Annotated Checklist of Vascular Flora, Bryce
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Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications Sherwin Carlquist Santa Barbara Botanic Garden
Aliso: A Journal of Systematic and Evolutionary Botany Volume 14 | Issue 1 Article 2 1995 Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications Sherwin Carlquist Santa Barbara Botanic Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Carlquist, Sherwin (1995) "Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 14: Iss. 1, Article 2. Available at: http://scholarship.claremont.edu/aliso/vol14/iss1/2 Aliso, 14(1), pp. 1-17 © 1995, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 WOOD ANATOMY OF CARYOPHYLLACEAE: ECOLOGICAL, HABITAL, SYSTEMATIC, AND PHYLOGENETIC IMPLICATIONS SHERWIN CARLQUIST1 Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara, CA 93105 ABSTRACT Wood of Caryophyllaceae is more diverse than has been appreciated. Imperforate tracheary elements may be tracheids, fiber-tracheids, or libriform fibers. Rays may be uniseriate only, multiseriate only, or absent. Roots of some species (and sterns of a few of those same genera) have vascular tissue produced by successive cambia. The diversity in wood anatomy character states shows a range from primitive to specialized so great that origin close to one of the more specialized families of Cheno podiales, such as Chenopodiaceae or Amaranthaceae, is unlikely. Caryophyllaceae probably branched from the ordinal clade near the clade's base, as cladistic evidence suggests. Raylessness and abrupt onset of multiseriate rays may indicate woodiness in the family is secondary. Successive cambia might also be a subsidiary indicator of secondary woodiness in Caryophyllaceae (although not necessarily dicotyledons at large). -
Wood Anatomy of Caryophyllaceae: Ecological, Habital, Systematic, and Phylogenetic Implications
Aliso, 14(1), pp. 1—17 © 1995, by The Rancho Santa Ana Botanic Garden, Claremont, CA 91711-3157 WOOD ANATOMY OF CARYOPHYLLACEAE: ECOLOGICAL, HABITAL, SYSTEMATIC, AND PHYLOGENETIC IMPLICATIONS SHERwIN CARLQuIsT’ Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara, CA 93105 ABSTRACT Wood of Caryophyllaceae is more diverse than has been appreciated. Imperforate tracheary elements may be tracheids, fiber-tracheids, or libriform fibers. Rays may be unisei-iate only, multiseriate only, or absent. Roots of some species (and stems of a few of those same genera) have vascular tissue produced by successive cambia. The diversity in wood anatomy character states shows a range from primitive to specialized so great that origin close to one of the more specialized families of Cheno podiales, such as Chenopodiaceae or Amaranthaceae, is unlikely. Caryophyllaceae probably branched from the ordinal dade near the dade’s base, as cladistic evidence suggests. Raylessness and abrupt onset of multiseriate rays may indicate woodiness in the family is secondary. Successive cambia might also be a subsidiary indicator of secondary woodiness in Caryophyllaceae (although not necessarily dicotyledons at large). Although a small shrub, Gyinnocarpos may be primitively woody, and because that genus appears primitive in many wood features the family as a whole may derive from woody ancestors, despite apparent secondary woodiness in many phylads. Systematic distribution of wood character states in the family corresponds closely to the Pax and Hoffmann system of three tribes and their progressive degrees of specialization (Paronychieae, Alsineae, Sileneae). Wood data support the idea that Caryophyllaceae represent a branch from near the base of the order Caryophyllales. -
Pima County Plant List (2020) Common Name Exotic? Source
Pima County Plant List (2020) Common Name Exotic? Source McLaughlin, S. (1992); Van Abies concolor var. concolor White fir Devender, T. R. (2005) McLaughlin, S. (1992); Van Abies lasiocarpa var. arizonica Corkbark fir Devender, T. R. (2005) Abronia villosa Hariy sand verbena McLaughlin, S. (1992) McLaughlin, S. (1992); Van Abutilon abutiloides Shrubby Indian mallow Devender, T. R. (2005) Abutilon berlandieri Berlandier Indian mallow McLaughlin, S. (1992) Abutilon incanum Indian mallow McLaughlin, S. (1992) McLaughlin, S. (1992); Van Abutilon malacum Yellow Indian mallow Devender, T. R. (2005) Abutilon mollicomum Sonoran Indian mallow McLaughlin, S. (1992) Abutilon palmeri Palmer Indian mallow McLaughlin, S. (1992) Abutilon parishii Pima Indian mallow McLaughlin, S. (1992) McLaughlin, S. (1992); UA Abutilon parvulum Dwarf Indian mallow Herbarium; ASU Vascular Plant Herbarium Abutilon pringlei McLaughlin, S. (1992) McLaughlin, S. (1992); UA Abutilon reventum Yellow flower Indian mallow Herbarium; ASU Vascular Plant Herbarium McLaughlin, S. (1992); Van Acacia angustissima Whiteball acacia Devender, T. R. (2005); DBGH McLaughlin, S. (1992); Van Acacia constricta Whitethorn acacia Devender, T. R. (2005) McLaughlin, S. (1992); Van Acacia greggii Catclaw acacia Devender, T. R. (2005) Acacia millefolia Santa Rita acacia McLaughlin, S. (1992) McLaughlin, S. (1992); Van Acacia neovernicosa Chihuahuan whitethorn acacia Devender, T. R. (2005) McLaughlin, S. (1992); UA Acalypha lindheimeri Shrubby copperleaf Herbarium Acalypha neomexicana New Mexico copperleaf McLaughlin, S. (1992); DBGH Acalypha ostryaefolia McLaughlin, S. (1992) Acalypha pringlei McLaughlin, S. (1992) Acamptopappus McLaughlin, S. (1992); UA Rayless goldenhead sphaerocephalus Herbarium Acer glabrum Douglas maple McLaughlin, S. (1992); DBGH Acer grandidentatum Sugar maple McLaughlin, S. (1992); DBGH Acer negundo Ashleaf maple McLaughlin, S. -
Vascular Plant and Vertebrate Inventory of Chiricahua National Monument
In Cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Chiricahua National Monument Open-File Report 2008-1023 U.S. Department of the Interior U.S. Geological Survey National Park Service This page left intentionally blank. In cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Chiricahua National Monument By Brian F. Powell, Cecilia A. Schmidt, William L. Halvorson, and Pamela Anning Open-File Report 2008-1023 U.S. Geological Survey Southwest Biological Science Center Sonoran Desert Research Station University of Arizona U.S. Department of the Interior School of Natural Resources U.S. Geological Survey 125 Biological Sciences East National Park Service Tucson, Arizona 85721 U.S. Department of the Interior DIRK KEMPTHORNE, Secretary U.S. Geological Survey Mark Myers, Director U.S. Geological Survey, Reston, Virginia: 2008 For product and ordering information: World Wide Web: http://www.usgs.gov/pubprod Telephone: 1-888-ASK-USGS For more information on the USGS-the Federal source for science about the Earth, its natural and living resources, natural hazards, and the environment: World Wide Web:http://www.usgs.gov Telephone: 1-888-ASK-USGS Suggested Citation Powell, B.F., Schmidt, C.A., Halvorson, W.L., and Anning, Pamela, 2008, Vascular plant and vertebrate inventory of Chiricahua National Monument: U.S. Geological Survey Open-File Report 2008-1023, 104 p. [http://pubs.usgs.gov/of/2008/1023/]. Cover photo: Chiricahua National Monument. Photograph by National Park Service. Note: This report supersedes Schmidt et al. (2005). Any use of trade, product, or firm names is for descriptive purposes only and does not imply endorsement by the U.S. -
Legumes of the North-Central States: C
LEGUMES OF THE NORTH-CENTRAL STATES: C-ALEGEAE by Stanley Larson Welsh A Dissertation Submitted, to the Graduate Faculty in Partial Fulfillment of The Requirements for the Degree of DOCTOR OF PHILOSOPHY Major Subject: Systematic Botany Approved: Signature was redacted for privacy. Signature was redacted for privacy. artment Signature was redacted for privacy. Dean of Graduat College Iowa State University Of Science and Technology Ames, Iowa I960 ii TABLE OF CONTENTS Page ACKNOWLEDGMENTS iii INTRODUCTION 1 HISTORICAL ACCOUNT 3 MATERIALS AND METHODS 8 TAXONOMIC AND NOMENCLATURE TREATMENT 13 REFERENCES 158 APPENDIX A 176 APPENDIX B 202 iii ACKNOWLEDGMENTS The writer wishes to express his deep gratitude to Professor Duane Isely for assistance in the selection of the problem and for the con structive criticisms and words of encouragement offered throughout the course of this investigation. Support through the Iowa Agricultural Experiment Station and through the Industrial Science Research Institute made possible the field work required in this problem. Thanks are due to the curators of the many herbaria consulted during this investigation. Special thanks are due the curators of the Missouri Botanical Garden, U. S. National Museum, University of Minnesota, North Dakota Agricultural College, University of South Dakota, University of Nebraska, and University of Michigan. The cooperation of the librarians at Iowa State University is deeply appreciated. Special thanks are due Dr. G. B. Van Schaack of the Missouri Botanical Garden library. His enthusiastic assistance in finding rare botanical volumes has proved invaluable in the preparation of this paper. To the writer's wife, Stella, deepest appreciation is expressed. Her untiring devotion, work, and cooperation have made this work possible. -
December 2012 Number 1
Calochortiana December 2012 Number 1 December 2012 Number 1 CONTENTS Proceedings of the Fifth South- western Rare and Endangered Plant Conference Calochortiana, a new publication of the Utah Native Plant Society . 3 The Fifth Southwestern Rare and En- dangered Plant Conference, Salt Lake City, Utah, March 2009 . 3 Abstracts of presentations and posters not submitted for the proceedings . 4 Southwestern cienegas: Rare habitats for endangered wetland plants. Robert Sivinski . 17 A new look at ranking plant rarity for conservation purposes, with an em- phasis on the flora of the American Southwest. John R. Spence . 25 The contribution of Cedar Breaks Na- tional Monument to the conservation of vascular plant diversity in Utah. Walter Fertig and Douglas N. Rey- nolds . 35 Studying the seed bank dynamics of rare plants. Susan Meyer . 46 East meets west: Rare desert Alliums in Arizona. John L. Anderson . 56 Calochortus nuttallii (Sego lily), Spatial patterns of endemic plant spe- state flower of Utah. By Kaye cies of the Colorado Plateau. Crystal Thorne. Krause . 63 Continued on page 2 Copyright 2012 Utah Native Plant Society. All Rights Reserved. Utah Native Plant Society Utah Native Plant Society, PO Box 520041, Salt Lake Copyright 2012 Utah Native Plant Society. All Rights City, Utah, 84152-0041. www.unps.org Reserved. Calochortiana is a publication of the Utah Native Plant Society, a 501(c)(3) not-for-profit organi- Editor: Walter Fertig ([email protected]), zation dedicated to conserving and promoting steward- Editorial Committee: Walter Fertig, Mindy Wheeler, ship of our native plants. Leila Shultz, and Susan Meyer CONTENTS, continued Biogeography of rare plants of the Ash Meadows National Wildlife Refuge, Nevada. -
Rubus Arcticus Ssp. Acaulis Is Also Appreciated
Rubus arcticus L. ssp. acaulis (Michaux) Focke (dwarf raspberry): A Technical Conservation Assessment Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project October 18, 2006 Juanita A. R. Ladyman, Ph.D. JnJ Associates LLC 6760 S. Kit Carson Cir E. Centennial, CO 80122 Peer Review Administered by Society for Conservation Biology Ladyman, J.A.R. (2006, October 18). Rubus arcticus L. ssp. acaulis (Michaux) Focke (dwarf raspberry): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http:// www.fs.fed.us/r2/projects/scp/assessments/rubusarcticussspacaulis.pdf [date of access]. ACKNOWLEDGMENTS The time spent and help given by all the people and institutions mentioned in the reference section are gratefully acknowledged. I would also like to thank the Wyoming Natural Diversity Database, in particular Bonnie Heidel, and the Colorado Natural Heritage Program, in particular David Anderson, for their generosity in making their records available. The data provided by Lynn Black of the DAO Herbarium and National Vascular Plant Identification Service in Ontario, Marta Donovan and Jenifer Penny of the British Columbia Conservation Data Center, Jane Bowles of University of Western Ontario Herbarium, Dr. Kadri Karp of the Aianduse Instituut in Tartu, Greg Karow of the Bighorn National Forest, Cathy Seibert of the University of Montana Herbarium, Dr. Anita Cholewa of the University of Minnesota Herbarium, Dr. Debra Trock of the Michigan State University Herbarium, John Rintoul of the Alberta Natural Heritage Information Centre, and Prof. Ron Hartman and Joy Handley of the Rocky Mountain Herbarium at Laramie, were all very valuable in producing this assessment. -
Fort Valley Experimental Forest-A Century of Research 1908-2008
Vascular Plant Checklist of the Chimney Spring and Limestone Flats Prescribed Burning Study Areas Within Ponderosa Pine Experimental Forests in Northern Arizona Catherine Scudieri, School of Forestry, Northern Arizona University, Flagstaff, AZ, USFS, Rocky Mountain Research Station, Flagstaff, AZ; James F. Fowler, Carolyn Hull Sieg, USFS, Rocky Mountain Research Station, Flagstaff, AZ; Laura Williams, Department of Biological Sciences, Northern Arizona University, Flagstaff, AZ; and Sally M. Haase, USFS, Pacific Southwest Research Station, Riverside, CA Abstract—This paper presents a vascular plant species list for two sites that are part of a long-term study exploring the effects of varying fire intervals on forest characteristics includ- ing the abundance and composition of understory vegetation. The Chimney Spring study area is on the Fort Valley Experimental Forest near Flagstaff, AZ, and the Limestone Flats study area is on the Long Valley Experimental Forest, 90 km (56 mi) southeast of Flagstaff. Since 1976 (Chimney Spring) and 1977 (Limestone Flats), three replicates of each of seven burn intervals (1, 2, 4, 6, 8, 10 years, plus unburned) have been maintained by the USFS Pacific Southwest Research Station. Each study area encompasses approximately 40 to 48 ha (99 to 119 acres) of dense ponderosa pine (Pinus ponderosa) forest. Our plant species list was generated through systematic sampling of the understory vegetation in 2006 and 2007 as well as surveys of the entire study areas for additional species. We documented a total of 147 species, with 96 species found at Chimney Spring and 123 species at Limestone Flats. There are eight introduced species on the list, with six introduced species found at Chimney Spring and seven found at Limestone Flats. -
Rare Plant Survey of San Juan Public Lands, Colorado
Rare Plant Survey of San Juan Public Lands, Colorado 2005 Prepared by Colorado Natural Heritage Program 254 General Services Building Colorado State University Fort Collins CO 80523 Rare Plant Survey of San Juan Public Lands, Colorado 2005 Prepared by Peggy Lyon and Julia Hanson Colorado Natural Heritage Program 254 General Services Building Colorado State University Fort Collins CO 80523 December 2005 Cover: Imperiled (G1 and G2) plants of the San Juan Public Lands, top left to bottom right: Lesquerella pruinosa, Draba graminea, Cryptantha gypsophila, Machaeranthera coloradoensis, Astragalus naturitensis, Physaria pulvinata, Ipomopsis polyantha, Townsendia glabella, Townsendia rothrockii. Executive Summary This survey was a continuation of several years of rare plant survey on San Juan Public Lands. Funding for the project was provided by San Juan National Forest and the San Juan Resource Area of the Bureau of Land Management. Previous rare plant surveys on San Juan Public Lands by CNHP were conducted in conjunction with county wide surveys of La Plata, Archuleta, San Juan and San Miguel counties, with partial funding from Great Outdoors Colorado (GOCO); and in 2004, public lands only in Dolores and Montezuma counties, funded entirely by the San Juan Public Lands. Funding for 2005 was again provided by San Juan Public Lands. The primary emphases for field work in 2005 were: 1. revisit and update information on rare plant occurrences of agency sensitive species in the Colorado Natural Heritage Program (CNHP) database that were last observed prior to 2000, in order to have the most current information available for informing the revision of the Resource Management Plan for the San Juan Public Lands (BLM and San Juan National Forest); 2. -
Vascular Plants and a Brief History of the Kiowa and Rita Blanca National Grasslands
United States Department of Agriculture Vascular Plants and a Brief Forest Service Rocky Mountain History of the Kiowa and Rita Research Station General Technical Report Blanca National Grasslands RMRS-GTR-233 December 2009 Donald L. Hazlett, Michael H. Schiebout, and Paulette L. Ford Hazlett, Donald L.; Schiebout, Michael H.; and Ford, Paulette L. 2009. Vascular plants and a brief history of the Kiowa and Rita Blanca National Grasslands. Gen. Tech. Rep. RMRS- GTR-233. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 44 p. Abstract Administered by the USDA Forest Service, the Kiowa and Rita Blanca National Grasslands occupy 230,000 acres of public land extending from northeastern New Mexico into the panhandles of Oklahoma and Texas. A mosaic of topographic features including canyons, plateaus, rolling grasslands and outcrops supports a diverse flora. Eight hundred twenty six (826) species of vascular plant species representing 81 plant families are known to occur on or near these public lands. This report includes a history of the area; ethnobotanical information; an introductory overview of the area including its climate, geology, vegetation, habitats, fauna, and ecological history; and a plant survey and information about the rare, poisonous, and exotic species from the area. A vascular plant checklist of 816 vascular plant taxa in the appendix includes scientific and common names, habitat types, and general distribution data for each species. This list is based on extensive plant collections and available herbarium collections. Authors Donald L. Hazlett is an ethnobotanist, Director of New World Plants and People consulting, and a research associate at the Denver Botanic Gardens, Denver, CO. -
Canyons of the Ancients National Monument Plant List by Genus
Canyons of the Ancients National Monument Plant List Please send all corrections and updates to Al Schneider, [email protected] Updated 6/2011 Scientific Name Common name Family Abronia fragrans Sand-verbena Nyctaginaceae Achillea lanulosa Western yarrow Asteraceae Achnatherum hymenoides Indian ricegrass Poaceae Achnatherum speciosum Showy needle grass Poaceae Acosta diffusa Tumble knapweed Asteraceae Acosta maculosa Spotted knapweed Asteraceae Acrolasia albicaulis Whitestem blazingstar Loasaceae Acroptilon repens Russian knapweed Asteraceae Adenolinum lewisii Blue Flax Linaceae Adiantum capillus-veneris Venus' hair fern Adiantaceae Agropyron cristatum Crested wheatgrass Poaceae Agrostis scabra Rough bentgrass Poaceae Agrostis stolonifera Redtop bentgrass Poaceae Allium acuminatum Tapertip onion Alliaceae Allium macropetalum Largeflower wild onion Alliaceae Allium textile Textile onion Alliaceae Alyssum minus Yellow alyssum Brassicaceae Amaranthus blitoides Prostrate pigweed Amaranthaceae Amaranthus retroflexus Redroot amaranth Amaranthaceae Ambrosia acanthicarpa Flatspine burr ragweed Asteraceae Ambrosia trifida great ragweed Asteraceae Amelanchier alnifolia? Saskatoon serviceberry Rosaceae Amelanchier utahensis Utah serviceberry Rosaceae Amsonia jonesii Jones's bluestar Apocynaceae Androsace occidentalis Western rockjasmine Primulaceae Androsace septentrionalis Pygmyflower rockjasmine Primulaceae Androstephium breviflorum Pink funnellily Alliaceae Anisantha tectorum Cheatgrass Poaceae Antennaria rosulata Rosy pussytoes Asteraceae -
Pamela Ricca Belter
University of Alberta Relationships among diverse root foraging behaviours: understanding plant behavioural types by Pamela Ricca Belter A thesis submitted to the Faculty of Graduate Studies and Research in partial fulfillment of the requirements for the degree of Master of Science in Ecology Department of Biological Sciences © Pamela Ricca Belter Spring 2014 Edmonton, Alberta Permission is hereby granted to the University of Alberta Libraries to reproduce single copies of this thesis and to lend or sell such copies for private, scholarly or scientific research purposes only. Where the thesis is converted to, or otherwise made available in digital form, the University of Alberta will advise potential users of the thesis of these terms. The author reserves all other publication and other rights in association with the copyright in the thesis and, except as herein before provided, neither the thesis nor any substantial portion thereof may be printed or otherwise reproduced in material form whatsoever without the author’s prior written permission. ABSTRACT Behaviours capture the functional response of plants to environmental factors. I explore behaviours for twenty co-occurring grassland species in response to common belowground environmental factors (competition, mycorrhizae, heterogeneous and high nutrients) and their relationship to plant outcomes. First, detailed plant response to neighbours was observed using a specialized apparatus which allowed for visualization of the root system in situ. Species varied in response from strongly aggregating to strongly segregating roots by adjusting overall size and occupation of the soil and/or by altering allocation to roots and the specific placement of roots. In the second study, associations among response to nutrients, mycorrhizae, neighbours, foraging precision, and soil exploration behaviours created behavioural types along two axes: assertiveness and focus.