Annotated Checklist of Vascular Flora, Bryce

Home , Ageratina herbacea, Allium brevistylum, Allium nevadense, Amauriopsis, Amauriopsis dissecta, Antennaria rosulata

National Park Service U.S. Department of the Interior

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Annotated Checklist of Vascular Flora Bryce Canyon National Park

Natural Resource Technical Report NPS/NCPN/NRTR–2009/153 ON THE COVER Matted prickly-phlox (Leptodactylon caespitosum), Bryce Canyon National Park, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Bryce Canyon National Park

Natural Resource Technical Report NPS/NCPN/NRTR–2009/153

Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Sarah Topp Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

January 2009

U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientiﬁ c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientiﬁ cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner.

The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientiﬁ c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientiﬁ c literature reviews; and peer- reviewed proceedings of technical workshops, conferences, or symposia.

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Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

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Please cite this publication as: Fertig, W., and S. Topp. 2009. Annotated checklist of vascular ﬂ ora: Bryce Canyon National Park. Natural Resource Technical Report NPS/NCPN/NRTR–2009/153. National Park Service, Fort Collins, Colorado.

NPS D-244, January 2009 ii Annotated Checklist of Vascular Flora: Bryce Canyon National Park Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justiﬁ cation ...... 1 1.2 Objectives ...... 1 1.1 Study area ...... 1 1.1.1 History and setting ...... 1 1.1.2 Geology ...... 2 1.1.3 Vegetation ...... 4 1.1.4 Previous botanical studies ...... 4

2 Methods ...... 9

3 Results ...... 11 3.1 Summary of the ﬂ ora of Bryce Canyon National Park ...... 11 3.2 Life form and biogeographic diversity ...... 12 3.3 Plants of special concern ...... 12 3.4 Non-native and noxious weeds ...... 12 4 Discussion ...... 17

5 Literature Cited ...... 19

Appendices ...... 23 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family and scientifi c name with taxonomic and biogeographic notes...... 25 Appendix B. Potential (unconfi rmed) taxa of Bryce Canyon National Park...... 79 Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park. .97 Appendix D. Confi rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes...... 104

Contents iii iv Annotated Checklist of Vascular Flora: Bryce Canyon National Park Figures

Figure 1.3.1. Location of Bryce Canyon National Park, Garﬁ eld and Kane counties, Utah...... 3 Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Bryce Canyon National Park by decade, 1920–2008...... 7

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Bryce Canyon National Park...... 5 Table 3.1. Statistical summary of the ﬂ ora of Bryce Canyon National Park...... 11 Table 3.3. Plant species of conservation concern from Bryce Canyon National Park...... 13 Table 3.4. Noxious weed species of Bryce Canyon National Park...... 15

Contents v vi Annotated Checklist of Vascular Flora: Bryce Canyon National Park Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Bryce Canyon National Park. This entailed a thorough review of existing literature, re-examination of specimens from the Bryce Canyon National Park herbarium and other state and national museum collections, and additional fi eld work to corrob- orate unverifi ed species reports or locate new species. Based on this study, 587 vascular plant taxa have been confi rmed or reliably reported as occuring in the park (553 are known from validated herbarium specimens and 34 from unvouchered literature reports). Another 76 species formerly considered part of the park fl ora have now been shown to be falsely reported. The current fl ora of Bryce Canyon National Park represents 16.3% of the known fl ora of Utah, and 36.5% of the fl ora of the Utah High Plateaus ecoregion. Although no species are federally listed as threatened or endangered, 50 plant taxa from the park were formerly candidates for potential listing, are considered sensitive by the U.S. Forest Service and Bureau of Land Management, or are considered species of concern by the state’s natural heritage program. Introduced plants make up 10.2% of the park fl ora (60 species), of which only 7 species are considered noxious weeds by the State of Utah. Further surveys in Bryce Canyon National Park are likely to fi nd additional new species (298 are known from similar habitats outside the park boundary), especially among late season fl owering, weedy, or wetland taxa.

Contents vii viii Annotated Checklist of Vascular Flora: Bryce Canyon National Park Acknowledgements

We would like to thank the following individuals for their assistance with this project. Botanist Dr. Angela Evenden, former leader of the Northern Colorado Plateau Network in Moab, Utah, was an active proponent of this project and helped secure funding for ﬁ eld work and herbarium studies in 2006. Kristin Legg, former Chief of Natural Resources for Bryce Canyon National Park (BRCA), assisted with herbarium access and permitting. Jan Stock, curator of museum collections for BRCA, provided access to all mounted and unmounted herbarium specimens from the park. Bruce Condie helped record specimen label data from all mounted collections in the BRCA herbarium. Libby Nance, former database specialist with the NCPN, assisted with development and quality control of the park’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the park. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University (BRY). Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hart- man and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Mark Madsen, botanist with Dixie National Forest, shared data from his Master’s thesis at Brigham Young University on the Claron ﬂ ora of the Paunsaugunt Plateau. Thanks to the technical reviewers (Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others whom we may have inadvertently forgotten, please accept our apolo- gies (and thanks).

Research was conducted under permit # BRCA-2006-SCI-0004 for study number BRCA-00018. Fund- ing was provided by the National Park Service through the Rocky Mountains Cooperative Ecosystem Studies Unit and the University of Wyoming.

Contents ix x Annotated Checklist of Vascular Flora: Bryce Canyon National Park (Evenden et al. 2002). This eff ort has entailed re- 1 Introduction viewing existing species checklists, corroborating the identifi cation of specimens in park museum In 2006, the National Park Service (NPS) estab- and herbarium collections, locating additional lished a cooperative agreement with the Univer- specimen records from other museums, mining sity of Wyoming, through the Rocky Mountains relevant literature and datasets, and conducting Cooperative Ecosystem Studies Unit, to produce fi eld surveys to locate new taxa or confi rm un- a series of technical reports summarizing the vas- substantiated reports.† All of these data have been cular plant fl ora of each of the 16 national park synthesized in NPSpecies, the park service’s stan- units of the Northern Colorado Plateau Network dardized database for documenting park status (NCPN). These reports are based on existing in- (present, probably present, unconfi rmed, falsely formation in the park service’s NPSpecies data- reported), abundance, residency, nativity, and base system and augmented by further data min- rarity. With the production of a technical series of ing, reviews of state and regional herbaria, and reports on the fl ora of each park, the NPSpecies new fi eld surveys to resolve the status of many data will now be available to a broader audience. reported and unconfi rmed species. The objec- tive of these reports is to summarize the current state of knowledge of the fl ora of each park for 1.2 Objectives the benefi t of NPS biologists and interpretive The objectives of this project were to: staff , outside researchers, and the general public. The following report addresses the vascular fl ora 1. Document the current status (present, of Bryce Canyon National Park (NP), in southern reported, potential, or falsely reported) of all Utah. Other NCPN park units are addressed in vascular plant species that have been cited as separate reports. occurring in Bryce Canyon National Park, based on a review of herbarium collections and past literature. 1.1 Background and justifi cation One of the primary missions of the NPS is “to 2. Develop an annotated checklist of the conserve unimpaired the natural and cultural fl ora of Bryce Canyon National Park, with resources and values of the national park system supplemental information on taxonomic for the enjoyment of this and future generations” synonyms, distribution, within-park abun- (NPS 2000). Due to their high degree of protec- dance, fl owering period, growth form, nativ- tion and distribution across a wide variety of hab- ity, habitat, and sources for each record. itat types, the nation’s national park units make a signifi cant contribution to the protection of bio- 1.1 Study area logical diversity (Noss and Cooperrider 1994). Unfortunately, the full contribution of the NPS 1.1.1 History and setting system is not known, as many park units have Native American cultures have occupied the area traditionally lacked comprehensive biological in- surrounding Bryce Canyon for several thousand ventories or monitoring programs to identify the years, but little physical evidence remains to composition and condition of their biota. Know- suggest that the high-elevation badlands of the ing the status of park fl ora and fauna is critical for park, itself, were utilized for anything more than eff ective park management and for systematically hunting and gathering. Following a period of re- identifying and fi lling gaps in the nation’s protect- gion-wide severe drought around 1200 A.D., the ed area network (Margules and Pressey 2000). valleys bordering the Paunsaugunt Plateau and Bryce Canyon were sparsely inhabited by transi- To rectify this shortcoming, Congress passed the tory bands of Paiutes. The Paiutes referred to the National Parks Omnibus Management Act in 1998 colorful hoodoos, spires, fi ns, and chimneys of to provide guidance and funding for an inventory the Bryce area as Angka-ku-wass-a-wits (“red- and monitoring program across the entire na- painted faces”) or Unka-timpe-wa-wince-pock- tional park system. Since 2000, the NCPN Inven- ich (“red rock standing like men in a bowl-shaped tory and Monitoring Program has been develop- recess”) (Davis and Pollock 2003). ing and verifying species lists of vertebrates and vascular plants for each park unit in the network

†Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 Mormon pioneers arrived in the area in the mid- erates laid down about 96 million years ago in 1870s and established several small settlements, a coastal environment of fl uctuating marine la- including Clifton (later abandoned), Tropic, Can- goons and river deltas (Davis and Pollock 2003). nonville, and Henrieville. Ebenezer Bryce was an Lying above the Dakota are beds of black or dark- early settler who built a road and ditch to water gray shales of the Tropic Formation (named for the fi elds of Clifton in the canyon that would bear the nearby town of Tropic) that were deposited in his name (Belknap 1958). Eventually, most of the deep marine waters during the last major incur- Paunsaugunt Plateau in the Bryce Canyon area sion of the inter-continental Cretaceous seaway would be transferred to the U.S. Forest Service that bisected North America. Younger deposits (as part of Powell National Forest, now known of the Straight Cliff s Formation form the “gray as Dixie National Forest), while the surrounding cliff s” of the Grand Staircase and can be as thick valleys were mostly converted to agriculture. as 1,700 feet (515 m) in the Bryce area (Davis and Pollock 2003). The Straight Cliff s consist of alter- J. W. Humphrey, an early forest supervisor who nating beds of sandstone and shale interspersed recognized the tourism potential of Bryce’s sce- with coal seams that were deposited in a coastal nic wonders, persuaded the Union Pacifi c Rail- area with periodically advancing and retreating road to construct roads into the area, along with marine waters. Topping the Straight Cliff s is the a visitor lodge near Sunset Point, in the late 1910s late Cretaceous Wahweap Sandstone, which was and early 1920s. Local settler Reuben “Ruby” formed in a river delta. Syrett also built a tourist lodge that remains a local landmark (Davis and Pollock 2003). In June 1923, Bryce’s pink cliff s consist of a thick mantle of President Warren Harding used the Antiquities Eocene-age limey sandstones of the Claron For- Act to create Bryce Canyon National Monument, mation (formerly called the Wasatch Formation). under the administration of the U.S. Forest Ser- The Claron was deposited 40–60 million years vice. Congress passed legislation in June 1924 to ago in a large freshwater lake (slightly larger than expand the monument and rename it Utah Na- modern Lake Erie) that extended from near the tional Park. The park was renamed Bryce Canyon Nevada state line to Boulder Mountain (Chronic National Park and permanently transferred to the 1988). Most of the Claron at Bryce Canyon con- National Park Service in February 1928. Over the sists of the lower, red member, with the upper, next several decades, the park’s boundaries were white member persisting only at the higher eleva- redrawn and expanded on several occasions, un- tions of the Paunsaugunt Plateau (most of it has til reaching its current size of 35,835 acres (14,502 eroded away elsewhere on the plateau). The red ha) (Evenden et al. 2002). member is not homogeneous, but consists of al- ternating bands of more-resistant limestone or Bryce Canyon National Park is located on the conglomerate. A small area of Oligocene or Mio- eastern fl ank of the Paunsaugunt Plateau in Gar- cene-age conglomerates deposited in a small lake fi eld and Kane counties, Utah (Figure 1.3.1). The persists on top of Boat Mesa. The park’s youngest park is located approximately 50 air miles east of strata consist of Quaternary alluvium in the major Cedar City and 30 air miles northeast of Kanab. drainages carving the eastern fl ank of the Paun- Elevations range from 6,100 feet (1,859 m) along saugunt Plateau (Davis and Pollock 2003). the eastern border to 9,100 feet (2,774 m) in the highlands at the park’s southern end. Annual pre- The fantastic hoodoos, rock spires, pinnacles, cipitation averages 16.3 inches (414 mm), with chimeys, fi ns, and windows that characterize the 6.7 inches falling as rain during the main grow- Bryce Canyon area were carved primarily by the ing season from June to September. Average an- freezing and thawing of water acting on the cracks nual air temperature is 41°F (5.2°C), with a low and fi ssures created by the regional faulting that of 9.3°F (–12.6°C) in January and a high of 77.8°F uplifted the Paunsaugunt Plateau along the Sevier (25.4°C) in July (Madsen 2001). and Paunsaugunt faults. Erosion by running water formed deep gullies and sliced the eastern wall 1.1.2 Geology of the plateau into a series of rock fi ns. The natural acidity of water acted to further erode the fi ns The oldest rocks exposed within Bryce Canyon into arch-like windows or solitary, chimney-like National Park are outcrops of the Cretaceous-age hoodoos, held in place by a caprock of less-resis- Dakota Sandstone and Tropic Shale found along tant limestone or conglomerate. Ultimately, even the park’s easternmost boundary. The Dakota these fall, but new erosional features continue to consists of sandstones, siltstones, and conglom- form as the Paria River and its tributaries march

2 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Bryce Canyon Nationalnal Park Pa rk Utah

12 Visitor Center

Tropic

Cannonville

Established June 8, 1923 Legend 1,859–2,774 m elevation Perennial streams Grand Staircase-Escalante NM (BLM) (6,100–8,000 ft elevation) Rivers 14,502 ha (35,835 ac) U.S. Bureau of Land Management Paved roads Private 0123 Kilometers Unpaved roads State Unimproved/4WD roads U.S. Forest Service 0123 Miles Park boundary (NPS) Water

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002) Figure 1.3.1. Location of Bryce Canyon National Park, Garﬁ eld and Kane counties, Utah.

Chapter 1: Introduction 3 headward against the red Claron beds at a rate of due to long-term fi re suppression. Montane for- four feet (1.2 m) per century toward its ultimate ests (with the exception of aspen groves) tend to capture of the East Fork of the Sevier River in 1-2 have a sparse understory of shade-tolerant shrubs million years (Davis and Pollock 2003). and forbs, such as common juniper (Juniperus communis), mountain snowberry (Symphoricar- 1.1.3 Vegetation pos oreophilus), and mountain-lover (Pachystima myrsinites) (Buchanan 1992). The vegetation of Bryce Canyon National Park has been described in detail elsewhere (Buchanan Due to the presenece of the Paunsaugunt Fault, 1960, 1971, Buchanan et al. 1981; NCPN in ed.; porous sandstone overlies relatively impervious Roberts et al. 1992). Buchanan (1992) presented Tropic Shale along the eastern margin of the park, a relatively simple vegetation classifi cation that creating suitable conditions for small seeps and subdivided the park’s fl ora into four main com- springs. Additional areas of riparian vegetation munities arranged within three elevational belts. occur in drainage bottoms and along the Tropic Ditch. Wetland vegetation in Bryce Canyon NP The harshest environments for plant growth is locally dominated by coyote willow (Salix ex- within Bryce Canyon are found on the badlands igua), yellow willow (S. lutea), narrow-leaved slopes and breaks along the eastern escarpment cottonwood (Populus angustifolia), Baltic rush of the Paunsaugunt Plateau, within the pinyon- (Juncus arcticus), and Nebraska sedge (Carex ne- juniper woodland belt (Buchanan 1992). Violent brascensis). summer rainstorms, high winds, and freeze-thaw in winter and spring conspire to prevent deep soils 1.1.4 Previous botanical studies from forming on barren slopes and favor cushion plants or perennial shrubs and trees. Many of the The Pink Cliff s area was explored by the Powell park’s Claron endemic plants are restricted to Survey in 1872, under the direction of John Wes- these colorful badland slopes. Western bristle- ley Powell’s brother-in-law, Almon Thompson. cone pine (Pinus longaeva) may be locally com- Botanist Lester Ward worked for later Powell mon on barren slopes or co-occur with ponde- surveys and collected extensively in the Aquarius rosa pine (P. ponderosa). Lower slopes and more and Fish Lake plateaus, but apparently did not sheltered sites may support open woodlands of collect specimens in the Bryce area as suggested two-needle pinyon (P. edulis), Utah juniper (Ju- by Evenden et al. (2002). Utah’s preeminent pio- niperus osteosperma) and scattered big sagebrush neer botanist, Marcus E. Jones, was apparently (Artemisia tridentata). the fi rst plant collector to explore the immediate vicinity of Bryce Canyon. Jones procured at least More-moderate slopes with better soil develop- two specimens from the “canyon above Tropic” ment characterize the submontane forest belt. near the northeast boundary of the park in 1894 This zone, located mostly below 8,500 feet, con- (Table 1.3.4). These specimens represented types sists of extensive forests of ponderosa pine with of silvery primrose (Primula incana) and King’s an open understory of greenleaf manzanita (Arc- fl ax (Linum kingii var. pinetorum, now considered tostaphylos patula), antelope bitterbrush (Purshia taxonomically negligible). Although Linum kingii tridentata), and Fendler’s mountain-lilac (Ceano- is known from several locations in the park today, thus fendleri). Ponderosa pine forests are replaced Primula incana has not been relocated. by montane sagebrush meadows dominated by black sagebrush (Artemisia nova), greenish rab- The fi rst collector to secure specimens within bitbrush (Chrysothamnus nauseosus), spineless Bryce Canyon National Park (earlier workers horsebrush (Tetradymia canescens), and a variety had collected in Red Canyon and other areas of of perennial bunchgrasses and forbs on sites with adjacent Dixie National Forest) was apparently deeper soils and cold air drainage (Buchanan Walter Cottam, of Brigham Young University, 1992). who discovered at least eight new species starting in 1926 (Table 1.3.4, Figure 1.3.4). Mildred Ma- Above 8,500 feet, the forests of Bryce Canyon NP thias collected the fi rst type specimen from the are dominated by Douglas-fi r (Pseudotsuga men- park when she discovered least lomatium (Loma- ziesii), Engelmann spruce (Picea engelmannii), and tium minimum) in 1929. George Goodman and white fi r (Abies concolor). Quaking aspen (Popu- C. Leo Hitchcock were the fi rst among a cadre lus tremuloides) may have been historically more of noteworthy taxonomists from outside Utah to prevalent in the area but has apparently declined collect within the park, adding at least four new

4 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Table 1.3.4. Chronology of plant-collecting efforts in Bryce Canyon National Park. # of new Year Collector(s) Comments taxa 1894 M. E. Jones 1 Marcus E. Jones collected the types of Primula incana and Linum kingii reported var. pinetorum from “canyon above Tropic” in the vicinity of Bryce Canyon National Park. Linum kingii has since been discovered within the park, but no confi rmed records have been found for Primula incana. 1912 W. W. Eggleston 0 W. W. Eggleston collected the type of Breaks bladderpod (Physaria [Lesquerella] rubicundula) from Red Canyon, NW of Bryce Canyon 1916 H. E. Malmsten 0 H. E. Malmsten, a U.S. Forest Service employee, collected the type of Podunk groundsel (Senecio malmstenii) from Little Podunk Creek, apparently just outside the boundary of Bryce Canyon NP. 1926– W. P. Cottam, F. W. 13 Walter Cottam made the earliest-known collections from within Bryce Canyon 1929 Hunnewell, & M. E. NP, documenting at least 8 species from 1926 to 1929. F. W. Hunnewell added Mathias at least 3 new species while collecting for the New York Botanical Garden in 1928. Mildred Mathias collected the holotype of Lomatium minimum from Bryce Canyon in 1929. 1930– G. Goodman, C. 152 George Goodman and C. L. Hitchcock discovered 4 new species for the park in 1935 L. Hitchcock, K. E. 1930. Kenneth Weight established the BRCA herbarium and initiated the fi rst Weight, J. A. Rowe, major fl oristic study of Bryce Canyon in 1931, documenting 144 new species F. M. Stone, M. C. over the next four years. Weight (1933) developed the fi rst checklist of the Ferguson, A. M. park and vicinity, listing 286 taxa. J. A. Rowe, F. M. Stone, M. C. Ferguson, Ottley, A. Eastwood, & and A. M. Ottley contributed 4 additional new species during this period. J. T. Howell In 1933, Alice Eastwood and John Howell collected the holotypes of Red Canyon penstemon (Penstemon bracteatus) and Bryce Canyon townsendia (Townsendia montana var. minima) from Red Canyon, just west of the park. 1938 W. S. Boyle 3 W. S. Boyle discovered 3 new species for the park. 1940– H. R. Shephard, E. 28 Bassett Maguire collected the type of Silene petersonii from Red Canyon 1946 Gifford, J. D. Slade, J. (outside Bryce Canyon NP) in 1940. H. R. Shephard, E. Gifford, J. D. Slade, W. Vogt, W. S. Boyle, and J. W. Vogt made more than 40 collections in the park and added 17 new O. Degener, L. Peiler, species to the fl ora in 1940–1941. W. S. Boyle contributed nearly 50 specimens & B. Maguire to the park herbarium and 9 new species in 1942. Otto Degener, Leroy Peiler, and L. J. Mitchell added two additional species from 1942 to 1945. 1947 D. Ripley & R. Barneby 0 Dwight Ripley and Rupert Barneby collected the holotype of Jones’ locoweed (Oxytropis oreophila var. jonesii) from Red Canyon, west of Bryce Canyon NP. 1950– H. H. Hall & B. F. 2 Heber Hall and Bertrand Harrison collected two new species for the park from 1954 Harrison 1950 to 1954. 1956– H. Buchanan 94 Hayle Buchanan, then a graduate student at the University of Utah, 1959 discovered 94 species in Bryce Canyon NP while conducting research on the vegetation of the park (Buchanan 1960). 1965 N. Holmgren & J. 2 Noel Holmgren and James Reveal collected the type of Reveal’s paintbrush Reveal (Castilleja parvula var. revealii) from Bryce Canyon NP. 1968 L. Higgins 0 Larry Higgins collected the type of yellowish cryptanth (Cryptantha ochroleuca) at Red Canyon, west of Bryce Canyon NP. 1969– H. Buchanan 3 Hayle Buchanan added 3 new species to the park fl ora while conducting 1970 research on forest ecology of Bryce Canyon. Buchanan and Nebeker (1971) compiled a checklist of the park fl ora citing 218 taxa. 1971– G. Stebbins & H. 4 Gary Stebbins discovered 3 new species in 1971–1972; Hayle Buchanan added 1973 Buchanan one new taxon in 1973. 1977 R. Foster & D. Foster 3 Robert Foster discovered 3 new species in 1977. 1978– H. Buchanan 9 Beginning in 1978, Hayle Buchanan conducted a follow-up study to his 1979 doctoral work in the park, examining successional patterns with Robert Graybosch and Kimball Harper.

Chapter 1: Introduction 5 Table 1.3.4. Chronology of plant collecting efforts in Bryce Canyon National Park, cont.

# of new Year Collector(s) Comments taxa 1980– R. Graybosch 138 + 10 Robert Graybosch collaborated with Hayle Buchanan to study vegetation and 1981 reported endemic plants of Bryce Canyon NP and added 148 new species to the park fl ora. Buchanan and Graybosch (1981) published a revised checklist with 446 species. 1982 R. Shaw 1 Richard Shaw added Colorado Plateau bladderpod (Lesquerella rectipes) to the park fl ora. 1986– L. Gottschalk & G. 8 Linda Gottschalk and Gregory Hallsten of Utah State University collected 8 1987 Hallsten new species for Bryce Canyon NP in 1986–1987. Hallsten and Roberts (1988) published a short paper listing 27 new species not previously reported for the park. 1989– C. Schelz, J. M. Passek, 6 Charles Schelz, J. M. Passek, and M. W. Britten added 6 new species to the 1991 and M. W. Britten park fl ora while conducting rare plant and invasive weed studies. 1993– R. Foster 24 Robert Foster discovered 24 new species for the park (specimens mostly 1994 at BRY), including the earliest collection of Barkley’s groundsel (Senecio werneriifolius var. barkleyi, described as a new taxon by Welsh et al. in 2003). 1998 B. Buchanan, K. 23 Beth Buchanan, Karen Kitchen, and Juli Ames-Curtis conducted a weed survey Kitchen, & J. Ames- and discovered 22 new non-native species for the park, including 4 new Utah Curtis state noxious weeds. 2004– S. Topp, K. Andersen, 40 + 12 Dewey and Andersen (2005) reported leafy spurge (Euphorbia esula) as a 2007 L. Ballenger, A. reported new invasive species for Bryce Canyon during their 2004 weed survey. Sarah Tendick, & W. Fertig Topp, Amy Tendick, and Liz Ballenger of the NCPN vegetation mapping crew vouchered 37 new species and reported 11 others (without vouchers) while conducting fi eld sampling from 2004 to 2007 (NCPN in ed., Topp 2007). W. Fertig added 3 new species in 2006 (Topp 2007).

species to the ﬂ ora in 1930. In subsequent years, while conducting research toward his doctoral Alice Eastwood, John T. Howell, Dwight Ripley, dissertation at the University of Utah on the veg- Rupert Barneby, Bassett Maguire, Larry Higgins, etation of Bryce Canyon (Buchanan 1960). Buch- Noel Holmgren, and James Reveal would visit anan returned to the park in the late 1960s and the area and collect holotypes of Claron endem- late 1970s to conduct follow-up research on suc- ics, though most of these were collected in Red cessional dynamics (Buchanan 1971, Buchanan et Canyon (on Dixie National Forest) rather than in al. 1981) and co-authored a revised park checklist Bryce Canyon National Park (Welsh 1982). in 1971 (Buchanan and Nebeker 1971).

Kenneth Weight initiated the fi rst major fl oristic From 1978 to 1981, Buchanan worked with grad- study of Bryce Canyon NP in 1931. Over the next uate student Robert Graybosch on a new study of four years, Weight collected over 180 specimens vegetation and endemic plants of Bryce Canyon for the nascent Bryce Canyon NP herbarium NP that resulted in the discovery of 157 new taxa (BRCA) and documented 144 new species for the and increased the park’s fl ora by nearly 33%. Bu- park. In 1933, Weight published the fi rst check- chanan and Graybosch (1981) published a revised list of the park’s fl ora and vicinity, listing 286 taxa checklist reporting 446 species for the park. (Weight 1933). Over the next decades, a number of other park employees and visitors (most nota- More than 100 new species have been added bly, W. S. Boyle, H. R. Shephard, and E. Giff ord) to the park fl ora in the past two decades. Linda contributed about 30 additional species to the Gottschalk and Gregory Hallsten, of Utah State park fl ora. University, discovered nine new species in the mid 1980s (Hallsten and Roberts 1988). Charles The next major fl oristic study of Bryce Canyon Schelz, Beth Buchanan, Karen Kitchen, Juli Ames- National Park began in 1956, with the work of Curtis, and other park employees discovered 29 Hayle Buchanan. During the late 1950s, Buch- new weed species during surveys in 1989–1991 anan discovered 94 new species for the park fl ora and 1998. John Spence conducted vegetation

6 Annotated Checklist of Vascular Flora: Bryce Canyon National Park monitoring studies in Bryce Canyon in the early 2004–2007 (NCPN in ed., Topp 2007). Dewey 1990s, and co-authored a revised plant checklist and Andersen (2005) added one additional spe- with Hayle Buchanan in 1993, listing 458 taxa and cies report while conducting a mapping survey of correcting some errors from previous lists (Spen- invasive weeds. Walter Fertig contributed three ce and Buchanan 1993). Robert Foster collected new species and relocated several historic re- extensively in 1993–1994 and added 24 new spe- cords in 2006. cies to the ﬂ ora. Most recently, Sarah Topp, Amy Tendick, Liz Ballenger, and colleagues from the Additional details on plant-collecting eﬀ orts at NCPN vegetation mapping team discovered or Bryce Canyon NP can be found in Table 1.3.4. reported 48 new taxa for Bryce Canyon NP from

600

500

400

300

200 Number of vascular taxa

100

0 1920 1930 1940 1950 1960 1970 1980 1990 2000 2008

Year

Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Bryce Canyon National Park by decade, 1920–2008.

Chapter 1: Introduction 7 8 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Each list was annotated with supplemental infor- 2 Methods mation on life form, geographic range (within the context of Utah), population size in the park (de- In April 2005, we reviewed all mounted specimens rived from NCPN in ed., and fi eld observations), in the Bryce Canyon National Park herbarium to fl owering period, distribution across general veg- determine whether each was correctly identifi ed etation types, nativity, data source (collector and to species, subspecies, or variety following the collection number from a sample specimen or lit- nomenclature of Welsh et al. (2003). As neces- erature citation), year of earliest collection (from sary, corrections and annotations were made to herbarium records), and relevant synonymy and each specimen and to the voucher records in the taxonomic problems (based on review of Bark- park’s Automated National Cataloging System worth et al. 2003, 2007; Barneby 1989; Cronquist (ANCS+) and NPSpecies database. Based on this 1994; Cronquist et al. 1972, 1977, 1984, 1997; herbarium work and a review of park references Dorn 2001; Flora of North America Editorial (Buchanan and Graybosch 1981; Hallsten and Committee 1993, 1997, 2000, 2002a, 2002b, 2003, Roberts 1988; Spence and Buchanan 1993), we 2005, 2006a, 2006b, 2006c; Holmgren et al. 2005; developed four lists: Weber and Wittmann 2001). The annotations to these lists and label data from the BRCA her- 1. A revised checklist of species that were pres- barium review were then incorporated into the ent in the park (confi rmed by an authenti- NPSpecies database for the park. cated voucher; included in Appendices A and D); Work in 2005–2007 focused on confi rmation of reported and potential species for the park fl ora 2. A list of species that were reported in the lit- through fi eld surveys and review of collections erature but not corroborated with a voucher from state and regional herbaria. We made several (equivalent to “probably present” in NPSpe- collecting trips to Bryce Canyon NP in the spring, cies terminology; included in Appendices A summer, and fall of 2005, 2006, and 2007 to target and D); missing species, emphasizing under-sampled wet- 3. A list of potential species (“unconfi rmed” land, weedy, and late-fl owering taxa. Herbarium in NPSpecies) A list of potential species studies were undertaken at Brigham Young Uni- (“unconfi rmed” in NPSpecies) known from versity (BRY), Utah State University (USU), and the vicinity of Bryce Canyon NP using an the University of Wyoming (RM) in March, June, unpublished list generated by Dr. N. Duane and October 2006 and February and March 2007. Atwood of Brigham Young University, Albee We also documented park specimens from the et al. (1988), Madsen (2001), Spence and Virtual Herbarium website of the New York Bo- Buchanan (1993), and Welsh et al. (1978) tanical Garden (http://sciweb.nybg.org/science2/ (Appendix B); and hcol/intf/index.asp). Finally, we found additional new species reports by examining unmounted 4. A list of species that were known to be mis- collections and reviewing plot data from recent identifi ed by Spence and Buchanan (1993) vegetation mapping studies (NCPN in ed.). or highly questionable (Appendix C).

Chapter 2: Methods 9 10 Annotated Checklist of Vascular Flora: Bryce Canyon National Park from collections at Brigham Young University, 3 Results Utah State University, and the New York Bo- tanical Garden Virtual Herbarium. Field surveys 3.1 Summary of the fl ora of Bryce from 2005 to 2007 netted another 37 new species (Topp 2007). Overall, the documented fl ora Canyon National Park of Bryce Canyon NP has increased by 86 species Of the 1,245 vascular-plant vouchers in the main (15.6%), bringing the total number of taxa to 553 BRCA herbarium and the park’s resource man- (Table 3.1, Appendices A, D). agement collection that we examined in 2005, 124 were misidentifi ed (10.0%), 54 were revised Currently, 34 additional taxa have been reported to a more current name (4.3%), 291 had variety for Bryce Canyon NP but have yet to be corrobo- or subspecies names added (23.4%), and 776 rated with a voucher specimen (Table 3.1, Appen- were found to be correctly determined (62.3%). dix A). Of these, 27 were reported by Buchanan Based solely on these collections, the confi rmed and Graybosch (1981), Spence and Buchanan fl ora of Bryce Canyon National Park stood at 467 (1993), or from observations by recent vegetation taxa (Fertig 2006). From 2005 to 2007, we located mapping and wetland research crews (NCPN in an additional 49 taxa (mostly species reported in ed.). Taken together, the number of confi rmed Spence and Buchanan 1993, but without a dupli- and reported vascular plant taxa in Bryce Canyon cate voucher deposited in the BRCA herbarium) NP stands at 587 taxa (Table 3.1, Appendix A).

Table 3.1. Statistical summary of the fl ora of Bryce Canyon National Park. Number Number Category confi rmed additionally Total in park reported for park Taxonomic diversity Total taxa 553 34 587 (including varieties and subspecies) Full species 526 27 553 (excluding varieties and subspecies) Families 73 0 73 Life form diversity Tree taxa 18 0 18 Shrub taxa 72 2 74 Perennial forb taxa 316 23 339 Annual forb taxa 58 6 64 Perennial graminoid taxa 77 3 80 Annual graminoid taxa 7 0 7 Fern taxa 5 0 5 Biogeographic diversity Introduced taxa 58 2 60 Native taxa Locally endemic taxa 19 3 22 Regionally endemic taxa 45 4 49 Disjunct taxa 3 0 3 Peripheral taxa 2 0 2 Sparse taxa 10 0 10 Widespread taxa 416 25 441 Total native taxa 495 32 527 The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003). See Appendices, “Codes and Defi nitions”, for an explanation of the various categories.

Chapter 3: Results 11 At the start of the 2006 fi eld season, we recognized 1° of latitude × 2° of longitude; see Appendices 327 taxa as potentially occurring in Bryce Canyon for more complete defi nitions of biogeographic NP, based on the proximity of the park to known categories). Regional endemics (largely restricted populations (Albee et al. 1988; Madsen 2001; Fer- to the Utah High Plateaus ecoregion or Colorado tig 2006). At least 29 of these species have recently Plateau) comprise an additional 9.3% of the park been found in the park based on fi eld collections fl ora (Table 3.1). Less than 3% of the park fl ora or relocated herbarium specimens. The current consists of species with disjunct, peripheral, or number of potential species that we are recogniz- sparse geographic distribution patterns. ing for Bryce Canyon NP is 298 (Appendix B).

Of the original pool of species documented or re- 3.3 Plants of special concern ported for the park, we found that 70 taxa were Bryce Canyon National Park has no plants listed falsely reported based on misidentifi ed voucher as threatened or endangered under the U.S. En- specimens in the BRCA and BRY herbaria and dangered Species Act. At least 21 park taxa were the work of Spence and Buchanan (1993). We formerly candidates for potential listing (Category consider an additional six species cited (but not 2 or 3C) under the act (Table 3.3; UDWR 1998). vouchered) by Spence and Buchanan (1993) and Another 12 species are currently listed as sensi- the NCPN vegetation mapping team (NCPN in tive by the U.S. Forest Service or Utah Bureau of ed.) to be questionable based on the absence of Land Management. The Utah Division of Wildlife suitable habitat or the distance of Bryce Canyon Resources Conservation Data Center (UTCDC) NP to the known or expected range of each spe- lists 50 species from Bryce Canyon NP as species cies (Appendix C). of concern based on state or range-wide rarity or lack of suffi cient data (UDWR 1998). Barkley’s The confi rmed and reported fl ora of Bryce Can- groundsel (Senecio werneriifolius var. barkleyi) is yon NP represents 16.3% of the 3,594 native and a local endemic that was described as a new taxon naturalized taxa documented for the state of Utah by Welsh et al. (2003), but currently is not tracked by Welsh et al. (2003) and 48.3% of the state’s 151 by the state’s natural heritage program and has no plant families. Within Garfi eld and Kane coun- formal legal status. ties, the park fl ora captures 34.1% of the 1,719 taxa reported by Welsh et al. (2003). At the ecore- Peabody (1995) conducted an extensive survey of gional scale, Bryce Canyon NP contains 36.5% rare plants of Bryce Canyon NP, focusing primar- of the 1,608 species reported for the Utah High ily on species being considered for listing under Plateaus (Shultz et al. 2006). the U.S. Endangered Species Act. In addition to plotting locations and describing habitat re- 3.2 Life form and biogeographic quirements, Peabody also established long-term monitoring plots to assess changes in population diversity size. Schelz (1990) compiled information on rare Although ecologically dominant, tree and shrub plants of Bryce Canyon NP from herbarium re- species make up only 15.7% of the confi rmed cords and data from the state’s natural heritage and reported plant taxa of Bryce Canyon NP program. Hallsten and Roberts (1990) conducted (Table 3.1). Perennial forbs are the most species- a detailed study of the autecology of the Paria rich life form in the park’s fl ora, with 339 known breadroot (Pediomelum pariense). and reported taxa (57.8% of the total fl ora). Only 12.1% of the fl ora is composed of annual forbs 3.4 Non-native and noxious weeds and grass-like plants. Like trees and shrubs, perennial graminoids are ecologically important Introduced plant species (not historically native but relatively species-poor, representing less than to Utah or North America) make up 10.2% of 14% of the total fl ora. The diversity of ferns and the confi rmed and reported fl ora of Bryce Can- fern-allies in Bryce Canyon NP is particularly low yon National Park (Table 3.1). This fi gure is lower (0.9%) (Table 3.1). than the statewide average of 13.5% non-native species (Fetig 2007, Welsh et al. 2003). Of the 60 Of the park’s 527 native taxa, 441 (83.7%) are wide- introduced plant species confi rmed or reported spread across Utah and western North America. for Bryce Canyon NP, seven are listed as noxious Only 22 species (4.2% of the park’s total native by the State of Utah (State of Utah Department of fl ora) have extremely limited global distributions Agriculture and Food 2007) (Table 3.4). Four of (local endemics found within an area of less than the park’s noxious weed species were discovered

12 Annotated Checklist of Vascular Flora: Bryce Canyon National Park during a survey of non-native plants by Beth Buchanan, Andersen (2005) also mapped invasive plants in 2004 and Karen Kitchen, and Juli Ames-Curtis in 1998. Dewey and reported one new noxious species.

Table 3.3. Plant species of conservation concern from Bryce Canyon National Park. TNC UTCDC Family Species Common name global Federal legal status status rank Asclepiadaceae Asclepias hallii Hall’s milkweed G3 Watch, S1 Boraginaceae *Cryptantha cinerea sand cryptanth G5T3? USFWS: former C2 Watch, S3? var. arenicola Boraginaceae Cryptantha ochroleuca yellowish G1? USFS Region 4: Sensitive; Rare, S1? cryptanth USFWS: former C2 Caryophyllaceae Silene petersonii Peterson’s campion G2G3 USFS Region 4: Sensitive; Watch, S2S3 USFWS: former C2 Compositae Artemisia ludoviciana Louisiana G5T5 Periph., S1 (Asteraceae) ssp. candicans wormwood Compositae Aster wasatchensis var. Markagunt aster G3?TNR Watch, S3? (Asteraceae) wasatchensis Compositae Chrysopsis jonesii Jones’ golden- G2 USFWS: former 3C Rare, S2 (Asteraceae) aster Compositae Erigeron abajoensis Abajo daisy G1G2 USFS Region 4: Sensitive Add. Data, (Asteraceae) S1S2 Compositae Erigeron canus hoary daisy G4 Periph., S2? (Asteraceae) Compositae Erigeron pumilus var. vernal daisy G4G5 Infreq., S2? (Asteraceae) condensatus T3T4 Compositae Haplopappus spindly G4TNR USFWS: former 3C Watch, S2S3 (Asteraceae) scopulorum var. goldenbush hirtellus Compositae Haplopappus zionis Cedar Breaks G2 BLM (UT): Sensitive; Rare, S2 (Asteraceae) goldenbush USFWS: former C2 Compositae Heterotheca jonesii Jones’ golden- G2 USFS Region 4: Sensitive; Rare, S2 (Asteraceae) aster USFWS: former 3C Compositae Senecio hartianus Hart’s groundsel G3G4 Add. Data, S1 (Asteraceae) Compositae Senecio werneriifolius Barkley’s groundsel G5TNR (Asteraceae) var. barkleyi Compositae Sphaeromeria capitata cluster-head G3 USFS Region 4: Sensitive Periph., S1 (Asteraceae) chicken-sage Compositae Townsendia montana Bryce Canyon G4T3 USFWS: former 3C Watch, S3 (Asteraceae) var. minima townsendia Compositae Viguiera longifolia var. southern G4G5 TNR Periph., S2? (Asteraceae) annua goldeneye Cruciferae Arabis demissa nodding rockcress G5T4 USFWS: former 3C Watch, S2? (Brassicaceae) Cruciferae Draba subalpina Breaks draba G3 USFWS: former 3C Watch, S3 (Brassicaceae) Cruciferae Lepidium montanum Claron pepperwort G5?T1? BLM (UT): Sensitive Rare, S1? (Brassicaceae) var. claronense Cruciferae Physaria arizonica Arizona G3? Watch, S2 (Brassicaceae) bladderpod

Chapter 3: Results 13 Table 3.3. Plant species of conservation concern from Bryce Canyon National Park, cont. TNC UTCDC Family Species Common name global Federal legal status status rank Cruciferae Physaria chambersii var. lepidote twinpod G3T2? Rare, S2? (Brassicaceae) membranacea Cruciferae Physaria chambersii var. Claron twinpod G4TNR Tax. Prob., (Brassicaceae) sobolifera S2? Cruciferae Physaria rubicundula Breaks bladderpod G3 USFWS: former 3C Watch, S3 (Brassicaceae) var. rubicundula Cruciferae Thelypodiopsis Palmer’s thelypody G4T2 USFWS: former 3C Rare, S2 (Brassicaceae) sagittata var. ovalifolia Cyperaceae Carex oederi var. green sedge G5 Periph., S2 viridula Hydrophyllaceae Phacelia Nipple Bench G3 USFWS: former 3C Watch, S3 mammillariensis phacelia Labiatae Hedeoma nanum dwarf pennyroyal G5 Periph., S1 (Lamiaceae) Leguminosae Astragalus hallii var. Hall’s milkvetch G4T4 Periph., S1 (Fabaceae) fallax Leguminosae Astragalus wardii Ward’s milkvetch G3 Watch, S3 (Fabaceae) Leguminosae Lupinus kingii var. King’s lupine G3G4 TNR Tax. Prob., S1 (Fabaceae) argillaceus Leguminosae Lupinus sericeus var. Marysvale lupine G5T3 Tax.Prob., S3 (Fabaceae) marianus Leguminosae Oxytropis oreophila Jones’ locoweed G5T3 USFWS: former 3C Watch, S3 (Fabaceae) var. jonesii Leguminosae Pediomelum pariense Paria breadroot G2G3 USFS Region 4: Sensitive; Watch, S2S3 (Fabaceae) USFWS: former C2 Liliaceae Leucocrinum star-lily G5 Periph., S2 (Anthericaceae) montanum Linaceae Linum australe small yellow ﬂ ax G5 Periph., S2 Polemoniaceae Phlox gladiformis daggerleaf phlox G3? USFWS: former 3C Watch, S2S3 Polygonaceae *Eriogonum aretioides Widtsoe wild G2 USFS Region 4: Sensitive; Rare, S2 buckwheat USFWS: former C2 Polygonaceae Eriogonum kidney-shaped G3G4 Watch, S3? subreniforme wild buckwheat Primulaceae Primula incana silvery primrose G4G5 Periph., S1 Pyrolaceae Pyrola picta whiteveined G4G5 Infreq., S1 (Ericaceae) wintergreen Rosaceae *Potentilla concinna modest cinquefoil G5?T4T5Q Tax. Ques., S? var. modesta Rosaceae Potentilla plattensis Platte River G4 Periph., S1 cinquefoil

Scrophulariaceae Castilleja parvula var. Reveal’s paintbrush G2 USFS Region 4: Sensitive; Rare, S2 revealii USFWS: former C2 Scrophulariaceae Orthocarpus violet owl-clover G3G4 Periph., S1S2 purpureoalbus Scrophulariaceae Penstemon bracteatus Red Canyon G2 USFS Region 4: Sensitive; Rare, S2 penstemon USFWS: former C2

14 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Table 3.3. Plant species of conservation concern from Bryce Canyon National Park, cont. TNC UTCDC Family Species Common name global Federal legal status status rank Scrophulariaceae Penstemon caespitosus painted G5T3? Watch, S2? var. desertipicti penstemon Scrophulariaceae Penstemon procerus smallﬂ ower G5T3? Add. Data, var. aberrans penstemon S3? Umbelliferae Cymopterus minimus least spring-parsley G1G2Q USFS Region 4: Sensitive; Rare, S1S2 (Apiaceae) USFWS: former C2 Umbelliferae Lomatium minimum least lomatium G3 USFWS: former 3C Watch, S3 (Apiaceae) This list is derived from the Utah Division of Wildlife Resources (1998) for UTCDC status.

All species are conﬁ rmed as present in the park except those indicated with an * (reported).

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Utah under the column UTCDC status.

Under legal status, USFWS = US Fish and Wildlife Service. C2 = category 2 candidate (a former category used for taxa that might warrant being pro- posed for Threatened or Endangered status following additional research). 3C = category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. BLM = U.S. Bureau of Land Management.

UTCDC status includes conservation categories adopted by the state natural heritage program to prioritize endemic and rare plant taxa (Utah Division of Wildlife Resources 1998). These categories include: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western US), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species appear rare, but more information needed before being assigned to a different category).

Table 3.4. Noxious weed species of Bryce Canyon National Park. Family Species Common name Park status Compositae (Asteraceae) Carduus nutans musk thistle Present Compositae (Asteraceae) Centaurea maculosa spotted knapweed Present Compositae (Asteraceae) Centaurea repens Russian knapweed Present Compositae (Asteraceae) Cirsium arvense Canada thistle Present Convolvulaceae Convolvulus arvensis ﬁ eld bindweed Present Cruciferae (Brassicaceae) Cardaria draba whitetop Present Euphorbiaceae Euphorbia esula leafy spurge Reported

This list is derived from the State of Utah Department of Agriculture and Food (2007) and Dewey and Andersen (2005).

Chapter 3: Results 15 16 Annotated Checklist of Vascular Flora: Bryce Canyon National Park ing season, or occur in undersampled wetland or 4 Discussion weedy habitats. A large number of weedy species that are known for the park from specimens in Since the publication of Weight’s 1933 checklist the resource management collection are not yet of 286 species, the confi rmed and reported fl ora represented in the offi cial herbarium and should of Bryce Canyon National Park has more than be the target of additional collection eff orts. Con- doubled to 587 taxa (Table 3.1, Figure 1.3.4, Ap- fi rmation of reported species should remain a pendix A). Based on available habitats and prox- high priority, as some of these taxa may ultimately imity, at least another 298 plant taxa may occur prove to be erroneous. Additional studies would within the park (Appendix B). Should all of these also be useful to better refi ne the abundance potential species be found, the documented fl ora and distribution of species of high management of the park would increase by nearly 33%. Many interest (especially rare plants and invasive non- of the “missing” potential or reported taxa con- natives). sist of cryptic species that fl ower late in the grow-

Chapter 4: Discussion 17 18 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Buchanan, H., and G. T. Nebeker. 1971. Check- 5 Literature Cited list of higher plants of Bryce Canyon Na- tional Park, Utah. Ogden, Ut.: Weber State Albee, B. J., L. M. Shultz, and S. Goodrich. 1988. College Printing Dept. 29 pp. Atlas of the vascular plants of Utah. Oc- Chronic, H. 1988. Pages of stone: Geology of casional Publ. 7, Utah Museum of Natural western national parks and monuments: History, Salt Lake City. 670 pp. Grand Canyon and the Plateau Country. Barkworth, M. E., K. M. Capels, S. Long, and M. Seattle, Wa.: The Mountaineers. 158 pp. B. Piep, eds. 2003. Magnoliophyta: Com- Cronquist, A. 1994. Volume 5, Asterales. In melinidae (in part): Poaceae, part 2. Flora Cronquist, A., A. H. Holmgren, N. H. of North America north of Mexico, Volume Holmgren, J. L. Reveal, and P. K. Holmgren, 25. New York: Oxford University Press. 783 eds. Intermountain fl ora: Vascular plants pp. of the Intermountain West, U.S.A. Bronx, Barkworth, M. E., K. M. Capels, S. Long, L. N.Y.: New York Botanical Garden. 496 pp. K. Anderton, and M. B. Piep, eds. 2007. Cronquist, A., A. H. Holmgren, N. H. Holmgren, Magnoliophyta: Commelinidae (in part): and J. L. Reveal, eds. 1972. Volume 1: Geo- Poaceae, part 1. Flora of North America logical and botanical history of the region, north of Mexico, Volume 24. New York: its plant geography and a glossary. The Oxford University Press. 911 pp. vascular cryptogams and the gymnosperms. Barneby, R. C. 1989. Volume 3, Part B. Fabales. Intermountain fl ora: Vascular plants of the In Cronquist, A., A. H. Holmgren, N. H. Intermountain West, U.S.A. Bronx, N.Y.: Holmgren, J. L. Reveal, and P. K. Holmgren, New York Botanical Garden. 270 pp. eds. Intermountain fl ora: Vascular plants Cronquist, A., A. H. Holmgren, N. H. Hol- of the Intermountain West, U.S.A. Bronx, mgren, J. L. Reveal, and P. K. Holmgren, N.Y.: New York Botanical Garden. 279 pp. eds. 1977. Volume 6, The monocotyledons. Belknap, W. 1958. Nature carves fantasites in Intermountain fl ora: Vascular plants of the Bryce Canyon. National Geographic Maga- Intermountain West, U.S.A. Bronx, N.Y.: zine 114(4):490–511. New York Botanical Garden. 584 pp. Buchanan, H. 1960. The plant ecology of Bryce ——. 1984. Volume 4: Subclass Asteridae (except Canyon National Park. PhD dissertation, Asteraceae). Intermountain fl ora: Vascular University of Utah, Salt Lake City. 136 pp. plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. ——. 1971. Changing conditions in the forests of 573 pp. Bryce Canyon National Park, Utah. Utah Academy Proceedings 48:31–36. Cronquist, A., N. H. Holmgren, and P. K. Holmgren, eds. 1997. Volume 3, Part A: ——. 1992. Wildfl owers of southwestern Utah: A Subclass Rosidae (except Fabales). In- fi eld guide to Bryce Canyon, Cedar Breaks termountain fl ora: Vascular plants of the and surrounding plant communities. Bryce Intermountain West, U.S.A. Bronx, N.Y.: Canyon National Park, Ut.: Bryce Canyon New York Botanical Garden. 446 pp. Natural History Association. 119 pp. Davis, G. H., and G. L. Pollock. 2003. Geology of Buchanan, H., and R. Graybosch. 1981. Revised Bryce Canyon National Park. Pages 37–60 checklist of the vascular plants of Bryce in Sprinkel, D. A., T. C. Chidsey, Jr., and P. Canyon National Park. Great Basin Natu- B. Anderson, eds. Geology of Utah’s parks ralist 41(1):109–120. and monuments. Utah Geological Associa- Buchanan, H., R. A. Graybosch, and K. T. Harp- tion Publication 28 (second edition). er. 1981. Plant community changes at Bryce Dewey, S., and K. Andersen. 2005. An inven- Canyon National Park from 1957 to 1980. tory of invasive non-native plants in Bryce Final report to the National Park Service, Canyon National Park (2004): Final report. Contract # CX-1200-8-B016. 68 pp. Logan, Ut.: Utah State University. Weed Sci- ence Research Project # SD0512A.

Chapter 5: Literature Cited 19 Dorn, R. D. 2001. Vascular plants of Wyoming, north of Mexico. Volume 19. New York: third edition. Cheyenne, Wy.: Mountain Oxford University Press. 579 pp. West Publishing. 412 pp. ——. 2006b. Magnoliophyta: Asteridae, part 7: Evenden, A., M. Miller, M. Beer, E. Nance, S. Asteraceae, part 2. Flora of North America Daw, A. Wight, M. Estenson, and L. Cudlip. north of Mexico. Volume 20. New York: 2002. Northern Colorado Plateau Vital Oxford University Press. 666 pp. Signs Network and Prototype Cluster, plan ——. 2006c. Magnoliophyta: Asteridae, part 8: for natural resources monitoring: Phase Asteraceae, part 3. Flora of North America I report, October 1, 2002 [two volumes]. north of Mexico. Volume 21. New York: National Park Service, Northern Colorado Oxford University Press. 616 pp. Plateau Network, Moab, Ut. 138 pp + app. Hallsten, G. P., and D. W. Roberts. 1988. Addi- Fertig, W. 2006. Herbarium and fi eld studies of tions to the vascular fl ora of Bryce Canyon vascular plant fl ora of BRCA for National National Park, Utah. Great Basin Naturalist Park Service Inventory and Monitoring 48:352. Program. Investigator’s annual report, Na- tional Park Service. 2 pp. ——. 1990. Analysis of community composition and habitat of Pediomelum pariense (Fa- ——. 2007. Introduced and naturalized plants of baceae) in Bryce Canyon National Park. 14 Utah. Sego Lily 30(5):7–11. pp. Flora of North America Editorial Committee. Holmgren, N. H., P. K. Holmgren, and A. Cron- 1993. Pteridophytes and Gymnosperms. quist. 2005. Volume 2, Part B: Subclass Flora of North America north of Mexico, Dilleniidae. Intermountain fl ora: Vascular Volume 2. New York: Oxford University plants of the Intermountain West, U.S.A. Press. 475 pp. Bronx, N.Y.: New York Botanical Garden. ——. 1997. Magnoliophyta: Magnoliidae and 488 pp. Hamamelidae. Flora of North America Madsen, M. 2001. A fl oristic study of the Tertiary north of Mexico, Volume 3. New York: Claron Formation in the Paunsaugunt Oxford University Press. 590 pp. Region of southern Utah. Master’s thesis, ——. 2000. Magnoliophyta: Alismatidae, Are- Department of Botany and Range Science, cidae, Commelinidae (in part), and Zin- Brigham Young University, Provo, Ut. giberidae. Flora of North America north Margules, C. R., and R. L. Pressey. 2000. Sys- of Mexico, Volume 22. New York: Oxford tematic conservation planning. Nature University Press. 352 pp. 405:243–253. ——. 2002a. Magnoliophyta: Commelinidae (in National Park Service. 2000. Management poli- part): Cyperaceae. Flora of North America cies. Washington, D.C.: U.S. Department of north of Mexico, Volume 23. New York: Interior, National Park Service. Publication Oxford University Press. 608 pp. # NPS D1416. 137 pp. ——. 2002b. Magnoliophyta: Liliidae: Liliales and Northern Colorado Plateau Network (NCPN). Orchidales. Flora of North America north in ed. Bryce Canyon National Park vegeta- of Mexico, Volume 26. New York: Oxford tion classifi cation and mapping project. University Press. 723 pp. Natural Resource Technical Report. Na- ——. 2003. Magnoliophyta: Caryophyllidae, part tional Park Service, Ft. Collins, Co. 1. Flora of North America north of Mexico. Noss, R. F., and A. Y. Cooperrider. 1994. Saving Volume 4. New York: Oxford University nature’s legacy: Protecting and restoring Press. 559 pp. biodiversity. Washington, D.C.: Island Press. ——. 2005. Magnoliophyta: Caryophyllidae, part 416 pp. 2. Flora of North America north of Mexico. Peabody, F. J. 1995. Rare plants of Bryce Canyon Volume 5. New York: Oxford University National Park, Utah: Final teport. De- Press. 656 pp. partment of Biology, University of South ——. 2006a. Magnoliophyta: Asteridae, part 6: Dakota, Vermillion. Asteraceae, part 1. Flora of North America

20 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Roberts, D. W., D. W. Wight, and G. P. Hallsten. Van Pelt, N., and C. D. Schelz. 1991. Survey and 1992. Plant community distribution and analysis of relict plant communities within dynamics in Bryce Canyon National Park. National Park Service units of the Colorado PX 1200-7-0966. 210 pp. Plateau. Cooperative Agreement 1440-8- 8001. National Park Service, Rocky Moun- Schelz, C. D. 1990. Bryce Canyon National Park tain Regional Offi ce, Lakewood, Co. 155 pp. rare plants. Report prepared for the Nation- al Park Service. 200 pp. Weber, W. A., and R. C. Wittmann. 2001. Colo- rado fl ora: Western Slope, third edition. Shultz, L. M., R. D. Ramsey, and W. Lindquist. Boulder: University Press of Colorado. 488 2006. Revised atlas of Utah plants. College pp. of Natural Resources, Utah State University, Logan, Ut. http://earth.gis.usu.edu/plants. Weight, K. E. 1933. Checklist of the fl owering plants of Bryce Canyon National Park. NPS Spence, J. R., and H. Buchanan. 1993. 1993 pamphlet. update: Checklist of the vascular plants of Bryce Canyon National Park, Utah. Great Welsh, S. L. 1982. Utah plant types: Historical Basin Naturalist 53(3):207–221. perspective, 1840–1981: Annotated list and bibliography. Great Basin Naturalist State of Utah Department of Agriculture and 42:1–44. Food. 2007. Utah noxious weed list. http:// ag.utah.gov/plantind/noxious_weeds.html. Welsh, S. L., N. D. Atwood, S. Goodrich, and L. C. Higgins. 2003. A Utah fl ora, third edi- Topp, S. 2007. Veg mappers fi nd “new” plants for tion, revised. Provo, Ut.: Brigham Young Utah national parks. Sego Lily 30(2):8–9. University. 912 pp. Utah Division of Wildlife Resources (UDWR). Welsh, S. L., N. D. Atwood, and J. R. Murdock. 1998. Inventory of sensitive species and 1978. Kaiparowits fl ora. Great Basin Natu- ecosystems in Utah. Endemic and rare ralist 38:125–179. plants of Utah: An overview of their distribution and status. Report prepared for the Utah Reclamation Mitigation and Conser- vation Commission and U.S. Department of the Interior. 566 pp + app.

Chapter 5: Literature Cited 21 22 Annotated Checklist of Vascular Flora: Bryce Canyon National Park 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Appendices Holmgren et al. 2005; Weber and Wittmann 2001). No ef- fort has been made to identify additional synonyms from The following appendices contain annotated checklists of monographs and fl oras preceding the early 1970s. the vascular fl ora of Bryce Canyon National Park. Appen- dix A lists all species that have been confi rmed for the fl ora Common name (Appendices A, B, C, and D) with an authenticated voucher specimen (including extant Common names are derived primarily from Welsh et al. and historical records) or which have been reliably report- (2003) and the Intermountain Flora (Barneby 1989; Cron- ed for the park in the literature (Buchanan and Graybosch quist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Hol- 1981; Dewey and Andersen 2005; Hallsten and Roberts mgren et al. 2005). 1988; NCPN in ed.; Spence and Buchanan 1993). Vouch- ers were examined from the BRCA herbarium, including Life form (Appendices A, B, C, and D) the separate resource management (BRCA-RM) reposi- Life form is based on the stature and growth habit of typi- tory), and collections from Brigham Young University cal members of a species at reproductive maturity. Trees (BRY), Utah State University (USU), the University of Wy- are woody perennials with a single stem or trunk and are oming (RM), and the digital museum database of the New typically over 3.5 m tall. Shrubs are woody perennials York Botanical Garden (NY). Species entries in Appendix with one to many trunks arising from near the base of the A are organized alphabetically by family and genus. plant and are usually less than 3.5 m tall. Perennial Forbs Appendix B includes species that may potentially occur (PerF) are broad-leaved dicots or monocots with non- within Bryce Canyon National Park based on the proxim- woody stems (at least above ground level) that live for mul- ity of the park to other populations or the presence of suit- tiple years. Annual Forbs (AnnF) are broad-leaved dicots able habitat. or monocots with non-woody stems that complete their life cycle (mature, fl ower, and die) in one year. Perennial Appendix C lists species that have been previously report- Graminoids (PerG) are linear-leaved, grass-like monocots ed for Bryce Canyon National Park but are now consid- that live for multiple years. Annual Graminoids (AnnG) ered erroneous because they are based on misidentifi ed are linear-leaved, grass-like monocots that complete their specimens or questionable literature records. life cycle (mature, fl ower, and die) in one year. Ferns and fern-allies (Fern) are non-fl owering vascular plants that Appendix D contains the same confi rmed, historical, and reproduce by spores. reported taxa as in Appendix A, but organized by life form rather than by family in order to better meet the needs of Range (Appendices A, B, and C) NPS interpretive specialists. Range represents the distribution of a species within Utah and the state’s contribution to the plant’s overall global The codes and references for each fi eld in the appropriate range. Disjunct (Disj) taxa have their Utah distribution appendices are summarized below. separated from the main, contiguous portion of their range by a gap of more than 800 km (ca 500 miles). Intro- Family (Appendices A, B, C, and D) duced (Intro) taxa are those that are not native to Utah Family concepts and nomenclature follow A Utah Flora, or North America but have become naturalized (breeding third edition (Welsh et al. 2003). Alternative family names on their own without human assistance). Local Endemics and treatments from other regional fl oras (Dorn 2001; (LocEn) are taxa whose entire global range is restricted Flora of North America Editorial Committee 1993, 2002b; to an area of less than 16,500 km2 (ca 6,370 mi2, or 1° of Holmgren et al. 2005; Weber and Wittman 2001) are listed latitude × 2° of longitude). Peripheral (Periph) taxa have a in parentheses. widespread global distribution but occur at the margin of their contiguous range in Utah and occupy less than 5% Species name (Appendices A, B, C, and D) of the state’s area (usually only within a few miles of the Nomenclature for scientifi c names and taxonomic con- state border). Regional Endemics (RegEn) are taxa with cepts are derived from Welsh et al. (2003). Scientifi c names a global range of 16,500–250,000 km2 (an area about the are listed in alphabetical rather than phylogenetic order. size of the state of Wyoming). Sparse taxa occur widely across Utah or North America but their range within Utah Synonyms/Taxonomic notes (Appendices A, B, and C) is small and patchy, with populations restricted to special- Pertinent synonyms are included for species with alter- ized or uncommon habitats. Widespread (Wide) taxa have native scientifi c names in other recent state and regional global ranges exceeding 250,000 km2 and occur over at fl oras (Barkworth et al. 2003, 2007; Barneby 1989; Cron- least 10% of the state. quist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of North America Editorial Committee 1993,

Chapter 6: Appendices 23 Park status (Appendices A, C, and D) management collection (BRCA-RM), Brigham Young Park status identifi es whether a species is Present (Pres), University (BRY), Utah State University (USU), New York Historical (Hist), or Reported (Rep) for Bryce Can- Botanical Garden (NY), and the University of Wyoming’s yon National Park. Species are considered Present if a Rocky Mountain Herbarium (RM). confi rmed voucher or documented observation has been Year doc. (Appendix A) made within park boundaries since 1970. Historical species have been confi rmed with a voucher from the park but Year documented indicates the fi rst year in which a spe- have not been relocated since before 1970. Reported species was confi rmed or reported to occur in Bryce Canyon cies are listed for the park in the literature (Buchanan and National Park. This year does not necessarily correspond Graybosch 1981; Dewey and Andersen 2005; Hallsten and with the date of the voucher specimen cited under Source. Roberts 1988; NCPN in ed.; Spence and Buchanan 1993), “NA” indicates that the fi rst year is not known (used most- but have not been corroborated with a voucher specimen. ly for ‘reported’ taxa). Species that may potentially occur in Bryce Canyon NP are listed in Appendix B. Rejected taxa (listed in Appendix Comments (Appendices A, B, C, and D) C) have either been falsely reported (FalsRep) in the liter- This column includes additional comments on the status ature based on misidentifi ed herbarium specimens or are of a particular species in the park, the place of origin for considered Questionable (Ques?) because park reports introduced species, legal status, and other data. are from well outside their known or expected range. Nativity (Appendix D) Pop. size (Appendix A) Nativity indicates whether a species is native to Utah or Population size refers to the abundance and frequency Introduced (Intro) based on Fertig (2007). with which a species occurs within Bryce Canyon NP. Abundance data are derived from Northern Colorado Pla- Flower time (Appendix D) teau Network (in ed.), and personal observations. Abun- Flower Time refers to the months in which a species is dant (Abun) taxa have large populations, have a broad most likely to be in fl ower. NA indicates a species that is a ecological amplitude (often being dominant in one to non-fl owering fern or gymnosperm. several vegetation types), or occur extensively across the park. Common (Com) taxa have large local populations Habitat (Appendix D) but may be restricted to a single vegetation type or do not The distribution of species across generalized habitats occur across the entire park. Uncommon (Unc) taxa have in Bryce Canyon National Park is indicated by an “x” in low to medium-sized populations, typically have a narrow the columns Sage/Grs through Wet and is derived from ecological amplitude (or at least contribute relatively little the vegetation classifi cation of Buchanan (1992), Madsen to total vegetative cover), or have a small range across the (2001), and Welsh et al. (2003). Sage/Grs=black sage- park. Rare taxa usually have low population numbers and brush (Artemisia nova) grasslands and montane meadows are restricted to a single location within the park. A “?” in- dominated by perennial grasses and herbaceous species. dicates that population size is unknown. PinJun=Pinyon-Juniper woodlands of two-needle pinyon (Pinus edulis) and Utah juniper (Juniperus osteosperma). Source (Appendix A) MontConf=montane brush and conifer forests of ponde- Sources are the evidence for the assigned park status. In rosa pine (Pinus ponderosa), Gambel oak (Quercus gam- the case of Present and Historical taxa, sources are her- belii), Douglas-fi r (Pseudotsuga menziesii), and white fi r barium records (with collector, collection number, and (Abies concolor). Wet=Wetlands (including wet meadows, repository) that were corroborated during this study. For ponds, swampy forests, and seeps and springs). Additional reported taxa, the source is a published or unpublished comments are included for species that are restricted to reference citing the species within the park. Specimen a particular soil/geologic type or are found mostly in dis- repositories include the Bryce Canyon National Park her- turbed sites. barium (BRCA) Bryce Canyon National Park resource

24 Annotated Checklist of Vascular Flora: Bryce Canyon National Park c name with Helper, Carbon Helper, Co., UT (Trelease s.n. MO) Comments doc. Year Year 1934 1981 1980 2007 1980 locality is Type 1932 1932 1931 1958 1932 2005 1931 1928 (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide PresPerF Unc Wide Buchanan 245 Pres Unc Graybosch 1361 AnnF Wide Hist ? B-32/295 Weight Shrub Wide Pres Com Graybosch 770 Shrub RegEn Pres Unc Buchanan 156 Shrub Wide Pres Com Graybosch 1354 Life form spreading dogbane common dogbane spider milkweed PerF Wide Pres Unc Graybosch 957 datil yucca Shrub Wide Presprostrate Rarepigweed ST09190702 Topp squawbush Shrub Wide Pres Com Graybosch 851 Harriman’s yuccaHarriman’s Shrub Wide Pres Unc Graybosch 932 Rocky Mountain maple bigtooth maple Treenarrowleaved Wideyucca Pres Unc Buchanan 371 Hall’s milkweedHall’s PerFshowy milkweed Sparse PerF Fremont’s Presmahonia Wide RareOregon-grape Pres ST08140506 Topp Rare Shrubwater birch Wide Gottschalk 6 (UTC)low cryptanth 1986 Pres Tree PerF Unc Wide RegEn Buchanan 136 Pres Pres Unc Unc Boyle 1117 (BRCA) Boyle 1155 (BRCA) 1932 1931 Common name

var.

var. var. & angustifolium Includes vars. cannabinum, glaberrimum, baccata A. graecizans R. trilobata trilobata angustissima tripartitum Includes vars. neomexicanum, douglasii, Synonyms/ notes Taxonomic

var.

var. var. asperula harimaniae androsaemifolium Apocynum cannabinum Asclepias asperula var. Asclepias hallii Yucca harrimaniae Yucca var. Amaranthus blitoides Rhus aromatica trilobata Apocynum androsaemifolium Acer grandidentatum angustissima Yucca angustissima Y. baccataYucca baccata Y. glabrum Acer glabrum Asclepias speciosa Mahonia repensBetula occidentalis Berberis repens B. fontinalis Mahonia fremontii Berberis fremontii Cryptantha abata vascular plant taxa of Bryce Canyon National Park, organized by family and scientiﬁ rmed, historical, and reported Apocynaceae Asclepiadaceae Asclepiadaceae Agavaceae Amaranthaceae Anacardiaceae Apocynaceae Aceraceae Agavaceae Agavaceae FamilyAceraceae Species name Asclepiadaceae Berberidaceae Betulaceae Berberidaceae Boraginaceae Appendix A. Conﬁ taxonomic and biogeographic notes.

Chapter 6: Appendices 25

eld Co., UT echinoides BRY) is from BRY) “near Bryce Canyon NP”. locality is Type Lone Pine Point, Johnson Canyon, Kane Co., UT (Higgins, Welsh, & Thorne 14297 USFWS: BRY). former C2 candidate Comments var. var. is Paria Canyon, Garfi (Jones 5297p POM) NA Foster (10866 doc. Year Year 1934 1980 1980 1932 1980 1932 locality of Type 1957 c name with taxonomic and (BRCA) (1991) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source (BRCA) Pop. size Park status Range PerF WideAnnF Pres Wide Unc Hist Graybosch 940 ? B-32/300 Weight Life form PerF RegEn Hist ? B-32/203 Weight PerF Wide Pres Unc Graybosch 758 Baker’s cryptanthBaker’s PerF RegEnsand cryptanth Pres PerF Unc LocEn Graybosch 756 Rep ? Pelt and Schelz Van Common name ashy cryptanth PerFbasin yellow Widecryptanth Pressand dune cryptanth Unc Graybosch 966 plateau cryptanth plateau yellow cryptanth dwarf cryptanth PerF Wide Pres Unc Buchanan 3046 ava C. atwoodii nana Includes Synonyms/ notes Taxonomic Oreocarya fulvocanescens, includes var. echinoides. ssp. avafl Oreocarya ora arenicola cinerea Cryptantha bakeri Oreocarya bakeri Cryptantha cinerea var. Cryptantha cinerea var. Cryptantha confertifl Cryptantha fendleri Cryptantha fl Cryptantha fulvocanescens Cryptantha humilis Oreocarya humilis rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Boraginaceae Appendix A. Confi FamilyBoraginaceae Species name biogeographic notes, cont. Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae

26 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Co., eld Red Canyon, Garﬁ UT (Higgins 1788 BRY). USFS: Region 4 Sensitive, USFWS: former C2 candidate Fish Lake, Sevier s.n. Co., UT (Ward GH) Comments doc. Year Year 1977 1934 1956 1940 1932 locality is Type 1931 locality is Type c name with taxonomic and 5529 (BRY) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Rare Foster 10796 (BRY) 1993 PerF Wide Pres Com Boyle 1114 (BRCA) 1938 PerF LocEn Pres Com Graybosch 891 PerF Intro Pres Unc Heyn s.n. (BRCA) 1998 Native to Eurasia Life form Arizona pincushion aspen bluebell PerF Wide Pres Unc Foster & western stickseed AnnF Wide Pres Unc Buchanan 70 cupseed stickseed AnnF Wide Pres Unc ST06040601 Topp showy stoneseed PerF Widesouthwestern stoneseed Pres Unc Shepherd s.n. yellowish cryptanth bristly cryptanth PerF RegEn Prescommon Unchound’s-tongue Buchanan 2974 claretcup PerF Wide Pres Unc Foster 11077 (BRY) 1994 Common name

Flora

Vars. Vars. var. C. by some var. Vars. not Vars. (2003) arizonica. Flora of North Escobaria vivipara var. not recognized in America of North America Includes var. Includes var. inermis. recognized in L. redowskii redowskii L. texana, redowskii cupulata, L. marginata Included in compacta authors Synonyms/ notes Taxonomic (2003).

var.

var. orum occidentalis cupulata cinale Echinocereus triglochidiatus melanacanthus Coryphantha vivipara arizonica Mertensia arizonica Lappula occidentalis var. Lithospermum incisum Lithospermum multifl Lappula occidentalis var. Cryptantha ochroleuca Cryptantha setosissima Cynoglossum offi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Cactaceae Cactaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Appendix A. Confi FamilyBoraginaceae Species name biogeographic notes, cont. Boraginaceae Boraginaceae

Chapter 6: Appendices 27 is Green River, is Green River, Emery Co., UT (Jones s.n. POM) Comments doc. Year Year 1931 2006 locality Type 1932 1931 1987 1980 2005 2006 1932 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (UTC) (BRCA) Plateau Network (in ed.) (BRCA) (BRCA) Source Pop. size Park status Range AnnF RegEn Pres Rare ST07280602 Topp PerF Wide Pres Unc Hallsten 2454 PerF Wide Pres Unc Graybosch 791a PerF Wide Hist ? B-32 /234 Weight Shrub Wide Pres Com Boyle 1132 (BRCA) 1931 Life form ower shhook cactus shhook blue elderberry Shrub Wide Pres Unc Buchanan 183 Palmer’s Palmer’s stinkweed yellow beeplant AnnF Wide Hist ? B-32 /238 Weight Parry’s harebellParry’s PerF Wide Pres Unc Graybosch 1043 small-ﬂ ﬁ Simpson’s Simpson’s hedgehog cactus plains pricklypear PerF Wide Rep Unc Northern Colorado brittle pricklypear PerF Wide Pres Rare ST06140601 Topp common pricklypear Common name mountain snowberry

var.

minor

parryi

var.

var. & (2003) ssp. orus, & spina simpsonii caerulea, S. cerulea S. parviﬂ S. whipplei intermedius Includes vars. & Flora of North America includes vars. ruﬁ trichophora O. polyacantha erinacea Synonyms/ notes Taxonomic S. rotundifolius, includes vars. parishii oreophilus according to some authors

var. roseus var. var. var. erinacea Sambucus caerulea S. nigra Cleomella palmeriana palmeriana Cleome lutea Campanula parryi C. parryi Sclerocactus whipplei Pediocactus simpsonii polyacantha Opuntia polyacantha Opuntia fragilis Opuntia erinacea var. utahensis Symphoricarpos oreophilus rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Caprifoliaceae Caprifoliaceae (Adoxaceae) Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Campanulaceae Cactaceae Cactaceae Appendix A. Conﬁ biogeographic notes, cont. Cactaceae Cactaceae FamilyCactaceae Species name

28 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld is Red Wasatch Plateau, Wasatch 1 mile above Baldy Ranger Station, Sanpete Co., UT (Maguire 20000 NY). Type locality of var. minor Canyon, Garﬁ Co., UT (Maguire 19550 NY). USFS: Region 4 Sensitive, USFWS: former C2 candidate Comments doc. Year Year 1957 1980 1980 locality is Type 1980 1938 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF RegEn Pres Unc Graybosch 1360 PerF Wide Pres Unc Buchanan 275 PerF Wide Pres Unc Graybosch 962 Life form y Drummond’s Drummond’s catchﬂ Peterson’s Peterson’s campion James’ chickweed PerF Wide Pres Unc Graybosch 1098 Creeping nailwort basin sandwort PerF Wide Pres Com Buchanan 2045 Common name

& S. minor &

includes & ssp. var. striata, drummondii striata, glabrescens, drummondii, petersonii kruckebergii, & Gastrolychnis drummondii vars. Includes vars. & clokeyi jamesiana Eremogone kingii var. E. kingii uintahensis, A. kingii uintahensis, compacta, plateauensis ssp. ssp. Synonyms/ notes Taxonomic ora glabrescens Paronychia sessilifl Silene petersonii Stellaria jamesiana Pseudostellaria Arenaria fendleri var. drummondiiLychnis Silene drummondii rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae Appendix A. Confi FamilyCaryophyllaceae (Alsinaceae) Species name biogeographic notes, cont.

Chapter 6: Appendices 29 Indian Creek, Mtns, Tushar Beaver Co., UT (Martin & Welsh 17148 BRY) Comments doc. Year Year 1980 1931 1980 2006 1980 2006 2006 Native to Eurasia 2005 Native to Eurasia 1931 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) EB07270601 (BRCA) (BRCA) (BRCA) (BRCA) Plateau Network (in ed.) (BRCA) (BRCA) Source Pop. size Park status Range AnnF Wide Hist ? B-31/137 Weight PerF Wide Pres Rare Graybosch 1114 Shrub Wide Pres Unc Graybosch 847 AnnF Wide Pres Unc Kitchen s.n. (BRCA) 1998 locality is Type Life form mountain-lover Shrub Wide Pres Unc Buchanan 132 basin saltbush Shrub Wide Pres Uncsummer-cypress Graybosch 946 AnnFlambsquarter Intro Pres AnnF Unc Intromountain goosefoot ST07280607 Topp Rep Unc Northern Colorado long-stalk starwort fourwing saltbush shadscale Shrub Wide Pres Unc orachPowell’s Ballenger AnnF Wide Pres Rare ST07280604 Topp smallhead goosefoot silvery goosefoot AnnF Wide Pres Unc Graybosch 955 Common name

Flora var.

var. var. ssp. in var. var. A. includes var. includes var. of North America Paxistima myrsinites, Pachistima myrsinites A. canescens A. tridentata, includes tridentata robusta sieversiana C. album C. fremontii atrovirens (2005) occidentalis monantha stricta, S. longipes Synonyms/ notes Taxonomic C. overi, foliosum C. incanum incanum, C. incanum occidentale

var.

var. album var. var. canescens utahensis longipes Pachystima myrsinites Atriplex canescens var. Atriplex confertifolia Atriplex gardneri var. Atriplex powellii powellii Bassia scopariaChenopodium Kochia scoparia, B. album Chenopodium atrovirens Chenopodium capitatum Stellaria longipes var. parvicapitatum Chenopodium fremontii incanum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Celastraceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix A. Conﬁ FamilyCaryophyllaceae (Alsinaceae) Species name biogeographic notes, cont. Chenopodiaceae

30 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 1978 2004 Native to Eurasia 1959 1931 Native to Asia 2006 2006 2006 1957 1980 1980 1980 1956 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) AT06290601 (BRCA) (BRCA) (BRCA) (BRCA) Graybosch (1981) (BRCA) (BRCA) (BRCA) Graybosch (1981) Source Pop. size Park status Range AnnF Wide Pres Unc Buchanan 2960 AnnF WideShrub Pres RegEn Rare Pres ST07280605 Topp Rare ST07280603 Topp Life form narrow-leaf goosefoot Russian thistle AnnF Intro Pres Torrey’s Uncseepweed ST07280608 Topp Plummer’s siltbush common yarrow PerF Wide Pres Unc Buchanan 254b halogetonwinterfat AnnF Intro Pres Shrub Wide Rare Pres ST07280609 Topp Uncgreasewood Buchanan 462 Shrub Wide Pres Unc Tendick orange agoseris PerFpurplish agoseris Wide PerF Rep Wide Unc Pres Buchanan & Unc Graybosch 1061 Common name cutleaf agoseris PerF Wide Pres Unc Buchanan 208 pale agoseris PerF Widelow pussytoes Pres PerF Unc Wide Graybosch 908 Rep ? Buchanan &

Vars. Vars. var.

Vars. Vars. var. var. ora (2006) (2003) Flora of North Flora of North pestifer, S. australis pestifer, S. moquinii, nigra, S. fruticosa Z. brandegeei plummeri, Grayia brandegei plummeri A. lanulosa, millefolium Ceratoides lanata, Eurotia lanata. occidentalis. not recognized in America not recognized in in America Synonyms/ notes Taxonomic A. parviﬂ

var. var. lanata var. lanulosa torreyana plummeri aurantiaca purpurea Chenopodium leptophyllum Halogeton glomeratus Krascheninnikovia lanata Salsola tragusSarcobatus vermiculatus S. kali, iberica, Suaeda torreyana var. Zuckia brandegei var. Achillea millefolium ssp. Agoseris aurantiaca var. Agoseris aurantiaca var. Agoseris glauca glauca Agoseris glauca laciniata Antennaria dimorpha rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Appendix A. Conﬁ FamilyChenopodiaceae Species name biogeographic notes, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae (Sarcobataceae) Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 31 Comments doc. Year Year 1942 2005 1959 1957 1956 1973 1998 Native to Eurasia 1980 1958 1980 1956 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Sparse Pres Unc ST08100502 Topp PerF Wide Pres Unc Buchanan 322 Shrub Wide Pres Unc Boyle 1167 (BRCA) 1942 PerF Wide Pres Unc Buchanan 369 PerF Wide Pres Unc Buchanan 16 Life form PerF Wide Pres Unc Buchanan 2951 Louisiana wormwood Louisiana wormwood Common name fringed sagebrush small-leaf pussyoes Chamisso’s arnicaChamisso’s PerF Wide Pres Unc Graybosch 1116 common pussytoes Breaks pussytoes PerFburdock Wide Pres Unc PerF Buchanan 3082 Intro Pres Unc Buchanan s.n. heartleaf arnica PerF Wide Pres Unc Graybosch 987 sand wormwood PerF Wide Pres Carruth’s wormwood Rare Graybosch 1101 tarragon PerF Wide Pres Unc Buchanan 525

var.

Flora Vars. Vars. ssp. in ssp. (2006) A. rosea cus, O. cus, cus cordata Flora of North Artemisia ludoviciana candicans Includes Synonyms/ notes Taxonomic A. chamissonis andina not Vars. recognized in of North America (2006). Misspelled var. Welsh et al. 2003 Welsh Oligosporus campestris paciﬁ paciﬁ Oligosporus dracunculus. not recognized in America

var.

var. var. var. foliosa scouleriana Artemisia ludoviciana latiloba Artemisia ludoviciana albula Antennaria microphylla Artemisia frigida Antennaria parvifolia Antennaria rosulata Arctium minus Arnica chamissonis var. Arnica cordifolia cordifolia Artemisia campestris var. Artemisia carruthii Artemisia dracunculus glauca rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name biogeographic notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

32 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 2006 1957 1931 1980 2005 1959 1932 1980 c name with taxonomic and AT07290601 AT07290601 (BRCA) (BRCA) (BRCA) (BRCA) Plateau Network (in ed.) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Shrub Wide Rep Unc Northern Colorado Shrub Wide Pres Unc Buchanan 540 Life form c aster PerF Wide Pres Com Boyle 1168 (BRCA) 1931 Siskiyou aster PerF Wide Preswestern aster Rare PerF Tendick Wide Pres Unc Buchanan 304 blueleaf aster PerF Wide Pres Unc Graybosch 1069 Paciﬁ asterEaton’s PerF Wide Pres Unc Graybosch 1070 Wyoming big sagebrush basin big sagebrush pygmy sagebrush Shrub Wide Pres Rare Graybosch 964 black sagebrush Shrub Wide Pres Com B-32/251 Weight Common name

var.

Flora

var. ssp. ssp. ssp. var. Vars. not Vars. hesperius, hesperium Symphyotrichum spathulatum var. Symphyotrichum lanceolatum glauca adscendens, A. adscendens, Symphyotrichum ascendens, Virgulaster ascendens eatonii, A. bracteolatus Eurybia glauca, Herrickia glauca wyomingense Seriphidium tridentatum tridentatum Artemisia arbuscula nova, Seriphidium novum. recognized in of North America pygmaeum Seriphidium tridentatum (2006) Synonyms/ notes Taxonomic

var. wyomingensis tridentata Aster spathulatus A. occidentalis, Aster hesperius A. lanceolatus Aster eatoniiAster glaucodes Symphyotrichum Eucephalus glaucus, Aster ascendens A. chilensis Artemisia tridentata var. Artemisia nova nova Artemisia pygmaea Seriphidium Artemisia tridentata var. rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name biogeographic notes, cont.

Chapter 6: Appendices 33 is Springdale, Co., Washington UT (Jones 5249u US). USFS: Region 4 Sensitive, USFWS: 3C American Fork, Jordan Valley, Utah Co., UT 493 YU) (Watson noxious weed. Native to Eurasia noxious weed. Native to Europe weed. Native to Eurasia Tate Mine near Tate Marysvale, Piute Co., UT (Jones 5861 POM). Comments doc. Year Year 1931 1980 locality Type 1959 locality is Type 1998 Utah state 1998 Utah state 1980 UT state noxious 1958 locality is Type 1940 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) Buchanan s.n. (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF LocEn Pres Rare Graybosch 924 Shrub Wide Pres Unc Buchanan 466 PerF Intro Pres Unc Kitchen & PerF Wide Pres Unc Boyle 1140 (BRCA) 1931 PerF Intro Pres Unc Graybosch 954 PerF Wide Pres Unc Buchanan 15 Life form rose-heath PerF Wide Pres Unc Buchanan 1122 Jones’ golden- aster Mohave brickellbush musk thistle PerFspotted Introknapweed Pres Unc Buchanan s.n. hoary dusty- maiden Russian knapweed Markagunt aster PerF LocEn Pres Unc Buchanan 403 arrowleaf balsamroot Common name

& micranthos, Leucelene ericoides, Aster arenosus Eurybia wasatchensis, Herrickia wasatchensis montana Includes ssp. leiophyllus macrolepis Centaurea stoebe ssp. Acosta maculosa Includes vars. douglasii Synonyms/ notes Taxonomic

var. wasatchensis Chaetopappa ericoides Chrysopsis jonesii Heterotheca jonesii linifolia Carduus nutans Centaurea maculosa C. biebersteinii, Centaurea repens Acroptilon repens Chaenactis douglasii Aster wasatchensis var. Balsamorhiza sagittata Brickellia oblongifolia rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name biogeographic notes, cont. Compositae (Asteraceae)

34 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 1931 2005 1931 2006 1950 2006 1980 1927 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Hutchings 2775 (BRY) Source Pop. size Park status Range Shrub Wide Pres Unc Buchanan 71 Shrub Sparse Pres Unc ST09070502 Topp Shrub Wide Pres Unc Boyle 1165 (BRCA) 1942 Shrub Wide Pres Unc Buchanan 2984 Shrub Wide Pres Unc ST07280601 Topp Shrub Wide Pres Com Fertig 23058 Shrub Wide Pres Com Fertig 23060 PerF Wide Pres Unc Graybosch 1090 Shrub Wide Pres Com Schelz s.n. (BRCA) 1989 Shrub Wide Hist ? Cottam & Life form lanceleaf rabbitbrush Parry’s Parry’s rabbitbrush Vasey’s Vasey’s rabbitbrush Parry’s Parry’s rabbitbrush spear-leaf spear-leaf rabbitbrush greenish rabbitbrush graystem rabbitbrush hispid golden- aster dwarf rabbitbrush glabrate rabbitbrush Common name

var.

var. var. included by some minor var. includes nis var. parryi afﬁ graveolens, hispida, hispida, H. oreophilus C. nauseosus Ericameria parryi var. Ericameria parryi var. Lorandersonia linifolia Ericameria nauseosa C. nauseosus Ericameria nauseosa var. E. nauseosa glabrata, consimilis, in hololeucus oreophilus authors C. villosa var. Heterotheca villosa var. villosa C. nauseosus var. according to some authors Synonyms/ notes Taxonomic

nis var. var. var. var. parryi affi orus var. var. minor lanceolatus Chrysothamnus viscidifl Chrysothamnus vaseyi Chrysothamnus parryi Chrysothamnus linifolius Chrysothamnus nauseosus consimilis Chrysothamnus nauseosus gnaphalodes Chrysothamnus nauseosus graveolens Chrysopsis villosa var. Chrysothamnus depressus Chrysothamnus parryi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) biogeographic notes, cont.

Chapter 6: Appendices 35 is nidulum var. var. Paria, Kane Co., UT (Jones 5290a POM). noxious weed. Native to Eurasia Comments doc. Year Year 2007 1998 Native to Eurasia 1956 locality of Type 1998 Utah state 1987 2005 1993 1959 1957 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (UTC) Plateau Network (in ed.) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Shrub Wide Pres Unc ST08250701 Topp PerF Wide Pres Rare Hallsten 2446 Shrub Wide Rep Rare Northern Colorado PerF Wide Pres Unc Graybosch 953 Life form green rabbitbrush Arizona thistle PerF Wide Pres Unc Graybosch 1068 chicory PerF Intro Pres Unc Buchanan s.n. Canada thistle PerF Intro Pres Unc Buchanan s.n. New Mexico thistle pretty thistle PerF Wide Pres Unc RS 9-21-04 (BRCA) 2004 slenderleaf rabbitbrush Tracy’s thistleTracy’s PerFwavy-leaved thistle RegEn Pres Rare Buchanan 512 elk thistle PerF Wide Presbull thistle Unc ST07110602 Topp PerF Intro Pres Unc Passek s.n. (BRCA) 1990 Native to Europe Common name

Flora Flora

by in & C.

var. var. & orus Includes vars. arizonicum nidulum bipinnatum Includes vars. horridum integrifolium, Breea arvensis not recognized Var. in Cronquist (1994). Included in arizonicum of North America (2006) Included in var. Included in var. viscidiﬂ some authors (2006) of North America Vars. not Vars. recognized in Synonyms/ notes Taxonomic C. tracyi C. undulatum

var.

var. var. orus orus orus pulchellum scariosum tracyi undulatum Cichorium intybus Cirsium arizonicum viscidifl Cirsium arvense Cirsium calcareum var. Cirsium neomexicanum neomexicanum Chrysothamnus viscidifl stenophyllus Chrysothamnus viscidifl Cirsium scariosum var. Cirsium undulatum var. Cirsium undulatum var. Cirsium vulgare rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) biogeographic notes, cont.

36 Annotated Checklist of Vascular Flora: Bryce Canyon National Park is Abajo Mtns, San Juan Co., UT (Rydberg & Garrett 9755 NY). USFS: Region 4 Sensitive Comments NA doc. Year Year 1959 1980 1927 locality Type 1980 1957 1940 1980 1957 c name with taxonomic and (BRCA) (BRCA) (BRY) Buchanan (1993) Graybosch (1981) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Graybosch 836 PerF Wide Rep Unc Buchanan & PerF Wide Pres Unc Graybosch 867 AnnF Wide Pres Unc Foster 11012 (BRY) 1993 Life form western hawksbeard gray hawksbeard PerF Wide Presglaucous Unchawksbeard Buchanan 461a Abajo daisy PerF LocEn Pres Unchairy daisy Atwood 6128 hoary daisy PerFcut-leaved daisy Wide PerF PerF Hist Disj Wide ? Rep Pres 6676 (BRY) Weight Unc ? 1935 Boyle 2289 (BRCA) Spence & 1931 hispidulous hawksbeard trailing daisy PerF Wide Pres Unc Buchanan 289 Wheeler’s thistleWheeler’s PerF Wide PresCanadian horseweed Com Boyle 1166 (BRCA) 1931 spreading daisy PerF daisyEaton’s Wide Pres PerF Unc Wide Graybosch 1095 Pres Unc Graybosch 838 Common name

Flora

Vars. Vars.

var. (2006) E. Flora of North Psilochenia occidentalis intermedia according to some authors Includes awapensis Vars. not Vars. recognized in of North America (2006) Includes var. Includes var. canadensis. not recognized in America Synonyms/ notes Taxonomic eatonii

var. var. var. agellaris occidentalis aphanactis glabratus glabrata divergens Crepis occidentalis var. Crepis runcinata glauca Erigeron aphanactis var. Crepis intermedia Psilochenia Erigeron canus Erigeron compositus var. Crepis runcinata hispidulosa Erigeron abajoensis Cirsium wheeleri wheeleri Conyza canadensis var. Erigeron eatoniiErigeron fl E. eatonii Erigeron divergens var. rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) biogeographic notes, cont. Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 37 Mount Tomasaki, Mount Tomasaki, La Sal Mtns, UT (Payson & Payson 4033 RM) Comments NA doc. Year Year 1932 1931 1980 1980 locality is Type 1956 1957 c name with taxonomic and Buchanan (1993) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Rep ? Spence & Shrub Wide Pres Com Buchanan 532 PerF Wide Hist ? Mitchell 16 (BRCA) 1946 Shrub Wide Pres Unc Buchanan 48 PerF Wide Pres Com Peabody 29 (BRCA) 1931 Life form eabane PerF Wide Pres Unc Foster 10775 (BRY) 1993 stemless goldenweed broom snakeweed vernal daisyaspen daisy PerF Widebear daisy PerF Pres Wide Unc Pres PerF s.n. (BRCA) Gifford daisyPayson’s Wide 1940 Com Buchanan 2915 Pres PerF Wide Unc Pres Graybosch 1120 curly-cup gumweed Unc Graybosch 1341 narrowleaf goldenweed longleaf daisy PerF Wide Pres Unc Buchanan 305 thrift goldenweed Navajo ﬂ Common name

var.

Flora Flora

var. var. var. includes var. subglaber linearis, canescens, acaulis Stenotus acaulis Vars. not Vars. recognized in of North America not Vars. recognized in of North America (2007) (2006) E. concinnus condensatus Trimorpha Trimorpha lonchophylla E. concinnus concinnus, var. H. macronema var. H. macronema var. Ericameria discoidea linearis, Macronema discoideum Synonyms/ notes Taxonomic Stenotus armerioides armerioides

var.

var. acaulis serrulata condensatus macranthus concinnus Gutierrezia sarothrae Haplopappus acaulis var. Erigeron ursinus ursinus Erigeron vagus Grindelia squarrosa var. Haplopappus macronema Erigeron pumilus var. Erigeron speciosus var. Erigeron lonchophyllus Erigeron pumilus var. Haplopappus armerioides armerioides rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name (Asteraceae) biogeographic notes, cont.

38 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Cedar Canyon, Iron Co., UT (Garrett 6051 US). USFWS: 3C Cedar Canyon, Iron Co., UT (Anderson, & Welsh, Chatterley 5094 UT BLM: BRY). Sensitive, USFWS: former C2 candidate 15 miles N of Orderville, Kane Co., UT (Maguire 18740 NY) Comments doc. Year Year 1959 locality is Type 1986 locality is Type 1977 locality is Type 1957 1978 1932 1931 c name with taxonomic and (BRCA) (BRY) 4363 (BRY) (BRCA) (BRY) (BRCA) (BRCA) Source Pop. size Park status Range Shrub RegEn Pres Unc Graybosch 1100 Shrub LocEn Pres Rare Gottschalk 10 PerF RegEn Pres Unc Foster & PerF Wide Pres Com Buchanan 209 AnnF Wide Pres Unc B-32/331 Weight PerF Wide Pres Com Graybosch 797 PerF Wide Pres Unc Peabody 13 (BRCA) 1991 Life form ower PerF Wide Pres Unc1022 Graybosch ower spindly goldenbush Cedar Breaks goldenbush little sunﬂ alpine hyalineherb Common name Arizona hymenoxys common sunﬂ common hyalineherb Great Basin hyalineherb

var. var. var. lifolius orus, (2006). arizonica paucifl lifolius lifolius lifolius Flora of North Hesperodoria scopulorum, Chrysothamnus scopulorum Synonyms/ notes Taxonomic Ssp. not recognized in America H. fi lugens, H. fi var. H. fi megacephalus, H. fi tomentosus Tetraneuris acaulis Tetraneuris var. ora

var. nudipes nanus cinereus var. var. var. lenticularis arizonica lifolius lifolius lifolius Haplopappus scopulorum hirtellus Haplopappus zionis Ericameria zionis Helianthella unifl Hymenopappus fi Hymenoxys acaulis var. Helianthus annuus var. Hymenopappus fi Hymenopappus fi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name biogeographic notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 39 graminea Willow Creek, San Juan Co., UT (Eastwood 45 CAS) “Southern Utah” (Thompson s.n. GH) reported by Doug Stone. Comments doc. Year Year 1956 1980 locality is Type 1957 1956 1978 1980 Native to Europe 1931 1956 Var. c name with taxonomic and (BRCA) Graybosch (1981) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range AnnF Wide Pres Unc Buchanan 2986 PerF RegEn Pres Unc Foster 10959 (BRY) 1993 locality is Type PerF Wide Pres Unc Boyle 1158 (BRCA) 1931 Life form ower showy rush-pink PerF Widethorny wire- Preslettuce Unc Cottam 2771 (BRY) 1927 aristate aster PerF RegEn Rephoary aster ? PerF Buchanan & Wide Pres Unc Buchanan 287 blue lettuce PerF Wide Pres Rare Buchanan 97 poverty-weed PerFprickly lettuce Wide AnnF Pres Intro Unc Pres Buchanan 2945 Unc Graybosch 1080 greenstem paperﬂ Colorado rubber- plant rock-goldenrod PerF Wide Pres Com Graybosch 1356 gumweed aster PerF Wide Pres Com Buchanan 396 Common name

var.

var. var. var. ora vacans, Dieteria canescens oribunda L. grandiﬂ spinosa, Pleiacanthus spinosus M. linearis, rigida, M. canescens var. canescens Aster canescens, Dieteria canescens var. aristata L. pulchella, oblongifolia, Mulgedium pulchellum axillaris Picradenia richardsonii ﬂ Synonyms/ notes Taxonomic Solidago petradoria Haplopappus nuttallii, Xanthisma grindelioides grindelioides

var.

var. var. var. var. var. ora ora ora pumila oribunda Lygodesmia spinosaLygodesmia Stephanomeria Machaeranthera canescens Machaeranthera canescens Machaeranthera grindelioides aristata canescens grandifl Lygodesmia Lygodesmia grandifl pulchella Lactuca serriola Lactuca tatarica Iva axillaris I. axillaris Hymenoxys richardsonii fl Petradoria pumila var. Psilostrophe sparsifl grindelioides rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name biogeographic notes, cont. Compositae (Asteraceae) Compositae (Asteraceae)

40 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Co., eld Sensitive Proctor Canyon, Garﬁ UT (Madsen & Madsen 516 BRY) Comments doc. Year Year 1998 Native to Europe 1980 USFS: Region 4 1931 1980 2006 1931 1931 1980 1980 1931 1998 Native to Europe c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) EB07290601 (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Kitchen s.n. (BRCA) Source Pop. size Park status Range PerF Disj Pres Rare Graybosch 1044 PerF LocEn Pres Rare Foster 11044 (BRY) 1994 locality is Type PerF Wide Pres Unc Graybosch 1107 PerF Wide PresAnnF Intro Com Buchanan 257 Pres Unc Ames-Curtis & Life form cluster-head cluster-head chicken-sage marsh sow-thistle PerF Intro Pres Unc Buchanan s.n. broom groundsel PerF Wide Pres Com B-31/151 Weight black groundsel PerF groundselHart’s Wide PerF Preswater groundsel RegEn Unc Pres PerF Buchanan 98 Wide Rarebasin groundsel Graybosch 896 Pres PerF Rare Wide Ballenger Pres Com Barkley’s Graybosch 1311 groundsel Parry’s Parry’s goldenweed coast goldenrod PerF Wide Pres Unc Buchanan 2962 three-nerve goldenrod spiny-leaf sow- thistle Common name

Vars. Vars.

Flora nana var. var. ssp. ora, (2006). ora Flora of North simplex Tanacetum Tanacetum capitatum uliginosus S. spartioides. not recognized in not Vars. recognized in of North America America (2006). Oreochrysum parryi includes vars. & S. velutina sparsiﬂ Synonyms/ notes Taxonomic glabrescens, S. arvensis

var. spartioides Sphaeromeria capitata Senecio atratus Senecio hartianus Packera hartiana Senecio hydrophilus Senecio multilobatus Packera multilobata Senecio spartioides var. Senecio werneriifolius barkleyi Solidago parryiSolidago simplexparryi, Haplopappus S. spathulata, Solidago velutina S. sparsifl Sonchus asper Sonchus uliginosus S. arvensis rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name biogeographic notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 41 eld Co., Utah is rim of Bryce Canyon, Bryce Canyon NP, Garﬁ (Eastwood & Howell 727 CAS). USFWS: 3C noxious weed. Native to Eurasia Comments doc. Year Year 1932 1940 1980 1957 Native to Europe 1956 1980 Utah state 1956 1980 Native to Eurasia 2007 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range AnnF Wide Hist ? B-32/343 Weight PerF LocEn Pres Unc Peabody 96 (BRCA) 1959 locality Type PerF WidePerF Pres Wide Unc Pres Buchanan 435 Unc Graybosch 901 PerF Wide Pres UncPerF Graybosch 1059 IntroShrub Pres Wide Pres Unc Graybosch 990 Unc Boyle 1160 (BRCA) 1931 AnnF Wide Pres Rare ST09190701 Topp Life form eld bindweed PerF Intro Pres Unc Graybosch 914 golden crownbeard Bryce Canyon townsendia yellow salsify PerF Intro Pres Unc Buchanan 196 stemless townsendia hoary townsendia showy goldeneye PerF Wide Pres Unc Buchanan 2987 ﬁ slender wirelettuce common dandelion spineless horsebrush southern goldeneye Common name

Flora

major

var. var. var. var. var. Vars. not Vars. ora Ximensia encelioides T. alpigena T. minima minima, T. V. annua, V. Heliomeris longifolia (2006) S. minor minor. recognized in of North America nevadensis Synonyms/ notes Taxonomic annua Heliomeris multiﬂ

ora cinale

var. var. var. annua nevadensis exauriculata Viguiera longifolia var. minima Townsendia incana Townsendia Townsendia montana dubiusTragopogon dubius T. Verbesina encelioides Townsendia exscapa Townsendia Convolvulus arvensis Taraxacum offi Taraxacum Tetradymia canescens tenuifolia Viguiera multifl var. Stephanomeria tenuifolia rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix A. Confi FamilyCompositae (Asteraceae) Species name biogeographic notes, cont.

42 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Weed. Native to Weed. Europe Comments doc. Year Year 1932 1932 Native to Europe 1980 Native to Europe 1958 USFWS: 3C 2005 1932 1931 1980 1980 2006 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) Plateau Network (in ed.) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Shrub Wide Pres Unc Buchanan 154 PerF Wide Pres Unc Graybosch 743 PerF Wide RepPerF Unc Wide Northern Colorado Hist ? B-32/181 Weight PerF Wide Pres Unc Graybosch 799 PerF Wide Hist ? B-31/5 Weight PerF Wide Pres Unc Graybosch 831 Life form red-osier dogwood nodding rockcress black mustard AnnFshepherd’s-purse Intro AnnF Intro Hist Pres ? Unc B-32/204 Weight Graybosch 1097 Drummond’s Drummond’s rockcress Fendler’s rockcress perennial rockcress Holboell’s Holboell’s rockcress Holboell’s rockcress rockcressSelby’s PerF Wide Pres Unc ST06150604 Topp Common name whitetop PerF Intro Pres Unc Kitchen s.n. (BRCA) 1998 UT State Noxious

var. draba var. languida, spatifolia, B. pinetorum, retrofracta, secunda, B. C. stolonifera, Swida sericea var. B. demissa draba demissa angustifolia, B. drummondii, B. stricta fendleri Boechera perennans Boechera fendleri var. A. fendleri, Boechera holboellii var. Name may be misapplied, according to Holmgren et al. (2005) A. holboellii var. Boechera holboellii var. retrofracta Synonyms/ notes Taxonomic

var. var. var. var. perennans Cornus sericea sericea Arabis demissa Boechera demissa Brassica nigraCapsella bursa- pastoris Sinapsis nigra Cardaria draba Lepidium draba, C. Arabis drummondii Boechera Arabis holboellii Arabis holboellii Arabis holboellii Arabis perennans var. Arabis selbyi Boechera selbyi fendleri pinetorum secunda rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix A. Conﬁ FamilyCornaceae Species name Cruciferae (Brassicaceae) biogeographic notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 43 Cedar Breaks, Cedar Breaks NM, Iron Co., UT (Goodman & Hitchcock 1622 MO). USFWS: 3C Comments NA doc. Year Year 1980 1998 1932 1957 Native to Europe 1934 locality is Type 1956 1980 1980 Native to Europe c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Buchanan (1993) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range AnnF Wide Pres Unc Graybosch 919 AnnF Wide Pres Unc Buchanan s.n. AnnF Wide Pres Unc Graybosch 860 AnnF Wide Rep ? Spence & AnnF Intro Pres Unc Graybosch 950 PerF Wide Pres Com Boyle 1107 (BRCA) 1934 Life form ower PerF Wide Pres Unc36 Buchanan ower ixweed AnnF Intro Pres Unc113 Buchanan sand dune wallfl prairie pepperwort western tansy- mustard western tansy- mustard fl Breaks draba PerF LocEn Pres Unc Buchanan 3051 mountain tansy- mustard shy wallfl spindlestem PerF hare’s-ear Widemustard Pres Unc Graybosch 862 Common name

var. var. ssp.

var.

D. var. ssp. by some by some var. arkansanum, included in var. capitatum authors Some authors include var. macrocarpum Included in pinnata osmiarum authors D. pinnata E. asperum halictorum incisa, D. incana incana D. richardsonii sonnei, D. incisa incisa, D. incana Synonyms/ notes Taxonomic

var. var. orum orum intermedia osmiarum purshii sonnei Descurainia pinnata var. Descurainia pinnata var. Descurainia sophia Draba subalpina Erysimum asperum var. densifl Lepidium densifl Erysimum inconspicuum Caulanthus crassicaulis Conringia orientalis crassicaulis Descurainia incana var. rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix A. Confi FamilyCruciferae (Brassicaceae) Species name biogeographic notes, cont. Cruciferae (Brassicaceae)

44 Annotated Checklist of Vascular Flora: Bryce Canyon National Park

eld Co., eld eld Co., UT eld Co., eld of var. of var. membranacea is Red Canyon, Garfi UT (Rollins & Chambers 2448 GH). is head of Sweetwater Creek, Escalante Mts E of Widtsoe, Garfi (Atwood 8164 BRY) Casto Canyon, Garfi UT (Atwood, Franklin, & Reynolds 13744 UT BLM: BRY). Sensitive Comments doc. Year Year 1980 1931 locality Type 1980 Native to Africa 1932 locality is Type c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Graybosch 796 PerF Wide Pres Com Boyle 1128 (BRCA) 1932 PerF RegEn Hist ? Stone 240 (BRY) 1934 PerF LocEn Pres Unc Foster 10968 AnnF Intro Pres Unc Kitchen s.n. (BRCA) 1998 Native to Europe Life form King’s King’s bladderpod Watson’s Watson’s bladderpod lepidote twinpod PerF LocEn Pres Unc Buchanan 3056 Claron twinpod PerF LocEn Pres Rare Foster 11052 (BRY) 1994 locality Type Arizona bladderpod African mustard AnnF Intro Pres Unc Graybosch 1075 Claron pepperwort clasping pepperwort Common name P.

var. var. var. . Lesquerella kingii. Holmgren et al. (2005) include wardii P. lepidota P. intermedia, L. alpina intermedia membranacea, lepidota P. lepidota arizonica Synonyms/ notes Taxonomic

var. membranacea sobolifera claronense Physaria kingii parvifolia Physaria chambersii var. Physaria chambersii var. Physaria intermedia Lesquerella Physaria arizonica Lesquerella Malcolmia africana Lepidium montanum var. Lepidium perfoliatum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix A. Conﬁ FamilyCruciferae (Brassicaceae) Species name biogeographic notes, cont. Cruciferae (Brassicaceae)

Chapter 6: Appendices 45 eld Co., UT eld Co., UT Panguitch Lake, Garfi (Jones 6015e US). USFWS: 3C Red Canyon, Garfi (Eggleston 8198 US). USFWS: 3C Comments doc. Year Year 1980 1932 2006 locality is Type 1974 Native to Europe 2006 1957 locality is Type 1932 1931 1931 1980 1980 1956 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF WidePerF Pres RegEn Unc Pres Buchanan 3333 Rare ST07110601 Topp PerF LocEn Pres Unc Graybosch 1335 PerF Wide Pres Unc Foster 10922 (BRY) 1932 PerF RegEn Pres Unc Shaw 2871 (BRY) 1982 Tree Wide Pres Com Buchanan 164 PerG Wide Pres Rare Graybosch 996 Life form ower eld pennycress AnnF Intro Pres Unc Graybosch 805 ax-leaved heart-leaf jewelfl Palmer’s thelypody alpine pennycress PerF Widecommon juniper Pres Shrub Wide Rare Pres Fertig 22534 Unc Buchanan 60 prince’s-plume PerF Wide Pres Unc Graybosch 943 fi Breaks bladderpod fl plainsmustard Colorado Plateau bladderpod Utah juniperRocky Mountain Shrub Widejuniper Pres Com Buchanan 226 slender-beaked slender-beaked sedge golden sedge PerGDouglas’ sedge Wide PerG Pres Wide Unc Pres Graybosch 865 Unc Buchanan 67 Common name

var. var. var. alpina montana S. cordatus cordatus Thelypodium sagittatum ovalifolium Noccaea montana var. Juniperus communis ssp. pinnata Lesquerella rubicundula, L. hitchcockii, P. rubicundula Sisymbrium linifolium Sabina osteosperma Sabina scopulorum Synonyms/ notes Taxonomic

var. montanum depressa rubicundula ovalifolia Streptanthus cordatus Thelypodiopsis sagitatta Thlaspi arvense Thlaspi montanum var. Juniperus communis var. Stanleya pinnata S. pinnata Physaria rubicundula var. Schoenocrambe linifolia Physaria rectipes Lesquerella rectipes Juniperus osteosperma Juniperus scopulorum Carex athrostachya Carex aureaCarex douglasii C. garberi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cupressaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix A. Conﬁ FamilyCruciferae (Brassicaceae) Species name biogeographic notes, cont. Cupressaceae Cupressaceae Cyperaceae Cyperaceae Cyperaceae

46 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 1980 1980 1969 1980 2006 1932 2006 2006 1980 1980 1980 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) ST06140601(BRCA) AT07290602 (BRCA) (BRCA) Graybosch (1981) Graybosch (1981) Source Pop. size Park status Range PerG Wide Pres Unc Graybosch 777 PerG Wide Pres Unc Gottschalk 4 (UTC) 1986 PerG Wide Pres Unc Graybosch 878 Life form ower few-ﬂ spikerush beaked sedge PerGvalley sedge Widecommon Pres PerGspikerush Wide Unc Pres Graybosch 1012 Unc Graybosch 915 Ross’ sedge PerG Wide Pres Com Buchanan 3054 green sedgewoolly sedge PerG Sparseblackcreeper PerG Pressedge Wide Rare Pres Graybosch 1125 Unc ST06280603 Topp small-wing sedge PerG Wide Pres Unc Graybosch 1013 narrowleaf sedge PerG Wide Presinland sedge Rare Topp PerG Wide Pres Rare Tendick Nebraska sedge PerG Wide Rep ? Buchanan & beaked spikerush PerG Wide Rep ? Buchanan & Common name

& E.

ssp. var. (sensu var. var. E. ora, ana exa microptera, E. pauciﬂ suksdorﬁ Includes macrostachya, E. erythropoda, uniglumis lato) boottii, C. brevipes, C. pityophila viridula var. C. festivella, macloviana microptera C. viridula eleocharis stenophylla, C. stenophylla Synonyms/ notes Taxonomic

var. ora Eleocharis quinquefl Carex vallicola Eleocharis palustris Carex utriculata C. rostrata Carex pellitaCarex praegracilis C. lanuginosa Carex rossii C. defl Carex interior Carex microptera C. microptera Carex nebrascensis Carex oederi viridula Carex duriuscula C. eleocharis, Eleocharis rostellata rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Appendix A. Confi FamilyCyperaceae Species name biogeographic notes, cont. Cyperaceae

Chapter 6: Appendices 47 upper valley of the Virgin, Kane Co., UT (Siler s.n. ISC) Comments doc. Year Year 1959 1934 1980 2004 Native to Europe 1959 1957 1931 locality is Type 1959 1957 1956 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Fern Sparse Pres Rare Buchanan 486 Shrub Wide Pres Com Buchanan 110 PerG Wide Pres Unc Graybosch 863 Fern Wide Pres Unc Buchanan 249 Shrub Wide PresShrub Wide Unc PresShrub Buchanan 528 RegEn Unc Pres Buchanan 348 Unc Graybosch 802 Life form eld horsetail Fern Wide Pres Unc Buchanan 463 Nelson’s Nelson’s scouring-rush greenleaf manzanita three-square bulrush smooth scouring- rush Russian olivesilver Treebuffaloberry IntroCanada Presbuffaloberry roundleaf Uncbuffaloberry ST06150603 Topp green ephedra Shrubﬁ Wide Pres Unc Foster 10973 (BRY) 1957 bearberry Shrub Wide Pres Unc Buchanan 59 Common name E.

Vars. Vars.

var. ssp. var. var. coactilis. polyphyllus, Hybrid between E. laevigatum variegatum A. patula platyphylla Scirpus pungens var. Schoenoplectus pungens longispicatus, Schoenoplectus pungens polyphyllus Hippochaeta laevigata, E. kansanum E. viridis Synonyms/ notes Taxonomic A. uva-ursi uva-ursi, A. uva-ursi ssp. not recognized in Holmgren et al. (2005)

var. var. nelsonii x Arctostaphylos patula Arctostaphylos uva- Scirpus pungens longispicatus Elaeagnus angustifolia Shepherdia argentea Shepherdia canadensis Shepherdia rotundifolia Ephedra viridis Equisetum laevigatum Equisetum viridis Equisetum arvense ursi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Ericaceae Ericaceae Appendix A. Conﬁ FamilyCyperaceae Species name biogeographic notes, cont. Elaeagnaceae Elaeagnaceae Elaeagnaceae Elaeagnaceae Ephedraceae Equisetaceae Equisetaceae Equisetaceae

48 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Co., UT noxious weed. Native to Europe Panguitch Lake, Garﬁ (Jones 6008 POM) Comments NA doc. Year Year 1980 2004 Utah state 1932 1978 1927 locality is Type 1932 1932 1956 c name with taxonomic and (BRCA) (2005) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Buchanan (1993) Source Pop. size Park status Range PerF Wide Pres Unc Buchanan 2943 PerF Wide Pres Unc Buchanan 83 Life form meadow gentian AnnF Wide Pres Uncelkweed Graybosch 1123 stork’s-bill PerF AnnF Wide Intro Pres Pres Unc Unc Boyle 1134 (BRCA) Kitchen s.n. (BRCA) 1934 1998 Native to Europe leafy spurge PerF Intro Repgolden corydalis PerF Unc Wide Dewey & Andersen Hist ? Stone 266 (NY) 1934 Gambel oak Tree Wide Pres Unc Buchanan 163 Engelmann’s gentian Jones’ gentian AnnF RegEn Pres Unc Graybosch 1029 shorthorn spurge PerF Wide Pres Unc Graybosch 768 Fendler’s spurgeFendler’s PerF Wide Pres Unc Buchanan 372 Rocky Mountain gentian Parry’s gentianParry’s PerF Wide Rep ? Spence & Common name

var. ssp.

var. elegans, robusta, nis Gentiana detonsa, Gentianopsis thermalis Gentianella detonsa var. esula, Tithymalus esula, Tithymalus esula C. curvisiliqua occidentalis Gentianella amarella heterosepala, Gentiana scopulorum E. robusta, brachycera var. E. montana, lurida, Tithymalus brachyceras Pneumonanthe afﬁ parryi Synonyms/ notes Taxonomic

var. var. nis elegans var. gambelii nis Swertia radiata Frasera speciosa Erodium cicutarium Quercus gambelii var. Corydalis aurea occidentalis Gentiana affi Gentianella heterosepala Gentianella tortuosa Gentiana tortuosa Gentianopsis detonsa Chamaesyce fendleri Euphorbia fendleri Euphorbia brachycera Euphorbia esulaesula Euphorbia affi Gentiana parryi Pneumonanthe rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Gentianaceae Geraniaceae Fagaceae Fumariaceae Gentianaceae Gentianaceae Gentianaceae Gentianaceae Euphorbiaceae Euphorbiaceae Appendix A. Confi FamilyEuphorbiaceae Species name biogeographic notes, cont. Gentianaceae

Chapter 6: Appendices 49 G. var. var. eld ed Aquarius Plateau, head of Poison Creek, Garﬁ Co., UT (Rydberg & Carlton 7401 NY) from BRCA are based on misidentiﬁ specimens of caespitosum marginale and Africa Comments doc. Year Year 2005 Previous reports 1956 Native to Eurasia 1956 Native to Eurasia 1980 1957 c name with taxonomic and Plateau Network (in ed.) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Com Boyle 392 (BRCA) 1931 locality is Type PerF Wide Rep Unc Northern Colorado PerG Intro Pres Unc Buchanan 56 Life form small-leaf geranium purple three-awn PerG Wide Pres Unc Graybosch 780 blue grama PerG Wide Pres Unc Buchanan 284 Richardson’s Richardson’s crane’s-bill crested wheatgrass redtop PerG Intro Pres Unc Graybosch 1126 Common name

ssp. var. var. var. ssp. cowenii, G. G. atropurpureum var. atropurpureum, G. caespitosum atropurpureum purpurea fendleriana, A. fendleriana, A. purpurea longiseta gracilie cristatum, A. cristatum pectinatum, A. pectinatum, A. sibiricum, A. desertorum, A. fragile, mongolicum Synonyms/ notes Taxonomic

var. marginale Geranium caespitosum Geranium richardsonii Agropyron cristatum A. cristatum Agrostis stolonifera A. alba, gigantea Aristida purpurea A. glauca, Bouteloua gracilis Chondrosum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Appendix A. Conﬁ FamilyGeraniaceae Species name biogeographic notes, cont. Geraniaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

50 Annotated Checklist of Vascular Flora: Bryce Canyon National Park is Springdale, Co., Washington UT (Jones 6075 POM) and Africa Comments doc. Year Year 1957 1980 Native to Eurasia 1980 1942 Native to Eurasia 1980 1931 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerG Wide Pres Unc Graybosch 1131 Life form smooth brome PerG Intro Pres Com Boyle 1130 (BRCA) 1942 Native to Eurasia slimstem reedgrass fringed brome PerG Wide Pres Unc Graybosch 939 Japanese chess AnnGcheatgrass Intro PresJones’ reedgrass Unc AnnG PerG Intro Wide Graybosch 1082 Pres Pres Unc Unc Boyle 1125 (BRCA) Boyle 1153 (BRCA) 1932orchard grass 1942 Native to Eurasia locality Type tufted hairgrass PerG Intro PerG Wide Pres Pres Unc Unc Buchanan 423 Graybosch 1129 mountain brome PerG Wide Pres Unc Graybosch 1021 nodding brome PerG Wide Pres Unc Buchanan 54 Common name

or . B. includes in oras is oras

& B. porteri Bromopsis inermis inexpansa richardsonii, Bromopsis canadensis B. commutatus Bromus frondosus, Bromus lanatipes, Bromopsis porteri. According to FNA (2007) material ascribed to anomalus mostly B. polyanthus, Ceratochloa carinata Includes vars. ciliatus regional ﬂ B. lanatipes Synonyms/ notes Taxonomic

var. Bromus inermis inermis Bromus japonicus B. arvensis, Bromus tectorum Anisantha tectorum Calamagrostis stricta C. neglecta, Dactylis glomerata Deschampsia cespitosa Calamagrostis scopulorum Bromus anomalus Bromus porteri, Bromus carinatus B. marginatus, Bromus ciliatus rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name biogeographic notes, cont.

Chapter 6: Appendices 51 top of Salina Pass, Sevier Co., UT (Jones 5447 POM) Comments doc. Year Year 1980 Native to Eurasia 1980 Native to Russia 1957 1931 locality is Type 1987 2006 1980 1998 1931 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (UTC) (BRCA) (BRCA) Buchanan s.n. (BRCA) (BRCA) Source Pop. size Park status Range PerG Intro Pres Unc Graybosch 895 PerG Wide Pres Unc Buchanan 326 PerG Wide Pres Unc Graybosch 1363 PerG Wide Pres Unc Ames-Curtis & Life form intermediate wheatgrass Russian wildrye PerG Intro Pres Unc Graybosch 921 Salina wildrye PerG Wide Pres Com Buchanan 300 thickspike wheatgrass Scribner wheatgrass blue wildrye PerG Wide Pres Rare Hallsten 2537 desert saltgrass PerGCanada wildrye Wide PerG Pres Wide Rare Pres ST07280606 Topp Unc Graybosch 1076 Great Basin wildrye squirreltail PerG Wide Pres Com Fertig 22542 Common name

var. var. var. var. var.

var. var. & riparium, thsii lanceolatus, dasystachyum hispidus, Agropyron intermedium, Elytrigia intermedia, Thinopyrum intermedium juncea var. Agropyron dasystachyum var. & var. A. albicans albicans, salinus, L. salina, ambiguus griffi glaucus stricta, D. stricta canadensis elymoides, Sitanion hystrix Synonyms/ notes Taxonomic Elymus junceusElymus lanceolatus Psathyrostachys E. lanceolatus Elymus salinus E. salina, Leymus Elymus scribneri Agropyron scribneri Elymus hispidus E. hispidus Elymus canadensis E. canadensis Elymus cinereus Leymus cinereus Elymus elymoides E. elymoides Elymus glaucus E. glaucus Distichlis spicata D. spicata rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi FamilyGramineae (Poaceae) Species name biogeographic notes, cont.

52 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 1981 1980 1957 1957 1980 1932 1956 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerG Wide Pres Unc Graybosch 1028 PerG Wide Pres Unc Buchanan 53 PerG Wide Pres Unc Boyle 1141 (BRCA) 1942 PerG Wide Pres Unc Boyle 1142 (BRCA) 1942 Life form six-weeks fescue AnnG Wide PresRocky Mountain Uncfescue Graybosch 1322 meadow fescue PerG Introfowl mannagrass Pres PerG Wide Unc Pres Boyle 1145 (BRCA)meadow barley 1942 Unc Native to Eurasia PerG Buchanan 303 Wide Pres Unc Graybosch 1127 galletafoxtail barley PerG Wide PerG Pres Wide Unc Pres Graybosch 845 Unc Buchanan 57 bluebunch wheatgrass slender wheatgrass western wheatgrass Common name

palmeri G.elata includes var. & ora, Vulpia Vulpia ora, ora inermis latiglumis, trachycaulus, octoﬂ octoﬂ ovina F. rydbergii, F. saximontana Schedonorus elatior pratensis, F. Includes Critesion brachyantherum Includes vars. glauca, hirtella, ssp. spicatum, Pseuodroegneria spicata, spicata var. E. alaskanus ssp. Agropyron caninum, A. subsecundum, A. trachycaulum, A. latiglume Pascopyrum smithii, includes vars. smithii Synonyms/ notes Taxonomic

var. ora Festuca ovina saximontana Festuca pratensis Lolium pratense, Glyceria striata Hilaria jamesiiHordeum Pleuraphis jamesii brachyantherum Hordeum jubatum Critesion jubatum Festuca octofl Elymus spicatus Agropyron Elymus trachycaulus E. trachycaulus Elymus smithii Agropyron smithii, rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi FamilyGramineae (Poaceae) Species name biogeographic notes, cont.

Chapter 6: Appendices 53 Comments doc. Year Year 1931 1998 1980 Native to Eurasia 1980 1980 Native to Eurasia 1931 1980 1980 2005 1956 1980 1956 1980 1957 1931 Native to Europe c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerG Wide Pres Unc Graybosch 1071 PerG Intro Pres Unc Buchanan 452 Life form muttongrass PerG Wide Pres Com Buchanan 130 witchgrasstimothy AnnG Widecommon reed Pres PerG PerGCanada bluegrass Unc PerG Intro Wide Buchanan s.n. Intro Pres Pres Pres Unc Unc Unc Graybosch 1103 Graybosch 1074 Graybosch 907 junegrass PerG Wide Pres Com ST08130504 Topp mountain muhly PerG Wide Pres Unc Buchanan 285 littleseed ricegrass foxtail muhlyscratchgrass PerG Widepullup muhly Pres PerG Wide Unc AnnG Pres Wide Graybosch 1077 Pressandhill muhly Unc Graybosch 1078 Unc PerGmat muhly ST09100502 Topp Wide Pres PerG Unc Wide Graybosch 1102 Pres Unc Buchanan 2949 interior bluegrass PerG WideKentucky Presbluegrass Unc Buchanan 282 Common name

ssp. ssp. var. M. liculmis longiligula fendleriana occidentale pyramidata, K. nitida Includes ﬁ micranthum nemoralis P. interior, P. interior P. interior, Synonyms/ notes Taxonomic

var. liformis Phleum pratense Phragmites australis communis P. Poa compressa Poa fendleriana longiligula, P. P. Koeleria macrantha K. cristata, Muhlenbergia andina Muhlenbergia asperifolia Muhlenbergia fi Muhlenbergia montana Muhlenbergia pungens Muhlenbergia richardsonis Oryzopsis micrantha Piptatherum Panicum capillare capillare P. Poa glauca glauca Poa pratensis agassizensis P. rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Gramineae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix A. Confi FamilyGramineae (Poaceae) Species name (Poaceae) biogeographic notes, cont.

54 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 1980 1981 1981 1998 Native to Eurasia 2006 1980 2006 1940 1931 1957 1957 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA (BRCA) AT07280601 (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerG Wide Pres Unc Graybosch 911 PerG Wide Pres Rare Graybosch 1336 PerG Wide Pres Com Buchanan 445 PerG Wide Pres Unc Graybosch 916 PerG Wide Pres Unc Buchanan 265 Life form Sandberg’s Sandberg’s bluegrass Nuttall’s Nuttall’s alkaligrass alkali sacaton PerG Wide Presneedle-and- Uncthread Graybosch 947 Indian ricegrass PerG Wide Pres Letterman’s needlegrass Com Buchanan 69 little bluestem PerG Widegreen bristlegrass Pres AnnG Intro Uncalkali cordgrass Pres Graybosch 1334 PerG Unc Wide Buchanan s.n. Pressand dropseed Rare PerG ST06150607 Topp Wide Pres Unc Tendick Nelson’s Nelson’s needlegrass Common name

var.

comata var. P. var. var. and all vars. P. secunda P. Includes gracillima, P. juncifolia, P. sandbergii, P. scabrella, canbyi, P. nevadensis, P. P. ampla of P. airoides P. Andropogon scoparius, S. scoparium S. airoides Hesperostipa comata hymenoides, Achnatherum hymenoides lettermanii neomexicanum Achnatherum nelsonii nelsonii, S. columbiana nelsonii Synonyms/ notes Taxonomic

var. var. airoides Poa secunda Puccinellia nuttalliana Schizachyrium scoparium Setaria viridis Spartina gracilis Sporobolus airoides var. Sporobolus cryptandrus Stipa comata comata Stipa hymenoides Oryzopsis Stipa lettermanii Achnatherum Stipa nelsonii nelsonii rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name biogeographic notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 55 eld Co., UT Nipple Bench, Kane Co., UT & Atwood (Welsh 9809 BRY). USFWS: Category 3C Panguitch Lake, Garﬁ (Jones 6023 POM) Comments doc. Year Year 1932 1972 2006 2006 locality is Type 1978 1988 locality is Type c name with taxonomic and (BRCA) (BRCA) personal obs. (BRCA) (BRCA) (1988) Source Pop. size Park status Range PerF Wide Pres Unc Stebbins s.n. AnnF LocEn Pres Unc ST06150606 Topp PerF Wide Pres Unc Boyle 1110 (BRCA) 1931 PerF Wide Pres Unc Kitchen s.n. (BRCA) 1998 PerG Wide Pres Unc Buchanan 2969 Life form Baltic rush PerG Wide Pres Unc Buchanan 64 Idaho blue-eyed grass Nipple Bench phacelia brittle phacelia AnnF RegEn Rep Rare Sarah Topp, Rocky Mountain iris western St. John’s-wort Bloomer’s Bloomer’s ricegrass pine needlegrass PerG Wide Rep Unc Hallsten & Roberts Common name

var.

and species P. var. Hybrids var. by some S. S. comata Stipa ater, J. balticus ater, montanus, bloomeri, balticus, J. arcticus ssp. includes var. mexicanus var. S. idahoense occidentale authors bloomeri. between hymenoides other demissa P. Included in crenulata x Achnatherum H. scouleri, formosum scouleri except pinetorum Synonyms/ notes Taxonomic var. var. bloomeri Stiporyzopsis x Juncus arcticus J. arcticus Sisyrinchium idahoense Phacelia demissa demissa Phacelia mammillarensis Iris missouriensis Hypericum formosum Stipa pinetorum Achnatherum Stipa rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Juncaceae Iridaceae Hydrophyllaceae Hydrophyllaceae Iridaceae Guttiferae (Hypericaceae) Gramineae (Poaceae) Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name biogeographic notes, cont.

56 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 1931 1957 1994 1932 Native to Eurasia 1980 1980 1980 1980 1980 1980 1931 1957 1998 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Graybosch 1000 PerF Intro Pres Unc Buchanan 543 PerG Wide Pres Unc Graybosch 866 PerF Wide Pres Unc Graybosch 936 Life form ower eld mint PerF Wide Pres Unc Schelz s.n. (BRCA) 1990 smallﬂ dragonhead mock pennyroyal PerF Widedwarf pennyroyal PerF Pres Periphcommon Pres Unchorehound Buchanan 331 Rare Foster 11076 toad rush AnnG Wide Pres Unc Buchanan 3342 seaside arrowgrass Tracy’s rushTracy’s PerG Wide Preslongstyle rush Unc PerG Graybosch 209 rushTorrey’s Wide Pres PerG Wide Uncmarsh arrowgrass Pres Graybosch 1063 PerG Wide Unc Pres Graybosch 959 Unc Graybosch 774 self-heal PerF Wide Pres Unc Buchanan 224 ﬁ stinking horsemint annual sage PerF Sparse Pres Unc Buchanan s.n. Common name

var.

var. J. & by var. T. brunnescens ora canadensis Moldavica parviﬂ M. arvensis Includes vars. bufonius, occidentalis, halophilus, ambiguus Included in ensifolius some authors, J. saximontanus Includes concinnum montanus forma M. odoratissima glauca, M. glauca Synonyms/ notes Taxonomic

var. var. var. exa orum elata Hedeoma drummondii Hedeoma nanum Hedeoma nana Marrubium vulgare Mentha arvensis glabrata Juncus bufonius Juncus ensifolius brunnescens Juncus longistylus Juncus torreyi maritima Triglochin var. palustris Triglochin Dracocephalum parvifl lanceolata Monardella odoratissima Prunella vulgaris Salvia refl rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Juncaceae Juncaceae Juncaceae Juncaginaceae Juncaginaceae Labiatae (Lamiaceae) Appendix A. Confi FamilyJuncaceae Species name biogeographic notes, cont. Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae)

Chapter 6: Appendices 57 eld Co., eld Panguitch Lake, Garﬁ UT (Jones 6023f POM) Johnson, Kane Co., UT (Jones s.n. POM) near Cisco, Grand Co., UT (Jones s.n. POM) Comments doc. Year Year 1957 locality is Type 1974 locality is Type 1931 1931 1979 1980 1932 1930 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRY) (BRCA) Source Pop. size Park status Range PerF RegEn Pres Unc Graybosch 989 PerF Wide Pres Unc Graybosch 844 PerF Wide Pres Unc Graybosch 1339 PerF RegEn Pres Unc Peabody 2 (BRCA) 1991 locality is Type PerF Wide Pres Unc Graybosch 853 PerF RegEn Pres Unc Foster 10785 (BRY) 1993 PerF RegEn Pres Unc Buchanan 338 PerF Wide Pres Unc Foster 10832 (BRY) 1993 Life form eld milkvetch PerF Wide Pres Unc Buchanan 3128 Panguitch milkvetch Johnson milkvetch milkvetchTorrey’s PerF Wide Pres Unc Buchanan 3116 lesser rushy milkvetch rimrock milkvetch ﬁ crescent milkvetch Sheldon’s Sheldon’s milkvetch groundcover milkvetch tall kentrophyta PerF Widemountain Preskentrophyta Unc Peabody 41 (BRCA) 1931 Hall’s milkvetchHall’s PerF RegEn Pres Unc Atwood 6130 Common name

var. var. by Included in var. Included in var. bisulcatus Barneby (1989) A. calycosus A. diversifolius campestris A. desperatus Synonyms/ notes Taxonomic A. impensus A. kentrophyta implexus

var.

var. var. var. var. var. var. var. var. major calycosus Astragalus argophyllus panguicensis Astragalus bisulcatus var. Astragalus calycosus var. Astragalus convallarius convallarius Astragalus desperatus desperatus Astragalus hallii Astragalus agrestis A. dasyglottis Astragalus amphioxys amphioxys Astragalus amphioxys vespertinus humivagans Astragalus humistratus Astragalus kentrophyta elatus Astragalus kentrophyta tegetarius fallax rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix A. Conﬁ FamilyLeguminosae (Fabaceae) Species name biogeographic notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

58 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Co., UT Aquarius Plateau, Garﬁ 424 GH) (Ward Frisco, Beaver Co., UT (Jones 180 MINN) “southern Utah”, (Parry 32 ISC) Cedar Canyon, Iron Co., UT (Parry 51 GH) Kanab, Kane Co., UT (Thompson s.n. GH) miles E of Zion Kane Co., UT NP, (Hitchcock 19013 isotype NY) Comments doc. Year Year 1931 1931 1980 locality is Type 1934 locality is Type 1934 1954 locality is 10 Type c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRY) Source Pop. size Park status Range PerF Wide Pres Unc Buchanan 2891 PerF RegEn Pres Rare Foster 11019 (BRY) 1994 locality is Type PerF Wide Pres Rare Graybosch 793 PerF Wide Pres Unc Buchanan 214 PerF Wide Pres Unc Buchanan 32 PerF Wide Pres Unc Foster 11015 (BRY) 1994 Life form northern sweetvetch pulse milkvetch PerF milkvetchWard’s Wide PerF Pres RegEn Pres Unc Rare Graybosch 1104 Foster 11041 (BRY) 1994 locality is Type cobwed milkvetch great rushy milkvetch great bladdery milkvetch Rydberg’s weedy Rydberg’s milkvetch woolly milkvetch PerF Wide Presstinking Uncmilkvetch Foster 11014 (BRY) 1994 locality is Type Zion sweetpea PerF RegEn Pres Rare Harrison 12318 Common name

var. Vars. Vars. Some authors include var. pabulare A. lentiginosus diphysus Includes var. Includes var. megacarpus. not recognized in Barneby (1989) Synonyms/ notes Taxonomic

var.

var. var. var. var. boreale zionis Astragalus wardii Hedysarum boreale var. Lathyrus brachycalyx var. Astragalus tenellus araneosus Astragalus lonchocarpus Astragalus megacarpus parryi Astragalus miser oblongifolius Astragalus mollissimus thompsoniae Astragalus praelongus praelongus Astragalus lentiginosus rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix A. Conﬁ FamilyLeguminosae (Fabaceae) Species name biogeographic notes, cont.

Chapter 6: Appendices 59 sericeus. of Carmel, Kane Co., UT (Ferguson & Ottley 5613 NY) represent var. kingii. Kane Co., UT (Siler s.n. GH) be based on var. locality is Type Piute Bullion Cr., Co., Rydberg & Carlton 7024 (NY) Comments doc. Year Year 1956 locality is W Type 1980 1980 1931 1977 locality is Type 1980 Reports may 1956 1958 Native to Europe 1959 Native to Europe c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) 4382 (BRY) Graybosch (1981) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Intro Pres Unc Buchanan 450 Life form silverleaf lupine PerF Wide Pres Unc Foster 10782 (BRY) 1993 sink lupine PerF RegEn Pres Unc Foster & silky lupine PerF Wide Pres Unc Buchanan 28 Utah trefoil PerF RegEnsilvery lupine Pressubalpine lupine PerF Unc PerF Wide Graybosch 887 Wide Pres Pres Unc Unc Graybosch 975 lupineKing’s Graybosch 982 AnnF lupineKing’s RegEn Pres AnnF ? Wide Pres Hall s.n. (BRY)Marysvale lupine Unc PerF Graybosch 974 LocEn 1934 BRCA reports may Rep ? Buchanan and black medickalfalfa AnnF Introyellow sweet- Presclover Unc PerF Buchanan 373 Intro Hist ? Rowe 178 (BRY) 1931 Native to Europe Common name

cutleri sativa Vars. Vars. var. var. var. L. caudatus, argenteus argophyllus, L. caudatus L. barbiger. not recognized by Barneby (1989) not Vars. recognized by Barneby (1989) Synonyms/ notes Taxonomic

var. var. var. var. var. cinalis argenteus rubricaulis argophyllus Lotus utahensis Lupinus argenteus var. Lupinus argenteus var. Lupinus caudatus var. Lupinus kingii argillaceus Lupinus kingii kingii Lupinus sericeus barbiger Lupinus sericeus marianus Lupinus sericeus sericeus Medicago lupulina Medicago sativaMelilotus offi M. sativa rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name Leguminosae (Fabaceae) biogeographic notes, cont.

60 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld eld Co., UT eld Co., UT Red Canyon, Garfi (Ripley & Barneby 8550 CAS). USFWS: 3C mile E of Bicknell, Co., UT Wayne & Moore (Welsh 13828 BRY) Aquarius Plateau, Garfi 541 G) (Ward Bryce Canyon Garfi NP, Co., UT (Welsh & Murdock 12859 BRY). USFS: Region 4 Sensitive, USFWS: C2 Comments doc. Year Year 1931 locality is Type 1980 locality is Type 1931 locality is Type 1957 Native to Europe 1932 1932 1980 c name with taxonomic and (BRCA) (BRCA) 12859 (BRY) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF RegEn Pres Unc Foster 11046 (BRY)PerF 1994 Wide locality is 1 Type Pres Unc Graybosch 889b PerF Wide Pres Unc Graybosch 1117 PerF Wide Pres Unc Foster 11021 (BRY) 1994 Life form Paria breadroot PerF LocEn Pres Unc & Murdock Welsh dwarf clover PerF Wide Pres Unc Britten s.n. (BRCA) 1990 Jones’ locoweed PerF RegEn Pres UncBicknell Graybosch 813 locoweed mountain locoweed white cloverAmerican vetch PerF PerF Intro oniontaper-tip Wide Pres PerF Pres Unc Wide Unc Graybosch 1011 Hist Graybosch 784 ? B-32/323 Weight nodding locoweed Lambert’s Lambert’s locoweed Common name

var. Psoralea pariensis gymnocarpon T. gymnocarpon Includes var. Includes var. lambertii Synonyms/ notes Taxonomic

var. exa jonesii juniperina oreophila sericea bigelovii Oxytropis oreophila var. Oxytropis oreophila var. Oxytropis oreophila var. Pediomelum pariense Trifolium gymnocarpon repens Trifolium Vicia americana americana Allium acuminatum Oxytropis defl var. Oxytropis lambertii var. rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Alliaceae) Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name biogeographic notes, cont. Leguminosae (Fabaceae)

Chapter 6: Appendices 61 Comments doc. Year Year 1932 1980 1980 Native to Eurasia 1931 1957 1940 1932 1957 1980 1931 1980 1941 1980 1940 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Graybosch 820 PerF Wide Pres Unc Buchanan 383 PerF Wide Pres Unc Graybosch 834 PerF WidePerF Pres Wide Unc Pres Graybosch 1081 Unc ST08240701 Topp Life form ax PerF Wide Pres Unc Boyle 1127 (BRCA) 1894 ax AnnF Sparse Pres Unc918 Graybosch ax PerF Wide Pres Unc29 Buchanan twincrest onion PerF Wide Hist ? B-32/155 Weight starry false Solomon’s-seal Nevada onion PerF Wideasparagus Pres Raresego-lily Graybosch 1015 PerFleopard-lily Intro Pres PerFstar-lily PerF Wide Unc Wide Pres Graybosch 1079 Pres Unc PerF Unc Buchanan 30 mountain death Periph Buchanan 140 camas Pres Unc Buchanan 3069 small yellow fl Common name scarlet globemallow common mallow AnnF Intro Pressmall-leaf Uncglobemallow Kitchen s.n. (BRCA)woodland 1998pinedrops Native to Eurasia King’s yellow fl King’s blue fl desert stickleaf PerF Wide Pres Unc Graybosch 965

A. var.

lewisii, & var. ora var. ora, Nuttallia ora, ora Includes vars. bisceptrum palmeri et al. Welsh (2003) include atrorubens stellatum cristatum Synonyms/ notes Taxonomic dissecta Includes ssp. coccinea, elata, L. lewisii Adenolinum lewisii multiﬂ multiﬂ

oramultifl M. ssp. cinalis Allium bisceptrum Allium nevadense Asparagus offi Calochortus nuttallii Fritillaria atropurpurea Leucocrinum montanum Smilacina stellata Maianthemum Zigadenus elegans Anticlea elegans Linum australe Mesynium australe Malva neglecta Sphaeralcea parvifolia Pterospora andromedea Sphaeralcea coccinea Linum kingii Mesyniopsis kingii Linum perenne lewisii Mentzelia multifl rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Liliaceae (Alliaceae) Liliaceae (Asparagaceae) Liliaceae (Calochortaceae) Liliaceae Liliaceae (Anthericaceae) Liliaceae (Convallariaceae, Ruscaceae) Liliaceae (Melanthiaceae) Appendix A. Confi FamilyLiliaceae (Alliaceae) Species name biogeographic notes, cont. Linaceae Malvaceae Malvaceae Monotropaceae (Ericaceae) Malvaceae Linaceae Linaceae Loasaceae

62 Annotated Checklist of Vascular Flora: Bryce Canyon National Park near Beaver City, near Beaver City, Beaver Co., UT (Palmer 404) Rabbit Valley, Rabbit Valley, Co., UT Wayne 526 US) (Ward Comments NA doc. Year Year 1932 1957 1980 1938 2006 2005 1980 1980 locality is Type 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) personal obs. Buchanan (1993) Plateau Network (in ed.) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres ? B-32/287 Weight AnnF WideAnnF Pres Wide Unc Pres Buchanan 3120 Unc Graybosch 902 PerF Wide Pres Unc Graybosch 755 PerF Wide Rep Unc Northern Colorado PerF Wide Pres Unc Graybosch 905 AnnF WidePerF Pres Wide Unc Pres Graybosch 782 Unc s.n. (BRCA) Gifford 1940 locality is Type Life form reweed PerF Wide Rep ?& Spence Hall’s willow- Hall’s herb raceme groundsmoke branching groundsmoke white-stem lavender-leaf lavender-leaf sundrops Paiute suncup AnnF Wide Rep Rare Sarah Topp, ﬁ narrowleaf four- o’clock fragrant sand- verbena low sand-verbena PerF RegEn Pres Unc Graybosch 858 evening-primrose Jones’ evening- primrose Common name

ssp. , ssp.

nana var. var. M. glabra canescens, C. harrisii lavandulifolius Oenothera scapoidea Chamerion angustifolium var. angustifolium circumvagum, C. danielsii C. hartwegii fragrans elliptica, A. nana var. Oxybaphus linearis, O. decumbens some authors include Synonyms/ notes Taxonomic

var. var. var.

var. var. var. Epilobium halleanum Gayophytum racemosum Gayophytum ramosissimum Oenothera albicaulis canescens Camissonia scapoidea scapoidea Epilobium angustifolium Calylophus lavandulifolius Abronia fragrans A. elliptica, Abronia nana A. nana Mirabilis linearis linearis Oenothera caespitosa crinita rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Nyctaginaceae Nyctaginaceae Appendix A. Conﬁ FamilyNyctaginaceae Species name biogeographic notes, cont. Onagraceae

Chapter 6: Appendices 63 Armstrong & White canyons (Natural Bridges NM), San Juan Co., UT (Rydberg & Garrett 9410 NY) Comments NA locality is Type NA doc. Year Year 1980 1980 1932 1931 1956 1957 1980 1958 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) Buchanan (1993) (BRCA) (BRCA) (BRCA) Buchanan (1993) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Graybosch 945 PerF Wide Pres Unc ST06290501 Topp PerF Wide Pres Unc Graybosch 881 PerF Wide PresPerF Unc Wide Buchanan 61 Pres Unc Buchanan 205 PerF Wide Rep ? Spence & PerF WidePerF Pres RegEn Unc Rep Graybosch 884 ? Spence & PerF Wide Pres Unc Buchanan 362 PerF Wide Hist ? Boyle 1162 (BRCA) 1931 Life form clustered broomrape slender bog orchid yellow evening- primrose pale evening- primrose spotted coral- root striped coral-root PerF Wide Pres white Lindley’s bog-orchid Unc Mueller s.n. (BRCA) 1980 bronze evening- primrose Bridges evening- primrose giant helleborine PerF Wide Pres Rare Graybosch 1055 long-tube evening-primrose hairy-throat evening-primrose Common name

striata var. var.

Vars. Vars. var. (2002) ora. ava leucostachys, P. leucostachys, P. Flora of North Aphyllon fasciculatum Platanthera sparsiﬂ not recognized in America O. ﬂ Welsh et al. (2003). Welsh Includes var. runcinata C. maculata maculata var. leucostachys O. cespitosa marginata Synonyms/ notes Taxonomic

var.

ava var. var. ora ora pallida ava Habenaria sparsifl Orobanche fasciculata Oenothera fl fl Oenothera howardii O. brachycarpa Oenothera longissima Oenothera pallida var. Corallorhiza maculata Corallorhiza striata C. striata Epipactis gigantea Habenaria dilatata Platanthera dilatata sparsifl Oenothera caespitosa marginata Oenothera coronopifolia rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Orchidaceae Orobanchaceae Onagraceae Onagraceae Onagraceae Onagraceae Orchidaceae Orchidaceae Orchidaceae Orchidaceae Onagraceae Appendix A. Confi FamilyOnagraceae Species name biogeographic notes, cont.

64 Annotated Checklist of Vascular Flora: Bryce Canyon National Park but introduced in Utah. and CO is Springdale, Co., Washington UT (Jones 5247 NY) Comments doc. Year Year 1956 Native to SW USA 1932 1957 State tree of UT 1927 1928 1957 1980 locality Type 1957 1980 1980 Native to Eurasia 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Tree Wide Pres Abun Cottam 4364 Tree Wide Pres Unc Buchanan 100 PerF Wide Pres Unc Graybosch 913 Life form r Tree Wide Pres Com106 Buchanan r Tree Wide Pres Com307 Buchanan California poppy AnnF Intro Pres ? Buchanan 13 white ﬁ blue sprucetwo-needle Treepinyon Wide Pres Unc Buchanan 133 limber pinewestern bristlecone pine Treeponderosa pine Wide Tree PresDouglas-ﬁ Wide Unc Pres Boyle Z283 (BRCA) Com 1938 Buchanan 251 shrubby gilia PerF RegEn Pres Unc Graybosch 1035 Common name woolly-base plantain scarlet gilia PerF Wide Pres Unc Graybosch 882 English plantain PerF Intro Presscarlet gilia Unc Graybosch 1072 PerF Wide Pres Com Boyle 1115 (BRCA) 1932

var. var.

ssp. ssp. maculata, I. E. californica californica Synonyms/ notes Taxonomic Gilia aggregata aggregata Gilia aggregata var. aggregata formosissima Gilia congesta frutescens . var glauca var. exilis aggregata maculata frutescens Eschscholzia californica Abies concolor Picea pungens Pinus edulis Pinus fl Pinus ponderosa Pseudotsuga menziesii scopulorum Pinus longaeva aristata P. Plantago eriopoda Plantago lanceolata Ipomopsis aggregata var. Ipomopsis aggregata var. Ipomopsis congesta var. rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Pinaceae Pinaceae Pinaceae Appendix A. Confi FamilyPapaveraceae Species name biogeographic notes, cont. Pinaceae Pinaceae Pinaceae Pinaceae Plantaginaceae Plantaginaceae Polemoniaceae Polemoniaceae Polemoniaceae

Chapter 6: Appendices 65

P. , and is Beaver Dam Mtns, Co., Washington UT (Fremont 447 US) Cedar Canyon, Iron Co., UT (Jones 5208c POM). USFWS: 3C in the BRCA herbarium & previous checklists. been called Phlox bryoides, diffusa P. austromontana Comments doc. Year Year 1932 1980 1980 locality Type 1970 locality is Type 1956 1940 This species has c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Hist ? Cottam 4399 (BRY) 1929 PerF Sparse Pres Unc Graybosch 1048 Life form long-leaf phlox PerF Wide Pres Unc Graybosch 752 carpet phlox PerF Wide Pres Unc Graybosch 886 desert phlox PerF Wide Pres Unc Graybosch 811 daggerleaf phlox PerF RegEn Pres Rare Buchanan 1720 Nuttall’s Nuttall’s linanthus prickly-phlox Shrub Wide Pres Unc Buchanan 26 matted prickly- phlox Common name

& var. var. ssp. stansburyi Includes vars. longifolia viscidula Var. not recognized Var. in Cronquist et al. (1984) Includes ssp. glabrata, hoodii, Leptodactylon pungens Linanthastrum nuttallii nuttallii, Leptosiphon nuttallii, Linanthus nuttallii nuttallii pungens, Linanthus pungens Linanthus caespitosus Synonyms/ notes Taxonomic

var. var. Phlox longifolia Phlox austromontana austromontana Phlox gladiformis caesia P. Phlox hoodii canescens Leptodactylon pungens Linanthastrum nuttallii Leptodactylon caespitosum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Appendix A. Conﬁ FamilyPolemoniaceae Species name biogeographic notes, cont.

66 Annotated Checklist of Vascular Flora: Bryce Canyon National Park

eld eld Co., UT panguicense Widtsoe, Garﬁ Co., UT (Ripley & Barneby 8570 CAS). USFS: Region 4 Sensitive var. var. is near Panguitch, Garﬁ (Jones s.n. POM). Comments NA locality is Type doc. Year Year 1957 1980 1928 2007 1931 1930 locality of Type 1931 c name with taxonomic and (BRCA) (BRCA) (BRCA) (1991) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Buchanan 318 PerF Wide PresPerF Unc Wide Graybosch 1005 Pres Unc Buchanan 166 PerF LocEn Rep ? Pelt and Schelz Van AnnF Wide Pres Rare ST08220701 Topp Shrub Wide Pres Unc Buchanan 542 Shrub Wide Pres Unc Foster 11009 (BRY) 1993 PerF RegEn Pres Unc Buchanan 2992 Life form PerF Wide Pres Unc Buchanan 35 slender wild buckwheat western Jacob’s- ladder winged wild buckwheat Widtsoe wild buckwheat nodding wild buckwheat crisp-leaf wild buckwheat slender wild buckwheat Panguitch wild buckwheat Common name redroot wild buckwheat

var.

Flora

& E. in ssp. & vars.

alatum, Includes E. microthecum foliosum Synonyms/ notes Taxonomic P. occidentale, P. occidentale, P. P. caeruleum amygdalinum var. Pterogonum alatum (2005) of North America Vars. not Vars. recognized in Includes var. velutinum lancifolium et al. 2003 Welsh alpestre panguicense E. racemosum

var. var. var. var. var. orum cernuum Eriogonum panguicense Eriogonum microthecum simpsonii pterospermum Polemonium caeruleum Eriogonum alatum E. alatum Eriogonum aretioides Eriogonum cernuum var. Eriogonum corymbosum corymbosum Eriogonum microthecum laxifl Eriogonum racemosum racemosum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Polygonaceae Polygonaceae Polygonaceae Polygonaceae Appendix A. Confi FamilyPolemoniaceae Species name biogeographic notes, cont. Polygonaceae Polygonaceae Polygonaceae Polygonaceae

Chapter 6: Appendices 67 is Virgin River Valley, Co., Washington UT (Parry 237 ISC) Comments NA Native to Europe doc. Year Year 1957 1980 1980 1980 1980 1980 locality Type 2005 1932 1932 Native to Eurasia 1940 c name with taxonomic and (BRCA) (BRCA) Graybosch (1981) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range AnnF Wide Pres Unc Graybosch 967 AnnF RegEn Pres Rare Graybosch 846 PerF RegEn PresPerF Unc Wide ST08110503 Topp Pres Unc Buchanan 88 AnnF Wide Pres Unc Buchanan 356 Life form ower alpine bistort PerF Wide Pres Unc Graybosch 1111 bracken fern Fern Wide Pres Rare Graybosch 1039 Sawatch knotweed curly dockwillow dock PerF PerF Intro Wide Presslender lip-fern Rep Unc Fern ? Foster 10841 (BRY) Wide 1932 Buchanan & Pres Native to Eurasia Unc Graybosch 769 kidney-shaped wild buckwheat yard knotweed AnnF Introblack-bindweed Pres AnnF Intro Unc Graybosch 1064 Pres ? Not provided Porter’s wild Porter’s buckwheat arid wild buckwheat pygmy-ﬂ rock-jasmine Common name

var.

& var. ssp. mexicanus, subilifera Bistorta vivipara aquilinum P. lanuginosum sawatchense R. mexicanus, R. salicifolius var. R. utahensis, triangulivalvis P. sawatchense P. P. aviculare P. aviculare Fallopia convolvulus Synonyms/ notes Taxonomic var. Includes ssp. puberulenta, subumbellata,

var. var. triangulivalvis pubescens johnstonii Polygonum viviparum Rumex crispus Cheilanthes feei Pteridium aquilinum var. Androsace Rumex salicifolius ssp. Polygonum douglasii var. Eriogonum subreniforme Eriogonum umbellatum porteri Eriogonum umbellatum subaridum Polygonum aviculare arenastrum, P. Polygonum convolvulus septentrionalis rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Polygonaceae Polygonaceae Polypodiaceae (Adiantaceae) Polypodiaceae (Dennstaedtiaceae) Primulaceae Polygonaceae Polygonaceae Appendix A. Conﬁ FamilyPolygonaceae Species name biogeographic notes, cont. Polygonaceae Polygonaceae Polygonaceae Polygonaceae

68 Annotated Checklist of Vascular Flora: Bryce Canyon National Park from “canyon is above Tropic” the holotype. The UT Conservation Data Center considers this location to be in Bryce Canyon NP, but exact site is not known Wasatch Peak, Wasatch Sanpete Co., UT 1788 (Tidestrom US) Comments doc. Year Year 1980 1980 1932 1980 1932 1957 1932 1894 Jones’ specimen 1940 locality is Type c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (POM) (BRCA) Source Pop. size Park status Range PerF WidePerF Pres Wide Unc Pres ST08090505 Topp Unc Graybosch 1108 PerF Wide Pres Unc Buchanan 161 PerF Wide Pres Unc Graybosch 980 PerF Wide Pres Unc Graybosch 874 Life form owered pink wintergreen PerF Wide Preswhiteveined Uncwintergreen Graybosch 1033 Columbian monkshood cutleaf anemone PerF Wide PresColorado Unccolumbine Graybosch 1027 green-ﬂ wintergreen dark-throat shooting-star silvery primrose PerF Wide Rep ? Jones 5312av Common name rock columbine PerF RegEn Pres Unc Graybosch 1349

ssp.

. huberi & picta, & orum. P. dentata P. var. da, saxicola, da, UT Flora stylosa asarifolia includes Includes var. ochroleucum A. tetonensis, includes vars. multiﬁ & Includes vars pinetorum ochroleuca Includes “subvars” from coriaceum, atwoodii, D. pauciﬂ Synonyms/ notes Taxonomic

var. var. var. tetonensis Pyrola chlorantha virens P. Pyrola pictaAconitum columbianum picta P. columbianum Anemone multifi var. Aquilegia coerulea Aquilegia scopulorum pulchellum Dodecatheon pulchellum Primula incana Pyrola asarifolia rotundifolia P. scopulorum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Appendix A. Confi FamilyPrimulaceae Species name biogeographic notes, cont. Primulaceae Pyrolaceae (Ericaceae)

Chapter 6: Appendices 69 Manti Canyon, Sanpete Co., UT (Jones s.n. POM) is Toquerville, Co., Washington UT (Jones 1716 GH) Comments doc. Year Year 1934 1932 1980 1932 1980 2005 1932 1978 1940 locality Type c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Com Graybosch 771 PerF Wide Pres Unc Buchanan 401 PerF WidePerF Pres Wide RareShrub Pres Wide ST09100501 Topp Unc PresShrub Wide Buchanan 272 Unc Pres Buchanan 2955 Com Boyle 1113 (BRCA) 1940 locality is Type Life form Columbian virgin’s-bower white virgin’s- bower Nuttall’s larkspurNuttall’s PerF Widemarsh buttercup PerF Pres Wide Unc Pres Foster 10894 (BRY)drab buttercup 1993 Unc Graybosch 772 PerF Wide Pres Unc Graybosch 875 pale larkspur PerF Wide Pres Unc Graybosch 848 alpine meadow- rue Fendler’s meadow-rue Fendler’s mountain-lilac Utah mountain- lilac Utah serviceberry Shrub Wide Pres Unc Buchanan 150 Common name

var. Includes ssp. alpeophilus cymbalaria, Atragene columbiana Delphinium nelsonii R. cymbalaria var. Halerpestes cymbalaria saximontana R. inamoenus inamoenus. var. D. scaposum Synonyms/ notes Taxonomic

var. Clematis columbiana C. pseudoalpina, Clematis ligusticifolia Delphinium andersonii Delphinium nuttallianum Ranunculus cymbalaria Ranunculus inamoenus Thalictrum alpinum Thalictrum fendleri Ceanothus fendleri Ceanothus martinii Amelanchier utahensis scaposum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Ranunculaceae Appendix A. Conﬁ FamilyRanunculaceae Species name biogeographic notes, cont. Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Thalictraceae) Ranunculaceae (Thalictraceae) Rhamnaceae Rhamnaceae Rosaceae

70 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Co., UT 10 km E of Oak Millard Co., City, UT (Holmgren & Holmgren 11298 NY) head of Sevier River 25 miles S of Panguitch, Garﬁ (Jones 6032 POM) Mt, Barrette near Marysvale, Piute Co., UT (Rydberg & Carlton 7261 NY) Comments doc. Year Year 1932 locality is ca Type 1940 1959 1980 1957 1932 1931 locality is Type 1957 1932 1970 2005 locality is Type c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Plateau Network (in ed.) Source Pop. size Park status Range Shrub Wide Pres Unc Buchanan 92 PerF WidePerF Pres RegEn Unc Rep Buchanan 1480 Unc Northern Colorado Shrub Wide Pres Unc Buchanan 115 PerF Wide Pres Unc Graybosch 877 Life form curl-leaf mountain mahogany bicrenate cinquefoil modest cinquefoil alder-leaf alder-leaf mountain mahogany Apache plume ShrubVirginia Widestrawberry Presbush oceanspray Unc Shrub Wide ivesiaGordon’s Graybosch 942 Pres ivesiaKing’s PerF Unc WideSevier ivesia Buchanan 217 PerF Hist Sparse ? Hist PerF RegEn ? Cottam 2782 (BRY) Pres 1927 common apple Unc B-32/327 Weight squaw-apple Graybosch 1046 Tree Introsilverweed Shrub Pres Wide Pres Rare PerF Kitchen s.n. (BRCA) Unc 1998 Wide Buchanan 219 Native to Eurasia Pres Unc Graybosch 775 Common name

glauca var. P.

in platypetala C. ledifolius intercedens Includes ssp. bicrenata P. concinna. P. Included in rubricaulis Holmgren et al. (1997) & Synonyms/ notes Taxonomic

var. bicrenata modesta Cercocarpus ledifolius intermontanus Cercocarpus montanus Fallugia paradoxa Fragaria virginiana Holodiscus dumosus Ivesia gordonii Ivesia sabulosa Malus pumila Pyrus malus Potentilla anserina Argentina anserina Potentilla concinna var. Potentilla concinna var. Ivesia kingii Peraphyllum ramosissimum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Appendix A. Conﬁ FamilyRosaceae Species name biogeographic notes, cont. Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae

Chapter 6: Appendices 71 Comments doc. Year Year 1931 1931 1931 2006 1980 1931 1931 1980 1980 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Shrub Wide Pres Unc Graybosch 979 Shrub Wide Pres Unc Buchanan 299 Shrub Wide Pres Unc Buchanan 165 PerF WidePerF Pres Disj Rare ST06280602 Topp Pres Unc Graybosch 1017 Shrub Wide Pres Com Boyle 1106 (BRCA) 1940 PerF Wide Pres Unc Boyle 1137 (BRCA) 1931 Life form Nootka rose roseWoods’ Shrub Wide Hist Shrubcommon red Wide ?raspberry Pres Unc Boyle Z295 (BRCA) Graybosch 910 1932 shrubby cinquefoil western chokecherry cliff-rose Shrub Wide Pres Unc ST06160601 Topp woolly cinquefoil PerF WidePennsylvania cinquefoil PresPlatte River Unccinquefoil Buchanan 2998 antelope bitterbrush hair-tuft hair-tuft cinquefoil Common name small bedstraw PerF Wide Pres Unc Graybosch 1113

var. var.

var. ssp. var. & R. manca orum oribunda, D. oribunda, fl melanocarpa dum oribunda, oribunda ultramontana. Includes R. idaeus aculeatissimus, R. ideaus melanolasius, R. idaeus sachalinensis Dasiphora fruticosa ssp. fl fruticosa, Pentaphylloides fl Padus virginiana ssp. stansburiana, P. Cowania mexicana hippiana Synonyms/ notes Taxonomic Includes ssp. columbianum, trifi subbifl

var. var. dum melanocarpa stansburyana Rubus idaeus strigosus Rosa nutkana hispida Rosa woodsii R. woodsii Potentilla fruticosa Pentaphylloides Potentilla hippiana hippiana P. Potentilla pensylvanica Potentilla plattensis Prunus virginiana var. Purshia mexicana var. Purshia tridentata Potentilla crinita Galium trifi rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Appendix A. Confi FamilyRosaceae Species name biogeographic notes, cont. Rubiaceae

72 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Comments doc. Year Year 2006 1959 2006 1931 1980 1980 1931 1932 1957 1958 1980 1957 1932 1994 c name with taxonomic and (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range Tree Wide Pres Unc ST06150601 Topp Tree Wide Pres Unc Buchanan 227 Tree Wide Pres Unc ST06150602 Topp PerF Wide Pres Unc Graybosch 824 PerF Wide Pres Unc Buchanan 301 Shrub Wide Pres Unc Foster 11083 Life form ax PerF Wide Pres Unc749 Graybosch lance-leaf cottonwood coyote willow Shrub Wide Pres Unc Buchanan 158 Bebb’s willowBebb’s Shrubshort-fruit willow Wide Shrubyellow willow Wide Pres Pres Shrub Unc Wide Rare Buchanan 465 Pres Foster 11006 (BRY) willowGeyer’s 1993 Unc Shrubbastard toadﬂ ST06270602 Wide Topp Pres Unc Graybosch 1008 milky kelloggia PerFnarrow-leaved Widecottonwood Pres Unc Graybosch 917 Fremont cottonwood quaking aspen Tree Wide Pres Unc Buchanan 308 slender woodlandstar Common name marsh grass-of- Parnassus golden currant Shrub Wide Hist ? s.n. (BRCA) Weight 1932 wax currantwhitestem Shrubgooseberry Wide Pres Unc Graybosch 1362 P.

tenuis var. Hybrid var. var. var. watsonii P. fremontii P. angustifolia S. exigua exigua S. lutea, rigida var. acuminata. P. deltoides P. acuminata, P. P. of fremontii L. tenella Synonyms/ notes Taxonomic Includes var. Includes var. aureum R. cereum pedicellare

var.

var. var. var. x fremontii montanensis Salix bebbiana Salix brachycarpa Salix exigua stenophylla Salix geyeriana Comandra umbellata pallida Salix eriocephala watsonii Kelloggia galioides Populus angustifolia Populus fremontii var. Populus tremuloides Populus intercurrens Lithophragma tenellum inebrians Parnassia palustris var. Ribes aureum Ribes cereum R. aureum Ribes inerme rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Salicaceae Salicaceae Salicaceae Salicaceae Santalaceae Salicaceae Salicaceae Appendix A. Conﬁ FamilyRubiaceae Species name biogeographic notes, cont. Salicaceae Salicaceae Salicaceae Saxifragaceae Saxifragaceae (Parnassiaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae)

Chapter 6: Appendices 73 ed eld Co., eld from BRCA were based on misidentifi specimens is along road to Bryce Point, Bryce Canyon Garfi NP, UT (Holmgren & Reveal 2017 NY). USFS: Region 4 Sensitive, USFWS: former C2 candidate Comments NA doc. Year Year 2005 Previous reports 1971 1980 1941 1980 1931 1980 1932 1965 locality Type 1957 c name with taxonomic and Plateau Network (in ed.) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) 2017 (NY) (BRCA) Buchanan (1993) Source Pop. size Park status Range AnnF Wide Rep Unc Northern Colorado PerF WideAnnF Pres Wide Unc Pres Graybosch 1099 Unc Graybosch 817 PerF LocEn Pres Unc Holmgren & Reveal PerF Wide Pres Com Buchanan 22 AnnF Wide Rep ? Spence and Life form ower ower ower Wright’s bird’s- Wright’s beak violet owl-clover AnnF RegEn Pres Rare Graybosch 1047 yellow monkeyfl reddish monkeyfl yellow owl-clover AnnF Wide Presdwarf lousewort Unc PerF Graybosch 1060 Wide Pres Unc Buchanan 3071 Reveal’s Reveal’s paintbrush desert paintbrush PerF WideWyoming Prespaintbrush Unc Graybosch 787 bird’s-beakKing’s AnnF RegEnsmallfl Presbird’s-beak Unc Buchanan 2963 Common name

var. martinii O. purpureo-albus C. revealii dubia, C. applegatei ssp. Synonyms/ notes Taxonomic orus revealii kingii Cordylanthus wrightii Mimulus guttatus Mimulus rubellus Orthocarpus luteus Orthocarpus purpureoalbus Pedicularis centranthera Castilleja linariifolia C. linariaefolia Castilleja parvula var. Cordylanthus kingii var. Cordylanthus parvifl Castilleja chromosa C. angustifolia rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix A. Confi FamilyScrophulariaceae Species name biogeographic notes, cont.

74 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld eld Co., UT eld Co., UT Mammoth Creek, Markagunt Plateau, Garfi Co., UT (Jones 6026b US) near Canaan Ranch, Rockville, Co., Washington UT (Jones 5262 POM) Aquarius Plateau, Garfi 462 GH) (Ward is lower Provo Canyon, Utah Co., 776 UT (Watson GH) is St. George, Co., Washington UT (Jones 5110h US) Red Canyon, Garfi (Eastwood & Howell 783 CAS). USFS: Region 4 Sensitive, US FWS: former C2 candidate Comments doc. Year Year 1930 locality is Type 1980 locality Type 1971 locality is Type c name with taxonomic and (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF LocEn Pres Unc Boyle 1131 (BRCA) 1931 locality is Type PerF Wide Pres Unc Foster 11038 (BRY) 1994 locality is Type PerF Wide Pres Unc Foster 11040 (BRY) 1994 locality Type PerF Wide Pres Unc Graybosch 1034 PerF LocEn Pres Unc Graybosch 791b PerF RegEn Pres Unc Boyle 1138 (BRCA) 1930 Life form recracker Markagunt penstemon Rockville penstemon dusty penstemon PerF RegEn Pres Eaton’s Uncfi ST08130503 Topp undulate penstemon Red Canyon penstemon painted penstemon Common name was “Penstemon brycensis” never validly published. Synonyms/ notes Taxonomic

var. var. var. eatonii undosus leiophyllus Penstemon pachyphyllus congestus Penstemon eatonii var. Penstemon eatonii var. Penstemon leiophyllus Penstemon comarrhenus Penstemon bracteatus Penstemon caespitosus desertipicti rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix A. Conﬁ FamilyScrophulariaceae Species name biogeographic notes, cont. Scrophulariaceae

Chapter 6: Appendices 75 is between & Wash Hatch’s Monticello, San Juan Co., UT (Eastwood s.n. CAS) America but introduced to UT America America, may be adventive in UT. Comments doc. Year Year 1980 1980 1986 1929 locality Type 1980 Native to Eurasia 1972 1998 Native to C North 1998 Native to South 1932 Native to C North 1980 Native to Eurasia 2006 2006 Native to Asia 1980 c name with taxonomic and (BRCA) (BRCA) (UTC) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) EB07290602 (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF RegEn Pres Unc Graybosch 1007 Shrub Intro Pres Unc Graybosch 944 PerF WidePerF Pres Wide Unc Pres Graybosch 1091 Rare Gottschalk 14 PerF Wide Pres UncAnnF Graybosch 1009 Intro PresAnnF Unc Wide Buchanan s.n. Hist ? B-32/290 Weight Life form ower ve-stamen smallﬂ penstemon ﬁ tamarisk beaked penstemon Rocky Mountain penstemon Utah penstemon PerF Wide Pres Unc Graybosch 849 woolly mullein PerFAmerican Introbrooklime Presbuffalobur Unc Graybosch 1093 AnnFground-cherry Intronightshade Prescut-leaf Uncnightshade Buchanan s.n. common cattail PerG WideSiberian elm Prespinnate angelica Rare Tree PerF Ballenger Intro Wide Pres Pres Unc Unc Fertig 22543 Graybosch 1110 Common name

var. S. villosum, physalifolium ramosissima P. bridgesii P. nitidibaccatum Synonyms/ notes Taxonomic orum orus aberrans Penstemon procerus var. Penstemon rostrifl Penstemon strictus Penstemon utahensis thapsus Verbascum americana Veronica Solanum rostratum Solanum sarrachoides Solanum trifl chinensisTamarix pentandra, T. T. latifolia Typha Ulmus pumila Angelica pinnata rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientifi Appendix A. Confi FamilyScrophulariaceae Species name biogeographic notes, cont. Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Solanaceae Tamaricaceae Typhaceae Ulmaceae Umbelliferae (Apiaceae)

76 Annotated Checklist of Vascular Flora: Bryce Canyon National Park C. Type Type var. var. eld Co., UT approaches purpureus purpureus. locality is Cedar Breaks, Cedar Breaks NM, Iron Co., UT (Mathias 703 MO). USFS: Region 4 Sensitive, USFWS: former C2 candidate. Bryce Canyon, Garﬁ (Mathias 670 MO). USFWS: 3C “Southern Utah” (Parry 85) Comments doc. Year Year 1987 1932 1979 1931 1929 locality is Type 1980 locality is Type 1979 1980 1959 c name with taxonomic and (BRY) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Source Pop. size Park status Range PerF Wide Pres Unc Hallsten 2556 PerF Wide Pres Com Buchanan 108 PerF Wide Pres Unc Buchanan 3064 PerF LocEn Pres Unc Peabody 28 (BRCA) 1931 BRCA material PerF RegEn PresPerF Com Wide Graybosch 1316 Pres Unc Graybosch 986 Life form cutleaf water- parsnip Lemmon’s spring- Lemmon’s parsley mountain valerian least spring- parsely Colorado Plateau spring-parsley southern ligusticum least lomatium PerF LocEn Pres lomatiumParry’s Unc PerF Buchanan 3359 Wide Pres Unctobacco-root Graybosch 922 prostrate vervain PerF PerF Wide Wide Pres Pres Unc Unc Graybosch 734 Buchanan 464 Common name

var. Pseudocymopterus montanus pubicarpa C. purpureus Synonyms/ notes Taxonomic

var. var. Berula erecta incisa Cymopterus lemmonii Cymopterus minimus Cymopterus purpureus Ligusticum porteri Lomatium minimum Lomatium parryi acutiloba Valeriana acutiloba V. edulis Valeriana bracteata Verbena purpureus rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Appendix A. Conﬁ FamilyUmbelliferae (Apiaceae) Species name biogeographic notes, cont. Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Valerianaceae Valerianaceae Verbenaceae

Chapter 6: Appendices 77

& Picea exilis (rarely or Pinus Juniperus Pinus edulis Pseudotsuga Abies Pinus fl species) on ponderosa on osteosperma on monophylla P. on menziesii on on Comments doc. Year Year 1980 parasitic Typically 1931 parasitic Typically 1959 parasitic Typically 1957 parasitic Typically 1932 1980 1980 1957 parasitic Typically c name with taxonomic and Graybosch (1981) (BRCA) (BRCA) (BRCA) (BRCA) (BRCA) Graybosch (1981) (BRCA) Source Pop. size Park status Range PerF Wide Rep ? Buchanan & PerF Wide PresPerF Unc Wide Buchanan 527 Pres Unc Buchanan 310 PerF Wide Pres Unc Graybosch 876 PerF Sparse Pres Unc Buchanan 306 Life form r southwestern dwarf-mistletoe juniper mistletoe PerF Wide Pres Unc Graybosch 319 pinyon dwarf- mistletoe Douglas-fi dwarf-mistletoe hook-spur violet PerFnorthern bog Wideviolet Pres Unc Graybosch 995 limber pine dwarf-mistletoe goose-foot violet PerF Wide Rep ? Buchanan & Common name nis affi var. cyanocarpum A. campylopodum var. Synonyms/ notes Taxonomic

var.

var. Arceuthobium vaginatum cryptopodum Phoradendron juniperinum Arceuthobium divaricatum Arceuthobium douglasii Viola aduncaViola nephrophylla montanensis sororia V. V. Viola purpurea venosa Arceuthobium cyanocarpum rmed, historical, and reported vascular plant taxa of Bryce Canyon National Park, organized by family scientiﬁ Viscaceae Viscaceae Viscaceae Viscaceae Violaceae Appendix A. Conﬁ FamilyViolaceae Species name biogeographic notes, cont. Violaceae Viscaceae

78 Annotated Checklist of Vascular Flora: Bryce Canyon National Park ) L. erecta Native race ( from the Rocky Mountains is probably extralimital Range Comments Life form PerF RegEn ower AnnF Wide ower stickseed ower PerF Wide Glen Canyon hedgehog European stickseed AnnF Intro Native to Eurasia. meadow popcorn-ﬂ salt heliotrope PerF Wide spindle-rooted bluebell PerF RegEn spotted stickseed PerF Wide smallﬂ showy stickseed PerF Wide slender cryptanth AnnF Wide yellow-eye cryptanth PerF Wide orange milkweedthick-sepaled cryptanth PerF AnnF Wide Wide pallid milkweed PerF Wide western poison ivy Shrub Wide box elder Tree Wide black twinberry Shrub Wide cactus Rocky Mountain beeplant AnnF Wide blunt-leaf sandwort PerF Wide red elderberry Shrub Wide

by S.

Flora of by some interius var. ssp. decumbens. ora squarrosa, L. interius, terminalis var. var. included in (2003) racemosa var. var. ssp. M. oblongifolia avoculata var. E. engelmannii not recognized in Vars. North America echinata Plagiobothrys scouleri hispidulus Included in some authors A. tuberosa A. cryptoceras Negundo aceroides S. microbotrys, authors racemosa

var.

var. var. var. patens var. var. oralaterifl Moehringia avoculatafl Oreocarya oribunda rmed) taxa of Bryce Canyon National Park. obovatum Echinocereus engelmannii variegatus Lappula squarrosa L. squarrosa Plagiobothrys scouleri penicillatus Heliotropium curassavicum var. Mertensia fusiformis Hackelia patens Hackelia micrantha H. jessicae Hackelia fl Cryptantha gracilis Cryptantha crassisepala Cryptantha fl elachantha Asclepias tuberosa interior Asclepias cryptoceras cryptoceras Toxicodendron rydbergiiToxicodendron radicans Rhus radicans, T. Acer negundo A. negundo Sambucus racemosa microbotrys Lonicera involucrata Distegia involucrata Cleome serrulata Arenaria laterifl Cactaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Asclepiadaceae Asclepiadaceae Anacardiaceae FamilyAceraceae Species name notes Synonyms/Taxonomic Common name Caprifoliaceae (Adoxaceae) Caprifoliaceae Capparaceae (Cleomaceae) Caryophyllaceae (Alsinaceae) Appendix B. Potential (unconfi

Chapter 6: Appendices 79 Range Comments Life form eld agoseris eld PerF Sparse eld chickweed eld PerF Wide showy pussytoes PerF Sparse short agoseris PerF Wide ﬁ broom seepweed AnnF Wide barbwire Russian thistle AnnF Intro Native to Eurasia Nuttall’s poverty-weedNuttall’s AnnF Wide winged pigweed AnnF Wide American bugseed AnnF Wide Fremont’s goosefootFremont’s AnnF Wide greenmollydesert goosefoot AnnF PerF Sparse Wide umbrella starwort PerF Wide alpine pearlwortMenzies’ campion PerF PerF Wide Wide Bering chickweed PerF Wide bur ragweedcommon ragweed AnnF AnnF Wide Wide ﬁ reddish sandwort PerF Wide shrubby sandwort PerF Periph by

var. ssp. menziesii earlei, C. elatior var. pulcherrima capillare ssp. var. dasycephala menziesii, ssp. var. americanum var. C. pratericola some authors Includes vars. rydbergii Includes C. fremontii Anotites menziesii Included in var. Included in var. beeringianum strictum rubella macradenia

var. rubella Minuartia rubella, Tryphane

agrestis var. var. var. rmed) taxa of Bryce Canyon National Park, cont. Agoseris glauca dasycephala Agoseris glauca Suaeda calceoliformis S. depressa Salsola paulsenii Monolepis nuttalliana Cycloloma atriplicifolia C. atriplicifolium Corispermum americanum Chenopodium desiccatum Chenopodium fremontii fremontii Bassia americana Kochia americana Silene menziesii Stellaria umbellata S. menziesii Sagina saginoides Ambrosia artemisiifoliaAntennaria pulcherrima A. artemisiifolia A. pulcherrima Arenaria macradenia Eremogone macradenia Ambrosia acanthicarpa Franseria acanthicarpa Cerastium beeringianum C. beeringianum Cerastium arvense C. strictum, arvense Arenaria rubella Compositae Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCaryophyllaceae (Alsinaceae) Species name notes Synonyms/Taxonomic Common name (Asteraceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Appendix B. Potential (unconﬁ

80 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Canyon, Zion NP, Canyon, Zion NP, Co., UT Washington & Hays 24379 (Welsh BRY). Range Comments Life form Bigelow’s sagebrushBigelow’s silver sagebrush Shrub Wide Shrub Wide mountain pussytoeshairy arnica PerFrayless arnica Wide biennial wormwood PerF PerF PerF Wide Wide Wide nodding bur-marigold PerF Wide rayless alkali aster AnnF Wide budsagemountain big sagebrush Shrub Wide Shrub Wide Louisiana wormwood PerF Wide Louisiana wormwood PerF Wide sand sagebrush Shrub Wide Welsh’s asterWelsh’s cutleaf PerF LocEn locality is Zion Type PerF Wide spruce aster PerF Sparse

var.

var. angusta ssp. ssp. biennis lifolius parryi var. var. A. tridentata A. ovata ora, Seriphidium ora, Includes viscidulum Includes Brachyactis ciliata pauciﬂ vaseyanum apricum

var. var. var. viscidula Seriphidium canum apricus Symphyotrichum foliaceum var. var. lifoliafi Oligosporus rmed) taxa of Bryce Canyon National Park, cont. Artemisia cana Artemisia fi Arnica mollis Arnica parryiArtemisia biennisArtemisia bigelovii A. parryi A. biennis Antennaria umbrinella Aster foliaceus Artemisia tridentata vaseyana Aster brachyactis Symphyotrichum ciliatum, Artemisia spinescens Picrothamnus desertorum Artemisia ludoviciana mexicana Artemisia ludoviciana ludoviciana Bidens cernua Aster welshiiBahia dissecta Symphyotrichum welshii Amauriopsis dissecta Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi

Chapter 6: Appendices 81 George, Washington George, Washington Co., UT (Palmer s.n. NY) weed. Native to Eurasia Lake, Sevier Co., UT (Jones 5715 NY) Creek, Grass Valley, Piute Co., UT (Welsh 14369 BRY). Range Comments Life form ower brickellbush ower PerF Wide wingless thistle PerF RegEn puberulent rabbitbrush Shrub Wide longbract rabbitbrush Shrub Wide smoothseed rabbitbrush Shrub RegEn locality is St. Type rush rabbitbrush Shrub RegEn Greene’s rabbitbrushGreene’s Shrub Wide squarrose knapweed PerFEastwood-plant Intro Utah state noxious PerF Wide longleaf brickellbush Shrub Wide California brickellbushtassel-ﬂ Shrub Wide spiny brickellbush Shrub Wide slender hawksbeard PerF Wide Fish Lake thistle thistleThorne’s PerF RegEnmountain hawksbeard locality is Fish PerF Type PerF LocEn locality is Otter Type Wide

juncea howardii var. var. var. var. atractyloides var. longifolia var. parva var. attenuatus, var. var. C. arizonicum bipinnatum Var. not recognized by Cronquist Var. (1994), C. parryi Ericameria parryi Ericameria nauseosa leiosperma Ericameria nauseosa C. triumfettii

var. var.

orus var.

var. var. var. var. ora puberulus clavatum Cirsium clavatum bipinnatum Cirsium calcareum Chrysothamnus viscidiﬂ var. Chrysothamnus parryi howardii Chrysothamnus nauseosus leiospermus Chrysothamnus nauseosus junceus Chrysothamnus greenei Brickellia longifoliaCentaurea virgata squarrosa B. longifolia Chamaechaenactis scaposa C. scaposa Brickellia grandiﬂ Brickellia californica Brickellia atractyloides B. atractyloides Crepis atribarba Psilochenia atribarba Cirsium scariosum thorneae Crepis acuminata Psilochenia acuminata Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

82 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Mountains, San Juan Co., UT (Newberry s.n. GH) NP, Washington Co., Washington NP, UT (Pilsbry s.n. PH). USFWS: former C2 candidate Kanab, Kane Co., UT (Thompson s.n. GH) “Southern Utah” (Parry 120 GH) Marysvale, Piute Co., UT (Jones 5386 POM) Range Comments Life form ower PerF RegEn locality is Type ower PerF RegEn locality is Abajo Type purpledisk sunﬂ orange sneezeweed PerF Wide meadow goldenweed PerF Wide Zion daisy PerF LocEn locality is Zion Type blanketﬂ Parry’s goldenweedClement’s PerF Wide Utah daisywhite thoroughwort PerF PerF Wide lowland cudweed RegEn locality is Type AnnF Wide Marysvale daisy PerF RegEn locality is Type smooth daisy PerF Wide beautiful daisy PerF Wide splendid daisy PerF Wide silver daisy PerF Wide sionis,

var. E. var. trilobatus by some authors var. included in agellaris hoopesii clementis E. ﬂ formosissimus

var. rmed) taxa of Bryce Canyon National Park, cont. Helianthella microcephala Helenium hoopesii Hymenoxys hoopesii, Dugaldia Haplopappus lanceolatus Pyrrocoma lanceolata Gaillardia parryi Haplopappus clementis Pyrrocoma clementis Erigeron utahensis utahensis Eupatorium herbaceum Ageratina herbacea Gnaphalium palustre Erigeron nauseosus Erigeron sionis E. proselyticus, sionis Erigeron leiomerus Erigeron formosissimus Erigeron eximius E. superbus, Erigeron argentatus Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconﬁ

Chapter 6: Appendices 83 eld the Muddy River, San the Muddy River, Rafael Reef, Emery Co., UT (Atwood & Goodrich 8652 BRY). USFWS: former C2 candidate Bromide Pass, Henry Mountains, Garfi Co., UT (Jones 5695y US) Range Comments Life form ower PerF Wide venerve sunfl venerve marsh-elder AnnF Wide Cooper’s hymenoxysCooper’s PerF Wide low hymenoxys PerF LocEn locality is E of Type lanate hymenoxys PerF Wide fi Bigelow’s asterBigelow’s PerF RegEn locality is Type woolly groundsel PerF Wide western groundsel PerF Wide cut-leaved groundsel PerF Wide Douglas’ groundsel PerF Wide

var. commixta var. accidus accidus commixta, lanata, Tetraneuris lanata, Tetraneuris var. caespitosa, T. caespitosa, T. var. var. accidus, S. fl nana fl var. H. acaulis acaulis brevifolia Dieteria bigelovii S. longilobus, fl var. douglasii

nana acaulis H. depressa, Tetraneuris var.

var. var. var. var. rmed) taxa of Bryce Canyon National Park, cont. Machaeranthera commixta M. bigelovii Iva xanthifolia Cyclachaena xanthifolia Hymenoxys cooperi Actinella cooperi Hymenoxys acaulis Hymenoxys acaulis Helianthella quinquenervis caespitosa Senecio douglasii longilobus Senecio canus Packera cana Senecio integerrimus exaltatus Senecio eremophilus kingii Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconﬁ

84 Annotated Checklist of Vascular Flora: Bryce Canyon National Park “Southern Utah” (Parry 109 GH) Podunk Creek”, Kane Co., UT (Malmsten 131 US) believed by Madsen (2001) to be just outside the SW corner of Bryce USFS: Canyon NP. Region 4 Sensitive, USFWS: former C2 candidate. Range Comments PerF Wide Life form showy goldeneye PerF Wide scapose greenthread PerF Wide locality is Type balsam goundselRocky Mountain groundsel PerF PerF Wide Wide New Mexico groundsel PerF Wide littleleaf horsebrush Shrub Wide baby goldenrodwhite-plume wire-lettuce AnnF PerF Wide Wide Podunk groundsel PerF LocEn locality is “Little Type Canadian goldenrodRocky Mountain PerFgoldenrod Wide Arizona mulesears PerF RegEn

var. Flora of Flora of var. var. var. ora Vars. not Vars. (2006). (2006). Flora of North exigua ssp. (2006) ora Vars. not recognized in Vars. North America Heliomeris multiﬂ multiﬂ Packera streptanthifolia streptanthifola. recognized in America S. altissima, lepida mutabilis Vars. not recognized in Vars. North America salebrosa Packera neomexicana

var. var.

var. var. var. var. ora rmed) taxa of Bryce Canyon National Park, cont. ora subnudum Viguiera multifl multifl Senecio streptanthifolius streptanthifolius Solidago canadensis salebrosa Senecio pauperculus Packera paupercula Thelesperma subnudum Stephanomeria exigua glabrata Tetradymia S. exigua Solidago multiradiata Solidago nana Senecio malmstenii scopulorum Wyethia arizonica Senecio neomexicanus mutabilis Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name notes Synonyms/Taxonomic Common name Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi

Chapter 6: Appendices 85 George, Washington George, Washington Co., UT (Jones 1636 NY) Range Comments Life form silver bladderpod PerF Wide hairy-pod pepperwortJones’ pepperwort AnnF Wide PerF Widesharp-leaf twinpod locality is St. Type PerF Wide slender whitlow-grassprairie pepperwort PerF Wide AnnF Wide watercress PerF Intro Native to Europe mountain draba AnnF RegEn wedgeleaf draba AnnF Wide golden draba PerF Wide California tansy-mustard PerF Wide blue mustard AnnF Intro Native to Asia heartleaf bittercress PerF Wide soldier rockcress PerF Wide lance-leaved stonecrop PerF Wide

var. macrocarpum lanceolatum Vars. not recognized in Vars. P. australis P. Includes var. Includes var. lemmonii. Holmgren et al. (2005) subalpinum includes ssp.

var. var.

var. var. nana D. albertina

lemmonii Boechera lemmonii var. var. var. var. orum cinalenasturtium-aquaticum Rorippa rmed) taxa of Bryce Canyon National Park, cont. Lepidium montanum Physaria ludoviciana Lesquerella ludoviciana Lepidium densifl Lepidium lasiocarpum lasiocarpum jonesii Nasturtium offi Physaria acutifolia acutifolia Draba stenoloba ramosum Draba rectifructa Draba cuneifolia cuneifolia Draba aurea Descurainia californica Chorispora tenella Cardamine cordifolia cordifolia Arabis lemmonii Sedum lanceolatum Amerosedum lanceolatum, Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) FamilyCrassulaceae Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconfi

86 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Co., UT (Ward Aquarius Plateau, Garﬁ 589 GH) Range Comments Life form false London rocket AnnF Intro Native to Eurasia Ward’s bladderpodWard’s PerF RegEn locality is Type tumble mustard AnnF Intro Native to Europe low yellowcress AnnF Wide alpine yellowcressblunt-leaf yellowcress PerF PerF Wide Wide hardstem bulrush PerG Wide Parish’s spikerushParish’s PerG Wide analogue sedge PerG Wide Liddon sedge PerG Wide western sedge PerG Wide blunt sedge PerG Wide Hayden’s sedgeHayden’s PerG Wide Eggleston’s sedgeEggleston’s PerG RegEn ebony sedge PerG Wide soft-leaved sedge PerG Wide northern bog sedge PerG Wide beautiful sedge PerG Wide water sedge PerG Wide entire-leaf thelypody PerF Wide P.

acutus ssp. var. var. by Holmgren included in aquatilis acutus, var. parvifolia ssp. var. et al. (2005) occidentalis, Schoenoplectus lacustris Schoenoplectus acutus gynocrates T. integrifolium T. kingii alpina R. alpina curvipes var. var. rmed) taxa of Bryce Canyon National Park, cont. integrifolium Thelypodium integrifolium var. Sisymbrium loeselii Physaria wardii Lesquerella wardii, Sisymbrium altissimum Rorippa curvipes Rorippa curvipes Rorippa tenerrima Scirpus acutus Schoenoplectus acutus Eleocharis parishii Carex simulata Carex petasata Carex occidentalis Carex obtusata Carex haydeniana Carex egglestonii Carex ebenea Carex disperma Carex dioica C. gynocrates, dioica Carex bella Carex aquatilis C. aquatilis Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) FamilyCruciferae (Brassicaceae) Species name notes Synonyms/Taxonomic Common name Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Appendix B. Potential (unconﬁ

Chapter 6: Appendices 87 “central Utah”, Emery Co?, UT (Parry 93 US). USFWS: 3C Africa W North America, but introduced in Utah. Range Comments Life form lanceleaf phacelia PerF Wide spike trisetum PerG Wide ticklegrass PerG Wide spike bentgrass PerG Wide Torrey’s ephedraTorrey’s Shrub Wide silverberry Shrub Periph alkali bulrush PerG Wide white scorpion-weed AnnF Wide European milfoil PerF Intro Native to Europe. Utah fescueperennial ryegrass PerG RegEnannual bluegrass PerG locality is Type Intro Native to Eurasia and AnnGweeping alkaligrass Intro Native to Eurasia and PerG Intro Native to Eurasia tall wheatgrass PerG Intro fescueThurber’s Native to Eurasia reed canary grass PerG Wide PerGbog bluegrass Wide PerG Wide timber oatgrass PerG Wide brookgrass PerG Wide shortawn foxtail PerG Wide

var. var.

& paludosus, alba var. ponticus, var. aequalis alpina hastata, var. & alaskanum var. congdonii, T. montanum congdonii, T. Schoenoplectus maritimus, Bolboschoenus maritimus paludosus Includes vars. leucophylla, exalbescens Includes ssp. Agropyron elongatum, Thinopyrum ponticum rmed) taxa of Bryce Canyon National Park, cont. Scirpus maritimus Scirpus maritimus Myriophyllum spicatum M. exalbescens, spicatum Alopecurus aequalis A. aequalis Agrostis scabra A. hiemalis, geminata Agrostis exarata Ephedra torreyana Elaeagnus commutata Phacelia hastata Phacelia alba neomexicana P. Trisetum spicatum Trisetum Festuca dasyclada Argillochloa dasyclada Lolium perenne Poa annua Puccinellia distans Elymus elongatus E. elongatus Festuca thurberi Phalaris arundinacea Phalaroides arundinacea Poa leptocoma Danthonia intermedia Catabrosa aquatica FamilyCyperaceae Species name notes Synonyms/Taxonomic Common name Haloragaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Ephedraceae Elaeagnaceae Hydrophyllaceae Hydrophyllaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix B. Potential (unconﬁ

88 Annotated Checklist of Vascular Flora: Bryce Canyon National Park eld Montezuma Canyon, San Juan Co., UT (Eastwood s.n. POM) miles E of Widtsoe Junction, Garﬁ 20666 Co., UT (Welsh USFS: Region BRY). 4 Sensitive, USFWS: former C2 candidate (near Alton), Valley Kane Co., UT (Jones s.n. POM) Loa Pass, 0.8 mi E of Piute Co line, Wayne & Isely, Co., UT (Welsh, Moore 6473 BRY) Range Comments Life form straggling milkvetch PerF RegEn locality is Type Table Cliff milkvetch Cliff Table PerF LocEn locality is 6 Type milkvetchSiler’s milkvetchWelsh’s PerF RegEn locality is Sink Type PerFSearls’ prairie-clover LocEn locality is W of Type PerF Wide chickpea milkvetch PerF Intro Native to Europe browse milkvetch PerF Wide nettle-leaf horseminthairy hedge-nettle PerF Wide PerF Wide knotted rush PerG Wide Nevada rush PerG Wide Mertens’ rush PerG Wide mountain rush PerG Wide wand phacelia PerF Wide

var. & var. J. tracyi,

by some & includes badius heterophylla pilosa A. limnocharis A. loanus var. by Barneby (1989) Included in montii not recognized by Barneby Vars. (1989) Included in authors nevadensis Includes vars. J. xiphoides J. saximontanus, Includes ssp. virgata

var. var. var.

pilosa S. pilosa var. var. rmed) taxa of Bryce Canyon National Park, cont. palans Astragalus lentiginosus Astragalus limnocharis tabulaeus Astragalus subcinereus silerianus Astragalus welshii Dalea searlsiae Astragalus cicer Stachys palustris Astragalus cibarius Agastache urticifolia Juncus nodosus Juncus nevadensis Juncus mertensianus Juncus ensifolius montanus Phacelia heterophylla Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Juncaceae Juncaceae Juncaceae Juncaceae FamilyHydrophyllaceae Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconﬁ

Chapter 6: Appendices 89 Range Comments PerF Wide Life form ax AnnF Wide ower sweetpea ower PerF RegEn Rusby’s stickleafRusby’s PerF Periph King’s cloverKing’s PerF RegEn strawberry clover PerF Intro Native to Europe silvery sophora PerF Wide Wyeth’s lupineWyeth’s PerF Wide longbracted trefoilUtah lupine PerFstemless lupine Wide PerF PerF Wide Wide New Mexico checker- mallow autumn willow-herb AnnF Wide Nuttall’s licoriceNuttall’s largeﬂ PerF Wide dandelion camissonia PerF Wide white checker-mallow PerF Wide downy ﬂ foothills death camas PerF Wide Geyer’s onionGeyer’s PerF Wide nodding onion PerF Wide short-style onion PerF Wide red clover PerF Intro Native to Europe Rusby’s cloverRusby’s PerF Wide

glabrata & & & paniculatum ora utahensis var. utahensis lepidota brachypus candida crenulata cernuum caespitosus, L. var. var. var. var. var. L. argenteus caespitosus L. wyethii neomexicana L. laetivirens Includes vars. Includes vars. Includes vars. obtusum

rusbyi longipes T.

var. var. utahensis L. lepidus kingii var. var. geyeri orabrevifl Oenothera var. var. var. rmed) taxa of Bryce Canyon National Park, cont. Mentzelia rusbyi Nuttallia rusbyi Trifolium longipes Trifolium Trifolium kingii Trifolium Trifolium fragiferum Trifolium Sophora stenophylla Lupinus caudatus utahensis Lupinus lepidus Lupinus polyphyllus humicola Lathyrus lanszwertii Lotus plebeius L. longibracteatus Epilobium brachycarpum E. paniculatum laetivirens Sidalcea neomexicana Camissonia brevifl Glycyrrhiza lepidota G. lepidota Sidalcea candida Linum puberulum Mesynium puberulum Zigadenus paniculatus paniculatum Toxicoscordion Allium geyeri Allium cernuum Allium brevistylum Trifolium pratense Trifolium Loasaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Onagraceae Malvaceae Onagraceae FamilyLeguminosae (Fabaceae) Species name notes Synonyms/Taxonomic Common name Malvaceae Linaceae Liliaceae (Melanthiaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconfi

90 Annotated Checklist of Vascular Flora: Bryce Canyon National Park River, Kane Co., UT River, (Cronquist 10202 MO) Range Comments PerF Wide Life form r Tree Wide at-topped broomrape at-topped PerF Wide little grapefernearly coralroot Fern Sparse PerF Wide western evening-primrose PerF Wide diffuse groundsmokediffuse Paria evening-primrose AnnF PerF Wide RegEn locality is Paria Type Watson’s prickly-phloxWatson’s PerF Wide shy giliascarlet gilia AnnF Wide PerF Wide small collomia AnnF Wide Tweedy’s plantainTweedy’s PerF Wide subalpine ﬁ Engelmann spruce Tree Wide ﬂ northern bog orchid PerF Wide Rocky Mountain willow- herb Hornemann’s willow-herbHornemann’s PerF Wide deceptive groundsmoke AnnF Wide glandular willow-herb PerF Wide Hooker’s wild buckwheatHooker’s AnnF Wide pretty Jacob’s-ladder PerF Wide Great Basin navarretia AnnF Sparse slender phlox AnnF Wide

var.

& includes orum, G. orum,

humilis var. propinqua parviﬂ glandulosum compositum inconspica ssp. var. ssp. hirsutissima var. ssp. Includes vars. tenebrosum lasiospermum Includes vars. sinuata, G. ophthalmoides, sinuata, G. clokeyi I. tenuituba, Gilia aggregata macrosiphon Platanthera aquilonis, P. Platanthera aquilonis, P. huronensis G. diffusum G. diffusum pulcherrimum Phlox gracilis P. pulcherrimum P.

var. var.

var. var. var. da rmed) taxa of Bryce Canyon National Park, cont. delicatum Corallorhiza triﬁ Habenaria hyperborea Limnorchis hyperborea, Botrychium simplex Oenothera caespitosa Oenothera elata O. elata navajoensis Microsteris gracilis Ipomopsis aggregata Leptodactylon watsonii Linanthus watsonii Gilia inconspicua macrosiphon Collomia linearis Plantago tweedyi Abies lasiocarpaPicea engelmannii A. bifolia Orobanche corymbosa Epilobium saximontanum Gayophytum diffusum strictipes Epilobium hornemannii hornemannii Gayophytum decipiens Epilobium glandulosum E. ciliatum Eriogonum hookeri Polemonium pulcherrimum var. humilior Navarretia intertexta N. intertexta Orchidaceae Orchidaceae Ophioglossaceae Onagraceae Onagraceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Plantaginaceae Pinaceae Pinaceae Orobanchaceae Onagraceae Onagraceae Onagraceae Onagraceae FamilyOnagraceae Species name notes Synonyms/Taxonomic Common name Polygonaceae Polemoniaceae Polemoniaceae Appendix B. Potential (unconﬁ

Chapter 6: Appendices 91 Range Comments Life form PerF Wide ower PerF Wide owered fame- owered ower cream wild buckwheat PerF Wide lady fern Fern Wide Oregon woodsiaRocky Mountain woodsia Fern Fern Periph Wide Douglas’ knotweed AnnF Wide brittle bladder fernmale fern Fern Wide Fern Wide water smartweed PerF Wide sulfur wild buckwheat PerF Wide Chamisso’s candy-fl Chamisso’s small-fl fl alpine bitter-root PerF Wide alpine sorrel PerF Wide Kellogg’s knotweedKellogg’s western dock AnnF Wide PerF Wide

Flora var. Some orus, P. P. orus, from kelloggii, P. kelloggii, P. subalpinum, californicum desereticum (2005) scopulina & fenestratus, R. L. nevadensis. oregana ssp. var. var. includes vars. occidentalis C. tenuis ssp. kelloggii aureum var. ssp. & ssp. var. C. utahensis, orus lix-femina Includes var. Includes var. Includes authors include reevesiana, P. douglasii P. dichrocephalum emersum, Polygonum coccineum, Persicaria coccinea, amphibia E. umbellatum E. subalpinum, of North America confertiﬂ kelloggii A. ﬁ aquaticus

var. var. var. var. orumparvifl Phemeranthus lix-mas lix-femina rmed) taxa of Bryce Canyon National Park, cont. Eriogonum umbellatum umbellatum Cystopteris fragilis scopulinaWoodsia Lewisia pygmaea scopulina W. Oreobroma pygmaea. Polygonum kelloggii polygaloides P. Dryopteris fi oregana Woodsia oregana W. Polygonum douglasii douglasii Polygonum amphibium Polygonum amphibium Eriogonum umbellatum majus Talinum parvifl Talinum Montia chamissoi Crunocallis chamissoi Athyrium fi Rumex occidentalisRumex paucifoliuscyclosorum R. aquaticus Acetosella paucifolia Polygonaceae Polypodiaceae (Aspleniaceae, Athyriaceae) Polypodiaceae (Aspleniaceae, Woodsiaceae) Portulacaceae Polygonaceae Polypodiaceae (Aspleniaceae, Aspidiaceae) Polypodiaceae (Aspleniaceae, Woodsiaceae) Polygonaceae Polygonaceae FamilyPolygonaceae Species name notes Synonyms/Taxonomic Common name Portulacaceae Portulacaceae Polypodiaceae (Aspleniaceae, Athyriaceae) Polygonaceae Polygonaceae Appendix B. Potential (unconfi

92 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Range Comments PerF Wide Life form ower meadow-rue ower PerF Wide few-ﬂ Saskatoon serviceberry Shrub Wide bur buttercup AnnF Intro Native to Eurasia Macoun’s buttercupMacoun’s PerF Wide sagebrush buttercup PerF Wide elegant columbine PerF Wide red baneberry PerF Wide heart-leaf buttercup PerF Wide thread water-buttercup PerF Wide plantain buttercup PerF Wide white marsh- marshmarigold subalpine larkspur PerF RegEn sidebells pyrola PerF Wide sheathed pondweed PerF Periph slenderleaf pondweed PerF Wide tall cinquefoil PerF Wide rock spiraea PerF Wide large-leaved avens PerF Wide

includes ssp. var.

eradicatus, R. trichophyllus, var. subrigidum, B. var. var. arguta ssp. liformis ssp. R. oreogenes orum ora, Psychrophila ora, saximontanum testiculata Includes R. circinatus, longirostris, trichophyllus trichophyllus Batrachium longirostre, B. circinatum trichophyllum leptosepala Stuckenia ﬁ vaginatus occidentalis Drymocallis arguta, arguta

var. var. var. var. var.

orum sparsifl T. var. liformis rmed) taxa of Bryce Canyon National Park, cont. Geum macrophyllum perincisum Amelanchier alnifolia Thalictrum sparsifl Ranunculus testiculatus Ceratocephala orthoceras, C. Ranunculus macounii Caltha leptosepala C. bifl Aquilegia elegantula Ranunculus glaberrimus ellipticus Ranunculus cardiophyllus Ranunculus aquatilis diffusus Ranunculus alismifolius montanus Delphinium barbeyi Actaea rubra A. rubra Pyrola secunda Orthilia secunda Potamogeton fi occidentalis Potamogeton vaginatus Stuckenia vaginata, Coleogeton Potentilla arguta convallaria Petrophyton caespitosum Spiraea caespitosa Rosaceae Rosaceae Ranunculaceae (Thalictraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Pyrolaceae (Ericaceae) FamilyPotamogetonaceae Species name notes Synonyms/Taxonomic Common name Potamogetonaceae Rosaceae Rosaceae Appendix B. Potential (unconfi

Chapter 6: Appendices 93 and possibly E North America Range Comments Life form brook saxifrage PerF Wide sticky currant Shrub Wide alpine prickly currant Shrub Wide trumpet gooseberry Shrub Wide Scouler’s willowScouler’s hairy alumroot Shrub Wide PerF Wide planeleaf willow Shrub Wide Booth’s willowBooth’s Shrub Wide sweet-scented bedstraw PerF Wide shrubby bedstraw PerF Wide cleavers AnnF Wide scarlet paintbrush PerF Wide annual paintbrush AnnF Wide Norwegian cinquefoil AnnFsheep cinquefoil Intro Native to Europe PerF Wide wedgeleaf cinquefoilsticky cinquefoil PerFglandular cinquefoil Wide PerF PerF Wide Wide biennial cinquefoil AnnF Wide pygmy saxifrage PerF Wide

elmeri debilis, , includes var. var. debilis monspeliensis ssp. includes vars. minor ssp. P. gracilis P. planifolia ssp. & intermedia. odontoloma monica H. versicolor G. spurium Included in Drymocallis glandulosa var. by some authors S. hyperborea

var. var.

var. var. var. ovina

var. var. var. orum orum rmed) taxa of Bryce Canyon National Park, cont. orum Saxifraga odontoloma S. arguta, Micranthes Ribes viscosissimum viscosissimum Ribes montigenum Heuchera rubescens Ribes leptanthum versicolor Salix scouleriana Salix planifolia S. phylicifolia, Salix boothii Galium trifl multifl Galium multifl echinospermon Castilleja miniata Potentilla ovina Galium aparine Potentilla glandulosa Potentilla gracilis brunnescens Potentilla norvegica P.norvegica pseudorupestris Potentilla diversifolia diversifolia Potentilla biennis Saxifraga rivularis S. debilis, rivularis Castilleja exilis C. minor Saxifragaceae Saxifragaceae Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae Saxifragaceae (Grossulariaceae) Salicaceae Salicaceae Salicaceae Rubiaceae Rubiaceae Scrophulariaceae Rosaceae Rubiaceae Rosaceae Rosaceae Rosaceae Rosaceae FamilyRosaceae Species name notes Synonyms/Taxonomic Common name Scrophulariaceae Appendix B. Potential (unconfi

94 Annotated Checklist of Vascular Flora: Bryce Canyon National Park America Lake Mountains, Sevier Co., UT (Ward 327 GH) dale, Washington Co., dale, Washington UT (Jones 5250 in part US) Kane Co., UT (Siler s.n. GH) miles N of Kanab, Kane Co., UT (Maguire, Maguire, & Piranian 12279 UTC) Kanab, Kane Co., UT (Thompson s.n. GH) Range Comments Life form giant burr-reed PerF Wide coyote tobacco AnnF Wide jimson weed AnnF Intro Native to tropical American alpine speedwell PerF Wide purslane speedwellthyme-leaved speedwell AnnF PerF Wide Wide water speedwell PerF Intro Native to Europe cutleaf synthyris PerF RegEn locality is Fish Type smooth penstemon PerF penstemonSiler’s RegEn locality is Spring- Type penstemonPalmer’s PerF RegEn PerF locality is Osmer, Type penstemonThompson’s RegEn locality is 2.5 Type PerF penstemonWhipple’s RegEn locality is near Type PerF Wide beardlip penstemon PerF Wide elephant-head PerF Wide butter-and-eggs PerF Intro Native to Eurasia Eastwood’s paintbrushEastwood’s PerF RegEn

var. humifusum laciniata torreyi ssp. var. var. da humifusa, Veronicastrum humifusa, Veronicastrum serpyllifolium P. barbatus P.

var.

var. var. var. var. rmed) taxa of Bryce Canyon National Park, cont. Sparganium eurycarpum Nicotiana attenuata Datura stramonium Veronica serpyllifolia Veronica wormskjoldiiVeronica serpyllifolia Veronica nutans V. Veronica peregrina Veronica xalapensis Veronica anagallis-aquatica Veronica Synthyris laciniata S. pinnatiﬁ Penstemon laevis Penstemon linarioides sileri Penstemon palmeri eglandulosus Penstemon thompsoniae Penstemon whippleanus Penstemon barbatus barbatus Pedicularis groenlandica Linaria vulgaris scabrida Castilleja scabrida Sparganiaceae Solanaceae Solanaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae FamilyScrophulariaceae Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconﬁ

Chapter 6: Appendices 95 (Albee et al. rmed for the park. Spe- Range Comments Atlas of the Vascular Flora of Utah Atlas of the Vascular Life form in the park, but are not included here. in the park, but are oung University in 2001, the puncture vine AnnF Intro Native to Eurasia alpine marsh violet PerF Wide Canadian white violet PerF Wide New Mexico vervain PerF Periph stinging nettle PerF Wide blunt sweet-cicely PerF Wide matted spring-parsley PerF Wide southern cattailNevada lomatium PerG PerF Wide Wide eld and Kane counties, Utah, but have not previously been reported or conﬁ been reported eld and Kane counties, Utah, but have not previously palustris & rugulosa, V. rugulosa, V. brevipes var. holosericea, U. holosericea ssp. ssp. Includes vars. scopulorum, V. rydbergii scopulorum, V. gracilis

var. occidentalis U. dioica var. rmed) taxa of Bryce Canyon National Park, cont. Tribulus terrestris Tribulus Viola palustris Viola canadensis canadensis V. Verbena macdougalii Verbena Urtica dioica Osmorhiza depauperata Lomatium nevadense Oreoxis alpina Cymopterus alpinus Typha domingensis Typha parishii Zygophyllaceae Violaceae Violaceae Verbenaceae Urticaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) FamilyTyphaceae Species name notes Synonyms/Taxonomic Common name Appendix B. Potential (unconﬁ 1988), Madsen (2001), and Spence and Buchanan (1993). Some species from Appendix C could be considered as potentially occurring Appendix C could be considered 1988), Madsen (2001), and Spence Buchanan (1993). Some species from The vascular plant species listed here are known from the vicinity of Bryce Canyon National Park in Garﬁ known from are The vascular plant species listed here N. Duane Atwood of Brigham Y by Dr. a checklist of potential species for NCPN parks prepared derived from cies on this list are

96 Annotated Checklist of Vascular Flora: Bryce Canyon National Park . E. C. USFWS:

. ed specimen of wasatchensis ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of . undulatum. . var. undulatum. erinacea glabrescens. ed specimen of cinerea. var. var. ed specimens of condensatus. var. . var. var. var. var. var. var. . var. var. based on misidentifi C. undulatum 3C on a misidentifi pumilus based on misidentifi Mahonia fremontii based on a misidentifi C. cinerea on misidentifi based on misidentifi C. fulvocanescens undulatum Comments Spence and Buchanan 1993. based on misidentifi A. wasatchensis based on misidentifi B. sagittata based on misidentifi O. erinacea based on misidentifi C. parryi based on misidentifi A. fendleri Park status Range Life form eabane PerF Wide FalsRep Report from BRCA herbarium based Rydberg’s thistleRydberg’s PerFtufted fl RegEn FalsRep Reports from BRCA herbarium are Fendler’s barberryFendler’s Shrub RegEn FalsRep Reports from BRCA herbarium are James’ cryptanth PerF Wide FalsRep Reports from BRCA herbarium are gray-green thistle PerF Periph FalsRep Reports from BRCA are based Virgin River cryptanth PerF RegEn FalsRep Reports from BRCA herbarium are Common name thick-stem wood-aster PerF Widecutleaf balsamroot FalsRep Reports from BRCA herbarium are PerF RegEn FalsRep Reports from BRCA herbarium are Nuttall’s sandwortNuttall’s PerF Wide Ques? Cited as outside range of BRCA in harebell PerF Periph FalsRep Reports from BRCA herbarium are Utah pricklypear PerF Wide FalsRep sandwortFendler’s Reports from BRCA herbarium are PerF Periph FalsRep Reports from BRCA herbarium are

var.

var. var. C. jamesii, Oreocarya suffruticosa canovirens Synonyms/ notes Taxonomic nuttallii, Minuopsis nuttallii O. polyacantha hystricina Eremogone fendleri

var. var. var. Cryptantha cinerea jamesii Cirsium rydbergii Erigeron caespitosus Berberis fendleri Cirsium canovirens C. cymosum Cryptantha virginensis Opuntia erinacea utahensis Arenaria fendleri fendleri Arenaria nuttalliiAster integrifolius Minuartia nuttallii Eurybia integrifolia Balsamorhiza macrophylla Campanula rotundifolia Boraginaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyBerberidaceae Species name Boraginaceae Cactaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Compositae (Asteraceae) Compositae (Asteraceae) Campanulaceae Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park. Appendix C. Rejected (falsely reported

Chapter 6: Appendices 97

ed var. var. as cited in . D. incana. A. pendulina . Welsh et al. . Welsh . ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of P. rubicundula P. . A. demissa A. demissa E. compositus and . . . ed specimen of is native to SW US but not , not Utah Spence and Buchanan 1993. based on misidentifi E. vagus based on misidentifi S. hartianus based on misidentifi S. velutina Spence and Buchanan (1993). based on misidentifi Solidago velutina based on misidentifi Sphaeromeria capitata bosch (1981) based on misidentifi specimens of (2003) do not recognize as distinct from based on misidentifi Streptanthus cordatus. Graybosch (1981) probably based on misidentifi D. obtusa known from Utah. based on misidentifi intermedia P. rubicundula Comments Park status Range Life form Holboell’s rockcressHolboell’s PerF Wide Ques? Native to Greenland, not known from arrowhead thelypody PerF Wide Ques? Cited as outside range of BRCA in alpine daisy PerF Wide FalsRep Reports from BRCA herbarium are Buek’s groundselBuek’s PerFMissouri goldenrod Wide PerF FalsRep Reports from BRCA herbarium are Wide FalsRep Reports from BRCA herbarium are western goldenrod PerF Wide FalsRep Reports from BRCA herbarium are silver chicken-sage PerF RegEn FalsRep Reports from BRCA herbarium are rabbit-ear rockcress PerF RegEn FalsRep Report from Buchanan and Gray- Cooper’s wild cabbageCooper’s AnnF Periph FalsRepblunt tansy mustard Reports from BRCA herbarium are PerF Wide FalsRep Report from Buchanan and Common name Newberry’s twinpodNewberry’s PerF RegEn FalsRep Reports from BRCA herbarium are

orus

var.

includes var. var. E. grandiﬂ extraria sagittatum buekii missouriensis, var. oxylobula Thelypodium sagittatum Synonyms/ notes Taxonomic Includes racemosa

var. sagittata Senecio cymbalarioides Packera subnuda, P. Solidago missouriensis S. missouriensis Arabis holboellii Arabis pendulina Boechera pendulina, A. holboellii Thelypodiopsis sagittata var. Erigeron simplex Euthamia occidentalis Solidago occidentalis nuttalliiTanacetum Sphaermeria argentea Caulanthus cooperi Descurainia obtusa Physaria newberryi newberryi P. Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park, cont.. FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

98 Annotated Checklist of Vascular Flora: Bryce Canyon National Park

P. glauca P. . . Native to Eurasia ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of . longispicatus ed specimen of var...... glauca. Comments based on misidentifi C. vallicola based on misidentifi S. pungens based on misidentifi A. uva-ursi based on misidentifi G. heterosepala based on misidentifi G. tortuosa based on misidentifi Phragmites australis based on misidentifi E. salinus based on misidentifi M. montana. on misidentifi var. based on misidentifi S. idahoense. based on misidentifi J. arcticus Park status Range Life form Common name hoary sedgeAmerican bulrush PerG Wide PerG FalsRep Wide Reports from BRCA herbarium are FalsRep Reports from BRCA herbarium are Mexican manzanita Shrub Wide FalsRep Reports from BRCA herbarium are northern gentian PerF Wide FalsRep Reports from BRCA herbarium are Lapland gentian AnnF Wide FalsRep Reports from BRCA herbarium are giant reed PerGNavajo muhly Intro FalsRep Reports from BRCA herbarium are nodding bluegrass PerG RegEn PerG FalsRep Wide Reports from BRCA herbarium are FalsRep Reports from BRCA herbarium based alkali wildrye PerG RegEn FalsRep Reports from BRCA herbarium are blue-eyed grass PerF Wide FalsRep Reports from BRCA herbarium are thread rush PerG Periph FalsRep Reports from BRCA herbarium are

simplex, var. amarella, var. var. acuta, Gentianella subliolacea, C. Synonyms/ notes Taxonomic canescens, C. canescens var. lapponica oplectus americanus Gentianella amarella ssp. amarella Gentianella acuta Comastoma tenellum Leymus simplex liformis exa Carex canescens C. canescens Scirpus americanus Scirpus olneyi, Schoen- Arctostaphylos pungens Gentianella amarella Gentiana amarella, Gentianella tenella Gentiana tenella, Elymus simplex E. simplex Arundo donax Muhlenbergia arsenei Poa reﬂ Sisyrinchium demissum Juncus ﬁ Cyperaceae Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park, cont.. FamilyCyperaceae Species name Ericaceae Gentianaceae Gentianaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Iridaceae Juncaceae

Chapter 6: Appendices 99 . Native to . Native to ed specimens of ed specimens of ed specimens of ed specimens ed ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of . parryi. convallarius. panguicensis. . . Native to Arizona var. var. var. var. var. var. cinalis oblongifolius . . var. Pediomelum pariense based on misidentifi Medicago lupulina California, Nevada, and Arizona, not known from Utah. based on misidentifi Melilotus offi based on misidentifi C. nuttallii based on misidentifi of based on misidentifi L. utahensis based on misidentifi A. megacarpus based on misidentifi A. miser Wyoming and Montana, not known from Utah. based on misidentifi A. argophyllus based on misidentifi H. drummondii and New Mexico, not known from Utah. based on misidentifi A. convallarius Comments Park status Range Life form liform clover AnnF Intro FalsRep Reports from BRCA herbarium are suckling cloverfi AnnF Introsinuous mariposa FalsRep Reports from BRCA herbarium are PerF Wide FalsRep Reports from BRCA herbarium are Beaver Dam breadrootDam Beaver PerF RegEn FalsRep Reports from BRCA herbarium are Wright’s trefoilWright’s PerF RegEn FalsRep Reports from BRCA herbarium are egg milkvetch PerF Wide FalsRep Reports from BRCA herbarium are timber milkvetch PerF RegEnBeaver Dam milkvetch FalsRep PerF Reports from BRCA herbarium are Wide FalsRep Reports from BRCA herbarium are Flagstaff pennyroyalFlagstaff PerF LocEn FalsRep Reports from BRCA herbarium are painted milkvetch PerF Wide FalsRep Reports from BRCA herbarium are Common name Synonyms/ notes Taxonomic

var. exuosus liforme castoreus ceramicus Trifolium dubium Trifolium ﬁ Trifolium Calochortus ﬂ Psoralea castorea Psoralidium castoreum Lotus wrightii Astragalus oophorus Astragalus miser decumbens Astragalus newberryi var. Hedeoma diffusum Astragalus ceramicus var. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Calochortaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park, cont.. FamilyLabiatae (Lamiaceae) Species name

100 Annotated Checklist of Vascular Flora: Bryce Canyon National Park

. P. . Not E. . Endemic and maculata var. . ed specimens of specimens ed ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of . USFWS: former C2. ed specimen of . . canescens ora var. var. Leptodactylon caespitosum based on misidentifi Leptodactylon caespitosum to Colorado and New Mexico. based on misidentifi of recognized as occurring in Utah et al. 2003. Welsh based on misidentifi Leptodactylon caespitosum hoodii based on misidentifi Leptodactylon caespitosum Comments based on misidentifi L. australe based on misidentifi M. multifl based on misidentifi Ipomopsis aggregata on misidentifi halleanum based on misidentifi G. ramosissimum based on misidentifi C. maculata based on misidentifi longaeva P. Park status Range Life form ax AnnF Wide FalsRep Reports from BRCA herbarium are broom-fl dwarf phloxspreading phlox PerF RegEn FalsRep PerF Reports from BRCA herbarium are Wide FalsRep Reports from BRCA herbarium are moss phlox PerF Wide FalsRep Reports from BRCA herbarium are cushion phlox PerF Wide FalsRep Reports from BRCA herbarium Common name Arapien stickleaf PerF LocEn FalsRep Reports from BRCA herbarium are Arizona scarlet gilia PerF Wide FalsRep Reports from BRCA herbarium are northern willow-herb PerF Wide FalsRep Reports from BRCA herbarium based low groundsmoke AnnF coralrootWister’s Periph FalsRep Reports from BRCA herbarium are PerFbristlecone pine Sparse FalsRep Reports from BRCA herbarium are Tree RegEn FalsRep Reports from BRCA herbarium are

var. ssp. ssp. muscoides, P. bryoides muscoides, P. pulvinata, P. caespitosa pulvinata, P. Synonyms/ notes Taxonomic ciliatum, E. watsonii, adenocaulon Phlox pulvinata sibirica P. Linum aristatum Mesynium aristatum Phlox condensata Phlox diffusa Phlox muscoides hoodii P. Mentzelia argillosa Nuttallia argillosa Epilobium ciliatum E. ciliatum Gilia arizonica Ipomopsis arizonica Gayophytum humile Corallorhiza wisteriana Pinus aristata Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park, cont.. FamilyLinaceae Species name Loasaceae Onagraceae Polemoniaceae Onagraceae Orchidaceae Pinaceae

Chapter 6: Appendices 101 F. S. P. S. . . ed specimens of ed specimens of ed specimens of ed specimens ed ed specimens of ed specimens of ed specimens of and may be from fremontii ed specimens of ed specimens of ed specimens of var. . . . R. glaberrimus Spence and Buchanan 1993. Comments based on misidentifi fremontii P. on misidentifi bebbiana. Plateau Network (in ed.), but BRCA is well outside the known or expected range of this species. on misidentifi bebbiana based on misidentifi Leptodactylon caespitosum based on misidentifi aviculare P. based on misidentifi of based on misidentifi virginiana based on misidentifi crinita P. on misidentifi hippiana outside the park based on misidentifi anserina P. Park status Range Life form Tree Wide FalsRep Reports from BRCA herbarium are ower cinquefoil PerF Wide FalsRep Reports from BRCA herbarium are sky pilot PerF Wide Ques? Cited as outside range of BRCA in Common name Rio Grande cottonwood Drummond’s willowDrummond’s Shrub Wide FalsRep Reports from BRCA herbarium based gray-leaf willow Shrub Wide Ques? Reported by Northern Colorado whiplash willow Shrub Wide FalsRep Reports from BRCA herbarium based mound phlox PerFbushy knotweed RegEn FalsRep Reports from BRCA herbarium are AnnF Wide FalsRep Reports from BRCA herbarium are woodland strawberry PerF Wide FalsRep Reports from BRCA herbarium are big-fl creeping buttercup PerF Wide FalsRep Reports from BRCA herbarium are beautiful cinquefoil PerF Wide FalsRep Reports from BRCA herbarium based near cinquefoil PerF RegEn FalsRep Reports from BRCA herbarium are

var.

ammula ssa var. var. var. ssp. cum, P. P. cum, prolifi reptans, R. reptans ammula liformis, R. fl Synonyms/ notes Taxonomic wislizeni caudata tumulosa ramosissimum P. var. ramosissimum ramosissimum R. fl var. pulcherrima P. fi P. proxima P. P. deltoides P.

var. var.

var. caudata S. lasiandra var. ammula ssp. ssafi Drymocallis ovalis Phlox tumulosaPolemonium viscosum griseola P. Polygonum ramosissimum Salix drummondiana Salix glauca Salix lucida pulcherrima Ranunculus fl var. bracteata Potentilla concinna proxima Potentilla fi Potentilla gracilis Populus fremontii wislizeni Fragaria vesca Polemoniaceae Polygonaceae Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park, cont.. FamilyPolemoniaceae Species name Salicaceae Salicaceae Salicaceae Ranunculaceae Rosaceae Rosaceae Rosaceae Salicaceae Rosaceae

102 Annotated Checklist of Vascular Flora: Bryce Canyon National Park

. ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of montanensis. e or Ques?). aberrans var. . . USFWS: 3C. var. var. . . Native to Wyoming and the based on misidentifi Angelica pinnata based on misidentifi C. minimus Garden, but BRCA is well outside the expected range of this taxon. based on misidentifi procerus P. based on misidentifi C. minimus based on misidentifi palustris P. Comments based on misidentifi L. porteri based on misidentifi L. parryi. based on misidentifi L. parryi western Great Plains (not known from Utah). Park status Range ure (Spence and Buchanan 1993) or from herbarium specimens, but are now herbarium specimens, but are (Spence and Buchanan 1993) or from ure PerF Wide FalsRep Reports from BRCA herbarium are Life form PerF Wide FalsRep Reports from BRCA herbarium are owered grass- owered basin white-cup spring-parsley water-hemlock PerF Wide FalsRep Reports from BRCA herbarium shortleaf penstemon PerF RegEn penstemonRydberg’s Ques? PerF Specimen cited at NY Botanical Periph FalsRep Reports from BRCA herbarium are small-fl of-Parnassus spring-parsleyRose’s PerF RegEn FalsRep Reports from BRCA herbarium are Common name fernleaf ligusticum PerF Wide FalsRep Reports from BRCA herbarium are King’s lomatiumKing’s PerF RegEn FalsRep Reports from BRCA herbarium are Nuttall’s lomatiumNuttall’s PerF RegEn FalsRep Reports from BRCA herbarium are

var. ora C. douglasii, occidentalis C. rosei Synonyms/ notes Taxonomic parviﬂ L. kingii nuttallii

var. var. cations (False Reports or FalsRep) because the park is well outside known expected range of species (Questionabl orapalustris P. licinum rosei alpinum Cicuta maculata angustifolia Cymopterus purpurascens Cymopterus purpureus var. brevifolius Penstemon humilis Penstemon rydbergii Parnassia parvifl Ligusticum fi Lomatium graveolens var. Lomatium nuttallii L. megarrhizum, Aletes Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Umbelliferae (Apiaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Bryce Canyon National Park, cont.. FamilySaxifragaceae (Parnassiaceae) Species name Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) The vascular plant species listed here have been reported for Bryce Canyon National Park from published and unpublished literat for Bryce Canyon National Park from have been reported The vascular plant species listed here because of misidentifi believed to be erroneous

Chapter 6: Appendices 103 x Wet Comments x x Conf Mont Jun Pin- xxx xxx Grs Sage/ Flower time Park status r Native Pres NA r Native Pres NA x common applenarrow-leaved cottonwood NativeFremont cottonwood Presquaking aspen Intro Native Apr–Junlance-leaf cottonwood PresSiberian elm Pres Native Mar–Jun MayRocky Mountain maple Native Presnarrowleaved yucca Pres Native Apr–Jundatil yucca Intro yuccaHarriman’s Pres Mar–Jun Native Pres x squawbush Apr–Jun Pres mahoniaFremont’s Mar–Apr Native May–Jul Native Pres x x Native Pres x Apr–May x x Native Pres x Cultivated Apr–Jun x x Pres Apr–Jul x x May–Jul x x x x x x bigtooth maplewater birchRocky Mountain juniperRussian olive Native NativeGambel oak Preswhite ﬁ Presblue spruce Native NAtwo-needle pinyon Apr–May Pres Introlmber pine Nativewestern bristlecone pine Apr–May Pres Nativeponderosa pine Pres Native Native Pres May–Jul PresDouglas-ﬁ Apr–Jun Pres x NA Native NA x x Native NA Pres Pres x x x NA x NA x x x x x x x Claron barrens Oregon-grapeblue elderberry Native Native Pres Pres Apr–Jun May–Jul x x x x

var.

gambelii

trilobata var. var. var. glabrum var. var. var. intercurrens x exilis Populus angustifolia Populus fremontii fremontii Populus Ulmus pumila Acer glabrum angustissima Yucca baccata Yucca harrimaniae Yucca harimaniae Mahonia fremontii Pseudotsuga menziesii Malus pumila Populus tremuloides Rhus aromatica Acer grandidentatum Betula occidentalis Juniperus scopulorum Elaeagnus angustifolia Quercus gambelii Abies concolor Picea pungens Pinus edulis Pinus ﬂ Pinus longaeva Pinus ponderosa scopulorum glauca Mahonia repens Sambucus caerulea taxa of Bryce Canyon National Park, organized by life form with ecological notes. rmed, historical, and reported Salicaceae Salicaceae Salicaceae Ulmaceae Shrubs Aceraceae Agavaceae Agavaceae Agavaceae Berberidaceae Berberidaceae Pinaceae Rosaceae Salicaceae Anacardiaceae FamilyTrees Aceraceae Betulaceae Cupressaceae Species nameElaeagnaceae Fagaceae Pinaceae Pinaceae Pinaceae Common namePinaceae Pinaceae Pinaceae Nativity Caprifoliaceae (Adoxaceae) Appendix D. Conﬁ

104 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Clay soils Clay soils Disturbed areas Claron Formation x x Wet Comments Conf Mont x Jun Pin- xx xx xx xx xx xx x xx x xxx xxx xxx xx Grs Sage/ Flower time Park status mountain-loverfourwing saltbushshadscalebasin saltbush Native Native Preswinterfat Pres May–Aug May–Sep greasewood Native x Native Pres Pres x Jun–Sep Native May–Sep x Native Pres x Pres May–Jul x May–Aug Clay soils mountain snowberry Native Pres Jun–Aug siltbushPlummer’s Native Pres Jun–Aug fringed sagebrush Native Pres Jul–Sep black sagebrush Native Pres Sep–Oct pygmy sagebrush Native Pres Aug–Sep dwarf rabbitbrush rabbitbrushspear-leaf Native Native Pres Pres Jul–Sep Jul–Sep basin big sagebrushWyoming big sagebrush Native NativeMohave brickellbush Pres Rep Jul–Sep Native Jul–Sep Pres Jun–Aug greenish rabbitbrush Native Pres Aug–Oct graystem rabbitbrush Native Pres Aug–Oct

var. var.

var.

var. var. nova var. var. var. var. utahensis Atriplex canescens canescens Atriplex gardneri utahensis Pachystima myrsinites Atriplex confertifolia Krascheninnikovia lanata lanata Sarcobatus vermiculatus Zuckia brandegei Symphoricarpos oreophilus var. plummeri Artemisia frigida Artemisia nova Artemisia pygmaea Artemisia tridentata Artemisia tridentata Brickellia oblongifolia Chrysothamnus depressus Chrysothamnus linifolius tridentata wyomingensis linifolia Chrysothamnus nauseosus consimilis Chrysothamnus nauseosus gnaphalodes rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Celastraceae Chenopodiaceae Chenopodiaceae Chenopodiaceae (Sarcobataceae) Chenopodiaceae Appendix D. Conﬁ FamilyShrubs, cont. Caprifoliaceae Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 105 Sandstone Claron barrens Wet Comments x x x x Conf Mont Jun Pin- xxx xxx xxx xxx xxx x xxx xx xx Grs Sage/ Flower time Park status spindly goldenbushCedar Breaks goldenbush Native Native Pres Pres Aug–Sep Jul–Aug Vasey’s rabbitbrushVasey’s Native Pres Jul–Sep broom snakeweednarrowleaf goldenweed Native Native Pres Pres Jul–Oct Jul–Sep spineless horsebrush Native PresUtah juniper Jun–Sep Native Pres NA x x x glabrate rabbitbrush rabbitbrushParry’s Native rabbitbrushParry’s Hist Native Aug–Oct Pres Nativelanceleaf rabbitbrush Jul–Sep Presslenderleaf rabbitbrush Native Jul–Sep Nativegreen rabbitbrush Pres Rep Jul–Sep Jul–Sep Native Pres Jul–Sep red-osier dogwoodcommon juniper Nativesilver buffaloberry Pres NativeCanada buffaloberryroundleaf buffaloberry May–Jul Presgreen ephedra Native Native Nativegreenleaf manzanita NA Pres Presbearberry Pres Apr–May Native May–Jul Mar–May Native Pres x Pres Mar–Jul x x NA Native x x Pres x x x May–Aug x x x x

var. var.

orus orus orus var. var. var. viridis sericea var. var. orus lanceolatus stenophyllus viscidifl nis Haplopappus scopulorum hirtellus Haplopappus zionis Chrysothamnus parryi Chrysothamnus parryi Chrysothamnus vaseyi Chrysothamnus viscidifl Chrysothamnus viscidifl Chrysothamnus viscidifl Gutierrezia sarothrae Haplopappus macronema canescens Tetradymia Cornus sericea Juniperus osteosperma Chrysothamnus nauseosus graveolens affi parryi var. var. var. Juniperus communis depressa Shepherdia argentea Shepherdia canadensis Shepherdia rotundifolia Ephedra viridis Arctostaphylos patula Arctostaphylos uva-ursi rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cornaceae Cupressaceae Appendix D. Confi FamilyShrubs, cont. Compositae (Asteraceae) Species name Common name Nativity Cupressaceae Elaeagnaceae Elaeagnaceae Elaeagnaceae Ephedraceae Ericaceae Ericaceae

106 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Rocky areas Wet Comments Conf Mont xx xx Jun Pin- xx xxx xxx xxx xxx xxx Grs Sage/ Flower time Park status Native Pres May–Jun Native Pres May–Jun slender wild buckwheat Native mountain-lilacFendler’s Utah mountain-lilac Pres NativeUtah serviceberry Jun–Oct Pres Native Jun–Aug Pres Native Pres Jun–Jul x Apr–Jun x x x x x x prickly-phloxcrisp-leaf wild buckwheat Native Pres Native Jul–Oct Prescurl-leaf mountain May–Julmahogany x x x alder-leaf mountain alder-leaf mahogany Apache plumebush oceanspraysquaw-appleshrubby cinquefoilwestern chokecherry Native Native Pres Pres Native Native Native Apr–Aug Pres Jun–Aug Pres Pres x Jun–Aug x May–Jul May–Jun x x x x x x x x x Rocky areas x cliff-rose Native Pres May–Jun antelope bitterbrushNootka rose roseWoods’ Nativecommon red raspberry Pres willowBebb’s Nativeshort-fruit willow Apr–Jul Nativeyellow willow Pres Native Hist x Jun–Aug Pres Native Native Jun–Jul x Pres Pres May–Jul Native x Jun–Jul May–Jun Pres Apr–Jun x x x x x x x x coyote willow willowGeyer’s golden currant Native Native Pres Native Pres Hist Apr–Aug May–Jun Mar–Jul x x wax currant Native Pres Apr–Aug

var. var. var.

watsonii var. var. hispida var. inebrians strigosus stenophylla var. var. var. var. orum Ceanothus martinii Amelanchier utahensis Cercocarpus ledifolius Eriogonum microthecum laxiﬂ Ceanothus fendleri Leptodactylon pungens Eriogonum corymbosum corymbosum intermontanus Holodiscus dumosus Peraphyllum ramosissimum Potentilla fruticosa Prunus virginiana melanocarpa Cercocarpus montanus Fallugia paradoxa Purshia mexicana stansburyana Rosa nutkana Rosa woodsii Rubus idaeus Salix bebbiana Salix brachycarpa Salix eriocephala Salix exigua Purshia tridentata Salix geyeriana Ribes aureum Ribes cereum rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Rhamnaceae Rosaceae Rosaceae Polygonaceae Rhamnaceae Appendix D. Conﬁ FamilyShrubs, cont. Polemoniaceae Polygonaceae Species name Common name Nativity Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Salicaceae Salicaceae Salicaceae Salicaceae Rosaceae Salicaceae Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae)

Chapter 6: Appendices 107 Sandy areas Wet Comments Conf Mont xxx Jun Pin- xxx xxx xx xxx xx xx Grs Sage/ Flower time Park status ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep x whitestem gooseberry Native Pres Apr–Jul ﬁ spreading dogbanecommon dogbanespider milkweed Native Pres milkweedHall’s Nativeshowy milkweed Jun–Aug Pres Nativelow cryptanth cryptanthBaker’s Pres Jun–Aug Nativesand cryptanth Native May–Jul Pres Pres Native Native Jun–Aug May–Aug Pres Pres Native x Rep May–Jun May–Jun x x x x Jun Claron barrens x x x x x x Disturbed areas ashy cryptanth Native Pres May–Aug basin yellow cryptanthplateau yellow cryptanth Nativeplateau cryptanth Nativedwarf cryptanth Pres Presyellowish cryptanth Apr–Junbristly cryptanth Native Apr–Juncommon hound’s-tongue Hist x Native Nativeshowy stoneseed x Introsouthwestern stoneseed Pres Pres May–Jun x Native x Presaspen bluebell Native Apr–Jun May–JunArizona pincushion x Pres x Native May–Jul Pres x x May–Aug Pres x Jun–Aug Native Native x x Apr–Jul x x Pres Pres x x x x x Apr–Aug Jun–Aug x x x Claron barrens x Disturbed areas x x x claretcup Native Pres Apr–Jun common pricklypearbrittle pricklypear Native Hist Native Pres May–Jun x Jun–Jul x x

var. erinacea orum ora var. var. var. var. cinale ava androsaemifolium melanacanthus Ribes inerme Apocynum andros-aemifolium var. Asclepias asperula asperula Asclepias speciosa Cryptantha abata Cryptantha bakeri Cryptantha cinerea arenicola Tamarix chinensis Tamarix Apocynum cannabinum Asclepias hallii Cryptantha cinerea cinerea Cryptantha fl Cryptantha fulvocanescens Cryptantha humilis Cryptantha ochroleuca Cryptantha setosissima Cynoglossum offi Lithospermum incisum Lithospermum multifl Mertensia arizonica Coryphantha vivipara arizonica Cryptantha confertifl Echinocereus triglochidiatus var. Opuntia erinacea Opuntia fragilis rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyShrubs, cont. Saxifragaceae (Grossulariaceae) Species name Common name Nativity Perennial Forbs Perennial Apocynaceae Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Tamaricaceae Apocynaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae Boraginaceae Cactaceae Cactaceae Cactaceae

108 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Wet Comments xx x Conf Mont Jun Pin- xxx xxx xxx xxx xxxx xxx xxx xx xx xx xxx xxx xx Grs Sage/ Flower time Park status Native Pres Apr–May y Native Pres May–Sep x x x shhook ower fi ower purplish agoseris Native Pres Jun–Aug pale agoseris Native Pres May–Sep low pussytoes Native Rep Apr–Jun cutleaf agoseris Native Pres May–Sep orange agoseris Native Rep Jun–Aug common yarrow Native Pres May–Sep long-stalk starwort Native Pres May–Aug Parry’s harebellParry’s basin sandwort catchfl Drummond’s creeping nailwort Native Native Pres campionPeterson’s PresJames’ chickweed Native Jun–Sep May–Aug Native Pres Pres Native May–Aug Pres Jun–Sep x Jun–Aug x Claron barrens cactus Simpson’s hedgehog cactusSimpson’s Nativesmall-fl Pres May–Jul x x x common pussytoesBreaks pussytoes Native Pres Native May–Jul Pres Jun–Aug plains pricklypear Native Rep May–Jun small-leaf pussyoes Native Pres Jun–Aug

var. var.

longipes ssp. var. var. glauca laciniata var. ora var. var. var. purpurea Agoseris glauca Agoseris glauca Antennaria dimorpha aurantiaca Agoseris aurantiaca lanulosa Agoseris aurantiaca Achillea millefolium Paronychia sessiliﬂ Arenaria fendleri glabrescens Stellaria jamesiana Campanula parryi drummondii Lychnis Silene petersonii Stellaria longipes Sclerocactus whipplei roseus Pediocactus simpsonii Antennaria parvifolia Antennaria rosulata Opuntia polyacantha polyacantha Antennaria microphylla rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Campanulaceae Caryophyllaceae Caryophyllaceae Caryophyllaceae (Alsinaceae) Cactaceae Cactaceae Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Cactaceae Species name Common name Nativity Compositae (Asteraceae)

Chapter 6: Appendices 109 Disturbed areas Disturbed areas Kaiparowits Formation Claron barrens x x x Wet Comments xx x x x Conf Mont xx Jun Pin- xx xx xx xxx xx x xxx xxxx xx xxx Grs Sage/ Flower time Park status c aster c Native Pres Jul–Oct spotted knapweedRussian knapweed Intro Pres Intro Jul–Sep Pres Jun–Sep Disturbed areas burdock Intro Pres Jul–Sep musk thistle Intro Pres Jun–Sep Chamisso’s arnicaChamisso’s heartleaf arnica Native Pres Native Jun–Aug Pres May–Aug sand wormwoodtarragon Native Pres Jul–Sep Native Pres Jul–Oct Carruth’s wormwoodCarruth’s Native Pres Aug–Sep Paciﬁ asterEaton’s blueleaf asterSiskiyou aster Nativewestern aster Pres NativeMarkagunt aster Pres Jul–Sep Nativearrowleaf balsamroot Jul–Sep Pres Native Native Native Jul–Sep Pres Pres Pres Jul–Sep Jul–Sep Apr–Jul Louisiana wormwoodLouisiana wormwood Native Pres Native Jul–Oct Pres Jul–Oct

foliosa var. var. var. var. cordifolia var. var. var. Centaurea repens Centaurea maculosa Arctium minus Carduus nutans Arnica chamissonis Arnica cordifolia Artemisia campestris Artemisia dracunculus scouleriana glauca wasatchensis Artemisia carruthii Artemisia ludoviciana Artemisia ludoviciana Aster ascendens Aster eatonii Aster glaucodes Aster hesperius Aster spathulatus Aster wasatchensis Balsamorhiza sagittata albula latiloba rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae

110 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Disturbed areas Kaiparowits Formation Disturbed areas x x x x Wet Comments x Conf Mont xx xx Jun Pin- xx xx xx xx xxx xxx xxx xxx xxx xxx Grs Sage/ Flower time Park status New Mexico thistleelk thistle Native Preswavy-leaved thistle May–Jun bull thistle Native Native Pres Pres Jun–Aug Jun–Sep Intro Pres Jul–Aug Tracy’s thistleTracy’s Native thistleWheeler’s Pres Jun–Sep Native Pres Jul–Sep gray hawksbeard Native Pres May–Jul rose-heathJones’ golden-asterhispid golden-aster Nativechicory Native Pres Native PresArizona thistle May–Sep Pres May–Sep Canada thistle May–Sep pretty thistle Native Intro Pres Intro Pres Jun–Aug Pres Jul–Oct Native Jul–Sep Pres Jun–Sep Disturbed areas hoary dusty-maiden Native Pres May–Sep western hawksbeard Nativehispidulous hawksbeard Pres Native May–Jul Pres May–Aug glaucous hawksbeard Native Rep May–Aug

var.

tracyi

minor wheeleri

glauca var. var. var. var. var. var. var. var. var. neomexicanum scariosum Cirsium scariosum Cirsium undulatum Cirsium undulatum undulatum Cirsium vulgare Cirsium wheeleri Crepis intermedia Crepis occidentalis Cirsium arizonicum Chaetopappa ericoides Chrysopsis jonesii Chrysopsis villosa Cichorium intybus Cirsium arvense Cirsium calcareum pulchellum Cirsium neomexicanum Chaenactis douglasii occidentalis Crepis runcinata Crepis runcinata hispidulosa rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 111 Claron barrens Claron barrens x Wet Comments x x Conf Mont xx Jun Pin- xxx xxx xxx xx xx xx xxx xx xxxx xxx xxx xxx xxx xx Grs Sage/ Flower time Park status ower Native Pres Jun–Aug eabane Native Pres May–Aug trailing daisylongleaf daisy Native Pres Native Pres May–Aug Jun–Sep Navajo ﬂ vernal daisy Native Pres May–Aug Payson’s daisyPayson’s curly-cup gumweed Native Native Pres Hist Jun–Aug Jul–Sep aspen daisybear daisy Native Pres Native Jun–Aug Pres Jun–Aug Abajo daisy Native Pres Jul–Sep hairy daisy Native Hist May–Jul hoary daisyspreading daisy Native daisyEaton’s Pres Native Jun–Jul Pres Native May–Sep Pres May–Jul cut-leaved daisy Native Rep May–Aug stemless goldenweed Native Rep May–Jul thrift goldenweedlittle sunﬂ Native Pres May–Jun common hyalineherb Native Pres May–Oct

var. var.

var. var. var. ursinus var. var. var. var. var. lifolius var. ora agellaris Erigeron lonchophyllus Erigeron pumilus concinnus Erigeron pumilus condensatus serrulata Erigeron speciosus Erigeron ursinus Erigeron vagus Grindelia squarrosa Haplopappus acaulis macranthus Erigeron abajoensis Erigeron aphanactis aphanactis divergens Erigeron canus Erigeron compositus Erigeron divergens Erigeron eatonii Erigeron fl glabratus acaulis Haplopappus armerioides Helianthella unifl armerioides Hymenopappus fi cinereus rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

112 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Claron barrens x x Wet Comments x x x Conf Mont xxx Jun Pin- xxx xx xxx xx xxx xx xxx xx xxx xxx xxx xxx Grs Sage/ Flower time Park status ower Native Pres May–Sep showy rush-pink Native Pres May–Jun greenstem paperﬂ black groundsel Native Pres Jul–Aug aristate asterhoary astergumweed aster Nativerock-goldenrod Rep Native Native Jul–Oct Pres Pres Native May–Sep May–Jul Pres Jul–Sep Arizona hymenoxys Nativepoverty-weed Presblue lettuce Apr–Jun Native Pres Native May–Sep Pres Jun–Sep Great Basin hyalineherb Nativealpine hyalineherb Pres May–Oct NativeColorado rubber-plant Pres Native May–Oct Pres May–Aug water groundselbasin groundselbroom groundsel Native Pres Native Jul–Aug Native Pres Pres Apr–Aug Jun–Oct Hart’s groundselHart’s Native Pres Jun–Jul groundselBarkley’s Native Pres Jun–Aug

var. var. var. var.

var. pumila pulchella var. ora var. ora var. lifolius lifolius var. ora aristata canescens grindelioides oribunda Senecio hartianus Machaeranthera canescens var. Machaeranthera canescens var. Machaeranthera grindelioides var. Petradoria pumila Psilostrophe sparsifl Senecio atratus grandifl Hymenopappus fi Hymenoxys acaulis arizonica Hymenoxys richardsonii Iva axillaris Lactuca tatarica grandifl Lygodesmia Hymenopappus fi nanus nudipes fl Senecio hydrophilus Senecio multilobatus Senecio spartioides spartioides Senecio werneriifolius barkleyi rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 113 Claron barrens Disturbed areas Claron barrens x Wet Comments x x x x Conf Mont xx xx xx xx Jun Pin- xxxx xx xx xxx xx xxx xxx xxx Grs Sage/ Flower time Park status eld bindweed Intro Pres Jun–Aug Disturbed areas common dandelionstemless townsendia Introhoary townsendia Native Pres Pres Apr–Dec Native May–Jun Pres Apr–Jul Bryce Canyon townsendia Native Pres Apr–Jun yellow salsifyshowy goldeneyeﬁ nodding rockcress Intro Native rockcressDrummond’s Pres Pres Native rockcressFendler’s May–Jul Jun–Sep Native Pres Rep May–Aug Native May–Sep Hist Apr–Aug three-nerve goldenrodmarsh sow-thistle Native Pres chicken-sagecluster-head Native Aug–Oct Introslender wirelettuce Pres Pres May–Jun Native Jul–Oct Pres Jun–Sep coast goldenrod Native Pres Jul–Sep Parry’s goldenweedParry’s Native Pres Jul–Sep Holboell’s rockcressHolboell’s rockcressHolboell’s Native Hist Native Pres May–Jul May–Jul

var.

fendleri secunda var. var. var. var. ora cinale Townsendia exscapa Townsendia incana Townsendia montana Townsendia minima Arabis demissa Tragopogon dubius Tragopogon Viguiera multifl nevadensis Convolvulus arvensis Arabis drummondii Arabis holboellii Arabis holboellii tenuifolia Solidago velutina Sonchus uliginosus Sphaeromeria capitata Stephanomeria tenuifolia offi Taraxacum Solidago simplex Solidago parryi pinetorum Arabis holboellii rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Cruciferae (Brassicaceae)

114 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Claron barrens Disturbed areas Claron barrens Claron barrens Claron barrens Wet Comments x x x x Conf Mont xx xx Jun Pin- xxx xx xxx xxx x xxx xxx xx xx xxx xxx Grs Sage/ Flower time Park status Native Pres Apr–Aug ower Native Pres Apr–Aug ower Native Pres Apr–Jul ax-leaved plainsmustard ax-leaved Native Pres Apr–Jul Breaks bladderpodfl Nativeprince’s-plume Pres May–Jul Native Pres Apr–Sep perennial rockcress rockcressSelby’s Nativewhitetop Presspindlestem Native Mar–Jul PresBreaks draba May–Aug Intro Native Pres Pres Native May–Aug Pres May–Jul May–Aug sand dune wallfl shy wallfl Claron pepperwort Native Pres Apr–Sep Arizona bladderpod Native Hist Apr–Jun lepidote twinpodClaron twinpod Native Pres Native May–Jul Pres Apr–Aug Watson’s bladderpodWatson’s bladderpodKing’s NativeColorado Plateau Presbladderpod Native May–Aug Pres Apr–Jul

var. purshii var.

var. var. var. var. parvifolia var. var. rubicundula Schoenocrambe linifolia Stanleya pinnata Arabis perennans perennans Arabis selbyi Cardaria draba Caulanthus crassicaulis crassicaulis Draba subalpina Erysimum asperum Erysimum inconspicuum Lepidium montanum claronense Physaria arizonica Physaria chambersii Physaria chambersii membranacea sobolifera Physaria intermedia Physaria kingii Physaria rectipes Physaria rubicundula rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 115 x x Wet Comments x Conf Mont Jun Pin- xx xxx xxx xxx xxxx Grs Sage/ Flower time Park status ower Native Pres Apr–Jul ower dragonhead ower Native Pres Jun–Aug x x x eld milkvetch eld Native Pres May–Aug eld mint eld Native Pres Jun–Sep x heart-leaf jewelfl self-heal Native Pres May–Sep annual sagefi Native Pres Jun–Sep x x Fendler’s spurgeFendler’s shorthorn spurgeleafy spurgegolden corydalis Native NativeRocky Mountain gentian Pres gentianParry’s Pres Native Native gentianEngelmann’s May–Sep Intro Pres May–Junelkweed Hist x Repsmall-leaf geranium x Native Jul–Sep Apr–Sep Native x Pres May–Aug crane’s-billRichardson’s x Rep Nativewestern St. John’s-wort x Native Jun–Aug x Pres Jul–Sep NativeRocky Mountain iris Rep Native xIdaho blue-eyed grass x Jun–Aug Pres Pres x Jun–Augsmallfl Nativemock pennyroyal May–Sep Native Jun–Aug x dwarf pennyroyal Pres x Pres xcommon horehound x x May–Jul x Disturbed areas fi May–Aug Nativestinking horsemint Native x Intro Pres Pres x Pres May–Aug Native May–Jun x May–Oct Pres x May–Aug x x x x x x x Disturbed areas Palmer’s thelypodyPalmer’s alpine pennycress Native Pres Native May–Aug Pres Apr–Aug

var. var. nis

orum

glabrata afﬁ

var. var. var. var. var. nis exa Streptanthus cordatus lanceolata Salvia refl Astragalus agrestis Monardella odoratissima Euphorbia brachycera Euphorbia esula Corydalis aurea occidentalis Gentiana parryi Gentianella heterosepala Swertia radiata Geranium caespitosum marginale Hypericum formosum Sisyrinchium idahoense Dracocephalum parvifl Hedeoma drummondii Hedeoma nanum Marrubium vulgare Mentha arvensis Prunella vulgaris Thelypodiopsis sagitatta Thlaspi montanum Chamaesyce fendleri Gentiana affi Geranium richardsonii Iris missouriensis ovalifolia montanum rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Labiatae (Lamiaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Euphorbiaceae Euphorbiaceae Fumariaceae Gentianaceae Gentianaceae Gentianaceae Geraniaceae Guttiferae (Hypericaceae) Iridaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Euphorbiaceae Gentianaceae Geraniaceae Iridaceae

116 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Wet Comments x Conf Mont x x x x Jun Pin- xx xxx xx xxx xx xxx xx xxx xxx xxx xx xx Grs Sage/ Flower time Park status Sheldon’s milkvetchSheldon’s Panguitch milkvetch NativeJohnson milkvetch Pres Native milkvetchTorrey’s Pres Mar–Jun Native May–Jul Pres Native May–Aug Pres Apr–Jul cobwed milkvetchgreat rushy milkvetch Native Native Pres Pres Apr–Jun May–Jul lesser rushy milkvetch Native Pres May–Aug Hall’s milkvetchHall’s groundcover milkvetchtall kentrophyta Native Native Pres Presmountain kentrophyta May–Sep May–Jul Native Native Pres Pres Jun–Sep Jun–Sep rimrock milkvetch Native Pres May–Jul crescent milkvetch Native Pres Mar–Jun woolly milkvetchstinking milkvetch Native Pres Native Pres Feb–Jun Apr–Jul great bladdery milkvetch Native weedy milkvetchRydberg’s Native Pres Pres Apr–Jul Jun–Aug

var. var. var. var. var. var. var. var. var. var. var. var. var. var.

fallax var. var. vespertinus panguicensis major Astragalus argophyllus Astragalus bisulcatus Astragalus calycosus calycosus Astragalus convallarius Astragalus lentiginosus araneosus Astragalus lonchocarpus convallarius humivagans Astragalus desperatus Astragalus hallii Astragalus humistratus Astragalus kentrophyta elatus Astragalus kentrophyta tegetarius desperatus amphioxys Astragalus amphioxys Astragalus miser Astragalus mollissimus thompsoniae Astragalus praelongus praelongus Astragalus amphioxys Astragalus megacarpus parryi oblongifolius rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae)

Chapter 6: Appendices 117 Wet Comments x x x Conf Mont Jun Pin- xxx xx xxx xxx xxx xxx xxx xxxxx x xxx xxx xx xx Grs Sage/ Flower time Park status Lambert’s locoweedLambert’s Native Pres Apr–Sep silky lupinealfalfayellow sweet-clover Nativenodding locoweed Intro Pres Pres May–Aug Native Intro Pres May–Oct Hist Jun–Aug May–Sep Disturbed areas Disturbed areas Marysvale lupine Native Rep May–Aug sink lupine Native Pres May–Aug silverleaf lupine Native Pres May–Aug Ward’s milkvetchWard’s Native PresUtah trefoil May–Sep silvery lupinesubalpine lupine Native Pres Native Native Pres May–Aug Pres May–Aug May–Aug pulse milkvetchnorthern sweetvetch NativeZion sweetpea Pres Native May–Aug Pres May–Aug Native Pres Apr–Jun Jones’ locoweedBicknell locoweedmountain locoweed Native Native Pres Native Pres May–Aug Pres May–Aug May–Aug

jonesii

sericea

var. var. var. var. sericeus marianus barbiger var. var. var. var. var. var. var. var. var. exa cinalis bigelovii Melilotus offi Oxytropis defl Medicago sativa Oxytropis lambertii Lupinus sericeus Lupinus sericeus Lupinus caudatus argophyllus argenteus rubricaulis Astragalus wardii Hedysarum boreale Lathyrus brachycalyx Lotus utahensis Lupinus argenteus Lupinus argenteus Lupinus sericeus Astragalus tenellus boreale zionis Oxytropis oreophila Oxytropis oreophila Oxytropis oreophila juniperina oreophila rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

118 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Claron barrens x x Wet Comments xx x Conf Mont xx Jun Pin- xxx xxx xxx xxx xxxx Grs Sage/ Flower time Park status ax Native Pres May–Aug x x xbarrens Claron ax Native Pres Apr–Aug x x x Paria breadroot Native Pres Jun–Jul dwarf cloverwhite cloverAmerican vetch Native oniontaper-tip Prestwincrest onion IntroNevada onion Native Apr–Aug asparagus Pres Pres Nativesego-lily Apr–Sep Native May–Aug Hist Hist Nativeleopard-lily May–Julstar-lily Pres May–Jul Intro xstarry false Solomon’s-seal Apr–Jul x Native Pres x Native x Pres x Nativemountain death camas Jun–Jul Pres x Pres x Apr–Jun x yellow fl King’s Native Native Jun–Jul blue fl Apr–Jul Pres Presdesert stickleafscarlet globe-mallow x Jun–Aug Apr–Jun small-leaf globemallow xwoodland pinedrops Native Native Native x fragrant sand-verbena Pres Pres Native Preslow sand-verbena Apr–Oct Native Mar–Oct Pres Apr–Sep Pres x x Jun–Sep x Native Apr–Sep x x Pres x x Apr–Sep x x x x narrowleaf four-o’ clocknarrowleaf four-o’ sundropslavender-leaf Native Rep Native Pres May–Aug May–Aug x x x x x Claron barrens linearis americana lewisii var. var. ora var. ssp. cinalis Pediomelum pariense Mentzelia multifl Allium bisceptrum Allium nevadense Asparagus offi Leucocrinum montanum Sphaeralcea coccinea Sphaeralcea parvifolia Pterospora andromedea Abronia nana Mirabilis linearis Trifolium gymnocarpon Trifolium repens Trifolium Vicia americana Allium acuminatum Calochortus nuttallii Fritillaria atropurpurea Smilacina stellata Zigadenus elegans Linum kingii Linum perenne Abronia fragrans Calylophus lavandulifolius rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Loasaceae Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Asparagaceae) Liliaceae (Anthericaceae) Malvaceae Malvaceae Monotropaceae (Ericaceae) Nyctaginaceae Nyctaginaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Alliaceae) Liliaceae (Calochortaceae) Liliaceae Liliaceae (Convallariaceae, Ruscaceae) Liliaceae (Melanthiaceae) Linaceae Linaceae Nyctaginaceae Onagraceae

Chapter 6: Appendices 119 x Wet Comments Conf Mont xx xx Jun Pin- xxx xxx xxx xxx xxx xxx Grs Sage/ Flower time Park status Native PresNative Apr–Jun Hist Jun–Aug reweed Native Rep Jun–Sep slender bog orchidclustered broomrapewoolly-base plantain NativeEnglish plantain Native Pres Native Pres Jun–Jul Pres Jun–Jul Intro May–Aug x Pres x x May–Aug x x scarlet giliascarlet gilia Native Pres Native May–Aug Pres May–Aug Hall’s willow-herbHall’s Nativehairy-throat evening- Presprimrose Jun–Aug x ﬁ Jones’ evening-primrose Nativelong-tube evening- primrose Pres Apr–Jun yellow evening-primrosebronze evening-primrose NativeBridges evening-primrose Native Prespale evening-primrose Native Presspotted coral-root Jun–Jul Rep Nativestriped coral-root May–Julgiant helleborine Jul–Sep Pres x white bog-orchidLindley’s x Native Native May–Jul Native Pres x Rep Native Pres x x Jun–Jul x Pres Jun–Aug Jun–Jul x x Apr–Aug x x x x x x matted prickly-phlox linanthusNuttall’s desert phlox Nativedaggerleaf phlox Pres Native Hist May–Jul Native Native x Jun–Aug Pres Pres x x Apr–Jun May–Jun x x x x x Claron barrens shrubby gilia Native Pres Jun–Sep

var.

pallida var. var. var. var. ava var. var. fl var. var. ora var. ava ora Plantago eriopoda Plantago lanceolata Ipomopsis aggregata Orobanche fasciculata Ipomopsis aggregata aggregata maculata Ipomopsis congesta frutescens Oenothera caespitosa Epilobium halleanum Oenothera caespitosa Oenothera coronopifolia Oenothera fl Epilobium angustifolium canescens crinita marginata Oenothera howardii Oenothera longissima Oenothera pallida Corallorhiza maculata Corallorhiza striata Epipactis gigantea Habenaria dilatata Habenaria sparsifl sparsifl Linanthastrum nuttallii Phlox austromontana austromontana Leptodactylon caespitosum Phlox gladiformis rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Plantaginaceae Plantaginaceae Polemoniaceae Orobanchaceae Polemoniaceae Polemoniaceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Appendix D. Confi FamilyPerennial Forbs, cont. Onagraceae Species name Common nameOnagraceae Onagraceae Onagraceae Nativity Orchidaceae Orchidaceae Orchidaceae Orchidaceae Orchidaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae

120 Annotated Checklist of Vascular Flora: Bryce Canyon National Park x x x Wet Comments x x xx x Conf Mont Jun Pin- xxx xxx xxx xxx xx xxx Grs Sage/ Flower time Park status Native Pres Jun–Aug owered Columbian monkshood Native Pres Jun–Aug whiteveined wintergreen Native Pres Jun–Aug green-ﬂ wintergreen dark-throat shooting-star Nativesilvery primrose Prespink wintergreen Mar–Aug Native Native Rep Pres Jun–Aug Jun–Aug x carpet phloxlong-leaf phloxwestern Jacob’s-ladder Native Native Native Pres Pres Pres Jun–Aug Apr–Jun Apr–Jul x x x x x x winged wild buckwheatWidtsoe wild buckwheat Nativeslender wild buckwheat Native Pres Native RepPanguitch wild buckwheat Native Jun–Oct Pres May–Aug Pres x Jun–Oct Jun–Sep x x x x x Claron barrens Claron barrens redroot wild buckwheat Native Pres Jun–Oct alpine bistortcurly dockwillow dock Native Pres Intro Native Jun–Sep Rep Pres May–Sep May–Sep x x x Disturbed areas Porter’s wild buckwheatPorter’s Nativearid wild buckwheat Pres Native Jul–Sep Pres Jun–Oct cutleaf anemone Native Pres Jun–Aug Colorado columbine Native Pres Jun–Aug

var. var. var. var. var. var. var.

var. ssp. canescens da var. Aconitum columbianum columbianum Pyrola picta Pyrola chlorantha pulchellum Pyrola asarifolia Primula incana Phlox hoodii Phlox longifolia Polemonium caeruleum pterospermum Eriogonum aretioides Eriogonum microthecum Eriogonum racemosum Eriogonum alatum simpsonii Eriogonum panguicense racemosum Rumex crispus Rumex salicifolius Eriogonum umbellatum Eriogonum umbellatum Polygonum viviparum triangulivalvis Dodecatheon pulchellum porteri subaridum Anemone multiﬁ tetonensis Aquilegia coerulea rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Ranunculaceae (Helleboraceae) Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae) Pyrolaceae (Ericaceae) Primulaceae Appendix D. Conﬁ FamilyPerennial Forbs, cont. Polemoniaceae Polemoniaceae Polemoniaceae Species name Common name Nativity Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Primulaceae Ranunculaceae Ranunculaceae (Helleboraceae)

Chapter 6: Appendices 121 Claron barrens x x Wet Comments xx Conf Mont xx x xx Jun Pin- xxx xxx xxx Grs Sage/ Flower time Park status ax Native Pres Apr–Aug slender woodlandstarmarsh grass-of-Parnassus Native Native Pres Pres Apr–Aug May–Oct x x x x rock columbineColumbian virgin’s-bower Nativewhite virgin’s-bowerpale larkspur Native Pres Native Pres larkspurNuttall’s May–Sep Pres Jun–Aug marsh buttercup xdrab buttercup Jul–Aug Native Nativealpine meadow-rue Pres Pres Native meadow-rueFendler’s x Apr–Jun Native Apr–Jul Pres NativeVirginia strawberry Native Pres Pres ivesiaGordon’s May–Sep Pres ivesiaKing’s Jul–Aug Jun–Sep NativeSevier ivesia Jun–Aug x silverweed Presbicrenate cinquefoil Native Jun–Jul Hist Native Native Native Jun–Aug Hist x cinquefoilhair-tuft x Pres Pres Nativewoolly cinquefoil Jun–AugPennsylvania cinquefoil Pres May–Jun Jun–Aug NativePlatte River cinquefoil x Native May–Augsmall bedstraw x Pres Nativemilky kelloggia Pres Native x x Pres xbastard toadﬂ Jul–Aug Pres Jul–Aug Jul–Aug x x Native Jun–Jul Native x Pres x Pres Jul–Aug x x Jun–Aug x x x x x modest cinquefoil Native Rep Jun–Aug

var.

var. var. var. var. dum Aquilegia scopulorum Lithophragma tenellum Parnassia palustris montanensis scopulorum Clematis ligusticifolia Delphinium andersonii Ranunculus inamoenus Thalictrum alpinum Ivesia gordonii Ivesia kingii Ivesia sabulosa Potentilla anserina Potentilla concinna bicrenata Potentilla hippiana Potentilla pensylvanica Potentilla plattensis Galium triﬁ Kelloggia galioides Comandra umbellata pallida Clematis columbiana scaposum Delphinium nuttallianum Ranunculus cymbalaria Thalictrum fendleri Fragaria virginiana Potentilla concinna Potentilla crinita modesta rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Forbs, cont. Ranunculaceae (Helleboraceae) Species name Common name Nativity Saxifragaceae Saxifragaceae (Parnassiaceae) Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Thalictraceae) Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rubiaceae Rubiaceae Santalaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Thalictraceae) Rosaceae Rosaceae Rosaceae

122 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Claron barrens x x Wet Comments x x Conf Mont Jun Pin- xxx xxx xxx xxx xx xxx xxx Grs Sage/ Flower time Park status Native Pres Jun–Aug Native Pres Apr–Jun ower Native Pres Jun–Aug x recracker Native Pres May–Jul x x ower penstemon ower Native Pres Jun–Aug desert paintbrushWyoming paintbrush Native Native Pres Pres Jul–Aug Apr–Jun x x x x x Reveal’s paintbrushReveal’s yellow monkeyfl dwarf lousewort NativeRed Canyon penstemonpainted penstemon Pres Native Nativedusty penstemon Jun–Aug Pres fi Eaton’s Native Pres May–Julundulate penstemon Pres Apr–Jun Native May–Jul Native Pres Pres x Jun–Aug x May–Jul x x Claron barrens x Claron barrens x Markagunt penstemon Native Pres Jun–Aug beaked penstemonRocky Mountain penstemon Utah penstemon Nativewoolly mullein PresAmerican brooklimepinnate angelica Jun–Aug Native Native Pres Intro Pres Native Apr–Jun x Pres Jul–Aug Pres x x Jun–Aug Jun–Aug x x x x x Rockville penstemonsmallfl Native Pres May–Jul cutleaf water-parsnip Native Pres Jun–Sep least spring-parsely Native Pres Jul–Aug Lemmon’s spring-parsleyLemmon’s Native Pres Jul–Aug Colorado Plateau spring- parsley

var.

var. var. var.

eatonii revealii var. var. var. orus var. incisa var. Castilleja chromosa Castilleja linariifolia Castilleja parvula Pedicularis centranthera Mimulus guttatus Penstemon bracteatus Penstemon caespitosus desertipicti Penstemon eatonii undosus Penstemon comarrhenus Penstemon eatonii Penstemon leiophyllus leiophyllus Penstemon strictus thapsus Verbascum americana Veronica Angelica pinnata Penstemon pachyphyllus Penstemon procerus Penstemon rostriﬂ Penstemon utahensis Berula erecta congestus aberrans Cymopterus lemmonii Cymopterus minimus Cymopterus purpureus purpureus rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Forbs, cont. Scrophulariaceae Scrophulariaceae Scrophulariaceae Species name Common name Nativity Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae)

Chapter 6: Appendices 123 Clay soils Claron limestone Wet Comments x x x x Conf Mont x Jun Pin- xxx xxx xxx xx xxx Grs Sage/ Flower time Park status Native Rep May–Jun Native Pres Mar–May Native Pres Aug r dwarf- r western stickseedyellow beeplant Native stinkweedPalmer’s Pres Native Native Apr–Jun Hist Pres Apr–Sep Apr–Oct cupseed stickseed Native Pres Apr–Jun juniper mistletoeprostrate pigweedsand dune cryptanth Native Pres Native Native Hist Jul–Sep Hist Jun–Sep May–Jul x x x x x Disturbed areas southwestern dwarf- mistletoe pinyon dwarf-mistletoeDouglas-ﬁ Nativemistletoe Pres Aug–Sep x least lomatium lomatiumParry’s mountain valerian Nativetobacco-root Native Presprostrate vervainhook-spur violet Native Pres May–Jun northern bog violet Pres Apr–Jun goose-foot violet Native Nativelimber pine dwarf- Jun–Jul Native Nativemistletoe Pres Pres Pres Pres Native May–Sep Jun–Aug Mar–Aug Rep Apr–Jul x x Mar–Aug x x x x x x x x x southern ligusticum Native Pres Jun–Aug Powell’s orachPowell’s summer-cypress Native Intro Pres Pres Jun–Sep Aug–Oct x x x x Clay soils Disturbed areas

var.

var. var. var. powellii venosa var. var. occidentalis Cleome lutea Cleomella palmeriana palmeriana cupulata Lappula occidentalis Amaranthus blitoides Cryptantha fendleri Lappula occidentalis Phoradendron juniperinum Arceuthobium vaginatum cryptopodum Arceuthobium douglasii Arceuthobium divaricatum Valeriana edulis Valeriana bracteata Verbena Viola adunca Viola nephrophylla Viola purpurea Arceuthobium cyanocarpum Lomatium minimum Lomatium parryi acutiloba Valeriana Ligusticum porteri Atriplex powellii Bassia scoparia rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Chenopodiaceae Boraginaceae Annual Forbs Amaranthaceae Boraginaceae Boraginaceae Viscaceae Viscaceae Viscaceae Viscaceae Valerianaceae Verbenaceae Violaceae Violaceae Violaceae Viscaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Valerianaceae Appendix D. Conﬁ FamilyPerennial Forbs, cont. Umbelliferae (Apiaceae) Species name Common name Nativity Chenopodiaceae

124 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Disturbed areas Disturbed areas Disturbed areas Clay soils Disturbed areas x x x Wet Comments x Conf Mont xx x x Jun Pin- x xx xxx xx x xx xx Grs Sage/ Flower time Park status ower Native Pres Jun–Sep mountain goosefoot Native Hist Jul–Sep x x x lambsquartersmallhead goosefootsilvery goosefoot Intro Native Rep Pres Native Jun–Sep Jun–Sep Pres Jun–Aug narrow-leaf goosefoothalogeton NativeRussian thistle seepweedTorrey’s PresCanadian horseweed Jul–Sep Native Intro Introcommon sunﬂ x Native Pres Pres Presprickly lettuce Pres Jun–Sep Jun–Sep Jul–Augthorny wire-lettuce Jul–Sep x spiny-leaf sow-thistle Intro Nativegolden crownbeard x Intro Pres Pressouthern goldeneye Pres Jul–Sep Jul–Sep Nativeblack mustard Jul–Oct Hist Nativeshepherd’s-purse Disturbed areas Disturbed areas Pres May–Oct Jun–Oct Intro Intro Hist Pres May–Aug Apr–Sep Disturbed areas hare’s-ear mustardhare’s-ear mountain tansy-mustard Intro Native Rep Pres Apr–Jul Apr–Aug

var.

var. annua sonnei

var.

var. var. var. var. var. var. var. Chenopodium capitatum Chenopodium atrovirens Chenopodium fremontii Chenopodium album album parvicapitatum incanum Halogeton glomeratus Salsola tragus Suaeda torreyana torreyana exauriculata Capsella bursa-pastoris Chenopodium leptophyllum Conyza canadensis glabrata Helianthus annuus lenticularis Lactuca serriola spinosa Lygodesmia Sonchus asper encelioides Verbesina Viguiera longifolia Brassica nigra Conringia orientalis Descurainia incana rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Appendix D. Conﬁ FamilyAnnual Forbs, cont. Chenopodiaceae Species name Common name Nativity Chenopodiaceae Chenopodiaceae Chenopodiaceae Cruciferae (Brassicaceae) Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 125 Disturbed areas Disturbed areas x Wet Comments xx Conf Mont x xx Jun Pin- xxx xx xxx xx xxx xx x Grs Sage/ Flower time Park status Native Pres Mar–Jun ax Native Pres Jun–Aug x x x ixweed Intro eld pennycress Pres Mar–Aug Intro Pres May–Aug Disturbed areas stork’s-billbrittle phaceliaNipple Bench phacelia lupineKing’s Native Native Pres Intro Rep Apr–Jun Pres Native May–Jun x Feb–Oct Pres x May–Aug x x Clay soils Shale Tropic Disturbed areas King’s lupineKing’s Native Pres May–Aug meadow gentian Native Pres Jul–Sep black medicksmall yellow fl common mallowPaiute suncup Intro Intro Pres Native Pres May–Oct Rep Apr–Oct May–Jun Disturbed areas fl clasping pepperwortAfrican mustard Introfi PresJones’ gentian Intro May–Jul Pres Apr–Jul Native Pres Jul–Aug x x Disturbed areas western tansy-mustardwestern tansy-mustard Native Pres Native Mar–Aug Presprairie pepperwort Mar–Aug Native Pres May–Jul raceme groundsmokebranching groundsmokewhite-stem evening- Native Nativeprimrose Pres Pres May–Aug May–Aug x x x x x x

var. var. var. var. demissa var. var. argillaceus kingii var. var. orum var. var. orum Phacelia demissa Phacelia mammillarensis Lupinus kingii Erodium cicutarium Lupinus kingii Malva neglecta elegans Camissonia scapoidea scapoidea Medicago lupulina Linum australe Gentianella tortuosa Thlaspi arvense Gentianopsis detonsa Descurainia pinnata Descurainia sophia Lepidium densifl Lepidium perfoliatum Malcolmia africana Descurainia pinnata intermedia osmiarum densifl Gayophytum ramosissimum Oenothera albicaulis Gayophytum racemosum rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Hydrophyllaceae Hydrophyllaceae Leguminosae (Fabaceae) Geraniaceae Leguminosae (Fabaceae) Malvaceae Onagraceae Leguminosae (Fabaceae) Linaceae Gentianaceae Cruciferae (Brassicaceae) Gentianaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Confi FamilyAnnual Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Onagraceae Onagraceae Onagraceae

126 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Wet Comments Conf Mont Jun Pin- xxx xx xx Grs Sage/ Flower time Park status Native Pres Apr–Oct ower Native Pres Apr–Jun x x x ower rock-jasmine ower Native Pres May–Aug x x x ower bird’s-beak ower Native Rep Aug–Sep x x King’s bird’s-beakKing’s smallfl bird’s-beakWright’s reddish monkeyfl Native Native Pres Rep Jun–Sep Jul–Sep x x x x x kidney-shaped wild buckwheat yard knotweedblack-bindweedSawatch knotweedpygmy-fl Intro Intro Native Pres Pres Pres May–Oct Jun–Aug May–Octyellow owl-cloverviolet owl-cloverbuffaloburground-cherry nightshade Native Introcut-leaf nightshade Native Pres Pres sedgeslender-beaked Pres Jul–Aug Nativegolden sedge Intro May–Oct Jul–AugDouglas’ sedge Disturbed areas Native Hist x Disturbed areas narrowleaf sedge Pres Pres May–Sepinland sedge Jun–Sepsmall-wing sedge May–Aug x Native xNebraska sedge Native x Native Presgreen sedge x x Pres Pres x woolly sedge Native Apr–Aug Nativeblackcreeper sedge Apr–Aug Jun–Jul Pres Native Pres Disturbed areas x Rep x Jun–Aug May–Jul Native Native x Native x May–Jul Pres x Pres Pres Disturbed areas May–Aug Jun–Aug May–Sep x x x x x x x nodding wild buckwheat Native Pres Apr–Oct California poppy Intro Pres Apr–May Cultivated Ross’ sedgebeaked sedge Native Native Pres Pres May–Aug Jun–Aug x x x x

kingii var. var. var. orus viridula var. orum Cordylanthus parvifl Cordylanthus wrightii Mimulus rubellus Cordylanthus kingii Orthocarpus luteus Polygonum convolvulus Polygonum douglasii johnstonii Orthocarpus purpureoalbus Solanum rostratum Solanum sarrachoides Solanum trifl Carex athrostachya Carex aurea Carex douglasii Carex duriuscula Carex interior Carex microptera Carex nebrascensis Carex oederi Carex pellita Carex praegracilis Carex rossii Eriogonum subreniforme Polygonum aviculare Androsace septentrionalis cernuum Eschscholzia californica Eriogonum cernuum Carex utriculata rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Polygonaceae Polygonaceae Scrophulariaceae Solanaceae Solanaceae Solanaceae Graminoids Perennial Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Polygonaceae Polygonaceae Primulaceae Appendix D. Confi FamilyAnnual Forbs, cont. Papaveraceae Polygonaceae Species name Common name Nativity Cyperaceae

Chapter 6: Appendices 127 x Wet Comments Conf Mont Jun Pin- Grs Sage/ Flower time Park status ower spikerush ower Native Pres Jun–Aug x valley sedgecommon spikerushfew-ﬂ Native Native Pres Pres May–Aug May–Jul x x x beaked spikerushthree-square bulrushcrested wheatgrass Native Nativeredtoppurple three-awn Pres Rep Introblue grama May–Aug Jun–Augnodding brome Presmountain brome Nativefringed brome Jun–Aug Pressmooth brome Intro NativeJones’ reedgrass x Native Mar–Sep Nativeslimstem reedgrass Pres Pres Pres x Presorchard grass x Nativetufted hairgrass Jun–Aug Jun–Sep Intro Jul–Sep x x Pres May–Aug Native xdesert saltgrass Native xCanada wildrye x Pres Pres x Jul–Sep Pres x Great Basin wildrye x Intro Jun–Aug Jul–Aug Native xsquirreltail Jul–Augblue wildrye Native x Pres Pres x x Native Nativeintermediate wheatgrass Pres Jul–SepRussian wildrye Jun–Aug Pres Intro Pres xthickspike wheatgrass x Jun–Sep x x Jun–AugSalina wildrye Pres Jul–Aug x Native x NativeScribner wheatgrass Native x Pres Jun–Aug x western wheatgrass Intro Pres Pres x bluebunch wheatgrass x x x May–Aug x Native Pres Jul–Augslender wheatgrass x Jun–Aug Native Native x Native x Pres x Jun–Jul Pres x x Pres Pres x Jul–Aug x x Native x Jun–Jul x x Jun–Aug Jun–Aug x Pres x x x x x x x Jun–Aug x x x x x x x x x x

ora inermis var. var. Carex vallicola Eleocharis palustris Eleocharis quinqueﬂ Eleocharis rostellata Scirpus pungens longispicatus Agrostis stolonifera Aristida purpurea Bouteloua gracilis Bromus anomalus Bromus carinatus Bromus ciliatus Bromus inermis Calamagrostis scopulorum Calamagrostis stricta Dactylis glomerata Deschampsia cespitosa Distichlis spicata Elymus canadensis Elymus cinereus Elymus elymoides Elymus glaucus Elymus hispidus Elymus junceus Elymus lanceolatus Elymus salinus Elymus scribneri Elymus smithii Elymus spicatus Elymus trachycaulus Agropyron cristatum rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Cyperaceae Cyperaceae Cyperaceae Species name Common name Nativity Cyperaceae Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

128 Annotated Checklist of Vascular Flora: Bryce Canyon National Park Wet Comments Conf Mont Jun Pin- xx Grs Sage/ Flower time Park status sand dropseedneedle-and-threadIndian ricegrass needlegrassLetterman’s needlegrassNelson’s Native Native Native Pres Pres Pres Native Native May–Jul Pres Jun–Aug Jul–Sep Pres x x May–Jul x Jun–Sep x x x x x x x x x x x Rocky Mountain fescuemeadow fescue Nativefowl mannagrassgalleta Presmeadow barley Jul–Aug Introfoxtail barley Nativejunegrass Pres Presfoxtail muhly Nativescratchgrass Jun–Aug Jul–Augmountain muhly Pres Native Nativesandhill muhly x Pres Presmat muhly Jun–Aug Native Nativelittleseed ricegrass May–Aug Native Jun–Aug Prestimothy Native Pres xcommon reed Pres x Native x Pres Jul–Sep Jun–AugCanada bluegrass Native Pres x Jul–Sep x muttongrass x Jul–Aug Native x Pres x xinterior bluegrass Jul–Sep Pres xKentucky bluegrass x Jun–Aug Intro Native x bluegrassSandberg’s Intro x Jul–Sep x Pres x x alkaligrassNuttall’s Pres Native Native Intro Preslittle bluestem x Native x Jul–Sep x Pres x Jun–Augalkali cordgrass Pres Pres Jun–Sep Pres Native x alkali sacaton x x Jun–Aug x May–Aug x Pres May–Aug Apr–Sep x x Native x x x May–Aug Native x x Pres x x x Pres Native x x x Jul–Sep x x Pres Jun–Sep x x x x x x Jun–Aug x x x x x pine needlegrass Native Rep Jun–Aug x x x

var. saximontana nelsonii comata glauca var. var. var. var. Stipa comata Stipa hymenoides Stipa lettermanii Stipa nelsonii Sporobolus cryptandrus Festuca ovina Festuca pratensis Glyceria striata Hilaria jamesii Hordeum brachyantherum Hordeum jubatum Koeleria macrantha Muhlenbergia andina Muhlenbergia asperifolia Muhlenbergia montana Muhlenbergia pungens Muhlenbergia richardsonis Oryzopsis micrantha Phleum pratense Phragmites australis Poa compressa Poa fendleriana Poa glauca Poa pratensis Poa secunda Puccinellia nuttalliana Schizachyrium scoparium Spartina gracilis Sporobolus airoides airoides Stipa pinetorum rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Species nameGramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Common nameGramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 129 Rock crevices x Wet Comments xx Conf Mont Jun Pin- xxx Grs Sage/ Flower time Park status eld horsetail eld Native Pres NA x x x Bloomer’s ricegrassBloomer’s Baltic rush rushTracy’s Native Pres Jun–Aug Native Native x Pres Pres x May–Jul Jun–Aug x longstyle rush rushTorrey’s seaside arrowgrassmarsh arrowgrasscommon cattail Native Native Native PresJapanese chess Native Prescheatgrass Pres Jun–Aug Pres Nativesix-weeks fescue May–Aug Jun–Augpullup muhly Pres Jun–Augwitchgrass Introgreen bristlegrass Jun–Aug Native Prestoad rush Intro Pres Native Jun–Augﬁ Pres Intro Apr–Jun Pressmooth scouring-rush x x Native x May–Jul scouring-rushNelson’s Pres x x Jun–Aug Presslender lip-fern x Native x Native x Jun–Sep Native x Jun–Sep Pres x Pres Pres x x NA x Jun–Aug x Native NA Pres x NA x Disturbed areas x x x bracken fern Native Pres NA

elata var.

var. var. liformis ora nelsonii x bloomeri x Stipa Juncus arcticus Juncus ensifolius brunnescens Equisetum laevigatum Juncus torreyi maritima Triglochin palustris Triglochin latifolia Typha Bromus japonicus Bromus tectorum Festuca octofl Muhlenbergia fi Panicum capillare Setaria viridis Juncus bufonius Equisetum arvense Equisetum Cheilanthes feei Juncus longistylus Pteridium aquilinum pubescens rmed, historical, and reported taxa of Bryce Canyon National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Juncaceae Juncaceae Species name Common name Nativity Equisetaceae Juncaceae Juncaginaceae Juncaginaceae Typhaceae Annual Graminoids Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Ferns and Allies Equisetaceae Equisetaceae Polypodiaceae (Adiantaceae) Juncaceae Polypodiaceae (Dennstaedtiaceae)

130 Annotated Checklist of Vascular Flora: Bryce Canyon National Park The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientiﬁ c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and afﬁ liated Island Communities.

NPS D-244, January 2009 National Park Service U.S. Department of the Interior

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