dubius Viguiera multifl sericea Arabis drummondiiArabis Boechera angustifolia, B. rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Cornaceae and biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 31 3C. Type locality is 3C. Type Zion Canyon, Co., Washington NP, UT (Jones s.n. POM). CEBR report may be based on a collection actually made in Zion NP Comments 3C. Type locality is 3C. Type Cedar Breaks, Breaks NM; Iron Co., UT (Goodman & Hitchcock 1622 MO) c name with taxonomic 2007 1934 1954 1934 1977 USFWS: Category Year Year doc. 1942 1930 USFWS: Category 1940 Reynolds 23344 (CEBR) Palmer 1076 (CEBR) (CEBR) (CEBR) s.n. (CEBR) Source (CEBR) (CEBR) Palmer 1067 (CEBR) Pop. size Park status Range Life form PerF Wide Unc Pres Fertig & PerF Wide Unc Pres Palmer & PerF RegEn Rare Pres Bruhn s.n. AnnF Wide Unc Pres Croft 32CB PerF Wide Unc Pres Jean 130 PerF Wide Unc Pres Palmer & ower Common name Fendler’s Fendler’s rockcress heartleaf bittercress heartleaf bittercress mountain tansy-mustard Zion draba PerF LocEn Rare Pres Dickman slender whitlow-grass Breaks draba PerF LocEn Unc Pres Boyle 1033 sand dune wallfl

var. var. var. ssp. fendleri,

var. by some included in var. var. Vars. not Vars. capitatum Synonyms/ notes Taxonomic A. holboellii Boechera fendleri spatifolia. recognized in Holmgren et al. (2005) incisa, D. incana incana D. zionensis arkansanum; var. D. richardsonii sonnei, D. incisa incisa, D. incana D. albertina authors E. asperum

var.

var. var. fendleri sonnei nana purshii Arabis holboellii var. Cardamine cordifolia cordifolia Cardamine cordifolia incana Descurainia incana var. Draba asprella zionensis Draba stenoloba var. Draba subalpina Erysimum asperum var. rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCruciferae (Brassicaceae) Species name Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

32 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument eld Co., eld eld Co., UT Type locality is Red Type Canyon, Garfi UT (Eggleston 8198 US) Aquarius Plateau, Garfi 589 GH) (Ward Comments c name with taxonomic 1989 USFWS: Category 3C. 1977 locality is Type 1935 1981 2005 Native to Europe 2007 1981 1935 1989 1989 Year Year doc. 1977 1981 (CEBR) s.n. (CEBR) 23330 (CEBR) Palmer 1181 (CEBR) ed.) AT09070701 AT09070701 (CEBR) Palmer 1227 (CEBR) s.n. (CEBR) 22732 (CEBR) 22979 (CEBR) (CEBR) Palmer 1229 (CEBR) Source Pop. size Park status Range PerF LocEn Unc Pres Jean 153 PerF RegEn UncPerF Pres Buchanan Wide Unc Pres Fertig PerF Wide Rare Pres Palmer & AnnF Intro Rare Rep NCPN (in Life form PerG Wide RarePerG Pres RegEn Unc Tendick Pres Palmer & PerF Wide Unc Pres Dickman Shrub Wide Unc Pres Fertig Tree Wide Unc Pres Fertig eld Breaks bladderpod Ward’s Ward’s bladderpod blunt-leaf yellowcress smooth yellowcress fi pennycress Common name northern bog sedge Eggleston’s sedge alpine pennycress common juniper Rocky Mountain juniper water sedge PerG Wide Unc Pres Harms s.n. golden sedge PerG Wide Unc Pres Palmer & alpina aquatilis glabra by some ssp. var. var. P. kingii P. gynocrates parvifolia P. rubicundula, P. Lesquerella rubicundula, L. hitchcockii included in var. authors Synonyms/ notes Taxonomic R. palustris Noccaea montana J. communis Sabina scopulorum ssp.

var. var. curvipes glabra montanum Physaria rubicundula rubicundula Physaria wardii Lesquerella wardii; Rorippa curvipes var. Rorippa islandica var. Thlaspi arvense Thlaspi montanum var. Carex aquatilis C. aquatilis Carex egglestonii Juniperus communis depressa Juniperus scopulorum Carex aureaCarex dioica C. garberi C. gynocrates, dioica rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCruciferae (Brassicaceae) Species name Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Cupressaceae Cupressaceae Cyperaceae Cyperaceae Cyperaceae

Chapter 6: Appendices 33 Comments c name with taxonomic 1952 Year Year doc. 1981 1981 1970 1981 1981 2005 1981 1981 1981 1981 1981 1989 (BRY) Source Palmer 1079 Palmer 1280 (CEBR) Palmer 1221 (CEBR) Palmer 1220 (CEBR) Palmer 1015 (CEBR) (CEBR) Palmer 1141 (CEBR) Palmer 1278 (CEBR) Palmer 1228 (CEBR) Palmer 1222 (CEBR) (CEBR) 22740 (CEBR) Pop. size Park status Range Life form PerG Wide Unc Pres Palmer & PerG Wide Unc Pres Palmer & PerG Wide Unc Pres Palmer & kobresia sedge PerG Wide Rare Hist Lewis 144 Common name blackened sedge inland sedge PerG Wide Rare Pres Palmer & small-wing sedge Scandinavian sedge western sedge PerG Widewoolly sedge Rare PerGliddon sedge Pres Wide Palmer & PerG Rare Wide Pres Unc Harms s.n. Pres Palmer & russet sedge PerGslender sedge Wide PerG Unc Sparsebeaked sedge Pres Unc PerG Palmer & Pres Wideblister sedge Palmer & Rare PerG Pres Wide Palmer & Unc Pres Harms s.n. Ross’ sedge PerG Wide Com Pres Fertig

var. laxa var. boottii, C. chalciolepis, ssp. var. var. exa Synonyms/ notes Taxonomic C. atrata C. chalciolepis microptera, C. festivella, C. macloviana microptera C. stevenii brevipes, C. pityophila chalciolepis stevenii scirpoidea Carex elynoides Carex heteroneura var. Carex interior Carex microptera C. microptera Carex norvegica var. Carex occidentalis Carex pellitaCarex petasata C. lanuginosa Carex rossii C. defl Carex scirpoidea var. Carex utriculata C. rostrata (sensu lato) Carex vesicaria Carex saxatilis C. saxatilis rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyCyperaceae Species name Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae

34 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Comments c name with taxonomic 1970 1981 1981 1989 1970 1981 1977 1981 1989 1981 Year Year doc. 2007 1946 1946 1934 (RM) Palmer 1013 (CEBR) Palmer 1071 (CEBR) (CEBR) (RM) (CEBR) s.n. (CEBR) Palmer 1012 (CEBR) (CEBR) Palmer 1016 (CEBR) 23339 (CEBR) Source s.n (CEBR) (CEBR) Palmer 1287 (CEBR) Pop. size Park status Range Fern SparseShrub Rare Wide Pres Com Nelson 9030 Pres Palmer & PerF Wide Com Pres Palmer & PerG Wide RarePerG Pres WideShrub Harms s.n. Rare Wide Pres Unc Nelson 8985 Pres Malm s.n. Fern Wide Unc Pres Palmer & Fern Disj Rare Pres Jean 1160 Life form PerF Wide Rare Pres Fertig PerF Wide Unc Pres Crawford PerF Wide Unc Pres Palmer & ower eld horsetail eld Fern Wide Unc Pres Buchanan Ferris’ scouring-rush greenleaf manzanita shorthorn spurge common spikerush few-fl spikerush Canada buffaloberry fi smooth scouring-rush northern scouring-rush Common name Gambel oak Tree Wide Unc Pres Palmer & golden corydalis Rocky Mountain gentian Parry’s gentianParry’s PerF Wide Rare Pres Bruhn s.n. Engelmann’s Engelmann’s gentian E. E. x nis

E. &

var. ssp. includes var. E. hyemale E. macrostachya, ora; ana robusta, Tithymalus robusta, Tithymalus laevigatum E. robusta, brachycera var. brachyceras uniglumis E. erythropoda, suksdorfi Includes E. paucifl Hippochaeta laevigata, E. kansanum E. variegatum variegatum, Hippochaete variegata Hybrid of Synonyms/ notes Taxonomic C. curvisiliqua occidentalis Pneumonanthe affi G. amarella heterosepala ferrisii nis x ora nis gambelii occidentalis affi Arctostaphylos patula Euphorbia brachycera Quercus gambelii Eleocharis quinquefl Shepherdia canadensis Equisetum arvense Equisetum laevigatum Eleocharis palustris Equisetum variegatum Equisetum var. Corydalis aurea var. var. Gentiana affi Gentiana parryi Pneumonanthe parryi Gentianella heterosepala rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Ericaceae Euphorbiaceae Fagaceae and biogeographic notes, cont. Cyperaceae Elaegnaceae Equisetaceae Equisetaceae Appendix A. Confi FamilyCyperaceae Species name Equisetaceae Equisetaceae Fumariaceae Gentianaceae Gentianaceae Gentianaceae

Chapter 6: Appendices 35 eld eld Co., UT Aquarius Plateau, head of Poison Creek, Garfi Co., UT (Rydberg & Carlton 7401 NY) and Africa Panguitch Lake, Garfi (Jones 6008 POM) Comments c name with taxonomic 2007 locality is Type 1970 1989 Native to Eurasia. 1954 1970 Native to Eurasia 2005 1940 locality is Type 1934 1954 1954 Year Year doc. Reynolds 23355 (CEBR) s.n. (CEBR) (CEBR) (CEBR) (CEBR) (CEBR) Palmer 1302 (CEBR) (CEBR) Palmer 1097 (CEBR) (CEBR) Source Pop. size Park status Range PerF Wide Rare Pres Fertig & PerF Wide Com Pres Buchanan PerG Intro Unc Pres Jean 157 PerG Wide Rare Pres Harms s.n. AnnF Wide Unc Pres Boyle 764 Life form small-leaf geranium Richardson’s crane’s-bill crested wheatgrass ticklegrass PerG Wide Rare Pres Bruhn s.n. redtop PerG Intro Unc Pres Jean 1116 Idaho bentgrass Jones’ gentian AnnF RegEn Unc Pres Palmer & meadow gentian elkweed PerF Wide Unc Pres Palmer & Common name pine dropseed PerG Wide Unc Pres Bruhn s.n.

var. var. pectinatum, A. elegans, Gentiana G. caespitosum atropurpureum A. cristatum cristatum, A. cristatum ssp. pectinatum, A. sibiricum, A. desertorum, fragile A. alba, gigantea A. idahoensis Gentiana tortuosa Gentianella detonsa var. detonsa, Gentianopsis thermalis Synonyms/ notes Taxonomic

var.

var. x marginale Geranium richardsonii Agropyron cristatum Agrostis scabra A. hiemalis, geminata Agrostis stolonifera Agrostis idahoensis Blepharoneuron tricholepis Gentianella tortuosa Gentianopsis detonsa elegans Swertia radiata Frasera speciosa Geranium caespitosum rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Geraniaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyGentianaceae Species name Gentianaceae Gentianaceae Geraniaceae

36 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument and Africa Comments c name with taxonomic 1981 1954 1954 1954 Native to Eurasia 1981 Native to Eurasia 1954 1954 1986 Native to Eurasia 1989 1935 1981 1935 Year Year doc. (CEBR) Palmer 1248 (CEBR) (CEBR) (CEBR) Palmer 1019 (CEBR) (CEBR) Palmer 1285 (CEBR) (CEBR) (CEBR) (CEBR) Palmer 1258 (CEBR) Palmer 1142 (CEBR) Source Pop. size Park status Range PerG Wide Unc Pres Jean 1139 PerG Wide Com Pres Palmer & PerG Wide Unc Pres Bruhn s.n. PerG Wide Unc Pres Palmer & PerG Wide Rare Pres Jean 1129 PerG Wide Com Pres Boyle 1049 PerG Wide Rare Pres Palmer & Life form nodding brome mountain brome fringed brome PerG Wide Com Pres Bruhn s.n. smooth brome PerGcheatgrass Intro Com AnnGJones’ Pres Introreedgrass Bruhn s.n. Unc Pres Palmer & slimstem reedgrass orchard grass PerGtimber Introoatgrass Unc Pres Johnson s.n. tufted hairgrass Great Basin wildrye squirreltail PerG Wide Com Pres Palmer & Common name

&

in ciliatus var. B. lanatipes. or oras is mostly According to B. anomalus B. porteri polyanthus, Ceratochloa carinata Includes vars. richardsonii, Bromopsis canadensis regional fl Bromopsis inermis C. neglecta, inexpansa D. caespitosa frondosus, Bromopsis porteri. Flora of North America (2007), material ascribed to elymoides, Sitanion hystrix Synonyms/ notes Taxonomic inermis Bromus carinatus B. marginatus, Bromus ciliatus Bromus inermis var. Bromus tectorum Anisantha tectorum Calamagrostis scopulorum Calamagrostis stricta Dactylis glomerata Danthonia intermedia Deschampsia cespitosa Bromus anomalus Bromus porteri, Elymus cinereus Leymus cinereus Elymus elymoides E. elymoides rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) and biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 37 Comments c name with taxonomic 1981 1989 1970 2006 1981 1935 1935 1989 Native to Eurasia Year Year doc. Palmer 1298 (CEBR) (CEBR) Palmer 1268 (CEBR) 22922 (CEBR) Palmer 1020 (CEBR) Palmer 1291 (CEBR) Palmer 1238 (CEBR) (CEBR) Source Pop. size Park status Range PerG Wide Unc Pres Jean 1116 PerG Wide UncPerG Wide Pres Palmer & UncPerG Pres Wide Fertig Unc Pres Palmer & PerG Wide Com Pres Palmer & Life form PerG Wide Unc Pres Palmer & PerG Intro Unc Pres Jean 1116 blue wildrye PerG Wide Unc Pres Palmer & thickspike wheatgrass Scribner wheatgrass western wheatgrass bluebunch wheatgrass slender wheatgrass Common name alpine sheep fescue intermediate wheatgrass

& &

var. var. includes

var. var. glaucus latiglumis, smithii ssp. inermis var. var. ssp. thsii riparium, A. albicans albicans, A. albicans griffi spicata palmeri Pascopyrum smithii; includes vars. Pseuodroegneria spicata; includes ssp. E. trachycaulus trachycaulus, E. alaskanus Agropyron caninum, A. subsecundum, A. trachycaulum, latiglume F. brachyphylla F. brachyphylla, F. Synonyms/ notes Taxonomic brachyphylla coloradensis hispidus, Agropyron intermedium, Elytrigia intermedia, Thinopyrum intermedium lanceolatus, Agropyron dasystachyum var. var. var. dasystachyum;

var. Elymus glaucus E. glaucus Elymus hispidus E. hispidus Elymus scribneri Agropyron scribneri Elymus smithii Agropyron smithii, Elymus spicatus Agropyron spicatum, Elymus trachycaulus Festuca ovina brevifolia Elymus lanceolatus E. lanceolatus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

38 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument and W North America Comments c name with taxonomic 1981 1935 1981 1989 1981 1989 1955 2006 1935 1954 Native to Eurasia 1954 2006 Native to Eurasia Year Year doc. Palmer 1124 (CEBR) Palmer 1171 (CEBR) Palmer 1118 (CEBR) (CEBR) Palmer 1299 (CEBR) (CEBR) H-185 (BRY) 22925 (CEBR) s.n. (CEBR) (CEBR) (CEBR) 22733 (CEBR) Source Pop. size Park status Range PerG Wide Unc Pres Palmer & PerG Wide Unc Pres Palmer & PerG Wide Rare Pres Fertig AnnG Intro Rare Pres Fertig PerG Wide Unc Pres Bruhn s.n. Life form fowl mannagrass meadow barley junegrass PerG Wide Unc Pres Palmer & oniongrassfoxtail muhly PerG PerG Wide Wide Rare Unc Pres Jean 1116 Pres Palmer & pullup muhly AnnG Wide Unc Pres Jean 1158 mat muhly PerG Wide Rare Pres Bruhn littleseed ricegrass alpine timothy PerGtimothy Wide Uncannual bluegrass PerG Pres Dickman Intro Unc Pres Bruhn s.n. Rocky Mountain fescue Common name rydbergii, F. rydbergii, F. G. elata var. Critesion brachyantherum K. cristata, pyramidata, K. nitida Piptatherum micranthum F. ovina F. saximontana Includes Synonyms/ notes Taxonomic

var. liformis Hordeum brachyantherum Koeleria macrantha Melica bulbosa Bromelica bulbosa Muhlenbergia andina Muhlenbergia fi Muhlenbergia richardsonis Oryzopsis micrantha Phleum alpinum commutatum P. Phleum pratense Poa annua Festuca ovina saximontana Glyceria striata rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) and biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name Gramineae (Poaceae)

Chapter 6: Appendices 39 Poa ed P. ” phase. includes the “ gracillima herbarium and by Roberts and Jean (1989) based on misidentifi specimen of secunda. Comments c name with taxonomic 1954 1954 1954 Native to Europe 1954 Material from CEBR 1940 Report from CEBR 1981 Native to Eurasia 1954 1989 1981 1935 Year Year doc. (CEBR) (CEBR) Palmer 1168 (CEBR) Palmer 1204 (CEBR) 9847 (BRY) Palmer 1087 (CEBR) Palmer 1063 (CEBR) Jean (1989) Palmer 1018 (CEBR) Palmer 1121 (CEBR) Source Pop. size Park status Range PerG Wide Unc Pres Bruhn s.n. PerG Wide Rare Pres Bruhn s.n. PerG Intro Com Pres Palmer & PerG Wide Unc Pres Palmer & PerG Wide ? Hist Harrison PerG Intro Unc Pres Palmer & Life form PerG Wide NA Rep Roberts & PerG Wide Unc Pres Palmer & PerG Wide Com Pres Palmer & interior bluegrass Wheeler bluegrass Kentucky bluegrass Sandberg’s Sandberg’s bluegrass Arctic bluegrass Canada bluegrass muttongrass PerG Wide Com Pres Palmer & Common name needle-and- thread Indian ricegrass Letterman’s Letterman’s needlegrass

P. var. and & & interior, interior,

rctica ssp. ssp. includes P. secunda P. P. cusickii P. P. gracillima, P. gracillima, P. P. P. nemoralis P. interior P. P. wheeleri P. juncifolia, P. sandbergii, juncifolia, P. scabrella, P. canbyi, P. P. ampla, nevadensis, P. Includes all vars. of grayana Includes vars. a fendleriana longiligula; epilis reports of Synonyms/ notes Taxonomic Hesperostipa comata comata Achnatherum hymenoides

var. var. var. glauca Poa nervosa wheeleri Poa pratensis agassizensis P. Poa secunda Stipa comata comata Poa compressa Poa fendleriana longiligula, P. P. Poa arctica Poa glauca Stipa hymenoides Oryzopsis hymenoides, Stipa lettermanii Achnatherum lettermanii rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyGramineae (Poaceae) Species name Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

40 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Comments c name with taxonomic 1981 1935 1981 1989 1981 1981 2005 2007 Year Year doc. 1981 2006 1970 Palmer 1259 (CEBR) Palmer 1093 (CEBR) Palmer 1250 (CEBR) Reynolds 23456 (CEBR) Palmer 1187 (CEBR) Palmer 1176 (CEBR) (CEBR) Reynolds 23328 (CEBR) Source Palmer 1008 (CEBR) 22911 (CEBR) Palmer 1232 (CEBR) Pop. size Park status Range PerG Wide Unc Pres Palmer & PerF Wide Unc Pres Fertig & Life form PerG Wide Unc Pres Palmer & PerG Wide Unc Pres Fertig & owered Nelson’s Nelson’s needlegrass spike trisetum PerG Wide Unc Pres Palmer & wand phacelia PerF Wide Unc Pres Palmer & Rocky Mountain iris Common name Mertens’ rush PerG Wide Rare Pres Palmer & small-fl woodrush Baltic rush PerG Wide Rare Pres Harms s.n. perplexing rush Tracy’s rushTracy’s PerG Wide Unc Pres Palmer & mountain rush PerG Widelongstyle rush Rare PerG Pres Wide Fertig Unc Pres Palmer &

includes nelsonii by some ater, J. ater, balticus, alaskanum mexicanus var. heterophylla montanus; J. ensifolius var. ssp. var. , J. saximontanus brunnescens virgata nelsonii, S. montanus congdonii, T. congdonii, T. Achnatherum nelsonii var. columbiana & montanum Includes ssp. Includes ssp. & var. Synonyms/ notes Taxonomic J. arcticus balticus includes var. includes var. Included in var. authors forma J. tracyi, xiphoides J. saximontanus;

var. ora brunnescens montanus Stipa nelsonii nelsonii Trisetum spicatum Trisetum Phacelia heterophylla Iris missouriensis Juncus arcticus J. arcticus mertensianus Luzula parvifl Juncus confusus Juncus ensifolius var. Juncus ensifolius var. Juncus longistylus Juncus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name Gramineae (Poaceae) Hydrophyllaceae Iridaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae

Chapter 6: Appendices 41 ed eld Co., UT Sensitive, USFWS: former C2 candidate. locality is Type Navajo Lake, Kane Co., UT (Maguire 19474 NY) Panguitch Lake, Garfi (Jones 6023f POM) Cedar Breaks vicinity Other by Welsh. reports are based on misidentifi specimens of var. elatus. Comments c name with taxonomic 1942 USFS Region 4: 1981 1930 1966 locality is Type 1970 1982 Reported from 1970 2007 1981 Year Year doc. (CEBR) Palmer 1108 (CEBR) s.n. (CEBR) 22989 (CEBR) Palmer 1300 (CEBR) 21286 (NY) Palmer 1243 (CEBR) Reynolds 23349 (CEBR) Palmer 1256 (CEBR) Source Pop. size Park status Range PerF LocEn Unc Pres Jean 317 PerF Wide RarePerF Pres Wide Palmer & Com Pres Dickman PerF RegEn UncPerF Pres Fertig Wide UncPerF Wide Pres Palmer & NA Rep Welsh PerF Wide Unc Pres Palmer & Life form Navajo Lake milkvetch great rushy milkvetch Rydberg’s weedy milkvetch Panguitch milkvetch tall kentrophyta mountain kentrophyta stinking horsemint self-heal PerF Wide Rare Pres Fertig & Common name pulse milkvetch PerF Wide Unc Pres Palmer &

var. var. A. kentrophyta implexus M. odoratissima glauca, M. glauca Synonyms/ notes Taxonomic

var. var. var. var. oblongifolius lanceolata limnocharis Astragalus lonchocarpus Astragalus miser var. panguicensis elatus tegetarius Astragalus kentrophyta Astragalus kentrophyta Astragalus limnocharis Monardella odoratissima Prunella vulgaris var. Astragalus argophyllus Astragalus tenellus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) FamilyLabiatae (Lamiaceae) Species name Labiatae (Lamiaceae) Leguminosae (Fabaceae)

42 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument ed eld Co., UT Carmel, Kane Co., UT (Ferguson & Ottley 5613 NY) (Ward 541 GH) (Ward from CEBR herbarium based on misidentifi specimens of var. rubricaulis. Comments Aquarius Plateau, Garfi c name with taxonomic 1981 1939 locality is W of Type 1930 1981 2006 Previous reports 1942 1954 2001 Year Year doc. 1989 1986 Native to Europe 1935 locality is Type Palmer 1144 (CEBR) Palmer s.n. (CEBR) Palmer 1040 (CEBR) Palmer 1016 (CEBR) 22928 (CEBR) Peiler s.n. (NY) (CEBR) Malm, & Alexander s.n. (CEBR) Source (CEBR) (CEBR) Palmer 1269 (CEBR) Pop. size Park status Range PerF RegEn Unc Pres Palmer & PerF Wide Unc Pres Palmer & Life form PerF Wide Com Pres Bruhn s.n. PerF Intro Unc Pres Johnson s.n. PerF Wide Unc Pres Palmer & ower largefl sweetpea Utah trefoil PerF RegEn Rare Pres Palmer & Common name Rusby’s cloverRusby’s PerF Wide Com Pres Palmer & American vetch silvery lupine PerF Wide Comsilvery lupine Pres Fertig PerF Widesubalpine lupine NA lupineKing’s Hist AnnF Degener & Wide Rare Pres Louie, stemless lupine PerF Wide Unc Pres Jean 1124 yellow sweet- clover mountain locoweed

utahensis brachypus

ssp. var. var. var. L. laetivirens Synonyms/ notes Taxonomic L. argenteus spathulatus T. longipes T. L. lepidus caespitosus, L. caespitosus

var.

var. var. argentatus rusbyi americana argenteus rubricaulis utahensis cinalis Lathyrus lanszwertii laetivirens Lotus utahensis Lupinus argenteus var. Trifolium longipes Trifolium var. Vicia americana var. kingii Lupinus argenteus var. Lupinus argenteus var. Lupinus kingii Lupinus lepidus var. Melilotus offi oreophila Oxytropis oreophila rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Chapter 6: Appendices 43 Pyrola ed ower of Utah from CEBR herbarium based on misidentifi specimens of chlorantha Comments c name with taxonomic 1935 1947 1981 State fl 1941 1954 1925 1981 1954 1987 Native to Eurasia 2007 Previous reports 2006 Year Year doc. Buchanan s.n. (CEBR) s.n. (BRY) Palmer 1028 (CEBR) (CEBR) Palmer 1049 (CEBR) Malm, & Alexander s.n. (CEBR) Palmer 1211 (CEBR) Palmer 1241 (CEBR) s.n. (CEBR) Reynolds photo (CEBR) 22912 (CEBR) Source Pop. size Park status Range PerF Wide NA Hist Woodbury PerF Wide UncPerF Wide Pres Palmer & UncPerF Pres Wide Louie, Unc Pres Palmer & AnnF IntroPerF Rare Wide Pres Rare Johnston Pres Fertig & Life form PerF Wide Rare Pres Fertig ax PerF Wide Unc Pres& Palmer reweed PerF Wide Com Pres& Dickman ax fi twincrest onion sego-lilyleopard-lily PerF Wide PerFstarry false UncSolomon’s-seal Wide Pres Uncmountain Palmer & death camas Pres Boyle 1038 yellow King’s fl blue fl common mallow woodland pinedrops Common name Hall’s willow- Hall’s herb

ssp. bisceptrum lewisii, var. canescens, C. palmeri. circumvagum, C. danielsii Includes vars. & L. lewisii Adenolinum lewisii Chamerion angustifolium var. angustifolium Synonyms/ notes Taxonomic

var. lewisii Allium bisceptrum Calochortus nuttallii Fritillaria atropurpurea Smilacina stellata Maianthemum stellatum Zigadenus elegans Anticlea elegans Linum kingii Mesyniopsis kingii Linum perenne ssp. Malva neglecta Pterospora andromedea Epilobium angustifolium canescens Epilobium halleanum rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Onagraceae FamilyLiliaceae (Alliaceae) Species name Liliaceae (Calochortaceae) Liliaceae Liliaceae (Convallariaceae or Ruscaceae) Liliaceae (Melanthiaceae) Linaceae Linaceae Malvaceae Monotropaceae (Ericaceae) Onagraceae

44 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Comments c name with taxonomic 1977 1954 1989 1981 1981 1988 Year Year doc. 1981 1989 1977 1981 1981 Palmer 1127 (CEBR) Palmer 1022 (CEBR) 22742 (CEBR) Palmer 1155 (CEBR) Palmer 1145 (CEBR) (CEBR) Source Palmer 1101 (CEBR) Louie, & Alexander s.n. (CEBR) Buchanan s.n. (CEBR) Palmer 1158 (CEBR) Palmer 1106 (CEBR) Pop. size Park status Range PerF Wide Unc Pres Palmer & PerF Wide Unc Pres Palmer & Life form PerF Wide Unc Pres Palmer & PerF Wide Rare Pres Palmer & PerF Wide Unc Pres Jean 101 Tree Wide Com Pres Malm, PerF Sparse Rare Pres Dickman & PerF Wide Unc Pres Palmer & PerF Wide Rare Pres Palmer & r Tree Wide Com Pres& Palmer r Tree Wide Com Pres Fertig Hornemann’s Hornemann’s willow-herb long-tube evening- primrose Common name subalpine fi yellow evening- primrose spotted coral- root giant rattlesnake- plantain Engelmann spruce white bog- orchid northern bog orchid slender bog orchid white fi

var. ora, maculata var. Platanthera ava Synonyms/ notes Taxonomic O. fl C. maculata Platanthera dilatata dilatata Limnorchis hyperborea; includes huronensis aquilonis, P. Platanthera sparsifl not recognized in Vars. Flora of North America 2003

var. ava var. var. ora ora ava fl dilatata Epilobium hornemannii hornemannii Oenothera caespitosa marginata var. Oenothera fl Abies lasiocarpa A. bifolia Corallorhiza maculata Goodyera oblongifolia Habenaria dilatata var. Picea engelmannii Habenaria hyperborea Habenaria sparsifl sparsifl Abies concolor rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyOnagraceae Species name Onagraceae Onagraceae Pinaceae Orchidaceae Orchidaceae Orchidaceae Pinaceae Orchidaceae Orchidaceae Pinaceae

Chapter 6: Appendices 45 CO Brigham Peak, Piute Co., UT (Jones 5949 NY) Comments c name with taxonomic 1989 State tree of UT and 1989 1970 1989 1989 1935 locality is Type 1940 1947 1989 1981 1981 1989 Year Year doc. 22980 (CEBR) Jean (1989) (RM) Louie, & Alexander s.n. (CEBR) 22982 (CEBR) s.n. (CEBR) s.n. (CEBR) (CEBR) 23310 (CEBR) Palmer 1236 (CEBR) Palmer 1194 (CEBR) Reynolds 23352 (CEBR) Source Pop. size Park status Range Tree Wide NA Rep Roberts & Tree Wide ComTree Pres Wide Malm, Unc Pres Fertig PerF LocEn Unc Pres Buchanan Life form PerF Wide Rare Pres Harms s.n. PerF Wide Unc Pres Palmer & r Tree Wide Unc Pres Fertig blue spruce Tree Wide Unc Pres Fertig two-needle pinyon limber pine Tree Wide Unc Pres Nelson 9031 western bristlecone pine ponderosa pine Douglas-fi Cedar Breaks gilia cushion phlox PerF Wide Unc Pres Dickman Common name western Jacob’s-ladder Tweedy’s Tweedy’s plantain small collomia AnnF Wide Unc Pres Palmer & scarlet gilia PerF Wide Unc Pres Fertig &

var.

pulvinata, ssp. ssp. tridactyla P. caespitosa P. P. occidentale, P. occidentale, P. P. caeruleum amygdalinum Synonyms/ notes Taxonomic Gilia aggregata aggregata Gilia tridactyla; I. spicata ssp.

. var. var. var exilis scopulorum Picea pungens Pinus edulis Pinus fl glauca Pinus longaeva aristata P. Pinus ponderosa var. Pseudotsuga menziesii Phlox pulvinata sibirica P. Polemonium caeruleum pterospermum Plantago tweedyi Collomia linearis Ipomopsis aggregata aggregata Ipomopsis tridactyla rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyPinaceae Species name Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Polemoniaceae Polemoniaceae Plantaginaceae Polemoniaceae Polemoniaceae Polemoniaceae

46 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument is Cedar alpestre Breaks NM; Iron Co., UT (Goodman & Hitchcock 1601) Comments c name with taxonomic 1935 1989 Native to Eurasia. 1934 1981 1954 1930 1930 locality of var. Type 1954 1946 1935 1954 Year Year doc. (CEBR) (CEBR) Palmer 1163 (CEBR) Palmer 1272 (CEBR) (CEBR) s.n. (CEBR) & Hitchcock 1601 (NY holotype) (CEBR) F-59 (CEBR) Palmer 1088 (CEBR) (CEBR) Source Pop. size Park status Range PerF Wide Rare Pres Bruhn s.n. PerF Wide Unc Pres Palmer & AnnF Wide Unc Pres Palmer & AnnF Wide Rare Pres Bruhn s.n. PerF RegEn Com Pres Buchanan PerF LocEn Com Pres Goodman PerF Wide Unc Pres Bruhn s.n. Life form sulfur wild buckwheat yard knotweed AnnF Intro Unc Pres Jean 121 American bistort Douglas’ knotweed Sawatch knotweed Porter’s wild Porter’s buckwheat Cedar Breaks wild buckwheat pretty Jacob’s- ladder Common name alpine bistort PerF Wide Rare Pres Crawford western dock PerF Wide Unc Pres Palmer & willow dock PerF Wide Unc Pres Bruhn s.n. parsley-fern Fern Wide NA Hist s.n. 1935

var.

var.

var.

aviculare var. var. ssp. occidentalis P. arenastrum, P. arenastrum, P. P. aviculare Bistorta bistortoides P. douglasii P. sawatchense P. sawatchense alpestre pulcherrimum E. chrysocephalum P. pulcherrimum P. Bistorta vivipara R. mexicanus, salicifolius mexicanus, R. utahensis, R. triangulivalvis C. acrostichoides Synonyms/ notes Taxonomic fenestratus, R. aquaticus ssp.

var.

var. var. var. var.

var. var. var. triangulivalvis umbellatum Polygonum aviculare Polygonum bistortoides douglasii johnstonii Polygonum douglasii Polygonum douglasii porteri Eriogonum umbellatum Eriogonum umbellatum Eriogonum panguicense alpestre delicatum Polemonium pulcherrimum Polygonum viviparum Rumex occidentalis R. aquaticus Rumex salicifolius ssp. Cryptogramma crispa acrostichoides rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae and biogeographic notes, cont. Appendix A. Confi FamilyPolemoniaceae Species name Polygonaceae Polygonaceae Polypodiaceae (Adiantaceae)

Chapter 6: Appendices 47 Gorge Wilderness Area (Fertig 23378) in 2007 Comments c name with taxonomic 1941 1942 2006 1954 1930 Located in Ashdown 1942 1942 1981 1977 1988 1981 Year Year doc. (CEBR) Palmer 1039 (CEBR) 22924 (CEBR) Buchanan s.n. (CEBR) & Hitchcock 1609 (NY) Palmer 1034 (CEBR) Palmer 1066 (CEBR) Palmer 1295 (CEBR) Buchanan s.n. (CEBR) (CEBR) Palmer 1289 (CEBR) Source Pop. size Park status Range PerF Wide Unc Pres Boyle 1030 PerF Wide Unc Pres Palmer & Fern Wide Rare Pres Fertig AnnF Wide UncPerF Pres Dickman & Wide NA Hist Goodman PerF Wide UncPerF Pres Wide Palmer & Unc Pres Palmer & PerF Wide Unc Pres Palmer & Life form PerF Wide Rare Pres Dickman & PerF Wide Unc Pres Jean 134 ower owered lanceleaf springbeauty alpine bitter- root brittle bladder fern pygmy-fl rock-jasmine alpine shooting-star dark-throat shooting-star Parry’s primrose pink wintergreen Common name green- fl wintergreen whiteveined wintergreen sidebells pyrola PerF Wide Com Pres Palmer &

&

Includes & var. ssp. C. rosea. var. C. tenuis & Welsh et al. Welsh picta; orum. C. utahensis, L. nevadensis P. dentata P. var. Includes Includes ssp. puberulenta, subumbellata, reevesiana, Includes C. lanceolata lanceolata. (2003) include subilifera D. paucifl “subvars” from UT Flora coriaceum, atwoodii, huberi asarifolia includes Synonyms/ notes Taxonomic

var. Lewisia pygmaea Oreobroma pygmaea. Androsace septentrionalis Claytonia lanceolata Cystopteris fragilis Dodecatheon alpinum Dodecatheon pulchellum pulchellum Primula parryi Pyrola asarifolia rotundifolia P. Pyrola chlorantha virens P. Pyrola picta picta P. Pyrola secunda Orthilia secunda rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Portulacaceae Primulaceae Portulacaceae FamilyPolypodiaceae (Aspleniaceae) Species name Primulaceae Primulaceae Primulaceae Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae)

48 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument is Toquerville, is Toquerville, Co., UT Washington (Jones 1716 GH) Comments c name with taxonomic 1935 1940 1942 1930 1981 1989 locality Type 1935 1988 Year Year doc. 1930 1981 1930 1929 Palmer 1166 (CEBR) Buchanan s.n. (CEBR) s.n. (CEBR) F-54 (CEBR) Palmer 1011 (CEBR) Reynolds 23346 (CEBR) (CEBR) (CEBR) Source (CEBR) Palmer 1014 (CEBR) (CEBR) F-13 (CEBR) Pop. size Park status Range PerF Wide Unc Pres Palmer & PerF Wide Unc Pres Dickman & Life form PerF WideShrub Unc Wide Unc PresShrub Crawford Wide Pres Palmer & NA Pres Fertig & PerF Wide Unc Pres Boyle 733 PerF Wide Unc Pres Croft 34CB PerF Wide Unc Pres Palmer & PerF RegEn Unc Pres Boyle 754 PerF Wide Unc Pres Crawford Columbian monkshood cutleaf anemone Common name drab buttercup PerF Wide Fendler’s Uncmeadow-rue Utah Presmountain-lilac Dickman Utah serviceberry Colorado columbine rock columbine PerF RegEn Rare Pres Jean 108 white marshmarigold Columbian virgin’s-bower subalpine larkspur plantain buttercup

includes var. pinetorum Includes var. includes vars. & da, saxicola, da, ora, Psychrophila ora, multifi stylosa Includes var. Includes var. ochroleucum A. tetonensis; vars. & Synonyms/ notes Taxonomic alpeophilus ochroleuca leptosepala C. pseudoalpina, Atragene columbiana R. inamoenus inamoenus.

var. var. var. da Aconitum columbianum columbianum Anemone multifi Aquilegia coeruleacaerulea; A. Thalictrum fendleri Ceanothus martinii Amelanchier utahensis Aquilegia scopulorum scopulorum Caltha leptosepala C. bifl Clematis columbiana Delphinium barbeyi Ranunculus alismifolius montanus Ranunculus inamoenus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyRanunculaceae (Helleboraceae) Species name Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Thalictraceae) Rhamnaceae Rosaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae

Chapter 6: Appendices 49 C. & (Jean C. ledifolius montanus 1103 CEBR) have been documented in CEBR. Comments c name with taxonomic 1935 1954 1946 1989 Hybrids between 1981 1981 2007 1935 1981 1981 1946 Year Year doc. 1047 (CEBR) (CEBR) Palmer 1151 (CEBR) 22913 (CEBR) Palmer 1140 (CEBR) Palmer 1007 (CEBR) Reynolds 23606 (CEBR) Palmer 1182 (CEBR) Palmer 1255 (CEBR) Palmer 1207 (CEBR) F-31 (CEBR) Source Pop. size Park status Range PerF WidePerF Unc Wide Pres Rare Palmer Pres Bruhn s.n. Shrub Wide Unc Pres Palmer & Shrub Wide Unc Pres Fertig PerF Wide RareShrub Wide Pres Unc Palmer & Pres Palmer & Shrub Wide Rare Pres Fertig & PerF Wide Unc Pres Palmer & PerF Wide Unc Pres Crawford Life form Virginia strawberry large-leaved avens shrubby cinquefoil curl-leaf mountain mahogany woolly cinquefoil antelope bitterbrush alder-leaf alder-leaf mountain mahogany prairie smoke PerF Wide Unc Pres Palmer & Sevier ivesia PerFwedgeleaf RegEncinquefoil Rare Pres Palmer & beautiful cinquefoil Common name

& ora

hippiana oribunda fl glauca var. var. ssp. oribunda, Dasiphora oribunda, platypetala C. ledifolius intercedens Includes ssp. Erythrocoma trifl fruticosa, Pentaphylloides fl fruticosa P. pulcherrima P. Synonyms/ notes Taxonomic

var.

var. var. orum ciliatum pulcherrima Geum macrophyllum perincisum Geum trifl var. Potentilla gracilis var. Cercocarpus ledifolius intermontanus Potentilla hippiana hippiana P. Purshia tridentata Cercocarpus montanus Fragaria virginiana Ivesia sabulosa Potentilla diversifolia diversifolia Potentilla fruticosa Pentaphylloides rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi Rosaceae Rosaceae Rosaceae FamilyRosaceae Species name Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae

50 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Sensitive, USFWS: 3C Comments c name with taxonomic 1989 2007 1989 1989 2006 1981 USFS Region 4: 1954 1981 1981 1989 1989 1988 Year Year doc. 22923 (CEBR) Reynolds 23394 (CEBR) 22986a (CEBR) 22741 (CEBR) 22986b (CEBR) Sidles 20137 (BRY) (CEBR) (CEBR) Palmer 1017 (CEBR) Jean (1989), Buchanan (1992) 22983 (CEBR) (CEBR) Source Pop. size Park status Range Shrub Wide Unc Pres Fertig PerF Wide Rare Pres Fertig & Tree Wide Unc Pres Fertig Tree Wide Unc Pres Fertig Shrub Wide Unc Pres Harms s.n. Life form Shrub Wide Unc Pres Jean 135 common red raspberry sweet-scented bedstraw narrow-leaved cottonwood quaking aspen Tree Wide Com Pres Fertig lance-leaf cottonwood Arizona willow Shrub RegEn Unc willowBooth’s Shrub Pres Wideshort-fruit Atwood & willow Unc Pres Bruhn s.n. coyote willow Shrub Wide Unc Pres Palmer & Nootka rose Shrub Wide ? Rep Roberts & Woods’ roseWoods’ Shrub Wide Unc Pres Fertig Common name willow P. & x

exigua hybrid var. var. var. P. fremontii P. melanolasius between acuminata, P. acuminata, P. P. angustifolia S. exigua ultramontana R. idaeus aculeatissimus, R. ideaus ssp. Synonyms/ notes Taxonomic

var. var. var. orum x Galium trifl Populus angustifolia Populus tremuloides Populus intercurrens Salix arizonica Salix boothii Salix brachycarpa Salix exigua stenophylla Rosa nutkana hispida Rosa woodsii R. woodsii strigosus Rubus idaeus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi Rubiaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salix scouleriana Scouler’s and biogeographic notes, cont. Appendix A. Confi FamilyRosaceae Species name Rosaceae Rosaceae

Chapter 6: Appendices 51 Sensitive, BLM (UT): Sensitive, USFWS: former C2 candidate. locality is S Fork Type Indian Canyon, Kane 1411 Co., UT (Tuhy NY). Comments c name with taxonomic 1989 USFS Region 4: 1981 1941 1954 1954 1988 1934 1942 1981 1954 Year Year doc. (CEBR) Palmer 1038 (CEBR) Palmer 1215 (CEBR) s.n. (CEBR) Palmer 1059 (CEBR) (CEBR) (CEBR) s.n. (CEBR) Palmer 1292 (CEBR) Palmer 1157 (CEBR) Source Pop. size Park status Range PerF Wide Unc Pres Palmer & Life form PerF Wide Unc Pres Palmer & Shrub Wide Com Pres Palmer & PerF Wide Unc Pres Harms s.n. PerF Wide Unc Pres Dickman PerF Wide Unc Pres Palmer & PerF Wide Com Pres Palmer & Zion jamesia Shrub LocEn Rare Pres Jean 1111 fringecup woodlandstar Common name marsh grass-of- Parnassus wax currant Shrub Wide Unc Pres Buchanan alpine prickly currant sticky currant Shrub Wide Unc Pres Jean 134 brook saxifrage diamondleaf saxifrage Wyoming paintbrush scarlet paintbrush pedicellare tenuis var. L. bulbifera, glabra Synonyms/ notes Taxonomic Includes var. Includes var. R. cereum S. arguta, Micranthes odontoloma Micranthes rhomboidea C. linariaefolia

var. var.

var. zionis montanensis Jamesia americana var. Lithophragma glabrum ramulosum Parnassia palustris var. Ribes cereum inebrians Ribes montigenum Ribes viscosissimum viscosissimum Saxifraga odontoloma Saxifraga rhomboidea Castilleja linariifolia Castilleja miniata rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilySaxifragaceae (Hydrangeaceae) Species name Saxifragaceae Saxifragaceae (Parnassiaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae Saxifragaceae Scrophulariaceae Scrophulariaceae

52 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument eld eld Co., UT Sensitive, USFWS: former C2 candidate. locality is road Type to Bryce Point, Garfi Canyon NP, Co., UT (Holmgren & Reveal 2017 NY). Comments Mammoth Creek, Markagunt Plateau, Garfi (Jones 6026b US) c name with taxonomic 1989 USFS Region 4: Year Year doc. 1989 1989 1954 1934 1954 1981 1930 locality is Type 1934 1930 (CEBR) Source 23333 (CEBR) Jean (1989) Palmer 1180 (CEBR) Palmer 1191 (CEBR) Malm, & Alexander s.n. (CEBR) Palmer 1009 (CEBR) (CEBR) & Dickman s.n. (CEBR) F-41 (CEBR) Pop. size Park status Range Life form PerF LocEn Rare Pres Jean 1111 PerF Wide Rare Pres Fertig PerF WidePerF NA Wide Rep Unc Roberts & Pres Palmer & PerF Wide Unc Pres Louie, PerF Wide UncPerF LocEn Pres Unc Palmer & PresPerF Boyle 732 RegEn UncPerF Pres Wide Buchanan Unc Pres Crawford ower Common name Reveal’s Reveal’s paintbrush rhexia-leaved paintbrush sulphur paintbrush yellow monkeyfl elephant-head PerF Wide Unc Pres Palmer & Parry’s Parry’s lousewort undulate penstemon Markagunt penstemon Rydberg’s penstemon Whipple’s penstemon mogollonica Synonyms/ notes Taxonomic C. revealii C. rhexifolia C. sulphurea Includes var. Includes var.

var. var. revealii rhexifolia sulphurea parryi undosus Castilleja parvula var. Castilleja rhexifolia var. Castilleja rhexifolia var. Mimulus guttatus Pedicularis groenlandica Pedicularis parryi var. aggregatus Penstemon eatonii var. Penstemon leiophyllus leiophyllus Penstemon rydbergii Penstemon whippleanus rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyScrophulariaceae Species name Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae

Chapter 6: Appendices 53 eld Lake Mts., Sevier Co., 327 GH) UT (Ward Known only from saplings Sensitive, USFWS: former C2 candidate. locality is Cedar Type Breaks, Cedar Breaks NM; Iron Co., UT (Mathias 703 MO). locality is Bryce Type Canyon, Garfi Co., UT (Mathias 670 MO). Comments c name with taxonomic 1941 locality is Fish Type 1954 1954 1981 2007 Native to Eurasia. 1946 1934 1929 USFS Region 4: 1935 1935 USFWS: Category 3C. Year Year doc. Palmer 1044 (CEBR) (CEBR) Palmer 1235 (CEBR) Palmer 1190 (CEBR) Reynolds 23397 (CEBR) Palmer 1219 (CEBR) Palmer 1086 (CEBR) Palmer 1056 (CEBR) Palmer 1096 (CEBR) Palmer 1271 (CEBR) Source Pop. size Park status Range PerF RegEn Unc Pres Palmer & PerF Wide Rare Pres Bruhn s.n. PerF Wide Unc Pres Palmer & PerF Wide Unc Pres Palmer & Shrub Intro Rare Pres Fertig & PerF Wide Unc Pres Palmer & PerF Wide Unc Pres Palmer & PerF LocEn Unc Pres Palmer & PerF Wide Com Pres Palmer & Life form ve-stamen cutleaf synthyris American brooklime thyme-leaved speedwell American alpine speedwell fi tamarisk pinnate angelica Lemmon’s Lemmon’s spring-parsley least spring- parsley southern ligusticum least lomatium PerF LocEn Unc Pres Palmer & Common name

var.

ssp. var. da laciniata Veronica serpyllifolia Veronica humifusa, Veronicastrum serpyllifolium humifusum V. nutans V. ramosissima Pseudocymopterus montanus Synonyms/ notes Taxonomic Synthyris laciniata S. pinnatifi Veronica Veronica americana Veronica serpyllifolia Veronica Veronica wormskjoldii Tamarix chinensisTamarix pentandra, T. T. Angelica pinnata Cymopterus lemmonii Cymopterus minimus Ligusticum porteri Lomatium minimum rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyScrophulariaceae Species name Scrophulariaceae Scrophulariaceae Scrophulariaceae Tamaricaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae)

54 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument

Picea (rarely or exilis Pseudotsuga Abies Pinus fl on menziesii on species) Comments c name with taxonomic 1979 1954 1954 1940 1977 1970 parasitic on Typically 2007 parasitic Typically Year Year doc. s.n. (CEBR) Palmer 1138 (CEBR) (CEBR) (CEBR) Palmer 1208 (CEBR) (RM) Reynolds 23608 (CEBR) Source Pop. size Park status Range PerF Wide Unc Pres Buchanan PerF Wide Rare Pres Bruhn s.n. PerF Wide Unc Pres Boyle 738 PerF Wide Unc Pres Palmer & PerF Sparse Unc Pres Nelson 9031 PerF Wide Rare Pres Fertig & Life form r blunt sweet- cicely tobacco-root PerFwestern Widevalerian Unc Pres Palmer & Canadian white violet northern bog violet limber pine dwarf- mistletoe Douglas-fi mistletoe Common name

var.

nis var. affi var. rugulosa, V. scopulorum, rugulosa, V. rydbergii V. A. campylopodium cyanocarpum Synonyms/ notes Taxonomic Osmorhiza depauperata Valeriana edulis Valeriana Valeriana occidentalis Viola canadensis canadensis V. Viola nephrophylla sororia V. Arceuthobium cyanocarpum Arceuthobium douglasii rmed, historical, and reported vascular plant taxa of Cedar Breaks National Monument, organized by family scientifi and biogeographic notes, cont. Appendix A. Confi FamilyUmbelliferae (Apiaceae) Species name Valerianaceae Valerianaceae Violaceae Violaceae Viscaceae Viscaceae

Chapter 6: Appendices 55

in Wilderness Area (Fertig 23377) Flora of North America (2003) Welsh et al. (2003) Welsh Lake; Iron Co., UT (Maguire 19472 NY). Considered a synonym of Minuartia macrantha Flora of North America (2005). Range Comments Life form cupseed stickseedshowy stoneseed AnnFwestern stoneseed Wide PerF PerF Wide Wide basin yellow cryptanth PerFshowy stickseedspotted stickseed Wide PerF PerF Wide Wide bigtooth maplebox elder Treeredroot pigweed Widesquawbush Occurs in Ashdown Gorge AnnF Tree Intro Wide Shrub Native to Central America Wide ashy cryptanth PerF Wide spindle-rooted bluebell PerFArizona pincushion RegEn PerF Wide not recognized in Vars. squawbushwestern poison ivyabortive cryptanth Shrub PerF Wide Shrub Wide Periph shrubby sandwort PerF Periph not recognized in Vars. Rydberg’s sandwortRydberg’s PerF Wide beach sandwort PerF RegEn locality is Navajo Type

by

by var. interius liorum; var. arizonica ssp. var. interius, simplicifolia trilobata var. var. var. M. oblongifolia C. atwoodii liorum, Minuartia fi cupulata, L. marginata Included in L. texana, redowskii Negundo aceroides R. trilobata some authors Escobaria vivipara Includes macradenia obtusiloba included in M. macrantha some authors

var. liorumfi A. var. fi ora patens var. var.

trilobata R. trilobata exus var. var. var. var. oribunda National Monument. rmed) vascular plant taxa of Cedar Breaks Lithospermum incisum Lithospermum ruderale Mertensia fusiformis Hackelia fl Hackelia patens Lappula occidentalis cupulata Acer grandidentatum Acer negundoAmaranthus retrofl Rhus aromatica simplicifolia Rhus aromatica A. negundo Cryptantha confertifl Arenaria macradenia Eremogone macradenia Coryphantha vivipara arizonica Toxicodendron rydbergiiToxicodendron Cryptantha cinerea abortiva radicans Rhus radicans, T. Cryptantha cinerea cinerea Arenaria obtusilobaArenaria rubella Minuartia obtusiloba, Lidia Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Aceraceae Amaranthaceae Anacardiaceae Anacardiaceae Boraginaceae Caryophyllaceae (Alsinaceae) FamilyAceraceae Species name notes Synonyms/Taxonomic Common name Cactaceae Anacardiaceae Boraginaceae Boraginaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Appendix B. Potential (unconfi

56 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument

Flora of North America (2005) Flora of North America (2005) Wilderness Area (Fertig 2007 observation) Wilderness Area (Fertig 23358) Range Comments Life form Shrub Wide Occurs in Ashdown Gorge Scouler’s campionScouler’s PerFumbrella starwortfourwing saltbush Periph PerF goosefootFremont’s Shrub Wide AnnF Wide poverty-weedNuttall’s AnnF Wide bur ragweed Wide mountain pussytoes PerF arnicaChamisso’s AnnF Wide Wide wormwoodCarruth’s PerF PerF Wide Wide Occurs in Ashdown Gorge Bering chickweed PerFnodding chickweed Wide AnnF not recognized in Vars. Sparse Not reported for UT in fringed sagebrush Shrub Wide mountain big sagebrush reddish sandwort PerF Wide black sagebrush Shrub Wide Eaton’s asterEaton’s cutleaf PerF Wide PerF Wide

Silene var. earlei, C. andina capillare by some ssp. occidentalis nova, var. var. var. var. hallii A. tridentata ssp. ora, Seriphidium ora, scouleri authors A. canescens C. fremontii UT material included in vaseyanum paucifl rubella beeringianum Seriphidium novum Includes bracteolatus

var.

foliosa A. chamissonis

var. rubella Minuartia rubella, Tryphane var. nova A. arbuscula var. pringlei var. var. ssp. rmed) taxa of Cedar Breaks National Monument, cont. Stellaria umbellata Atriplex canescens canescens Chenopodium fremontii fremontii Monolepis nuttalliana Ambrosia acanthicarpaAntennaria umbrinella Franseria acanthicarpa Arnica chamissonis Artemisia carruthii Arenaria rubella Cerastium nutans Silene scouleri Artemisia frigida Artemisia nova Cerastium beeringianum C. beeringianum Artemisia tridentata vaseyana Aster eatoniiBahia dissecta Symphyotrichum eatonii, A. Caryophyllaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Caryophyllaceae Caryophyllaceae Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi FamilyCaryophyllaceae (Alsinaceae) Species name notes Synonyms/Taxonomic Common name Caryophyllaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 57 Canyon; Iron Co., UT (Atwood & Higgins 5918 BRY) Sevier Co., UT (Jones 5715 NY) Washington Co., UT Washington (Palmer s.n. NY) Native to Eurasia Wilderness Area (Fertig 2007 observation) in Ashdown Gorge Wilderness Area (Fertig 23390) Range Comments Shrub RegEn locality is St. George, Type Life form PerF Wide ower Margakunt thistle PerF LocEn locality is Cedar Type glabrate rabbitbrush Shrub Wide smoothseed rabbitbrush Fish Lake thistle PerF RegEn locality is Fish Lake, Type Vasey’s rabbitbrushVasey’s lanceleaf rabbitbrush Shrub Wide Shrub Wide Canada thistle PerF Intro Utah state noxious weed. green rabbitbrushchicory Shrub Wide PerF Intro Native to Eurasia Canadian horseweed AnnF Wide tassel-fl brickellbush arrowleaf balsamroot PerF Wide Occurs in Ashdown Gorge elk thistlebull thistle PerF Wide PerF Intro Native to Europe. Occurs

var. & include

var. var. oreophilus horridum var. canadensis some authors Ericameria nauseosa leiosperma C. nauseosus glabrata, integrifolium, Breea arvensis Includes vars. Ericameria nauseosa graveolens, E. nauseosa Includes var.

var. var.

orus orus

var. var. var. var. ora orus rmed) taxa of Cedar Breaks National Monument, cont. lanceolatus viscidifl markaguntensis Cirsium scariosum Chrysothamnus nauseosus leiospermus clavatum Cirsium clavatum Chrysothamnus vaseyi Chrysothamnus viscidifl var. Chrysothamnus viscidifl var. Cirsium clavatum Cichorium intybus Cirsium arvense Balsamorhiza sagittata Brickellia grandifl Chrysothamnus nauseosus graveolens Conyza canadensis glabrata scariosum Cirsium vulgare Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi

58 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wilderness Area (Fertig 22736) Wilderness Area (Fertig 23370) Canyon; Iron Co., UT (Garrett 6051 US) Wilderness Area (Fertig & Reynolds 23452) Range Comments Life form ower PerF Wide Arizona hymenoxys PerF Wide gray hawksbeard PerFsilver daisy Widespreading daisy Occurs in Ashdown Gorge daisyEaton’s PerF PerF Wide Wide Occurs in Ashdown Gorge PerF Wide Hopi blanketfl curly-cup gumweed PerFglabrate goldenweed PerF Wide Wide thrift goldenweedspindly goldenbush PerF Shrub Wide RegEn locality is Cedar Type Great Basin hyalineherb PerF Wide lanate hymenoxys PerF Wide Colorado rubber-plant PerF Widepoverty-weed Occurs in Ashdown Gorge blue lettuce PerF Wide PerF Wide hoary aster PerF Wide

var. arizonica var. var. lanata, Tetraneuris lanata, Tetraneuris caespitosa, T. caespitosa, T. var. var. H. acaulis Stenotus armerioides armerioides Hesperodoria scopulorum Tetraneuris acaulis Tetraneuris acaulis brevifolia Picradenia richardsonii richardsonii Aster canescens

var. var. var.

var. var.

var. var. pulchella L. pulchella, oblongifolia var. var. var. da lifolius var. rmed) taxa of Cedar Breaks National Monument, cont. canescens Hymenoxys acaulis caespitosa Crepis intermediaErigeron argentatus Psilochenia intermedia Erigeron divergens divergens Erigeron eatonii Gaillardia pinnatifi Grindelia squarrosa serrulata Haplopappus acaulis glabratus Haplopappus armerioides armerioides Haplopappus scopulorum hirtellus Hymenopappus fi nanus Hymenoxys acaulis arizonica Hymenoxys richardsonii richardsonii Iva axillaris Lactuca tatarica Machaeranthera canescens var. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 59 in Ashdown Gorge Wilderness Area (Fertig 23362) in Ashdown Gorge Wilderness Area (Fertig 23391) Range Comments Life form PerF Wide ixweed AnnF Intro Native to Europe. Occurs western tansy-mustard AnnFfl Wide bracteate astermountain tarweeed PerF AnnF RegEn Wide western tansy-mustard AnnF Wide two-leaf groundsel PerF RegEn blue mustardCalifornia tansy- mustard AnnF Intro Native to Asia coast goldenrodspineless horsebrush PerF Shrubstemless townsendia Wide Wide PerFredpod stonecrop Wide rockcressHolboell’s PerF PerF Wide shepherd’s-purse Wide AnnF Intro Native to Europe. Occurs

var. (not

var. UT var. according to , ) paysonii halictorum includes vars. M. canescens Name may be ssp. D. pinnata by some authors by some authors simplex & dimorphophylla; D. pinnata Included in Packera dimorphophylla var. canescens material is var. material is var. Cronquist (1994) nana Included in osmiarum rhodantha Boechera holboellii pinetorum. misapplied, according to Holmgren et al. (2005) dimorphphyllus

var.

var. var.

var. rmed) taxa of Cedar Breaks National Monument, cont. monticola osmiarum Descurainia sophia Machaeranthera canescens var. Madia glomerata Senecio dimorphophyllus dimorphophyllus Descurainia pinnata Solidago simplex S. spathulata; Chorispora tenella Descurainia californica Descurainia pinnata intermedia Tetradymia canescens Tetradymia exscapa Townsendia Sedum rhodanthumArabis holboellii Rhodiola rhodantha, Clementsia Capsella bursa-pastoris pinetorum Cruciferae (Brassicaceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Crassulaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix B. Potential (unconfi

60 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wilderness Area (Fertig 23365) City; Iron Co., UT (Parry 16 GH) Range Comments PerF LocEn locality is near Cedar Type PerF Wide Life form ower PerF Wide Cedar Canyon pepperwort prairie pepperwort AnnF Wide Occurs in Ashdown Gorge clasping pepperwort AnnFChambers’ twinpod Intro bladderpodWatson’s PerF Native to Europe PerF Wide bladderpodKing’s Wide PerF Wide fern-leaf false candytuft heart-leaf jewelfl beautiful sedgehair sedgeDouglas’ sedge PerGebony sedge Wide PerG PerG Wide PerG Wide Wide golden drabasnowbed whitlow-grass PerF PerFwedgeleaf draba Wide Wide AnnF Wide Nebraska sedgedunhead sedgevalley sedge PerG spikerushParish’s PerGbeaked spikerush Wide Wide PerG PerG PerG Wide Wide Wide

Holmgren et with var. with var. P. wardii P. var. cordatus var. include

orum macrocarpum intermedia heterophyllum L. integrifolium var. al. (2005) Welsh et al. 2003 combine Welsh var. densifl

var. var. var.

var. parvifolia Lesquerella kingii; var. orum var. rmed) taxa of Cedar Breaks National Monument, cont. orum Lepidium montanum heterophyllum Lepidium perfoliatum Physaria chambersii chambersii Physaria intermediaPhysaria kingii Smelowskia calycina Lesquerella intermedia, L. alpina americana Streptanthus cordatus Carex bella Carex capillaris S. cordatus Carex douglasii Carex ebenea Draba aurea Draba crassifolia Draba cuneifolia cuneifolia Lepidium densifl densifl Carex nebrascensis Carex phaeocephala Carex vallicola Eleocharis parishii Eleocharis rostellata Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Cyperaceae Cyperaceae Cyperaceae Appendix B. Potential (unconfi FamilyCruciferae (Brassicaceae) Species name notes Synonyms/Taxonomic Common name Cyperaceae

Chapter 6: Appendices 61 Wilderness Area (Fertig 23388) Wilderness Area (Fertig 23364) Wilderness Area (Fertig observation, 2007) Range Comments PerF Wide Occurs in Ashdown Gorge Life form ower eld mint eld PerF Wide lanceleaf phaceliaIdaho blue-eyed grass PerF rushParry’s PerF Wide nettle-leaf horsemint Wide PerF Wide PerG Occurs in Ashdown Gorge Wide fi smallfl dragonhead mountain muhlyfalse buffalograssbog bluegrass PerGwhite scorpion-weed AnnG Wide AnnF Wide PerG Wide Wide motherwortcommon horehound PerF PerF Intro Intro Native to Eurasia Native to Asia three-square bulrush PerG Wide spike bentgrassmarsh foxtailpurple reedgrass PerGRussian wildryemeadow fescue Wide PerG PerGravine fescue Wide PerG Intronorthern mannagrass PerG Intro Native to Eurasia foxtail barley PerG Intro Native to Russia Wide PerG Native to Eurasia Sparse PerG Wide Occurs in Ashdown Gorge milkvetchMartin’s PerF Wide

var. polyphyllus, alba var. occidentale ora var. canadensis alpina hastata, var. polyphyllus liculmis & var. var. M. fi Includes vars. leucophylla, Scirpus pungens Schoenoplectus pungens longispicatus, Schoenoplectus pungens pratensis Includes

var. orumparvifl Moldavica glabrata M. arvensis

var. var. rmed) taxa of Cedar Breaks National Monument, cont. Sisyrinchium idahoense Juncus parryi Agastache urticifolia S. idahoense Dracocephalum parvifl Munroa squarrosaPoa leptocoma Phacelia albaPhacelia hastata Monroa squarrosa neomexicana P. Leonurus cardiaca Marrubium vulgare Mentha arvensis Scirpus pungens longispicatus Agrostis exarata Alopecurus geniculatus Calamagrostis purpurascens Elymus junceusFestuca pratensisFestuca sororia Glyceria borealis Hordeum jubatum Psathyrostachys juncea Lolium pratense, Schedonorus Muhlenbergia montana Critesion jubatum Astragalus argophyllus martinii Iridaceae Juncaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Gramineae (Poaceae) Gramineae (Poaceae) Hydrophyllaceae Hydrophyllaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyCyperaceae Species name notes Synonyms/Taxonomic Common name Leguminosae (Fabaceae) Appendix B. Potential (unconfi

62 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument eld Co., UT Kane Co., UT (Jones s.n. POM) Wilderness Area (Fertig 23387) Canyon; Iron Co., UT (Parry 51 GH) Piute Co., UT (Jones 5355 NY) UT (Siler s.n. GH) in Ashdown Gorge Wilderness Area (Fertig 23384) Canyon, Garfi (Ripley & Barneby 8550 CAS) Range Comments Life form PerF Wide locality is Cedar Type PerF Wide scarlet globe-mallow PerFdandelion camissonia Wide PerF Wide Johnson milkvetch PerF Wide locality is Johnson, Type browse milkvetch PerF Wide groundcover milkvetch PerFgreat bladdery RegEn milkvetch white checker-mallow PerF Wide Occurs in Ashdown Gorge Sevier milkvetch PerF RegEn locality is Marysvale, Type sink lupine PerF RegEn locality is Kane Co., montane golden pea Type PerFwhite clover Wide PerF Intro Native to Europe. Occurs rough-fruited fairy- bells wild hollyhock PerF Wide nodding locoweedJones’ locoweed PerF Wide PerF RegEn locality is Red Type

& by . Vars. . Vars. glabrata rivularis T. & & ora ovata bisulcatus includes diversa candida coccinea, elata, megacarpus var. Vars. not recognized Vars. montana; Included in var. dissecta Barneby (1989) Includes var. Includes var. not recognized by Barneby (1989) Includes ssp. by Barneby (1989) Includes vars. Includes vars. var. rhombifolia

var. var. sericea var. var. barbiger L. barbiger. var. orabrevifl Oenothera var. exa rmed) taxa of Cedar Breaks National Monument, cont. major Camissonia brevifl Astragalus bisulcatus Astragalus cibarius Astragalus humistratus humivagans Astragalus megacarpus parryi Sphaeralcea coccinea Astragalus piutensisLupinus sericeus A. marianus Trifolium repens Trifolium Oxytropis defl Disporum trachycarpumIliamna rivularis Prosartes trachycarpa Sidalcea candida Thermopsis montana rhombifolia pinetorum, T. T. Oxytropis oreophila jonesii Onagraceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconfi FamilyLeguminosae (Fabaceae) Species name notes Synonyms/Taxonomic Common name Leguminosae (Fabaceae) Malvaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Calochortaceae) Malvaceae Malvaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Chapter 6: Appendices 63 . ed specimen of Wilderness Area (Fertig 23385) Co., UT Mtns., Washington (Fremont 447 US) Canyon; Iron Co., UT (Jones 5208c POM) Valley, Wayne Co., UT Wayne Valley, 526 US) (Ward herbarium was based on a misidentifi edulis Valeriana Range Comments Life form AnnF Wide PerF Wide PerF WidePerF locality is Rabbit Type Wide PerF Wide One report from CEBR long-leaf phloxcushion milkwort PerF PerF Wide Wide nodding wild buckwheat autumn willow-herb AnnFRocky Mountain Widewillow-herb Occurs in Ashdown Gorge Jones’ evening- primrose white bog- Lindley’s orchid clustered broomrape PerFscarlet gilia Wide scarlet gilia PerF Wide PerFdaggerleaf phlox Wide PerF RegEn locality is Cedar Type winged wild buckwheat Great Basin navarretia AnnFdesert phlox Sparse PerF Wide locality is Beaver Dam Type naked broomrape PerF Sparse

var.

& var. arizonica paniculatum propinqua var. var. occidentalis, orum alatum, var. longifolia var. var. ora stansburyi Platanthera dilatata leucostachys leucostachys, P. I. tenuituba, Gilia aggregata macrosiphon Includes vars. Pterogonum alatum Aphyllon unifl Gilia aggregata

var. var. var. var. var. var. ora unifl O. rmed) taxa of Cedar Breaks National Monument, cont. Polygala subspinosa Eriogonum alatum E. alatum Eriogonum cernuum cernuum Epilobium brachycarpum E. paniculatum Epilobium saximontanum Oenothera caespitosa crinita leucostachys Orobanche fasciculataOrobanche unifl Aphyllon fasciculatum Ipomopsis aggregata macrosiphon Navarretia intertexta N. intertexta Phlox gladiformis Phlox longifolia Phlox austromontana austromontana Habenaria dilatata Ipomopsis aggregata arizonica Polygalaceae Polygonaceae Polygonaceae Onagraceae Onagraceae Orobanchaceae Orobanchaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae FamilyOnagraceae Species name notes Synonyms/Taxonomic Common name Polemoniaceae Orchidaceae Polemoniaceae Appendix B. Potential (unconfi

64 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Flora of (2005). Washington Co., UT (Jones Washington 6091 POM). Considered a local endemic restricted to the Shivwits area of Co. in Washington North America in Ashdown Gorge Wilderness Area (Fertig 23371) Range Comments Life form PerF Wide Fern Wide PerF Wide owered fame- owered ower redroot wild buckwheat golden wild buckwheat Shrub RegEn locality is St. George, Type arid wild buckwheat PerFcurly dock Wide PerF Intro Native to Eurasia. Occurs holly-fern Fern Wide creeping buttercup PerF Wide marsh buttercup PerF Wide small-fl fl grass pondweedwoodnymph PerF larkspurNuttall’s Wide heart-leaf buttercup PerF PerF PerF Wide Wide Wide Rocky Mountain woodsia bracken fern Fern Wide Ssp.

Flora of & orus, P. P. orus, liformis, cymbalaria. scopulina. lanuginosum fi reptans, R. 1997 var. maximus ssp. var. glutinosum var. var. orus ora ammula ammula E. racemosum Includes var. Includes var. R. fl R. fl reptans Vars. not recognized in Vars. North America confertifl gramineus not recognized in Welsh et al. not recognized in Welsh (2003) aquilinum P. Includes vars.

var.

var. var. var. var. orumparvifl Phemeranthus oraunifl P. ammula rmed) taxa of Cedar Breaks National Monument, cont. Eriogonum racemosum racemosum Eriogonum umbellatum subaridum Rumex crispus Eriogonum corymbosum aureum Polystichum lonchitis scopulinaWoodsia scopulina W. ovalis Ranunculus fl Moneses unifl Delphinium nuttallianumRanunculus cardiophyllus D. nelsonii Ranunculus cymbalaria R. cymbalaria Pteridium aquilinum pubescens parvifl Talinum Potamogeton gramineus Polygonaceae Polygonaceae Polygonaceae Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae) Appendix B. Potential (unconfi FamilyPolygonaceae Species name notes Synonyms/Taxonomic Common name Ranunculaceae Pyrolaceae (Ericaceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Polypodiaceae (Densstaedtiaceae) Portulacaceae Potamogetonaceae

Chapter 6: Appendices 65 Wilderness Area (Fertig 23374) Peak, Piute Co., UT (Rydberg & Carlton 7369 NY) Wilderness Area (Fertig 23376) Range Comments Life form western chokecherry Shrub Wide Occurs in Ashdown Gorge creeping sibbaldiacleavers PerFtwin-leaf bedstraw Wide AnnF Wide AnnF Wide glabrate cinquefoilPennsylvania cinquefoil PerF PerF Wide Wide shrubby bedstraw PerF Wide alpine avensrock spiraeasilverweednear cinquefoil PerF PerF Wide PerF cinquefoilhair-tuft PerF Wide RegEn Wide PerF locality is Mt. Belknap Type Wide small bedstrawmilky kelloggiaplaneleaf willow PerF Wide PerF Shrub Wide Wide blister buttercupbur buttercup AnnF mountain-lilacFendler’s Wide ShrubSaskatoon serviceberry Wide AnnF Shrub Intro Widewoodland strawberry Occurs in Ashdown Gorge Native to Eurasia PerF Wide turbinata

ssp. orum ssp. includes vars. nuttallii, P. gracilis nuttallii, P. columbianum, subbifl . Vars. not recognized . Vars. var. planifolia & & glabrata dum melanocarpa G. spurium P. gracilis P. var. Padus virginiana P. proxima P. by Holmgren et al. (1997) Includes ssp. monica Hecatonia scelerata testiculata trifi

var.

var. var.

bracteata vesca F. var. var. var. turbinatum Acomastylis rossii var. orum var. dum rmed) taxa of Cedar Breaks National Monument, cont. orum dus Sibbaldia procumbens Galium aparine echinospermon Galium bifolium Galium multifl Potentilla pensylvanica Prunus virginiana melanocarpa fastigiata multifl Galium trifi Petrophyton caespitosumPotentilla anserina Spiraea caespitosa Potentilla concinna proxima Argentina anserina Potentilla crinita Potentilla gracilis Kelloggia galioides Salix planifolia S. phylicifolia; Ranunculus sceleratus multifi Ranunculus testiculatusCeanothus fendleri Ceratocephala orthoceras, C. Amelanchier alnifolia Fragaria vesca Geum rossii Rosaceae Rubiaceae Rubiaceae Rubiaceae Rosaceae Rosaceae Rubiaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rubiaceae Salicaceae Ranunculaceae Rhamnaceae Rosaceae Rosaceae Rosaceae FamilyRanunculaceae Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconfi

66 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument eld Co., UT Kane Co., UT (Siler s.n. GH) Wilderness Area (Fertig observation, 2007) Wilderness Area (Fertig & Reynolds 23357). Type locality is near Beaver, Mtns., Beaver Co., Tushar UT (Palmer s.n. GH) Plateau, Garfi 462 GH) (Ward Juab Co., UT (Jones s.n. GH) Range Comments Life form ax PerF Wide recracker PerF Wide Siler’s penstemonSiler’s penstemonPalmer’s PerF PerF RegEn Wide locality is Osmer, Type Eastwood’s paintbrushEastwood’s PerFblue-eyed Mary RegEn AnnF Wide Occurs in Ashdown Gorge King’s bird’s-beakKing’s dwarf lousewort penstemonTushar AnnF RegEn PerF PerF Wide LocEn Occurs in Ashdown Gorge dusty penstemon PerFmistaken penstemon PerF RegEn locality is Aquarius Type fi Eaton’s RegEn locality is Detroit, Type bastard toadfl pygmy saxifragewhitestem gooseberry PerF Shrubdesert paintbrush Wide Wide PerF Wide hairy alumrootslender woodlandstar PerF PerF Wide Wide debilis, var. dubia, C. debilis var. martinii ssp. Intermountain Vars. not Vars. ssp. (1997) P. tusharensis P. H. versicolor. H. versicolor. recognized in Flora S. hyperborea applegatei

kingii

var. var. eatonii var.

var. var. var. var. var. var. ora rmed) taxa of Cedar Breaks National Monument, cont. Penstemon linarioides sileri Penstemon palmeri palmeri scabrida Collinsia parvifl Cordylanthus kingii Pedicularis centranthera Penstemon caespitosus suffruticosus Penstemon comarrhenus Penstemon confusus Penstemon eatonii Comandra umbellata pallida Ribes inerme Castilleja chromosaCastilleja scabrida C. angustifolia Heuchera rubescens Lithophragma tenellumSaxifraga rivularis L. tenella S. debilis, rivularis versicolor Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Saxifragaceae (Grossulariaceae) Scrophulariaceae Scrophulariaceae Appendix B. Potential (unconfi FamilySantalaceae Species name notes Synonyms/Taxonomic Common name Saxifragaceae Saxifragaceae Saxifragaceae

Chapter 6: Appendices 67 (Albee rmed for the monument. Species Utah” (Parry 85) Beaver Co., UT (Jones 1864 US) in Ashdown Gorge Wilderness Area (Fertig, 2007 observation) Atlas of the Vascular Flora of Utah Atlas of the Vascular Range Comments PerF RegEn Life form ere. University in 2001 or from the University in 2001 or from have not previously been reported or confi been reported have not previously ower penstemon ower PerF RegEn stinging nettlehook-spur violet PerF PerF Wide Wide Fendler’s ground-cherryFendler’s PerF Wide ground-cherryPalmer’s PerFnarrowleaved burr-reed RegEn PerFColorado Plateau spring-parsley Wide rough lomatiumwestern sweet-cicely PerF PerF RegEn locality is Frisco, Type Wide Nevada lomatium lomatiumParry’s PerF Wide PerF Wide locality is “Southern Type smallfl woolly mulleintwo-lobe speedwell AnnF PerF Intropurslane speedwell Intro Native to Asia. Occurs coyote tobacco Native to Eurasia AnnF Wide AnnF Wide

var. var. palmeri var. holosericea, U. holosericea ssp. ssp. fendleri hederaefolia P. C. purpureus gracilis cordifolia, P. hederaefolia cordifolia, P. P. fendleri, P. hederifolia fendleri, P. P.

var.

var. var. var. var. var. var. occidentalis U. dioica var. rmed) taxa of Cedar Breaks National Monument, cont. Viola adunca Physalis hederifolia palmeri Sparganium angustifolium Cymopterus purpureus purpureus Lomatium nevadense Lomatium scabrum scabrum Osmorhiza occidentalis Urtica dioica parishii Lomatium parryi Penstemon procerus aberrans thapsus Verbascum bilobaVeronica peregrina Veronica xalapensis Pocilla biloba Nicotiana attenuata Physalis hederifolia fendleri Violaceae Solanaceae Sparganiaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Urticaceae Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae FamilyScrophulariaceae Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconfi The vascular plant species on this list are known from the vicinity of Cedar Breaks National Monument in Iron County, Utah, but County, National Monument in Iron the vicinity of Cedar Breaks known from The vascular plant species on this list are N. Duane Atwood of Brigham Young by Dr. a checklist of potential species for NCPN parks prepared derived from on this list are et al. 1988). Some species from Appendix C could be considered as potentially occurring in the monument, but are not included h as potentially occurring in the monument, but are Appendix C could be considered et al. 1988). Some species from

68 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument

A. A. D. E. M. C. C. E. . USFS S. E. speciosus

Vars. not Vars. . ed specimens of and callianthemus. ed specimen of ed specimen of ed specimen of ed specimen of ed specimen of nana. ursinus wheeleri. ed specimens of ed specimens of ed specimens of ed specimens of wheeleri. . ursinus var. var. . Not native to Utah. var. var. var. ed specimen of var. . var. var. var. var. . mollis misidentifi jamesiana on a misidentifi pulcherrima on a misidentifi mollis on a misidentifi Machaeranthera commixta Region 4: Sensitive, USFWS: former C2 candidate on a misidentifi stenoloba on misidentifi arizonica on misidentifi wheeleri on misidentifi wheeleri Comments recognized in Intermountain Flora, 1994. based on misidentifi E. ursinus on misidentifi ursinus var. var. on a misidentifi peregrinus Park status Range Life form boreal chickweed PerFpearly-everlasting Wide PerF FalsRepbroadleaf arnica Report from CEBR based on a Wide FalsRep PerF Report from CEBR herbarium based Wide FalsRep Report from CEBR herbarium based King’s asterKing’s PerF LocEn FalsRep Report from CEBR herbarium based mountain draba AnnF RegEn FalsRep Report from CEBR herbarium based Common name streamside bluebell PerF Wide FalsRep Reports from CEBR herbarium based Caineville thistle PerF LocEn FalsRep Reports from CEBR herbarium based pretty thistle PerF Wide FalsRep Reports from CEBR herbarium based Engelmann’s daisyEngelmann’s PerF Wide FalsRep Reports from CEBR herbarium are beautiful daisy PerF Wide FalsRep Reports from CEBR herbarium based smooth daisy PerF Wide FalsRep Report from CEBR herbarium based var. var. glabellus

2006) 2006) Flora of Flora of var. ( ( var. var. C. C. Includes var. Includes var. kingii. barnebyana, Tonestus barnebyana, Tonestus kingii Synonyms/ notes Taxonomic Included in arizonicum bipinnatum North America Included in arizonicum bipinnatum North America E. formosissimus formosissimus

var. calcareum pulchellum Stellaria borealis Anaphalis margaritacea Arnica latifolia latifolia Aster kingii Machaeranthera Cirsium calcareum var. Draba rectifructa Cirsium calcareum var. Mertensia ciliata Erigeron engelmannii Erigeron formosissimus Erigeron glabellus E. glabellus Caryophyllaceae (Alsinaceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Brassicaceae Compositae (Asteraceae) FamilyBoraginaceae Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) vascular plant taxa of Cedar Breaks National Monument. or questionable) vascular plant taxa of Cedar Breaks Appendix C. Rejected (falsely reported

Chapter 6: Appendices 69 C. C. S. D. C. R. H. in CEBR USFS Region . Not native to ed specimens of ed specimens ed cinereus stevenii ed specimen of ed specimen of ed specimen of ed specimen of ed specimen of ed specimen of scirpoidea. var. var. curvipes zionensis. in CEBR herbarium. . var. var. var. var. ed specimens of . var. var. var. var. lifolius H. fi on a misidentifi occidentalis. on a misidentifi scirpoidea herbarium. CEBR is well outside the known range of this species. Comments based on misidentifi H. clementis and Buchanan (1992) based on misidentifi on a misidentifi malmstenii on a misidentifi asprella 4: Sensitive, USFWS: former C2 candidate Utah. on a misidentifi norevegica clementis based on misidentifi of on a misidentifi curvipes Park status Range Life form PerF RegEn FalsRep Not native to Utah; falsely reported PerF Wide FalsRep Not native to Utah; falsely reported PerF Wide FalsRep Report from CEBR herbarium based Common name Hood’s sedgeHood’s spike-rush sedge PerG Wide PerG FalsRep Wide Report from CEBR herbarium based FalsRep Report from CEBR herbarium based manystemmed goldenweed thickleaf groundsel PerF Wide Ques? Reported by Buchanan (1992), but Chalk Hills hyalineherb meadow goldenweed PerF Wide FalsRep Report from CEBR herbarium montane groundsel Per F Wide FalsRep Report from CEBR herbarium based creeping draba PerFsouthern marsh LocEnyellowcress FalsRep Report from CEBR herbarium based white and black sedge PerG Wide FalsRep Report from CEBR herbarium based Flora (2006) Vars. Vars. werneriifolia, S. Synonyms/ notes Taxonomic Pyrrocoma integrifolia Pyrrocoma lanceolata Packera werneriifolia var. werneriaefolius. not recognized in of North America

var.

var. Haplopappus integrifolius Carex hoodii Carex scirpoidea pseudoscirpoidea Haplopappus lanceolatus Senecio crassulus Senecio werneriifolius werneriifolius Hymenopappus tenuifolius Draba sobolifera Rorippa teres Radicula obtusa Carex albonigra Cyperaceae FamilyCompositae (Asteraceae) Species name Cyperaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Appendix C. Rejected (falsely reported or questionable) taxa of Cedar Breaks National Monument, cont..

70 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Occurs L. E. E. S. P. E. in CEBR . . ed specimens of ed specimens of ed specimens of hornemannii. alpestre alpestre . ed specimen of . Utah state noxious rubricaulis. ed specimen of ed specimen of ed specimen of ed specimens of var. var. oblongifolius var. var. var. var. Type locality is Panguitch Type . eld Co., UT (Jones 6023 var. var. ed specimen of var. var. attributed to Utah and CEBR based on misidentifi panguicense on misidentifi lettermanii. Lake, Garfi POM) based on misidentifi A. miser herbarium. in Ashdown Gorge Wilderness Area (Fertig 23389) misidentifi panguicense Comments based on misidentifi G. heterosepala based on misidentifi E. trachycaulus weed. Native to Eurasia on misidentifi compressa on a misidentifi argenteus on misidentifi hornemannii herbarium is well beyond known range and probably erroneous. Park status Range Life form PerF Wide FalsRep Report from CEBR based on Common name buckwheat Colorado buckwheat PerF LocEn FalsRep Endemic to Colorado; falsely pine needlegrass PerG Wide FalsRep Report from CEBR herbarium based halfmoon milkvetch PerF Wide FalsRep Not native to Utah, falsely reported quackgrass PerG Intro FalsRep Reports from CEBR herbarium are northern gentian PerF Wide FalsRep Report from Roberts and Jean (1989) Patterson’s bluegrassPatterson’s PerG Sparse FalsRep Report from CEBR herbarium based sand lupine PerF RegEn FalsRep Report from CEBR herbarium based northern willow-herb PerF Wide FalsRep Reports from CEBR herbarium based dwarf wild buckwheat PerF LocEn Ques? Historical report for CEBR from NYBG matted wild

ssp. amarella, var. var. acuta, Gentianella Synonyms/ notes Taxonomic Gentiana amarella, Gentianella amarella ssp. amarella Elytrigia repens Gentianella acuta pattersonii L. ammophilus ciliatum, E. watsonii, adenocaulon E. grayi

var. ammophilus Gentianella amarella Eriogonum coloradense Elymus repens Agropyron repens, Stipa pinetorumpinetorum Achnatherum Astragalus allochrous Lupinus polyphyllus var. Epilobium ciliatum E. ciliatum Eriogonum brevicaule nanum Poa pattersonii abbreviata P. Eriogonum caespitosum Appendix C. Rejected (falsely reported or questionable) taxa of Cedar Breaks National Monument, cont.. FamilyGentianaceae Species name Polygonaceae Gramineae (Poaceae) Gramineae (Poaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Onagraceae Polygonaceae Gramineae (Poaceae) Polygonaceae

Chapter 6: Appendices 71 n- C. P. P. P. Geum S. Type Type Aconitum R. ed specimens of perincisum. columbianum. leiophyllus. leiophyllus. leiophyllus. ed specimen of ed specimen of ed specimen of ed specimen of ed specimen of . var. var. var. ed specimen of . var. var. var. var. var. ed specimens of ed specimens of . leiophyllus and Buchanan (1992) based on misidentifi on a misidentifi miniata on a misidentifi leiophyllus on a misidentifi leiophyllus Comments columbianum on misidentifi boothii. Co., locality is Springdale, Washington UT (Jones 5250 in part US). on a misidentifi based on misidentifi S. brachycarpa and Buchanan (1992) based on misidentifi on a misidentifi macrophyllum inamoenus Park status terature (Buchanan 1992; Roberts and Jean 1989) or from herbarium (Buchanan 1992; Roberts and Jean 1989) or from terature Range Life form PerF Wide FalsRep Reports from CEBR herbarium Common name western larkspur PerF Wide FalsRep Report from CEBR herbarium Raft River penstemon PerF RegEn FalsRep Report from CEBR herbarium based western paintbrush PerF Periph FalsRep Report from CEBR herbarium based smooth penstemon PerF RegEn FalsRep Report from CEBR herbarium based Eschscholtz’s Eschscholtz’s buttercup Bebb’s willowBebb’s Shrub Wide FalsRep Reports from CEBR herbarium based showy penstemon PerF Periph FalsRep Report from CEBR herbarium based gray-leaf willow Shrub Wide FalsRep Reports from CEBR herbarium are tall cinquefoil PerF Wide FalsRep Report from CEBR herbarium is based cations (False Reports or FalsRep) because the monument is well outside known expected range of species (Questio D. barbeyi, arguta x Synonyms/ notes Taxonomic D. glaucum D. glaucum P. holmgrenii P. Drymocallis arguta; includes ssp.

var. var. var. convallaria Delphinium occidentale Penstemon cyananthus subglaber Castilleja occidentalis Penstemon laevis Ranunculus eschscholtzii eschscholtzii Salix bebbiana Salix glauca Penstemon speciosus villosa Potentilla arguta var. FamilyRanunculaceae (Helleboraceae) Species name Scrophulariaceae Scrophulariaceae Scrophulariaceae Ranunculaceae Salicaceae Salicaceae Scrophulariaceae Rosaceae Appendix C. Rejected (falsely reported or questionable) taxa of Cedar Breaks National Monument, cont.. able or Ques?). The vascular plant species on this list have been reported for Cedar Breaks National Monument from published and unpublished li National Monument from for Cedar Breaks The vascular plant species on this list have been reported because of misidentifi now believed to be erroneous specimens, but are

72 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status Native Pres Apr–Jun X r Native Pres NA X r Native Pres NA X r Native Pres NA X silver sagebrush Native Pres Aug–Sep X X X mountain-lover Native Pres May–Aug X X mountain snowberry Native Pres Jun–Aug X X red elderberry Native Pres Jun–Jul X X X blue elderberry Native Pres May–Jul X black twinberry Native Pres May–Jul X X Oregon-grape Native Pres Apr–Jun X X X Rocky Mountain maple Native Pres Apr–Jun X X lance-leaf cottonwood Native Pres Apr–Jun X quaking aspen Native Pres Mar–Jul X X X Douglas-fi narrow-leaved cottonwood ponderosa pine Native Pres NA X western bristlecone pine Native Pres NA X X limber pine Native Pres NA X X two-needle pinyon Native Rep NA X blue spruce Native Pres NA X subalpine fi Engelmann spruce Native Pres NA X white fi Gambel oak Native Pres Apr–Jun X X Rocky Mountain juniper Native Pres NA X X water birch Native Pres Apr–May X .

var

var.

var. glabrum var. var. var. intercurrens x exilis utahensis Artemisia cana viscidula Pachystima myrsinites Symphoricarpos oreophilus var. Sambucus racemosa microbotrys Sambucus caerulea Lonicera involucrata Mahonia repens Acer glabrum Populus Populus tremuloides Pseudotsuga menziesii glauca Populus angustifolia Pinus ponderosa scopulorum Pinus longaeva Pinus fl Pinus edulis Picea pungens Picea engelmannii Abies lasiocarpa Abies concolor Quercus gambelii gambelii Juniperus scopulorum Betula occidentalis National Monument organized by life form with ecological vascular plant taxa of Cedar Breaks rmed, historical, and reported Compositae (Asteraceae) Celastraceae Caprifoliaceae Caprifoliaceae (Adoxaceae) Caprifoliaceae (Adoxaceae) Caprifoliaceae Berberidaceae Shrubs Aceraceae Salicaceae Salicaceae Pinaceae Salicaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Fagaceae Cupressaceae FamilyTrees Betulaceae Species name Common name Nativity Appendix D. Confi notes.

Chapter 6: Appendices 73 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status Native Pres May–Jun X Native Pres May–Jun X X Rocky sites Scouler’s willowScouler’s Native Pres Apr–Jun X coyote willow Native Pres Apr–Aug X short-fruit willow Native Pres Jun–Jul X Booth’s willowBooth’s Native Pres May–Jun X Arizona willow Native Pres Jun–Jul X common red raspberry Native Pres Jun–Aug X X Woods’ roseWoods’ Native Pres May–Jul X X X X Nootka rose Native Rep Jun–Jul X antelope bitterbrush Native Pres Apr–Jul X shrubby cinquefoil Native Pres Jun–Aug X X X alder-leaf mountain alder-leaf mahogany curl-leaf mountain mahogany Utah serviceberry Native Pres Apr–Jun X X Utah mountain-lilac Native Pres Jun–Jul X greenleaf manzanita Native Pres Mar–Jul X Canada buffaloberry Native Pres May–Jul X X common juniper Native Pres NA X X X Cedar Breaks goldenbush Nativered-osier dogwood Pres Jul–Aug Native Pres May–Jul Claron barrens X narrowleaf goldenweed Native Pres Jul–Sep X X X Barrens broom snakeweed Native Pres Jul–Oct X Nevada rabbitbrush Native Pres Jul–Sep X greenish rabbitbrush Native Pres Aug–Oct X

var. var. var. sericea hispida strigosus

var. var. var. var. . consimilis Salix scouleriana Salix exigua stenophylla Salix brachycarpa Salix boothii Salix arizonica Rubus idaeus Rosa woodsii Rosa nutkana Purshia tridentata Potentilla fruticosa Cercocarpus montanus Cercocarpus ledifolius intermontanus Amelanchier utahensis Ceanothus martinii Arctostaphylos patula Shepherdia canadensis Cornus sericea Juniperus communis depressa Haplopappus zionis Haplopappus macronema Gutierrezia sarothrae Chrysothamnus parryi nevadensis Chrysothamnus nauseosus var rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rhamnaceae Ericaceae Elaeagnaceae Cornaceae Cupressaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyShrubs, cont. Compositae (Asteraceae) Species name Common name Nativity Appendix D. Confi

74 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status y Native Pres May–Sep X X ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep X purplish agoseris Native Pres Jun–Aug X common yarrow Native Pres May–Sep X X X X long-stalk starwort Native Pres May–Aug X X James’ chickweed Native Pres Jun–Aug X X Peterson’s campionPeterson’s Native Pres Jun–Sep Claron barrens Menzies’ campion Native Pres Jun–Aug X X Drummond’s catchfl Drummond’s sprawling sandwort Native Hist May–Aug X basin sandwort Native Pres May–Aug X X X Eastwood’s sandwortEastwood’s Native Pres May–Sep X Parry’s harebellParry’s Native Hist Jun–Sep X aspen bluebell Native Pres Jun–Aug X X X southwestern stoneseed Native Pres Jun–Aug X X common dogbane Native Rep Jun–Aug X spreading dogbane Native Pres Jun–Aug X X fi sticky currant Native Pres May–Aug X X alpine prickly currant Native Pres Apr–Aug X X X X wax currant Native Pres Apr–Aug X X X Zion jamesia Native Pres May–Aug X Claron barrens

orum var. var. var. var. var. var. var. var. var. inebrians var. Agoseris aurantiaca purpurea Achillea millefolium lanulosa Stellaria longipes longipes Stellaria jamesiana Silene petersonii Silene menziesii Lychnis drummondii Lychnis Arenaria lanuginosa Arenaria fendleri glabrescens Arenaria fendleri eastwoodiae Campanula parryi Mertensia arizonica Lithospermum multifl Apocynum cannabinum Apocynum androsaemifolium androsaemifolium Tamarix chinensis Tamarix Ribes viscosissimum viscosissimum Ribes montigenum Ribes cereum Jamesia americana zionis rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Campanulaceae Boraginaceae Boraginaceae Apocynaceae Perennial Forbs Perennial Apocynaceae Tamaricaceae Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Appendix D. Confi FamilyShrubs, cont. Saxifragaceae (Hydrangeaceae) Species name Common name Nativity

Chapter 6: Appendices 75 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status c aster c Native Pres Jul–Oct X X elds agoseris elds Native Pres May–Sep X Welsh’s asterWelsh’s Native Pres Aug–Oct X Markagunt aster Native Pres Jul–Sep Claron barrens western aster Native Pres Jul–Sep X Parry’s asterParry’s Native Pres Jul–Sep X Canby’s asterCanby’s Native Pres Jul–Sep X X X Pacifi Louisiana wormwood Native Pres Jul–Oct X X tarragon Native Pres Jul–Oct X X rayless arnica Native Pres Jun–Aug X hairy arnica Native Pres Jul–Aug X X heartleaf arnica Native Pres May–Aug X X Breaks pussytoes Native Pres Jun–Aug X X showy pussytoes Native Pres Jul–Sep X X common pussytoes Native Pres May–Jul X X small-leaf pussyoes Native Pres Jun–Aug X short agoseris Native Pres May–Sep X X fi

ssp. var.

var. parryi canbyi var. var. var. var. var. Aster welshii Aster wasatchensis wasatchensis Aster spathulatus Aster foliaceus Aster foliaceus Aster ascendens Artemisia ludoviciana ludoviciana Artemisia dracunculus glauca Arnica parryi Arnica mollis Arnica cordifolia cordifolia Antennaria rosulata Antennaria pulcherrima Antennaria parvifolia Antennaria microphylla Agoseris glauca dasycephala Agoseris glauca agrestis rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Appendix D. Confi

76 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status eabane Native Pres Jul–Aug X X X ower Native Pres Jun–Aug X X common hyalineherb Native Pres May–Oct X X X orange sneezeweedlittle sunfl Native Pres Jun–Aug X X Clement’s goldenweedClement’s Native Pres Jun–Sep X X X Payson’s daisyPayson’s Native Pres Jun–Aug Claron barrens bear daisy Native Pres Jun–Aug X X hairy pretty daisy Native Pres Jul–Aug X aspen daisy Native Pres Jun–Aug X Cedar Breaks daisy Native Pres Apr–Sep X Claron Formation wandering fl trailing daisy Native Pres May–Aug X cut-leaved daisy Native Pres May–Aug X X X Wheeler’s thistleWheeler’s Native Pres Jul–Sep X X X Arizona thistle Native Pres Jun–Aug X X X Barrens hispid golden-aster Native Pres May–Sep X X X hoary dusty-maiden Native Pres May–Sep X alpine hyalineherb Native Pres May–Oct X

var.

var.

var. var. var. var. lifolius lifolius var. ora var. var. agellaris nudipes cinereus Hymenopappus fi var. Helianthella unifl var. Hymenopappus fi Helenium hoopesii Haplopappus clementis Erigeron vagus ursinus Erigeron ursinus Erigeron speciosus mollis Erigeron speciosus macranthus trilobatus Erigeron sionis Erigeron peregrinus callianthemus Erigeron fl Erigeron compositus glabratus Cirsium wheeleri wheeleri Cirsium arizonicum Chrysopsis villosa minor Chaenactis douglasii rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity

Chapter 6: Appendices 77 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status Native Pres Jul–Aug X X X ower Native Hist Jun–Sep X yellow salsify Intro Pres May–Jul X X Bryce Canyon townsendia Native Pres Apr–Jun Claron barrens common townsendia Native Pres Jun–Jul X three-nerve goldenrodcommon dandelion Native Pres Intro Aug–Oct Pres X Apr–Dec X X X X X Parry’s goldenweedParry’s Native Pres Jul–Sep X X Rocky Mountain goldenrod broom groundsel Native Hist Jun–Oct X basin groundsel Native Pres Apr–Aug X X X Podunk groundsel Native Pres Jun–Aug Claron barrens western groundsel Native Pres Jun–Aug X X Hart’s groundselHart’s Native Pres Jun–Jul X cut-leaved groundsel Native Pres Jul–Sep X X woolly groundsel Native Pres May–Aug X X black groundsel Native Pres Jul–Aug X Barrens montane conefl Bigelow’s asterBigelow’s Native Pres Jul–Aug X X X

var. var. var. var. var. cinale Tragopogon dubius Tragopogon Townsendia montana Townsendia minima Taraxacum offi Taraxacum leptotes Townsendia Solidago velutina Solidago parryi Solidago multiradiata scopulorum spartioides Senecio spartioides Senecio multilobatus Senecio malmstenii Senecio integerrimus exaltatus Senecio hartianus Senecio eremophilus kingii Senecio canus Senecio atratus Rudbeckia montana Machaeranthera commixta rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Appendix D. Confi

78 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status ower Native Pres Apr–Aug X X golden corydalis Native Pres Apr–Sep X shorthorn spurge Native Pres May–Jun X X X alpine pennycress Native Pres Apr–Aug X X smooth yellowcress Native Pres Jun–Sep X blunt-leaf yellowcress Native Pres May–Aug X Ward’s bladderpodWard’s Native Pres Jun–Jul X sand dune wallfl Breaks bladderpod Native Pres May–Jul X X Claron Formation Breaks draba Native Pres May–Aug X X slender whitlow-grass Native Pres Jun–Aug X X Zion draba Native Pres Apr–Jun X heartleaf bittercress Native Pres Jun–Sep X heartleaf bittercress Native Pres Jun–Sep X elkweed Native Pres Jun–Aug X X Fendler’s rockcressFendler’s Native Pres Apr–Aug X Engelmann’s gentianEngelmann’s Native Pres Jun–Aug X Drummond’s rockcressDrummond’s Native Pres May–Sep X X Parry’s gentianParry’s Native Pres Jul–Sep X showy goldeneye Native Pres Jun–Sep X X X Rocky Mountain gentian Native Pres Jul–Sep X X

nis

var. var. var.

nana

affi var.

var. var.

var. var. var. var. var. var. ora var. nis ora Corydalis aurea occidentalis Euphorbia brachycera montanum Thlaspi montanum Rorippa islandica glabra Rorippa curvipes curvipes Physaria wardii Physaria rubicundula Erysimum asperum purshii rubicundula Draba subalpina Draba stenoloba Draba asprella zionensis Cardamine cordifolia incana Cardamine cordifolia cordifolia Swertia radiata Arabis holboellii fendleri Gentianella heterosepala Arabis drummondii Viguiera multifl Gentiana affi Gentiana parryi multifl rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Fumariaceae Euphorbiaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Gentianaceae Cruciferae (Brassicaceae) Gentianaceae Cruciferae (Brassicaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Gentianaceae Gentianaceae

Chapter 6: Appendices 79 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status ower sweetpea ower Native Pres May–Aug X X X subalpine lupine Native Pres May–Aug X X silvery lupine Native Hist May–Aug X silvery lupine Native Pres May–Aug X X X Utah trefoil Native Pres May–Aug X largefl pulse milkvetch Native Pres May–Aug X Rydberg’s weedy milkvetchRydberg’s Native Pres Jun–Aug X X X great rushy milkvetch Native Pres May–Jul X Barrens Navajo Lake milkvetch Native Pres Jun–Sep X X Claron barrens mountain kentrophyta Native Rep Jun–Sep X tall kentrophyta Native Pres Jun–Sep X X X Barrens Panguitch milkvetch Native Pres May–Jul X X self-heal Native Pres May–Sep X stinking horsemint Native Pres May–Aug X X Claron barrens Rocky Mountain iris Native Pres May–Jul X wand phacelia Native Pres Jun–Aug X Richardson’s crane’s-billRichardson’s Native Pres Jun–Aug X X small-leaf geranium Native Pres Jun–Sep X stemless lupine Native Pres May–Aug X

var. var. var.

var.

var. var. var. var. var. var. tegetarius elatus Lupinus argenteus rubricaulis Lupinus argenteus argenteus Lupinus argenteus argentatus Lotus utahensis Lathyrus lanszwertii laetivirens Astragalus tenellus Astragalus miser oblongifolius Astragalus lonchocarpus Astragalus limnocharis limnocharis Astragalus kentrophyta var. Astragalus kentrophyta var. Astragalus argophyllus panguicensis lanceolata Prunella vulgaris Monardella odoratissima Iris missouriensis Phacelia heterophylla Geranium richardsonii Geranium caespitosum marginale Lupinus lepidus utahensis rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Iridaceae Hydrophyllaceae Geraniaceae FamilyPerennial Forbs, cont. Geraniaceae Species name Common name Nativity Appendix D. Confi

80 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status Native Pres Apr–Jun X ax Native Pres May–Aug X X ax Native Pres Apr–Aug X reweed Native Pres Jun–Sep X X Barrens white bog-orchid Native Pres Jun–Aug X giant rattlesnake-plantain Native Pres Jul–Aug X X spotted coral-root Native Pres Jun–Jul X long-tube evening- primrose yellow evening-primrose Native Pres Jun–Jul X X Hornemann’s willow-herbHornemann’s Native Pres Jun–Aug X Hall’s willow-herbHall’s Native Pres Jun–Aug X fi woodland pinedrops Native Pres Jun–Sep X X blue fl King’s yellow fl King’s mountain death camas Native Pres Jun–Aug X X X starry false Solomon’s-seal Native Pres Apr–Jun X X X leopard-lily Native Pres Apr–Jul X sego-lily Native Pres Jun–Jul X twincrest onion Native Hist May–Jul Barrens American vetch Native Pres May–Aug X Rusby’s cloverRusby’s Native Pres May–Aug X X X mountain locoweed Native Pres May–Aug X X X northern bog orchid Native Pres Jun–Aug X yellow sweet-clover Intro Pres May–Oct X

var.

ava fl var.

lewisii var. var. var. var. ssp. cinalis ava hornemannii canescens Habenaria dilatata dilatata Goodyera oblongifolia Oenothera fl Corallorhiza maculata Oenothera caespitosa marginata Epilobium hornemannii var. Epilobium halleanum Epilobium angustifolium var. Pterospora andromedea Linum perenne Linum kingii Zigadenus elegans Smilacina stellata Fritillaria atropurpurea Calochortus nuttallii Allium bisceptrum Vicia americana americana Trifolium longipes Trifolium rusbyi Melilotus offi Oxytropis oreophila oreophila Habenaria hyperborea rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Orchidaceae Orchidaceae Orchidaceae Onagraceae Orchidaceae Onagraceae Onagraceae Onagraceae Onagraceae Monotropaceae (Ericaceae) Linaceae Linaceae Liliaceae (Melanthiaceae) Liliaceae (Convallariaceae or Ruscaceae) Liliaceae Liliaceae (Calochortaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Leguminosae Species name(Fabaceae) Common name Nativity

Chapter 6: Appendices 81 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status Native Pres Jun–Aug X X Native Pres Jun–Sep X X Claron barrens owered green-fl wintergreen pink wintergreen Native Pres Jun–Aug X X Parry’s primroseParry’s Native Pres Jun–Sep X X dark-throat shooting-star Native Pres Mar–Aug X alpine shooting-star Native Hist Apr–Sep X alpine bitter-root Native Pres May–Aug X lanceleaf springbeauty Native Pres Apr–Jul X willow dock Native Pres May–Sep X western dock Native Pres May–Aug X alpine bistort Native Pres Jun–Sep X X American bistort Native Pres Jul–Sep X sulfur wild buckwheat Native Pres Jun–Sep X X Porter’s wild buckwheatPorter’s Native Pres Jul–Sep X X pretty Jacob’s-ladder NativeCedar Breaks wild buckwheat Pres Jun–Aug X X western Jacob’s-ladder Native Pres Jun–Aug X cushion phlox Native Pres Jun–Aug X Cedar Breaks gilia Native Pres Jun–Aug X Barrens scarlet gilia Native Pres May–Aug X Tweedy’s plantainTweedy’s Native Pres Jun–Aug X slender bog orchid Native Pres Jun–Jul X

var.

var. ssp.

ora var. ora alpestre pulchellum umbellatum porteri pterospermum Pyrola chlorantha Pyrola asarifolia Primula parryi Dodecatheon pulchellum var. Dodecatheon alpinum Lewisia pygmaea Claytonia lanceolata Rumex salicifolius triangulivalvis Rumex occidentalis Polygonum viviparum Polygonum bistortoides Eriogonum umbellatum var. Eriogonum panguicense var. Eriogonum umbellatum var. Polemonium pulcherrimum delicatum Polemonium caeruleum var. Phlox pulvinata Ipomopsis tridactyla Ipomopsis aggregata aggregata Plantago tweedyi Habenaria sparsifl sparsifl rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae) Primulaceae Primulaceae Primulaceae Portulacaceae Portulacaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Plantaginaceae FamilyPerennial Forbs, cont. Orchidaceae Species name Common name Nativity Appendix D. Confi

82 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status woolly cinquefoilsweet-scented bedstraw Native Native Pres Pres Jul–Aug Jul–Aug X X beautiful cinquefoil Native Pres Jun–Aug X wedgeleaf cinquefoil Native Pres Jun–Aug X X X Sevier ivesia Native Pres Jun–Aug Claron barrens large-leaved avensprairie smoke Native Pres May–Aug Native Pres Jun–Aug X X Virginia strawberry Native Pres Jun–Jul X X X Fendler’s meadow-rueFendler’s Native Pres Jun–Aug X X X drab buttercup Native Pres Jun–Sep X X plantain buttercup Native Pres May–Jul X X subalpine larkspur Native Pres Jul–Aug X X Columbian virgin’s-bower Native Pres May–Sep X X white marshmarigold Native Pres May–Jul X rock columbine Native Pres Jun–Aug Claron barrens Colorado columbine Native Pres Jun–Aug X cutleaf anemone Native Pres Jun–Aug X X X Columbian monkshood Native Pres Jun–Aug X X X sidebells pyrola Native Pres Jun–Aug X X whiteveined wintergreen Native Pres Jun–Aug X X

var. var. var.

var. da var. orum orum montanus columbianum Galium trifl Potentilla hippiana Potentilla gracilis pulcherrima Potentilla diversifolia diversifolia Geum trifl Ivesia sabulosa Geum macrophyllum ciliatum perincisum Fragaria virginiana Thalictrum fendleri Ranunculus inamoenus Ranunculus alismifolius var. Delphinium barbeyi Clematis columbiana Caltha leptosepala Aquilegia scopulorum scopulorum Aquilegia coreulea Anemone multifi Aconitum columbianum var. Pyrola secunda Pyrola picta rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Rubiaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Ranunculaceae (Thalictraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Helleboraceae) Pyrolaceae (Ericaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Pyrolaceae (Ericaceae) Species name Common name Nativity

Chapter 6: Appendices 83 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status ower Native Pres Jun–Aug X Lemmon’s spring-parsleyLemmon’s Native Pres Jul–Aug X X X Barrens pinnate angelica Native Pres Jun–Aug X X X American alpine speedwell Native Pres Jul–Aug X X thyme-leaved speedwell Native Pres Jun–Aug X American brooklime Native Pres Jul–Aug X cutleaf synthyris Native Pres Jun–Aug X X Whipple’s penstemonWhipple’s Native Pres Jul–Aug X Rydberg’s penstemonRydberg’s Native Pres Jun–Jul X Markagunt penstemon Native Pres Jun–Aug X X undulate penstemon Native Pres May–Jul X Parry’s lousewortParry’s Native Pres Jul–Aug X X elephant-head Native Pres Jun–Aug X yellow monkeyfl sulphur paintbrush Native Rep Jun–Aug X X rhexia-leaved paintbrush Native Pres Jul–Aug X Barrens Reveal’s paintbrushReveal’s Native Pres Jun–Aug Claron barrens scarlet paintbrush Native Pres Jul–Aug X X X Wyoming paintbrush Native Pres Jul–Aug X diamondleaf saxifrage Native Pres May–Sep X brook saxifrage Native Pres May–Sep X marsh grass-of-Parnassus Native Pres May–Oct X fringecup woodlandstar Native Pres Mar–Aug X X

var.

var.

var. var. var. var. var. var. ramulosum Cymopterus lemmonii Angelica pinnata Veronica wormskjoldii Veronica Veronica serpyllifolia Veronica Veronica americana Veronica Synthyris laciniata Penstemon whippleanus Penstemon rydbergii aggregatus Penstemon leiophyllus leiophyllus Penstemon eatonii undosus Pedicularis parryi parryi Pedicularis groenlandica Mimulus guttatus Castilleja rhexifolia sulphurea Castilleja rhexifolia rhexifolia revealii Castilleja parvula Castilleja miniata Castilleja linariifolia Saxifraga rhomboidea Saxifraga odontoloma Parnassia palustris montanensis Lithophragma glabrum var. rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Saxifragaceae Saxifragaceae Saxifragaceae (Parnassiaceae) FamilyPerennial Forbs, cont. Saxifragaceae Species name Common name Nativity Appendix D. Confi

84 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status Native Pres Aug X X r mistletoe r Native Pres Mar–May X eld pennycress eld Intro Rep May–Aug X yard knotweed Intro Pres May–Oct X Disturbed small collomia Native Pres May–Aug X common mallow Intro Pres Apr–Oct X Disturbed meadow gentian lupineKing’s Native Pres Native Jul–Sep Pres May–Aug X X Jones’ gentian Native Pres Jul–Aug X Barrens fi mountain tansy-mustard Native Pres Apr–Jul X smallhead goosefoot Native Pres Jun–Sep X mountain goosefoot Native Pres Jul–Sep X X Douglas-fi lambsquarter Intro Pres Jun–Sep X Disturbed limber pine dwarf- mistletoe northern bog violet Native Pres Apr–Jul X Canadian white violet Native Pres Apr–Aug X X western valerian Native Pres May–Jul X X tobacco-root Native Pres Jun–Aug X blunt sweet-cicely Native Pres Jun–Jul X X X least lomatium Native Pres May–Jun X X Claron barrens southern ligusticum Native Pres Jun–Aug X X X least spring-parsley Native Pres Jul–Aug X X Claron barrens

var. var. var. kingii var. parvicapitatum Polygonum aviculare Collomia linearis Lupinus kingii Malva neglecta Gentianopsis detonsa elegans Gentianella tortuosa Thlaspi arvense Descurainia incana sonnei Chenopodium capitatum var. Chenopodium atrovirens Chenopodium album album Arceuthobium douglasii Arceuthobium cyanocarpum Viola nephrophylla Viola canadensis Valeriana occidentalis Valeriana Valeriana edulis Valeriana Osmorhiza depauperata Lomatium minimum Ligusticum porteri Cymopterus minimus rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Polygonaceae Polemoniaceae Leguminosae (Fabaceae) Malvaceae Gentianaceae Gentianaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Chenopodiaceae Chenopodiaceae Annual Forbs Chenopodiaceae Viscaceae Viscaceae Violaceae Violaceae Valerianaceae Valerianaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Umbelliferae (Apiaceae) Species name Common name Nativity

Chapter 6: Appendices 85 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status ower rock-jasmine ower Native Pres May–Aug X X X ower spikerush ower Native Pres Jun–Aug X Idaho bentgrass Native Pres Jul–Aug X redtop Intro Pres Jun–Sep X X ticklegrass Native Pres Jul–Aug X X crested wheatgrass Intro Pres Jun–Aug X few-fl common spikerush Native Pres May–Aug X blister sedge Native Pres Jun–Aug X beaked sedge Native Pres Jun–Aug X slender sedge Native Pres Jun–Aug X russet sedge Native Pres Jun–Aug X Ross’ sedge Native Pres May–Aug X X liddon sedge Native Pres May–Jul X woolly sedge Native Pres May–Sep X western sedge Native Pres Jun–Aug X Scandinavian sedge Native Pres Jun–Aug X small-wing sedge Native Pres Jun–Aug X inland sedge Native Pres May–Jul X blackened sedge Native Pres Jul–Aug X kobresia sedge Native Hist Jun–Aug X Eggleston’s sedgeEggleston’s Native Pres Jul–Aug X northern bog sedge Native Pres Jun–Jul X golden sedge Native Pres Apr–Aug X water sedge Native Pres Jun–Aug X pygmy-fl Sawatch knotweed Native Pres Jun–Aug X Douglas’ knotweed Native Pres Jun–Oct X

var. var.

ora var. var. var. idahoensis x Agrostis Agrostis stolonifera Agrostis scabra Agropyron cristatum Eleocharis quinquefl Eleocharis palustris Carex vesicaria Carex utriculata Carex scirpoidea scirpoidea Carex saxatilis Carex rossii Carex petasata Carex pellita Carex occidentalis Carex norvegica stevenii Carex microptera Carex interior Carex heteroneura chalciolepis Carex elynoides Carex egglestonii Carex dioica Carex aurea Carex aquatilis Androsace septentrionalis Polygonum douglasii johnstonii Polygonum douglasii douglasii rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Perennial Graminoids Perennial Cyperaceae Primulaceae Polygonaceae FamilyAnnual Forbs, cont. Polygonaceae Species name Common name Nativity Appendix D. Confi

86 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status littleseed ricegrass Native Pres Jun–Aug X mat muhly Native Pres Jul–Sep X X foxtail muhly Native Pres Jul–Sep X oniongrass Native Pres May–Aug X junegrass Native Pres Jun–Aug X X meadow barley Native Pres Jun–Aug X X fowl mannagrass Native Pres Jul–Aug X Rocky Mountain fescue Native Pres Jul–Aug X alpine sheep fescue Native Pres Jul–Aug X slender wheatgrass Native Pres Jun–Aug X X X X bluebunch wheatgrass Native Pres Jun–Aug X western wheatgrass Native Pres Jun–Aug X Scribner wheatgrass Native Pres Jul–Aug X X thickspike wheatgrass Native Pres Jun–Aug X intermediate wheatgrass Intro Pres Jun–Aug X Disturbed blue wildrye Native Pres Jul–Aug X squirreltail Native Pres May–Aug X X X X Great Basin wildrye Native Pres Jun–Aug X X tufted hairgrass Native Pres Jul–Sep X X timber oatgrass Native Pres Jul–Aug X orchard grass Intro Pres Jun–Aug X slimstem reedgrass Native Pres Jul–Aug X Jones’ reedgrass Native Pres Jul–Aug X X Barrens smooth brome Intro Pres Jun–Aug X X fringed brome Native Pres Jul–Sep X X mountain brome Native Pres May–Aug X X X nodding brome Native Pres Jun–Aug X pine dropseed Native Pres Jul–Sep X X

var. var. var. Oryzopsis micrantha Muhlenbergia richardsonis Muhlenbergia andina Melica bulbosa Koeleria macrantha Hordeum brachyantherum Glyceria striata Festuca ovina saximontana Festuca ovina brevifolia Elymus trachycaulus Elymus spicatus Elymus smithii Elymus scribneri Elymus lanceolatus Elymus hispidus Elymus glaucus Elymus elymoides Elymus cinereus Deschampsia cespitosa Danthonia intermedia Dactylis glomerata Calamagrostis stricta Calamagrostis scopulorum Bromus inermis inermis Bromus ciliatus Bromus carinatus Bromus anomalus Blepharoneuron tricholepis rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Confi FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity

Chapter 6: Appendices 87 Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status owered woodrush owered Native Pres Jun–Aug X X eld horsetail eld Native Pres NA X northern scouring-rush Native Pres NA X smooth scouring-rush Native Pres NA X fi annual bluegrass Intro Pres Apr–Sep X cheatgrasspullup muhly Intro Native Pres Pres Jun–Aug May–Jul X X X Disturbed small-fl Mertens’ rush Native Pres Jul–Aug X longstyle rush Native Pres Jun–Aug X mountain rush Native Pres Jun–Aug X Tracy’s rushTracy’s Native Pres Jun–Aug X perplexing rush Native Pres Jun–Aug X X Baltic rush Native Pres May–Jul X spike trisetum Native Pres Jul–Sep X X X X Letterman’s needlegrassLetterman’s needlegrassNelson’s Native Pres Native Jul–Sep Pres X Jun–Sep X X X Indian ricegrass Native Pres May–Jul X needle-and-thread Native Rep May–Jul X Sandberg’s bluegrassSandberg’s Native Pres Apr–Sep X X X Kentucky bluegrass Intro Pres May–Aug X X X X Wheeler bluegrass Native Pres Apr–Aug X X muttongrassinterior bluegrass Native Native Pres Pres Jun–Aug May–Aug X X X X X Canada bluegrass Intro Pres Jun–Aug X Arctic bluegrass Native Hist Jul–Aug X X timothy Intro Pres Jun–Sep X X alpine timothy Native Pres Jun–Aug X X

nelsonii var. var. comata wheeleri liformis glauca var. var. ora var. var. Equisetum variegatum Equisetum laevigatum Equisetum arvense Muhlenbergia fi Poa annua Bromus tectorum Luzula parvifl Juncus mertensianus Juncus longistylus Juncus ensifolius montanus Juncus ensifolius brunnescens Juncus confusus Juncus arcticus Stipa nelsonii spicatum Trisetum Stipa lettermanii Stipa hymenoides Stipa comata Poa secunda Poa pratensis Poa glauca Poa nervosa Poa fendleriana Poa compressa Poa arctica Phleum pratense Phleum alpinum rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Equisetaceae Equisetaceae Ferns and Allies Equisetaceae Gramineae (Poaceae) Gramineae (Poaceae) Annual Graminoids Gramineae (Poaceae) Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Appendix D. Confi

88 Annotated Checklist of Vascular Flora: Cedar Breaks National Monument Wet Comments Sub Mdw Fir White Fir Sub Flower time Park status brittle bladder fern Native Pres NA X X parsley-fern Native Hist NA X Talus Ferris’ scouring-rush Native Pres NA X

var. ferrisii x Cystopteris fragilis Cryptogramma crispa acrostichoides Equisetum rmed, historical, and reported taxa of Cedar Breaks National Monument, organized by life form with ecological notes, cont. Polypodiaceae (Aspleniaceae) Polypodiaceae (Adiantaceae) Appendix D. Confi FamilyFerns and Allies, cont. Equisetaceae Species name Common name Nativity

Chapter 6: Appendices 89

The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientifi c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and affi liated Island Communities.

NPS D-225, Month 2009 National Park Service U.S. Department of the Interior

Natural Resource Program Center 1201 Oak Ridge Drive, Suite 150 Fort Collins, Colorado 80525 www.nature.nps.gov

EXPERIENCE YOUR AMERICA™