The Vercors (By Plane)

France - The Vercors (By Plane)

Naturetrek Tour Report 21 - 28 May 2017

Common Redstart Green Hairstreak

Narcissus poeticus Scarce Swallowtail

Report compiled by Paul Harmes Images courtesy of Clive Pickton

Naturetrek Wolf’s Lane Chawton Alton Hampshire GU34 3HJ UK T: +44 (0)1962 733051 E: [email protected] W: www.naturetrek.co.uk

Tour Report France - The Vercors

Tour Participants: Paul Harmes (leader) and and seven Naturetrek clients.

Day 1 Sunday 21st May

London to Lyon St. Exupery – La Chapelle en Vercors Seven group members met with Paul at Lyon St. Exupery Airport, after catching flight BA360 from London Heathrow, Terminal 3. After completing immigration formalities and collecting our luggage, we made our way to the car park, where our vehicle was waiting. With luggage loaded, we set off travelling towards Grenoble, on the A43 and A48. Along the way White Stork, several Black Kites and Common Buzzard were noted. We stopped for our picnic lunch at a motorway rest area, Aire Burcin. With our lunch completed, we had a look for some butterflies, finding Common Blue, Heath Fritillary, Small White, Small Heath and Black-veined White. We also found Burnet Companion moths. For the birds, there was Common Chaffinch, Spotted Flycatcher and Great Tit.

Continuing on down the A48, we left the motorway, and made our way the area immediately around Grenoble Isère Airport. Here we found Common Buzzard, European Stonechat, Serin and Common Kestrel. Along the arable margins we found Cruciata laevipes (Crosswort), Gallium mollugo (Hedge Bedstraw) and Papaver rhoeas (Common Poppy).

We decided to take the inland route onwards, so we drove up through Saint-Marcellin, from where we made our way up onto the Vercors plateau, via Pont-en-Royans and the spectacular Petit Goulets. At this point, we stopped to check out a verge bank near the hamlet of Toutre. Eight species of Orchid were found, including Orchis simia (Monkey Orchid), Cephalanthera longifolia (Sword-leaved Helleborine) and Ophrys fuciflora (Late Spider Orchid). In addition, there was the curious Asarum europaeum (Asarabacca), and in a nearby field, Sooty Copper butterfly.

We now moved on to La Chapelle en Vercors and the Hotel Bellier, our base for the week. After settling into our rooms, we met up, when Paul outlined the week and the hotel logistics, before it was time for our evening meal.

Day 2 Monday 22nd May

Font d’Urle – Col de la Machine – Route Combe Laval - St. Jean enRoyans We started the day with a short excursion to Vassieux en Vercors, where we made a short stop at the cemetery where the victims of the Nazi massacre at that village, during the Second World War, are buried. Whilst we were there we saw two Short-toed Snake Eagles, one adult and one juvenile, two Booted Eagles and about eight Griffon Vultures. Blackbird, Goldfinch and Skylark were also recorded.

Moving on, we made a second stop on the roadside of the Col de la Chau. Here we had views across to Le Grand Veymont in the east, the Vercors’ highest peak at 2,341 metres. Tau Emporer, Small Yellow Underwing, Burnet Companion and Speckled Yellow moths were seen, as well as the curious Ascalaphid, a relative of the Ant-lions. For the botanists there was Lactuca perennis (Blue Lettuce), Saponaria ocymoides (Rock Soapwort), Erinus alpinus (Fairy Foxglove) and Amelanchier ovalis (Juneberry).

France - The Vercors Tour Report

We now moved to the Font d’Urle, an area of open limestone crags and meadows used for skiing in winter. We began by exploring a bank beside the approach. Here we found the last few remains of Soldanella alpina (Alpine Snowbell, and the adjacent slope had Erythronium dens-canis (Dog-tooth Violet), Viola calcarata (Long-spurred Pansy), Gentiana verna (Spring Gentian) and Geum montanum (Alpine Avens). Moving on, a number of Alpine Choughs were circling above us, and we had distant views of Water Pipit, female Northern Wheatear, Linnet and Black Redstart. We also had several sightings of Alpine Marmot, and there was evidence of the winter tunnels of Snow Vole. As we progressed, we added Narcissus pseudonarcissus (Wild Daffodil), Narcissus poeticus (Pheasant’s-eye Daffodil), Draba aizoides (Yellow Whitlow-grass), Ranunculus amplexicaule (Amplexicaule Buttercup), both colour forms of Dactylorhiza sambucina (Elder-flowered Orchid), Orchis pallens (Pale-flowered Orchid), Crocus caeruleus (White Crocus) and Saxifraga moschata subsp. delphinensis (a Musky Saxifrage), before we returned to the vehicle.

Leaving the Font d’Urle, we made our way to the Forêt de Bauvante, where we set up our picnic among a fine stand of Orchis mascula (Early Purple Orchid). As we ate, Skylark was singing, and Des spotted a distant Roe Deer.

We now continued along the precipitous Route Combe Laval and on down to St. Jean en Royans, where we visited the hypermarket to do the week’s picnic shopping. Whilst our leader was shopping, we explored a small, rough field, sometimes use to graze horses. Here we found Baton Blue, Heath Fritillary, Large Copper, Large White, Small Heath and Common Blue butterflies, before setting off to retrace our steps towards the viewpoint at the Col de la Machine. We made a brief stop on a verge above St. Jean to admire some fine specimens of Orchis purpurea (Lady Orchid), Ophrys insectifera (Fly Orchid) and Cephalanthera longifolia (Sword-leaved Helleborine), before stopping at the Col to enjoy the magnificent scenery. Hornungia petraea (Hutchinsia), Lamium garganicum (Large Red Dead-nettle), Botrychium lunaria (Moonwort), Valeriana tripteris (Three-leaved Valerian) and Kernera saxatilis (Kernera) were all noted, before we returned to La Chapelle.

Day 3 Tuesday 23rd May

Col de Rousset – Above Les Nonnières – Vallon de Combeau – Cirque d’Archiane Our first stop today was at the head of the spectacular Col de Rousset – there are truly magnificent views from here. A few distant Griffon Vultures were seen, and Chiffchaff was calling. Anthyllis montana (Mountain Kidney- vetch) and Artemisia absinthium (Wormwood) were also recorded, together with Humming-bird Hawk-moth.

Moving on, we made our way to the bottom of the valley and turned east through the town of Die and on through Pont de Quart and Châtillon en Diois. We continued on through Menée and up to Les Nonnières, where we parked by some meadows. In some woodland we found Cypripedium calceolus (Lady’s Slipper Orchid), Cephalanthera longifolia (Sword-leaved Helleborine), Neottia ovata (Twayblade) and Equisetum telmateia (Great Horsetail). Great Tit and Common Chiffchaff were heard. An adjacent meadow produced Latticed Heath and Burnet Companion moths, Small Heath, Silver-studded Blue, Red-underwing Skipper, Grizzled Skipper and Wood White butterflies. We also found a Field Cricket in search of a mate.

Retracing our route, we turned up into the Vallon de Combeau. Here we made our way up the valley and parked by a ford in the mountain stream, where we set up the picnic. A few Griffon Vultures were seen soaring over a distant crag, and Berger’s Clouded Yellow, Dingy Skipper, Chequered Skipper and Scarce Swallowtail were seen

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France - The Vercors Tour Report

taking salts in the stream gravels. In a nearby ditch there was Pinguicula grandiflora (Large-flowered Butterwort) and close by, Euphorbia cyparissias (Cypress Spurge), Pinus uncinata (Mountain Pine), Meum athamanticum (Spignel) and Gentiana angustifolia (Narrow-leaved Gentian). We now moved down the valley and parked in a layby, in order to search a verge bank. Here we found Arctostaphylos uva-ursi (Bearberry), and Orchis spitzelii (Spitzel’s Orchid), which has a restricted distribution. The group now took a little time descending the valley on foot, enjoying the wildlife and the spectacular views, before rejoining the vehicle, in order that we could move south to the Cirque d’Archiane. We parked in a layby, south east of the hamlet of Archiane. Griffon Vultures were seen high on the ridges, as well as Aphyllanthes monspeliensis (Blue Aphyllanthes), Globularia cordifolia (Matted Globularia), Globularia punctata (Common Globularia) and Lavandula officinalis (Lavender). Little Blue, Wall Brown, Heath Fritillary, Scarce Swallowtail and Berger’s Clouded Yellow butterflies were also added.

We began our journey back to La Chapelle, stopping to have refreshments outside Die.

Day 4 Wednesday 24th May

Reserve Naturelle.des Hauts Plateaux du Vercors Soon after breakfast, we left La Chapelle and made our way down to St Agnan, and up to the Hauts Plateux, parking at the car park at La Coche, where we prepared for our morning walk. Along the way, we stopped to admire the views up the valley and to photograph Anemone alpina (Alpine Pasqueflower) and Ribes alpina (Mountain Currant). Leaving La Coche, we began exploring the mixed coniferous and deciduous woodland and found that Abies alba (Silver Fir) and Picea abies (Norway Spruce) and Fagus sylvatica (Beech) were the predominant trees. Soon after we had left the car park, we came across a small group of Crossbills, which we watched for a while, and Paul found Soldanella alpina (Alpine Snowbell) and Orchis mascula subsp. speciosa (an Early Purple Orchid). We soon added Common Firecrest, calling Dunnock, Coal Tit and a distant Song Thrush. Continuing on, we added Arctostaphylos uva-ursi (Bearberry) and Viola tricolor subsp. subalpina (Alpine Heartsease). As we were arriving at the open plateau, we had some very brief views of Alpine Marmot. As we progressed, a few Crocus caeruleus (White Crocus) were seen, as well as Daphne cneorum (Garland Flower), Tulipa sylvestris subsp. sylvestris (Wild Tulip) and Narcissus pseudonarcissus (Wild Daffodil). Before we set off on the return journey to La Coche, we saw Crested Tits with young, and a second group of Crossbills. Butterflies were well represented, with Grizzled Skipper, Orange Tip, Dingy Skipper, Swallowtail, Northern Wall Brown, Berger’s Clouded Yellow and Brimstone.

We set up our picnic in a shaded clearing near the car park, where Chiffchaff and Chaffinch were heard. After lunch, we rejoined our vehicle for the return journey on the Route Forestère des Charbonniers (Charcoal-makers route), around the back of the plateau, which afforded great views of the Grand Veymont. As we went we found more Anemone alpina (Alpine Pasqueflower).

Before rejoining the main road, we stopped to look at a verge bank near Les Meunières, which had a good number of Orchis anthropophora (Man Orchid) and Ophrys insectifera (Fly Orchid). Our final stop of the day was on a verge to the south-east of La Chapelle en Vercors. Here we saw Neotinea ustulata (Burnt Orchid), Ophrys fuciflora subsp. fuciflora (Late Spider Orchid), Orchis militaris (Military Orchid) and Orchis anthropophora (Man Orchid).

France - The Vercors Tour Report

Day 5 Thursday 25th May

Col de la Bataille – Col de Bacchus - Gigors Today we travelled down towards the Col de la Machine then on over the Col de la Bataille, where we stopped to enjoy the magnificent views in all directions. A few plants were added, including Centaurea montana (Perennial Cornflower), Rumex scutatus (Shield Dock) and Alyssum montanum (Mountain Alyssum). Continuing on towards the village of Leoncel, we turned left and headed south through La Vacharie to the Col de Bacchus. Here we made our first stop to explore a broad road verge and adjacent wood, finding the remnants of Anacamptis morio (Green-winged Orchid), Neotinea ustulata (Burnt Orchid), Neottia nidus-avis (Bird's-nest Orchid), Dactylorhiza sambucina (Elder-flowered Orchid) in its yellow colour form, Orchis militaris (Military Orchis), Silene italica (Italian Catchfly), and Cephalanthera damasonium (White Helleborine). We also recorded Escher’s Blue, Common Blue and Small Heath Butterflies.

We now continued on south through Pain de Baix towards Beaufort sur Gervanne. Along the way, we saw Stonechat and Red-backed Shrike. We paused on a wide verge, just north of Beaufort to photograph Platanthera bifolia (Lesser Butterfly Orchid), Ophrys apifera (Bee Orchid) and Orobanche gracilis (Slender Broomrape), before continuing south. Near Beaufort, we turned west to an area of open, scrubby land with marginal woodland, near Gigors, for our lunch stop. A brief exploration before lunch produced Cephalanthera rubra (Red Helleborine), Ophrys scolopax (Woodcock Orchid), a fine stand of Limodorum abortivum (Violet Bird’s-nest Orchid), Ophrys x flavicans (Drome Orchid), Ruscus aculeatus (Butcher’s-broom), Ajuga chamaepitys (Ground-pine) and Malva setigera (Hairy Mallow). Butterflies included Glanville Fritillary, Common Blue, Silver-studded Blue, Spotted Fritillary, Heath Fritillary, Scarce Swallowtail and Painted Lady. Western Bonelli’s Warbler was calling throughout our stay in the area, but could not be seen.

From here we moved back towards Leoncel, stopping at a second site on the Col de Bacchus. Here we added Astragalus monspesulanus (Montpellier Milk-vetch), Orchis simia (Monkey Orchid), Neotinea tridentata (Toothed Orchid), Neotinea ustulata (Burnt Orchid) and a good population of Ophrys x flavicans (Drome Orchid).

We made a stop in Leoncel for group members to enjoy refreshments, before we made our way to the hotel.

Day 6 Friday 26th May

Gorges de la Bourne - Les Merciers – Les Bruyeres Today we made our way up to the northern end of the Vercors Plateau. Our main objective was to try to find a Wallcreeper, but we found interesting things en route, as well. Leaving La Chapelle, we drove north to the villages of St Martin en Vercors, St. Julian en Vercors and into the spectacular Gorges de la Bourne, where we made our first stop. Along the way we saw Common Buzzard, Black Redstart and two Red Squirrels. Our target in the gorge was White-throated Dipper, but they were not feeding where we stopped. However, we did find Saxifraga rotundifolia (Round-leaved Saxifrage), Gymnocarpium robertianum (Limestone Fern), Valeriana tripteris (Three-leaved Valerian) and Kernera saxatilis (Kernera), and we also saw Crag Martin and House Martin. As we continued north through the gorge, we passed several stands of Lunaria redivida (Perennial Honesty).

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France - The Vercors Tour Report

We continued on to a point just north of Lans en Vercors, to the hamlet of St. Merciers. Parking, we set off to explore. Among the plants, we found Saxifraga paniculata (Live-long Saxifrage), Anthyllis montana (Mountain Kidney-vetch), Orchis mascula (Early Purple Orchid), Dactylorhiza sambucina (Elder-flowered Orchid), in the yellow colour form, Ranunculus aconitifolius (Aconite-leaved Buttercup) and the leaves of Gentiana cruciata (Cross Gentian). Insects were plentiful in the morning sunshine, with Red-underwing Skipper, Adonis Blue, Sooty Copper, Apollo, Queen of Spain Fritillary, Heath Fritillary and Berger’s Clouded Yellow, all recorded. We searched the cliff walls and crags, but were not able to find a Wallcreeper. However, we did see Black Redstart, Crag Martin, Jackdaw, Common Buzzard and Rock Bunting, a scarce bird in this area. Common Cuckoo, Common Chiffchaff and Robin were also heard. We now returned to our minibus in order to prepare our picnic.

After eating, we made our way to some wet meadows at Les Bruyeres, where we found Tephroseris helenitis (Swiss Ragwort), Valeriana dioica (Marsh Valerian), Trollius europaeus (Globeflower) and Dactylorhiza majalis (Broad-leaved Marsh Orchid). A small pond produced a good number of Rana kl. esculenta (hybrid Edible Frog), and we also added Azure Bluet and Large Red Damselflies and Broad-bodied Chaser dragonfly.

It was now time to start our homeward journey. We made two more stops in the Gorges de la Bourne, but were still unable to locate White-throated Dipper, so we made our way back to La Chapelle for dinner.

Day 7 Saturday 27th May

Canyons des Écourges – Forêt Dominiale des Coulmes – Pont-en-Royans We began once again today by driving north through St. Martin en Vercors and St. Julian en Vercors and into the Gorges de la Bourne, where we turned left towards Rencurel, and over the Col de Romeyere and down to the Canyon des Écourges, where we planned to spend some time looking for the Wallcreeper. As we went, we saw a solitary Common Buzzard. As we were preparing to leave the car park, Eurasian Blackcap and Eurasian Wren were heard and Grey Wagtail seen. We began by exploring the rock verges, finding Primula auricula (Bear’s-ear), Centaurea montana (Perennial Cornflower), Cardamine heptaphylla (Seven-leaflet Bittercress), Narcissus poeticus (Pheasant’s-eye Daffodil), Cardamine pentaphyllos (Five-leaved Bittercress) and Lamiastrum galeobdolon subsp. montanum (Yellow Archangel). Making our way to a vantage point, we spent time scanning the cliff walls for the Wallcreeper. Crag Martin, Common Wood Pigeon and Raven were seen, and also Wood White butterfly, but, unfortunately, no Wallcreeper. A wet area adjacent to the car park produced Chrysosplenium oppositifolium (Opposite-leaved Golden Saxifrage) and Chrysosplenium alternifolium (Alternate-leaved Golden Saxifrage).

Re-joining the vehicle, we made our way, via the Col de Romeyere, into the Forêt Comunuale des Coulmes. In the forest we took our picnic lunch at Petrouze. Local exploration produced Paris quadrifolia (Herb-Paris) and emerging Lilium martagon (Martagon Lily), and, in some adjacent clearings, Pearl-bordered Fritillary, Weaver’s Fritillary, Little Blue, Queen of Spain Fritillary, Wall Brown, Wood White and Mazarine Blue butterflies, and Mother Shipton, Burnet Companion, Grass Rivulet and Speckled Yellow moths.

It was now time to move down through the magnificent Rocher de Presles, a splendid limestone cliff-face overlooking the valley of the River Bourne. We parked a little further down the road, to admire the splendid views, before continuing down the Route de Presles, and made our way into Pont-en-Royans, where we parked. We walked along the river and up into the town, making for the café Restaurant Picard, famous for its ice-

France - The Vercors Tour Report

creams, where we all indulged unreservedly. In the gorge, Grey Wagtails were rather active, and a fleeting White- throated Dipper was glimpsed. Leaving Pont-en-Royans, we made our way back to the river bridge below Châtelus where our fortunes changed as we found an adult Dipper feeding two young.

It was now time to return to La Chapelle en Vercors, to pack before dinner.

Day 8 Sunday 28th May

La Chapelle en Vercors to Lyon St. Exupery Airport We left La Chapelle en Vercors at 7.15am, bidding farewell to Mme. Bellier and her daughters, before making our way via the Grand Goulets Tunnel and Pont-en-Royans and onto the A49, A48 and A43 motorways to Lyon St. Exupery Airport. En route we had plenty of time to admire the scenery and spot some birds from the vehicle including Black Kite, Grey Heron and Common Kestrel. At Heathrow, we said our farewells, and travelled homewards, after a wonderful week in the Vercors.

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Tulipa sylvestris subsp. sylvestris

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France - The Vercors Tour Report

Species Lists

Plants (* = Endemic or restricted distribution)

Nomenclature contained within this list follows the taxonomic amendments, based on DNA analysis, made by the Royal Botanic Gardens, Kew and the Missouri Botanic Garden, and following The Plant List, an internationally accepted list of vascular plant families, published and maintained by these two Institutions.

Scientific name Common name Location

PTERIDOPHYTES FERNS & ALLIES

Calamophytes Horsetails

Equisetaceae Horsetail Family Equisetum arvense Common Horsetail La Chapelle en Vercors Equisetum hyemale Dutch Rush Canyons des Ecourges Equisetum palustre Marsh Horsetail Canyons des Ecourges Equisetum telmateia Great Horsetail Above Les Nonnieres

Eusporangiate Ferns Adder's-tongues & Moonworts

Ophioglossaceae Adder's-tongue family Botrychium lunaria Moonwort Col de la Machine

Leptosporangiate Ferns True Ferns

Aspleniaceae Spleenwort Family Asplenium fontanum Smooth Rock Spleenwort Col de la Machine Asplenium ruta–muraria Wall Rue Font d'Urle Asplenium scolopendrium Hart’s-tongue Pont-en-Royans Asplenium trichomanes subsp. quadrivalens Maidenhair Spleenwort Font d'Urle Asplenium viride Green Spleenwort Hauts Plateaux

Cystopteridaceae Bladder-fern Family Cystopteris fragilis Brittle Bladder-fern Hauts Plateaux Gymnocarpium robertianum Limestone Fern Gorges de la Bourne

Dennstaedtiaceae Bracken Family Pteridium aquilinum Bracken Foret des Coulmes

Dryopteridaceae Buckler-fern Family Dryopteris dilitata Broad Buckler-fern Foret des Coulmes Dryopteris filix-mas Male-fern Les Merciers Polystichum aculeatum Hard Shield-fern Hauts Plateaux Polystichum lonchitis Holly Fern Font d'Urle

GYMNOSPERMS CONIFERS

Cupressaceae Juniper Family Juniperus communis Juniper Font d'Urle Juniperus thurifera French Alpine Juniper Gorges de la Bourne

Pinaceae Pine Family

France - The Vercors Tour Report

Scientific name Common name Location

Abies alba European Silver Fir Foret de Beauvente Picea abies subsp. abies Spruce Foret de Beauvente Pinus nigra subsp. nigra Austrian or Black Pine Hauts Plateaux Pinus sylvestris Scots Pine Valle de Combeau Pinus uncinata Mountain Pine Valle de Combeau

ANGIOSPERMS FLOWERING PLANTS

Pre-Dicots Primitive Angiosperms

Aristolochiaceae Birthwort Family Asarum europaeum Asarabacca Near Toutre

Eu-Dicots True Dicotyledons

Adoxaceae Moschatel Family Sambucus ebulus Dwarf Elder Les Merciers (leaves) Sambucus nigra Elder Roadsides Sambucus racemosus Red-berried Elder Canyons des Ecourges

Amaranthaceae Pigweed Family Chenopodium bonus-henricus Good King Henry Font d'Urle

Anacardiaceae Sumach Family Cotinus coggygria Smoke Bush Route Presles

Apiaceae Carrot Family Aegopodium podagraria Ground Elder Les Merciers Angelica sylvestris Wild Angelica Gorges de la Bourne Anthriscus sylvestris Cow Parsley Col de Bacchus Eryngium c.f. campestre Field Eryngo Gigors (leaves) Heracleum sphondylium Hogweed Font d'Urle Laserpitium siler a Sermountain Route Presles Meum athamanticum Spignel Valle de Combeau Orlaya grandiflora Orlaya Gigors Peucedanum carvifolia - Route Presles Pimpinella major Greater Burnet Saxifrage Canyons des Ecourges Pimpinella saxifraga Burnet Saxifrage Les Merciers Sanicula europaea Sanicle Above Les Nonnieres

Apocynaceae Periwinkle Family Vincetoxicum hirundinaria Swallow-wort Les Meuniers

Aquifoliaceae Holly Family Ilex aquifolium Holly Les Merciers

Araliaceae Ivy Family Hedera helix agg. Ivy Pont-en-Royans

Betulaceae Birch Family Corylus avellana Hazel Above Les Nonnieres Boraginaceae Forget-me-not Family

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France - The Vercors Tour Report

Scientific name Common name Location

Cerinthe minor Lesser Honeywort Valle de Combeau Echium vulgare Viper’s-bugloss Above St. Jean en Royans Myosotis alpestris Alpine Wood Forget-me-not Font d'Urle Myosotis arvensis Field Forget-me-not Les Merciers Myosotis ramosissima Early Forget-me-not Near Toutre Myosotis sylvatica Wood Forget-me-not Hauts Plataeu Pulmonaria montana Mountain Lungwort Font d'Urle

Brassicaceae Cabbage Family Alliaria petiolata Garlic Mustard Gigors Alyssum montanum Mountain Alyssum Font d'Urle Arabis alpina Alpine Rock-cress Font d'Urle Arabis hirsuta Hairy Rock-cress Les Merciers Arabis turrita Tower-cress Les Merciers Barbarea vulgaris Common Winter-cress Font d'Urle Bunias orientalis Warty Cabbage La Chapelle en Vercors Capsella bursa-pastoris Shepherd’s-purse La Chapelle en Vercors Capsella rubella - Les Merciers Cardamine heptaphylla Seven-leaflet Bitter-cress Foret de Beauvente Cardamine impatiens Narrow-leaved Bitter-cress Gorges de la Bourne Cardamine pentaphyllos Five-leaved Bitter-cress Hauts Plataeu Draba aizoides Yellow Whitlow-grass Font d'Urle Erophila verna Common Whitlow-grass Col de la Bataille Erysimum ochroleucum Decumbent Treacle-mustard Font d'Urle Hesperis matronalis Dame's Violet Near St. Martin en Vercors Hornungia petraea Hutchinsia Col de la Machine Kernera saxatilis Kernera Col de la Machine Lepidium campestre Field Pepperwort Cirque d'Archiane Lunaria redivida Perennial Honesty Gorges de la Bourne Microthlaspi perfoliata Perfoliate Penny-cress Near Toutre Sinapis arvensis Charlock Gigors Thlaspi montanum Mountain Penny-cress Font d'Urle Turritis (Arabis) glabra Tower Mustard Gigors

Buxaceae Box Family Buxus sempervirens Box Above St. Jean en Royans

Campanulaceae Bellflower Family Campanula cochlearifolia Fairy's Thimbles Gorges de la Bourne (leaves) Campanula rhomboidalis a Bellflower Above St. Jean en Royans Phyteuma orbiculare Round-headed Rampion Above Les Nonnieres

Caprifoliaceae Honeysuckle Family Centranthus ruber Red Valerian Col de Bacchus Dipsacus fullonum Teasel Cirque d'Archiane Knautia arvensis Field Scabious Hauts Plateaux Knautia dipsacifolia Wood Scabious Above St. Jean en Royans Lonicera alpigena Alpine Honeysuckle Hauts Plateaux Lonicera xylosteum Fly Honeysuckle Above Les Nonieres Valeriana dioica Marsh Valerian Les Bruyeres Valeriana montana Mountain Valerian Hauts Plateaux Valeriana tripteris Three-leaved Valerian Col de la Machine

France - The Vercors Tour Report

Scientific name Common name Location

Valerianella eriocarpa - Above Les Nonnieres Valerianella locusta Common Cornsalad Near Toutre

Caryophyllaceae Pink Family Arenaria serpyllifolia subsp. serpyllifolia Thyme-leaved Sandwort Les Merciers Cerastium arvense Field Mouse-ear Col de la Chau Cerastium tomentosum Snow-in-Summer Col de Rousset Dianthus sylvaticus Wood Pink Near Menee Minuartia verna Spring Sandwort Font d'Urle Moehringia muscosa - Canyons des Ecourge Saponaria ocymoides Rock Soapwort Col de la Chau Silene dioica Red Campion Les Merciers Silene (Lychnis) flos-cuculi Ragged Robin Les Bruyeres Silene italica Italian Catchfly Cirque d'Archiane Silene latifolia White Campion La Chapelle en Vercors Silene vulgaris Bladder Campion Above St. Jean en Royans Stellaria holostea Greater Stitchwort Canyons des Ecourges Stellaria media Chickweed La Chapelle en Vercors

Cistaceae Rock-rose Family Helianthemum apenninum White Rock-rose Gigors Helianthemum nummularium Common Rock-rose Above Les Nonnieres H.oelandicum subsp. incanum Hoary Rock-rose Gigors

Compositae (Asteraceae) Daisy Family Achillea millefolium Yarrow Les Merciers Bellidiastrum michelii (Aster bellidiastrum) False Aster Gourges de la Bourne Bellis perennis Daisy La Chapelle en Vercors Buphthalmum salicifolium Yellow Ox-eye Col de Bacchus Centaurea montana Perennial Cornflower Col de la Bataille Cirsium palustre Marsh Thistle Canyons des Ecourges Homogyne alpina Alpine Colts-foot Hauts Plateaux (leaves) Lactuca perennis Blue or Mountain Lettuce Col de la Chau Leucanthemum vulgare Ox-eye Daisy Col de la Chau Mycelis muralis Wall Lettuce Pont-en-Royans Petasites albus White Butterbur Above Les Nonnieres Senecio vulgaris Groundsel La Chapelle en Vercors Taraxacum agg. Dandelion La Chapelle en Vercors Tephroseris helenitis Swiss Ragwort Les Bruyeres Tragopogon dubius a Goat's-beard Col de la Chau Tussilago farfara Colt’s-foot Font d'Urle

Convolvulaceae Bindweed Family Cuscuta europaeum Greater Dodder Les Merciers

Cornaceae Dogwood Family Cornus sanguinea Dogwood Cirque d'Archiane

Crassulaceae Stonecrop Family Sedum album White Stonecrop Gorges de la Bourne (leaves) Sedum dasyphyllum Fat Stonecrop Route Presles Sedum forsterianum Rock Stonecrop Hauts Plateaux (leaves)

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France - The Vercors Tour Report

Scientific name Common name Location

Sedum sexangulare Tasteless Stonecrop Hauts Plateaux (leaves) Sempervivum tectorum Common Houseleek Hauts Plateaux

Cucurbitaceae Cucumber Family Bryonia dioica White Bryony Canyons des Ecourge

Ericaceae Heath family Arctostaphylos uva-ursi Bearberry Valle de Combeau Orthilia secunda Nodding Wintergreen Hauts Plateaux (leaves) Pyrola minor Common Wintergreen Hauts Plateaux Vaccinium myrtillus Bilberry Font d'Urle Vaccinium vitis-idaea Cowberry Font d'Urle

Euphorbiaceae Spurge Family Euphorbia amygdaloides Wood Spurge Above St. Jean en Royans Euphorbia cyparissias Cypress Spurge Above Les Nonnieres Euphorbia dulcis Sweet Spurge Near Toutre Euphorbia helioscopia Sun Spurge Cirque d'Archiane Euphorbia verrucosa - Hauts Plateaux Mercurialis perennis Dog’s Mercury Hauts Plateaux

Fagaceae Oak & Beech Family Fagus sylvatica Beech Foret de Beauvente Quercus pubescens Downy Oak Gigors

Gentianaceae Gentian Family Gentiana angustifolia Narrow-leaved Gentian Valle de Combeau Gentiana clusii Clusius’ Gentian Font d'Urle Gentiana cruciata Cross Gentian Les Merciers Gentiana lutea Yellow Gentian Les Merciers (leaves) Gentiana verna Spring Gentian Font d'Urle

Geraniaceae Geranium Family Geranium columbinum Long-stalked Crane’s-bill Foret des Coulmes Geranium molle Dove’s-foot Crane’s-bill Les Merciers Geranium nodosum Knotted Crane's-bill Hauts Plateaux (leaves) Geranium pyrenaicum Hedgerow Crane’s-bill Col de la Chau Geranium robertianum Herb Robert St. Jean en Royans Geranium sylvaticum Wood Crane’s-bill Hauts Plateaux

Grossulariaceae Gooseberry Family Ribes alpinum Mountain Currant Hauts Plateaux Ribes uva-crispa Wild Gooseberry Foret de Lente

Juglandaceae Walnut Family [Juglans regia] Walnut Planted

Lamiaceae Dead-nettle Family Ajuga chamaepitys Ground-pine Gigors Ajuga genevensis Blue Bugle Above St. Jean en Royans Ajuga reptans Bugle Les Merciers Clinopodium alpinum Alpine Basil-thyme Gorges de la Bourne

France - The Vercors Tour Report

Scientific name Common name Location

Glechoma hederacea Ground Ivy Canyons des Ecourges Lamiastrum galeobdolon subsp. montanum Yellow Archangel Gorges de la Bourne Lamium garganicum Large Red Dead-nettle Col de la Machine Lavandula officinalis Lavender Cirque d'Archiane Melittis melissophyllum Bastard Balm Above St. Jean en Royans Origanum vulgare Wild Marjoram Route Presles Salvia pratensis Meadow Clary Roadsides Stachys recta Yellow Woundwort Gigors Stachys sylvatica Hedge Woundwort Canyons des Ecourges Teucrium chamaedrys Wall Germander Route Presles (leaves) Teucrium polium Felty Germander Gigors (leaves) Thymus polytrichus Wild Thyme Above Les Nonnieres Thymus serpyllum A Thyme Rocher de Presles

Lentibulariaceae Bladderwort Family Pinguicula grandiflora Large-flowered Butterwort Valle de Combeau

Legouminosae (Fabaceae) Pea Family Anthyllis montana Mountain Kidney-vetch Col de Rousset Anthyllis vulneraria agg. Kidney-vetch Above Les Nonnieres Astragalus glycyphyllos Wild Liquorice Gigors (leaves) Astragalus monspessulanus Montpellier Milk-vetch Col de Bacchus Bituminaria bituminosa Pitch Trefoil Gigors (leaves) Coronilla scorpoides Annual Scorpion-vetch Gigors (leaves) Coronilla vaginalis A Scorpion-vetch Route Presles Dorycnium pentaphyllum - Gigors Genista sericea a Greenweed Font d'Urle Hippocrepis comosa Horseshoe Vetch Above St. Jean en Royans Hippocrepis emerus False Senna Foret de Beauvente Laburnum alpinum Alpine Laburnum Below the Col de Rousset Laburnum anagyroides Laburnum Below the Col de Rousset Lathyrus cicera - Gigors Lathyrus latifolius Broad-leaved Everlasting Pea Route Presles Lathyrus pratensis Yellow Vetchling Les Merciers Lathyrus setifolius - Gigors Lathyrus vernus Spring Vetchling Col de Bacchus Lotus alpinus Alpine Bird’s-foot Trefoil Col de la Bataille Lotus edulis Edible Lotus Gigors Lotus (Tetragonolobus) maritimus Dragon’s-teeth Below the Grand Goulets Medicago lupulina Black Medick Above Les Nonnieres Onobrychis arenaria a Sainfoin Above Les Nonnieres Ononis fruticosa Shrubby Restharrow Below Chamaloc [Robinia pseudoacacia] False Acacia Roadsides Trifolium campestris Hop Trefoil Gigors Trifolium medium Zigzag Clover Gigors Trifolium montanum Mountain Clover Col de Bacchus Trifolium pratense Red Clover Valle de Combeau Trifolium repens White Clover Font d'Urle Vicia cracca Tufted vetch Gigors Vicia sativa subsp. sativa Cultivated Vetch Grenoble Isere Airport Vicia sepium Bush Vetch Canyons des Ecourges Linaceae Flax Family

12 © Naturetrek October 17

France - The Vercors Tour Report

Scientific name Common name Location

Linum catharticum Purging or Fairy Flax Gigors Linum perenne agg. Perennial Flax Font d'Urle

Malvaceae Mallow Family Malva setigera (Althaea hirsuta) Hairy Mallow Gigors Tilia cordata Small-leaved Lime Pont-en-Royans Moraceae Mulberry Family Ficus carica Fig Pont-en-Royans

Oleaceae Olive Family Fraxinus excelsior Ash Valle de Combeau Ligustrum vulgare Wild Privet Gigors

Onagraceae Willowherb Family Epilobium (Chamerion) angustifolium Rosebay Willowherb Canyons des Ecourges (leaves)

Orobanchaceae Broomrape Family Orobanche gracilis Slender Broomrape Above Beaufort sur Gervanne Pedicularis gyroflexa Tufted Lousewort Font d'Urle (leaves) Rhinanthus aristatus agg. Greater Yellow-rattle Cirque d'Archiane Rhinanthus minor agg. Yellow-rattle Hauts Plateaux

Oxalidaceae Wood-sorrel Family Oxalis acetosella Wood-sorrel Hauts Plateaux

Papaveraceae Poppy Family Chelidonium majus Greater Celandine LA Chapelle en Vercors Fumaria officinalis agg. Common Fumitory Cirque d'Archiane Papaver rhoeas Corn Poppy Grenoble Isere Airport

Plantaginaceae Plantain Family Antirrhinum latifolium Broad-leaved Snapdragon Cirque d'Archiane Cymbalaria muralis Ivy-leaved Toadflax LA Chapelle en Vercors Erinus alpinus Fairy Foxglove Col de la Chau Globularia cordifolia Matted Globularia Cirque d'Archiane Globularia nudicaulis Leafless-stemmed Globularia Hauts Plateaux Globularia punctata Common Globularia Cirque d'Archiane Plantago lanceolata Ribwort Plantain Font d'Urle Plantago major Greater Plantain Canyons des Ecourges Plantago media Hoary or Lamb’s-tail Plantain Above Les Nonnieres Plantago sempervirens - Font d'Urle Veronica austriaca agg. Large Speedwell Les Merciers Veronica beccabunga Brooklime Canyons des Ecourges Veronica chamaedrys Germander Speedwell Font d'Urle Veronica urticifolia Nettle-leaved Speedwell Canyons des Ecourges (leaves)

Plumbaginaceae Thrift Family Armeria alpina Mountain Thrift Fonte d'Urle

Polygalaceae Milkwort Family Polygala alpestris Mountain Milkwort Fonte d'Urle Polygonaceae Knotweed Family

France - The Vercors Tour Report

Scientific name Common name Location

Fallopia convolvulus Black Bindweed Persicaria bistorta Bistort Near Pont-en-Royans Rumex acetosa Sorrel Font d'Urle Rumex obtusifolius Broad-leaved Dock Canyons des Ecourges Rumex pulcher Fiddle Dock Aire Burcin Rumex scutatus French Sorrel Col de la Bataille

Primulaceae Primrose Family Androsace villosa Woolly Rock-jasmine Font d'Urle Lysimachia nummularia Creeping Jenny Les Bruyeres (leaves) Primula auricula Bear’s-ear Primrose Canyons des Ecourges Primula veris Cowslip Font d'Urle Soldanella alpina Alpine Snowbell Font d'Urle

Ranunculaceae Buttercup Family Aconitum napellus Common Monkshood Canyons des Ecourges (leaves) Anemone (Pulsatilla) alpina Alpine Pasqueflower Hauts Plateaux Anemone ranunculoides Yellow Anemone Font d'Urle Aquilegia atrata Dark-flowered Columbine N of Les Barraques Aquilegia vulgaris Columbine Above St. Jean en Royans Caltha palustris Kingcups Valle de Combeau Helleborus foetidus Stinking Hellebore Col de la Machine Ranunculus aconitifolius Aconite-leaved Buttercup Les Merciers Ranunculus acris Meadow Buttercup Above Les Nonnieres Ranunculus amplexicaulis Amplexicaule Buttercup Font d'Urle Ranunculus bulbosus Bulbous Buttercup Near Toutre Ranunculus kuepferi Kuepfer's Buttercup Font d'Urle Ranunculus montanus Mountain Buttercup Font d'Urle Ranunculus repens Creeping Buttercup Foret des Coulmes Ranunculus trichophyllus Thread-leaved Water-crowfoot Les Bruyeres Thalictrum minus Lesser Meadow Rue Les Merciers (leaves) Trollius europaeus Globeflower Font d'Urle

Rhamnaceae Buckthorn Family Rhamnus alpinus Alpine Buckthorn Above St. Jean en Royans

Rosaceae Rose Family Alchemilla alpina Alpine Lady’s-mantle Font d'Urle Alchemilla conjuncta - Font d'Urle Alchemilla glaucescens Silky Lady's-mantle Font d'Urle Alchemilla glabra Smooth Lady’s-mantle Font d'Urle Amelanchier ovalis Juneberry Col de la Chau Cotoneaster tomentosus Cotoneaster Hauts Plateaux Crataegus monogyna Hawthorn Col de la Chau Filipendula ulmaria Meadowsweet Les Bruyeres Fragaria moschata Hautbois Strawberry Above St. Jean en Royans Geum montanum Alpine Avens Font d'Urle Geum rivale Water Avens Les Bruyeres Geum urbanum Wood Avens or Herb Bennet Foret des Coulmes Potentilla crantzii Alpine Cinquefoil Font d'Urle Potentilla rupestris Rock Cinquefoil Hauts Plateau Prunus avium Wild Cherry Col de Bacchus

France - The Vercors Tour Report

Scientific name Common name Location

Prunus mahaleb St. Lucie's Cherry Above Les Nonnieres Rosa canina agg. Dog Rose Vallee de Combeau Rosa spinosissima (R. pimpinellifolia) Burnet Rose Les Merciers Rubus caesius Dewberry Vallee de Combeau Rubus fruticosus agg. Bramble St. Jean en Royans Rubus idaeus Raspberry Col de la Bataille Rubus saxatile Stone Bramble Hauts Plateaux Sanguisorba minor (Poterium s.) agg. Salad Burnet Hauts Plateaux Sorbus aria agg. Whitebeam Col de la Chau Sorbus aucuparia Rowan Col de la Chau

Rubiaceae Bedstraw Family Cruciata laevipes Crosswort Grenoble Isere Airport Galium mollugo Hedge Bedstraw Grenoble Isere Airport Galium odoratum Sweet Woodruff Above Les Nonnieres Rubia peregrina Wild Madder Gigors Sherardia arvensis Field Madder Les Merciers

Salicaceae Willow & Poplar Family Populus tremula Aspen Les Merciers Salix caprea Goat Willow Col de la Chau Salix triandra Almond Willow Valle de Combeau Salix viminalis Basket Willow Valle de Combeau

Santalaceae Sandalwood Family Thesium alpinum Alpine Bastard Toadflax Canyons des Ecourges

Sapindaceae Maple Family Acer campestre Field Maple La Chapelle en Vercors Acer opalus Italian Maple Above St. Jean en Royans Acer pseudoplatanus Sycamore Hauts Plateaux

Saxifragaceae Saxifrage Family Chrysosplenium alternifolium Alternate-leaved Golden Saxifrage Canyons des Ecourges Chrysosplenium oppositifolium Opposite-leaved Golden Saxifrage Canyons des Ecourges Saxifraga exarata subsp. delphinensis Musky Saxifrage Font d'Urle Saxifraga granulata Meadow Saxifrage Near Toutre Saxifraga oppositifolia Purple Saxifrage Font d'Urle (leaves) Saxifraga paniculata Live-long Saxifrage Col de la Machine Saxifraga rotundifolia Round-leaved saxifrage Gorges de la Bourne Saxifraga tridactylites Rue-leaved Saxifrage Col de la Machine

Scrophulariaceae Figwort Family Buddleja davidii Butterfly-bush Pont-en-Royans Scrophularia canina subsp. hoppii Alpine Figwort Cirque d'Archiane Scrophularia nodosa Common Figwort Canyons des Ecourge

Thymelaeaceae Mezereon Family Daphne cneorum Garland Flower Hauts Plateaux Daphne laureola Spurge Laurel Col dde Bacchus Daphne mezereum Mezereon Font d'Urle Ulmaceae Elm Family

France - The Vercors Tour Report

Scientific name Common name Location

Ulmus glabra Wych Elm Gorges de la Bourne

Urticacae Nettle Family Parietaria judaica Pellitory-of-the-Wall Pont-en-Royans Urtica dioica Common Nettle St. Jean en Royans

Violaceae Violet Family Viola arvensis Field Pansy Foret des Coulmes Viola calcarata Long-spurred Pansy Font d'Urle Viola riviniana Common Dog Violet Hauts Plateaux Viola tricolor subsp. subalpina Alpine Heartsease Font d'Urle

Viscaceae Mistletoe Family Viscum album subsp. album Mistletoe Vallee de Combeau

Monocots Monocotyledons

Amaryllidaceae Daffodil Family Narcissus poeticus Pheasant’s-eye Daffodil Font d'Urle Narcissus pseudonarcissus Wild Daffodil Font d'Urle

Araceae Arum Family Arum maculatum Lords-and-Ladies Foret des Coulmes

Asparagaceae Asparagus Family Anthericum liliago St. Bernard’s Lily Gigors Aphyllanthes monspeliensis Blue Aphyllanthes Cirque d'Archiane Convallaria majalis Lily-of-the -Valley Hauts Plateaux Leopoldia (Muscari) comosa Tassel Hyacinth Grenoble Isere Airport Muscari neglectum Grape Hyacinth Col de Bacchus Ornithogalum angustifolium Common Star of Bethlehem Above Les Nonnieres Ornithogalum pyrenaicum Bath Asparagus Route Presles Polygonatum odoratum Angular Solomon’s-seal Above St. Jean en Royans Polygonatum verticillatum Whorled Solomon’s-seal Hauts Plateaux Ruscus aculeatus Butcher’s-broom Gigors

Colchicaceae Meadow Saffron Family Colchicum autumnale Autumn Crocus Font d'Urle (in fruit)

Cyperaceae Sedge Family Carex caryophyllea Spring Sedge Font d'Urle Carex disticha Brown Sedge Les Bruyeres Carex flacca Glaucous Sedge Above Les Nonnieres Carex ornithopoda Bird's-foot Sedge Hauts Plateaux

Iridaceae Iris Family Crocus caeruleus (C.vernus subsp. albiflorus) White Crocus Font d'Urle Gladiolus italicus Field Gladiolus Above Beafort sur Gervanne

Juncaceae Rush Family Juncus inflexus Hard Rush Les Bruyeres Luzula luzuloides White Woodrush Hauts Plateaux

France - The Vercors Tour Report

Scientific name Common name Location

Luzula nivea Snowy Woodrush Aire Burcin

Liliaceae Lily Family Erythronium dens-canis Dog’s-tooth Violet Font d'Urle Gagea liotardii (G. fragifera, G. fistulosa) a Yellow Star of Bethlehem Font d'Urle Lilium martagon Martagon Lily Below Col de la Bataille Tulipa sylvestris subsp. sylvestris Wild Tulip Font d'Urle

Melanthiaceae Herb-Paris Family Paris quadrifolia Herb-Paris Foret des Coulmes Veratrum album White False Helleborine Les Merciers (leaves)

Orchidaceae Orchid Family Anacamptis morio Green-winged Orchid Col de Bacchus Anacamptis pyramidalis Pyramidal Orchid Gigors Cephalanthera damasonium White Helleborine Col de Bacchus Cephalanthera longifolia Sword-leaved Helleborine Near Toutre Cephalanthera rubra Red Helleborine Gigors Coeloglossum viride Frog Orchid Hauts Plateaux Cypripedium calceolus Lady’s Slipper Orchid Above Les Nonnieres Dactylorhiza alpestris Alpine Broad-leaved Marsh Orchid Above Les Nonnieres Dactylorhiza fuchsii Common Spotted Orchid Below Col de la Bataille Dactylorhiza majalis Broad-leaved Marsh Orchid Les Bruyeres Dactylorhiza sambucina (Pink) Elder-flowered Orchid Font d'Urle Dactylorhiza sambucina (Yellow) Elder-flowered Orchid Font d'Urle Himantoglossum hircinum Lizard Orchid Gigors (leaves) Limodorum abortivum Violet Bird's-nest Orchid Gigors Neottia nidus-avis Bird's-nest Orchid Above Les Nonnieres Neottia ovata Twayblade Near Toutre Neotinea tridentata Toothed Orchid Col de Bacchus Neotinea tridentata x Neotinea ustulata a Hybrid Orchis Col de Bacchus Neotinea ustulata Burnt Orchid Near La Chapelle en Vercors Ophrys apifera Bee Orchid Above Beaufort sur Gervanne Ophrys x flavicans (O. drumana)* Drome Bee Orchid Gigors Ophrys fuciflora subsp. fuciflora (O. linearis) Late Spider Orchid Near Toutre Ophrys fuciflora x (Ophrys x flavicans)* Hybrid Ophrys Gigors Ophrys insectifera Fly Orchid Near Toutre Ophrys insectifera x (Ophrys x flavicans)* a Hybrid Ophrys Gigors Ophrys scolopax Woodcock Orchid Gigors Orchis anthropophora Man Orchid Near Toutre Orchis mascula Early Purple Orchid Near Toutre Orchis mascula subsp. speciosa (O. ovalis) an Early Purple Orchid Hauts Plateaux Orchis militaris Military Orchid Near Toutre Orchis pallens Pale-flowered Orchid Font d'Urle Orchis purpurea Lady Orchid Above St. Jean en Royans Orchis simia Monkey Orchid Near Toutre Orchis spitzelii Spitzel’s Orchid Valle de Combeau Platanthera bifolia Lesser Butterfly Orchid Avove Beaufort sur Gervanne

Poaceae Grass Family

France - The Vercors Tour Report

Scientific name Common name Location

Alopecurus aequalis Orange Foxtail Les Bruyeres Alopecurus pratensis Meadow Foxtail Les Bruyeres Anthoxanthum odoratum Sweet Vernal Grass Les Bruyeres Briza media Quaking Grass Gigors Poa bulbosa var. vivipara Bulbous Meadow-grass Col de Bacchus Poa nemoralis Wood Meadow-grass Gigors Sesleria caerulea Blue Moor-grass Font d'Urle

LICHENS: Lobaria pulmonaria Tree Lungwort Hauts Plateaux Peltigera sp. Hauts Plateaux Pseudevernia furfuracea Valle de Combeau

FUNGI: Morchella esculenta Common Morel Canyons des Ecourges Pycnoporus cinnabarinus an Orange Bracket Hauts Plateaux

Birds (=recorded but not counted; H = heard only) May Common name Scientific name 21 22 23 24 25 26 27 28 1 Mute Swan Cygnus olor   2 White Stork Ciconia ciconia  3 Grey Heron Ardea cinerea  4 Griffon Vulture Gyps fulvus   5 Short-toed Snake Eagle Circaetus gallicus   6 Booted Eagle Hieraaetus pennatus    7 Black Kite Milvus migrans    8 Common Buzzard Buteo buteo     9 Eurasian Curlew Numenius arquata  10 Rock Dove (Feral) Columba livia        11 Common Wood Pigeon Columba palumbus      12 Eurasian Collared Dove Streptopelia decaocto    13 Common Cuckoo Cuculus canorus H H H H H 14 Eurasian Scops Owl Otus scops H H 15 Tawny Owl Strix aluco H 16 Alpine Swift Tachymarptis melba  17 Common Swift Apus apus        18 Great-spotted Woodpecker Dendrocopos major  H  19 European Green Woodpecker Picus viridis H H H 20 Common Kestrel Falco tinnunculus    21 Red-backed Shrike Lanius collurio   22 Eurasian Jay Garrulus glandarius        23 Eurasian Magpie Pica pica        24 Alpine Chough Pyrrhocorax graculus    25 Western Jackdaw Coloeus monedula   26 Rook Corvus frugilegus  27 Carrion Crow Corvus corone        28 Northern Raven Corvus corax    29 Coal Tit Periparus ater       30 European Crested Tit Lophophanes cristatus  

France - The Vercors Tour Report

May Common name Scientific name 21 22 23 24 25 26 27 28 31 Marsh Tit Poecile palustris H  32 Willow Tit Poecile montanus 33 Eurasian Blue Tit Cyanistes caeruleus      34 Great Tit Parus major  H      35 Eurasian Skylark Alauda arvensis   36 Barn Swallow Hirundo rustica       37 Eurasian Crag Martin Ptyonoprogne rupestris   38 Common House Martin Delichon urbicum  H      39 Long-tailed Tit Aegithalos caudatus H  40 Common Chiffchaff Phylloscopus collybita H H H H H H H 41 Western Bonelli’s Warbler Phylloscopus bonelli H 42 Eurasian Blackcap Sylvia atricapilla H H H   H  43 Common Firecrest Regulus ignicapilla H  H 44 Eurasian Wren Troglodytes troglodytes H H H H H  45 Common Starling Sturnus vulgaris      46 Common Blackbird Turdus merula        47 Song Thrush Turdus philomelos H H H 48 Mistle Thrush Turdus viscivorus     49 Spotted Flycatcher Muscicapa striata   50 European Robin Erithacus rubecula  H  H  H 51 Common Nightingale Luscinia megarhynchos H 52 Black Redstart Phoenicurus ochruros        53 Common Redstart Phoenicurus phoenicurus      54 European Stonechat Saxicola rubicola   55 Northern Wheatear Oenanthe oenanthe   56 White-throated Dipper Cinclus cinclus   57 House Sparrow Passer domesticus        58 Dunnock Prunella modularis H 59 Grey Wagtail Motacilla cinerea    60 White Wagtail Motacilla alba         61 Water Pipit Anthus spinoletta  62 Common Chaffinch Fringilla coelebs        63 Eurasian Bullfinch Pyrrhula pyrrhula    H 64 European Greenfinch Chloris chloris H  H H H 65 Common Linnet Linaria cannabina     66 Red Crossbill Loxia curvirostra  67 European Goldfinch Carduelis carduelis       68 European Serin Serinus serinus        69 Yellowhammer Emberiza citrinella H H 70 Rock Bunting Emberiza cia  71 Cirl Bunting Emberiza cirlus  H

Mammals (D=Dead; S=Signs of) 1 Common Shrew Sorex araneus D 2 European Mole Talpa europaea S 3 Daubenton's bat Myotis daubentonii  4 Greater Mouse-eared Bat Myotis myotis  5 Common Pipistrelle Bat Pipistrellus pipistrellus  6 Pine Marten Martes martes S 7 Wild Boar Sus scofa S S S S

France - The Vercors Tour Report

May Common name Scientific name 21 22 23 24 25 26 27 28 8 European Roe Deer Capreolus capreolus  

9 Alpine Ibex Capra ibex  10 Red Squirrel Sciurus vulgaris    11 Alpine Marmot Marmota marmota   12 Brown Rat Rattus norvegicus D 13 Snow Vole Chionomys nivalis S 14 European Hare Lepus europaeus   15 Rabbit Oryctolagus cuniculus S S

Amphibians & Reptiles 1 Edible-Pool Frog hybrid Pelophylax kl. esculentus  2 Western Green Lizard Lacerta bilineata   3 Common Wall Lizard Podarcis muralis     

Fish 1 Brown Trout Salmo trutta 

Butterflies (LO=Leader only) 1 Swallowtail Papilio machaon   2 Scarce Swallowtail Iphiclides podalirius      3 Apollo Parnassius apollo  5 Black-veined White Aporia crataegi    6 Large White Pieris brassicae     7 Small White Artogeia rapae       8 Orange Tip Anthocharis cardamines        9 Clouded Yellow Colias crocea   10 Berger’s Clouded Yellow Colias alfacariensis     11 Brimstone Gonepteryx rhamni        12 Cleopatra Gonepteryx cleopatra  13 Wood White Leptidea sinapis       15 Green Hairstreak Callophrys rubi    16 Small Copper Lycaena phlaeas  17 Large Copper Lycaena dispar  18 Sooty Copper Lycaena tityrus    19 Little Blue Cupido minimus      20 Baton Blue Pseudophilotes baton  21 Silver-studded Blue Plebejus argus    22 Brown Argus Aricia agrestis  23 Mazarine Blue Cyaniris semiargus  24 Escher’s Blue Agrodiaetus escheri  25 Provence Chalk-hill Blue Lysandra hispana  26 Adonis Blue Lysandra bellargus      27 Common Blue Polyommatus icarus      29 Southern White Admiral Limenitis reducta   30 Red Admiral Vanessa atalanta    31 Painted Lady Vanessa cardui    32 Small Tortoiseshell Aglais urticae        33 Queen of Spain Fritillary Issoria lathonia LO   34 Pearl-bordered Fritillary Clossiana euphrosyne  

France - The Vercors Tour Report

May Common name Scientific name 21 22 23 24 25 26 27 28 35 Weaver's Fritillary Clossiana dia  36 Glanville Fritillary Melitaea cinxia  37 Knapweed Fritillary Melitaea phoebe  38 Spotted Fritillary Melitaea didyma   46 Heath Fritillary Mellicta athalia       48 Meadow Brown Maniola jurtina   49 Small Heath Coenonympha pamphilus       50 Pearly Heath Coenonympha darwiniana    51 Darwin's Heath Coenonympha arcania   52 Wall Brown Lasiommata megera     53 Northern Wall Brown Lasiommata petropolitana    55 Grizzled Skipper Pyrgus malvae       56 Rosy Grizzled Skipper Pyrgus onopordi  57 Red-underwing Skipper Spialia sertorius   58 Dingy Skipper Erynnis tages     59 Chequered Skipper Carterocephalus palaemon  60 Small Skipper Thymelicus sylvestris  61 Large Skipper Ochlodes venatus 

Micro Moths (C=caterpillar) 1 Moth (Long Antennae) Adela reaumurella  2 Arched Marble Olethreutes arcuella  3 Box Tree Moth Cydalima perspectalis C C

Macro Moths (B=Bivouac) 1 Six Spot Burnet Zygaena filipendulae  2 Five Spot Burnet Zygaena trifolii  3 Tau Emperor Aglia tau   4 Lewes Wave Scopula immortata  5 Small Argent & Sable Epirrhoe tristata  6 Common Carpet Epirrhoe alternata  7 Grass Rivulet Perizoma albulata     8 Latticed Heath Semiothisa clathrata    9 Speckled Yellow Pseudopanthera macularia      10 Common Heath Ematurga atomaria   11 Black Mountain Moth Glacies coracina  12 Hummingbird Hawk-moth Macroglossum stellatarum   13 Broad-bordered Bee Hawk-moth Hemaris fuciformis   14 Pine Processionary Thaumetopoea pityocampa B B B 15 Noctuid Actinotia radiosa  16 Small Yellow Underwing Panemeria tenebrata  17 Mother Shipton Callistege mi     18 Burnet Companion Euclidia glyphica       

Other Invertebrates 1 Mayfly Ephemeroptera sp    2 Azure Bluet Coenagrion puella  3 Large Red Damselfly Pyrrhosoma nymphula  4 Broad-bodied Chaser Libellula depressa 

France - The Vercors Tour Report

May Common name Scientific name 21 22 23 24 25 26 27 28 5 Stonefly Dinocras cephalotes  6 Wart-biter Cricket Decticus verrucivorus N 7 Field Cricket Gryllus campestris H H  H H H H 8 Ear-wig Anechura bipunctata  9 Shield Bug Graphosoma lineatum  10 Forest Bug Pentatoma rufipes   11 Green Shield Bug Palomena prasina  12 Assasin Bug Rhinocoris iracundus   13 Common Pond Skater Gerris lacustris        14 Froghopper (Red & Black) Ceropis vulnerata    15 Ascalaphid Libelloides coccajus     16 Scorpion Fly Panorpa communis  17 Marsh Crane-fly Tipula oleracea  18 St. Mark's Fly Bibio marci   19 Bee-fly Bombylius sp.     20 Wood Ant Formica rufa     21 Hornet Vespa crabro  22 European Paper Wasp Polistes gallicus    23 Carpenter Bee Xylocopa sp.       24 Honey Bee Apis millifera        25 Garden Bumblebee Bombus hortorum  26 Tree Bumblebee Bombus hypnorum  27 White-tailed Bumblebee Bombus lucorum  28 Early Bumblebee Bombus pratorum  29 Red-tailed Bumble Bee Bombus lapidarius     30 Cullams Bumblebee Bombus cullumanus  31 Common Carder Bee Bombus pascuorum   32 Common Ground Beetle Pterostichus sp.   33 Green Ground Beetle Carabus auratus  34 Dor Beetle Geotrupes stercorarius  35 Cock Chafer Melolontha melolontha      36 Rose Chafer Cetonia aurata    37 Click Beetle Ampedus cinnabarinus  38 Soldier Beetle Cantharis rustica    39 Soldier Beetle Trichodes alvearius   40 Seven-spot Ladybird Coccinella 7-punctata   41 Two-spot Ladybird Adalia bipunctata  42 Green 'Fat-thighed' Beetle Oedemera nobilis  43 a Longhorn Beetle Agapanthia villosoviridescens  44 Wasp Beetle Clytra arietis  45 a Longhorn Beetle Rhagium inquisitor  46 a Longhorn Beetle Rhagium mordax  47 a Longhorn Beetle Leptura rubra  48 Lily Beetle Lilioceris lillii  49 Leaf Beetle Chrysolina geminata  50 Weevil sp. -   