In Memory of PETER LENK

SALAMANDRA 42 4 193-196 Rheinbach, 20 November 2006 ISSN 0036-3375

In Memoriam PETER LENK (26 March 1964 – 23 November 2005)

It is really hard to find words for an obitu- ary for a friend who has left us far too pre- maturely. But PETER LENK deserves to be remembered long into the future. I first made PETER’s acquaintance soon after I had begun tutoring at the Technical University of Darmstadt in the late 1980’s. Lower Franconian by birth, he had origi- nally studied landscaping at the TU Munich, and then switched to biology in Darmstadt. He was an enthusiastic terrarium keeper and field herpetologist, and dragonflies were his second passionate focus. As the subject of his dissertation, we se- lected a comparative study of the population genetics of the Aesculapian snake, Elaphe longissima (now Zamenis longissimus), from throughout its range. One of the enduring results of this research was the realization that the southern Italian Aesculapian snake was a separate species, Elaphe lineata (today Zamenis lineatus). While working in my laboratory, PETER met a young graphic designer who was com- pleting her practicum at the Hessian State Museum in 1991. It must have been love at first sight. SABINE and PETER made a hand- analysis he had been trained in during a stay some and very harmonious couple and were with ROGER THORPE in Bangor (Wales/UK), married in 1995. he also worked out an innovative RNA-RNA PETER worked on his doctoral thesis at the hybridisation method that would permit the Institute for Pharmacology and Molecular identification of genetic haplotypes. As a Biotechnology in Heidelberg. He participat- man of many interests, PETER also contrib- ed in a research project on the phylogeogra- uted to other projects: he established a DNA phy of European reptiles that was headed by family tree for the viperids, and he substi- MICHAEL WINK and myself. His part of the tuted for me in a research trip to Vietnam in project focussed on the intraspecific differ- a joint effort with our Russian colleagues, entiation of the European pond turtle, Emys NATALIA ANANJEVA and NIKOLAI ORLOV. It was orbicularis. an adventurous field trip, with PETER’s leg PETER was a perfectionist who always being pierced by an arrow that, although not remained open to criticism. His methodologi- poisoned, caused a wound that healed badly. cal flexibility was remarkable. Not only did Moreover he was infected with Malaria. PE- he adapt the complex methods of statistical TER graduated summa cum laude in 1997, and

© 2006 Deutsche Gesellschaft für Herpetologie und Terrarienkunde e.V. (DGHT) http://www.salamandra-journal.com 193 ULRICH JOGER his work on the European pond turtle can be special person. We will miss his humour, considered as trail blazing. As recently as his multifaceted knowledge, his uncompro- 2004, UWE FRITZ used it as a basis for the mising love for truth, and even his usually definition of a new species, Emys trinacris, justified criticism. His friends will not forget from Sicily and Calabria. him. PETER was also a nature conservation ac- tivist and contributed to the establishment of Bibliography of PETER LENK a Red List for the amphibians and reptiles of Bavaria. After his relocation to Kahl am 1. LENK, P. (1988): Eine ungewöhnlich helle Gelb- Main, he became a founding member of the bauchunke (Bombina variegata LINNAEUS 1758) Workgroup Amphibian and Reptile Conser- bei Aschaffenburg – Nachrichten des Naturwis- vation in Hesse (AGAR), later becoming a senschaftlichen Museums der Stadt Aschaffen- member of its board. He was a critical thinker burg, 95: 67-69. who often wanted all the details before he 2. DROBNY, M., D. WENGER, P. LENK & O. ASSMANN would approve of a project. The popular (1989): Untersuchungen zur Aktivitätsdynamik claim of Hessian nature activists that sur- und Habitatwahl einer Population der Äskulapnat- viving autochthonous pond turtles had been ter, Elaphe longissima (LAURENTI, 1768) mit Hilfe der Radiotelemetrie. – Unveröffentlichter Bericht found in the Rhine/Main region was criti- für das Landratsamt Altötting, 52 pp. cised by him as not sufficiently proven. 3. LENK, P. (1989): Ein Fremdling am Unter- Fate hit PETER hard after Christmas of main: Die Mauereidechse (Podarcis muralis ssp.) 1999. Having meanwhile become a father of [(Reptilia; Bay.: Unterfranken)]. [Abb. 1 auf S. two and working in a postdoctoral position at 102, Abb. 2 und 4 auf der unpaginierten S. 113]. Aventis Pharma in Frankfurt/M., he was di- – Nachrichten des Naturwissenschaftlichen Muse- agnosed with a brain tumour. The following ums der Stadt Aschaffenburg, 96: 02-113. years were a constant alternation between 4. LENK, P. (1991): Verzeichnis der Amphibien und phases of recovery and hope for complete Reptilien des Naturwissenschaftlichen Museums convalescence and ever recurring periods of Aschaffenburg. – Nachrichten des Naturwissen- deterioration. Surgery failed to completely schaftlichen Museums der Stadt Aschaffenburg, remove the tumorous tissue, and chemo- and 98: 101-106. radiotherapies only managed to stabilize its 5. LENK, P. (1992): Die Libellen der Stadt Aschaf- growth temporarily. PETER fought his disease, fenburg. – Nachrichten des Naturwissenschaft- lived his life with his family, worked as a lichen Museums der Stadt Aschaffenburg, 99: bioinformatics specialist at Aventis, and kept 1-24. on publishing. 6. LENK, P. (1992): Die Fröhlich’sche Odonaten- His last personally authored publication, Sammlung im Naturwissenschaftlichen Museum on the discovery of a parthenogenetic mo- der Stadt Aschaffenburg – Nachrichten des Na- nitor lizard, appeared two weeks after his turwissenschaftlichen Museums der Stadt Aschaf- fenburg, 99: 25-36. death on 23 November 2005. A paper on the area-determining effect of insolation that was 7. LENK, P. (1992): Die Amphibien der Stadt Aschaffenburg. – Mitteil. Landesverband f. Am- intended for the proceedings of the sympo- phibien- u. Reptilienschutz Bayern, 12: 10-27. sium on “Ranges and Distribution Patterns” of the DGHT Workgroup Field Herpetolo- 8. LENK, P. (1993): Untersuchungen zur inner- artlichen Gliederung der Äskulapnatter, Elaphe gy in 2006, remained a nearly completed longissima (LAURENTI, 1768). – Diplomarbeit, manuscript. PETER was also a co-author of Technische Hochschule Darmstadt, 87 pp. the paper “Phylogeographie, Artbildung und 9. LENK, P. & U. JOGER (1994): Genetic relation- postpleistozäne Einwanderung mitteleuropä- ships between populations and intraspecific sub- ischer Reptilien” that was published in the division of Elaphe longissima (LAURENTI, 1768) same proceedings volume. as suggested by plasma protein electrophoresis With PETER LENK we have lost a very and DNA fingerprinting. – Amphibia-Reptilia, 15:

194 In Memory of PETER LENK

363-373. 21. LENK, P. & M. WINK (1997): A RNA/RNA 10. FRITZ, U., P. LENK & S. LENK (1995): Sumpf- heteroduplex cleavage analysis to detect rare mu- schildkröten (Emys orbicularis galloitalica) tations in populations. – Molecular Ecology, 6: aus Südfrankreich und Latium. – Herpetofauna 233-237. (Weinstadt), 17(97): 13-20. 22. FRITZ, U., A. PIEH, P. LENK, J. MAYOL, B. 11. LENK, P. (1995): Zum Fund einer leukistischen SÄTTELE & M. WINK (1998): Is Emys orbicula- Larve der Knoblauchkröte (Pelobates fuscus) aus ris introduced on Majorca? – Mertensiella, 10: Alzenau, Unterfranken. – Zeitschrift für Feldher- 123-133. petologie, 2: 221-223. 23. JOGER, U., T. AMANN, P. LENK & U. WILLAND 12. MALKMUS, R. & P. LENK (1995): Libellen. (1998): Molekulare Merkmale und das phyloge- – Schriftenreihe Flora und Fauna im Landkreis netische Artkonzept. – Zoologische Abhandlun- Main-Spessart, Bd. 2. 112 pp. gen, Staatliches Museum für Tierkunde Dresden, 50/Suppl. „100 Jahre Artkonzepte in der Zoolo- 13. LENK, P. (1996): Book review of: Das Terra- gie“: 109-123. rium für den Anfänger. H.-J. HERRMANN. 1994. – Copeia, 1996(2): 506-507. 24. LENK, P., U. JOGER, U. FRITZ, P. HEIDRICH & M. WINK (1998): Phylogeographic patterns in the 14. JOGER, U. & P. LENK (1997): Entnahme und mitochondrial cytochrome b gene of the European Behandlung von Blutproben für molekulargeneti- pond turtle, Emys orbicularis (LINNAEUS, 1758): sche Untersuchungen im Freiland. – Mertensiella First results. – Mertensiella, 10: 159-175. (Rheinbach), 7: 329-340. 25. HERRMANN, H.-W., U. JOGER, P. LENK & M. 15. JOGER, U., P. LENK, I. BARAN, W. BÖHME, T. WINK (1999): Morphological and molecular phy- ZIEGLER, P. HEIDRICH & M. WINK (1997): The logeny of Viperines: Conflicting evidence? – Kau- phylogenetic position of Vipera barani and V. pia, 8: 21-30. nikolskii within the Vipera berus complex. – pp. 185-194 in BÖHME, W., W. BISCHOFF & T. ZIEGLER 26. JOGER, U., H.-W. HERRMANN, S. KALYABINA, (eds.): Herpetologia Bonnensis. Proceedings of P. LENK & M. WINK (1999): Molecular phylo- the 8th Ordinary General Meeting of the Societas genetics and systematics of true vipers (Reptilia: Europaea Herpetologica, Bonn 1995. Viperinae). – Zoology, 102, Suppl. II: 7. 16. JOGER, U., M. MEDER & P. LENK (1997): Split- 27. LENK, P., H.-W. HERRMANN & U. JOGER (1999): ter oder Lumper – wieviele Arten umfaßt der Vi- Phylogeny and systematics of Bitis (Viperidae) pera xanthina-Komplex? – Zusammenfassungen based on molecular evidence. – Kaupia, 8: 31- DGHT Jahrestagung 1997: 15. 38. 17. LENK, P. (1997): Molekularbiologische Unter- 28. LENK, P., U. JOGER, U. FRITZ & M. WINK suchungen zur Mikroevolution der Europäischen (1999): Mitochondrial phylogeography of the Eu- Sumpfschildkröte Emys orbicularis (LINNAEUS, ropean pond turtle, Emys orbicularis (LINNAEUS 1758). – Dissertation, Technische Universität 1758). – Molecular Ecology, 8: 1911-1912. Darmstadt, 178 pp. 29. LENK, P., U. JOGER, U. FRITZ, P. HEIDRICH & 18. LENK, P. (1997): Die Gestreifte Quelljungfer M. WINK (1999): Phylogeographic patterns in the (Cordulegaster bidentatus SELYS 1843) im Spes- mitochondrial cytochrome b gene of the European sart. – Nachrichten des Naturwissenschaftlichen pond turtle, Emys orbicularis (LINNAEUS, 1758). Museums der Stadt Aschaffenburg, 104: 34-39. – Mertensiella (Rheinbach), 10: 159-175. 19. LENK, P., S. HANKA, U. FRITZ, U. JOGER & M. 30. LENK, P. & W. WÜSTER (1999): A multivariate WINK (1997): Die Europäische Sumpfschildkröte approach to the systematics of Italian ratsnakes im Enkheimer Ried bei Frankfurt/M: Nachweis of the Elaphe longissima complex (Reptilia: Col- für Einbürgerung. – Elaphe, n.F., 5: 70-75. ubridae): Revalidation of Camerano’s Callopeltis longissimus var. lineata. – Herpetological Journal, 20. LENK, P., U. JOGER & M. WINK (1997): Phy- 9: 153-162. logeography of mitochondrial genotypes in the European pond turtle (Emys orbicularis). – in 31. LENK, P. (2001): Libellenbeobachtungen in ROCEK, Z. & HART, S. (eds.): Herpetology `97 der Kahler Seenlandschaft. – Unser Kahlgrund. – Abstracts of the Third World Congress of Herpe- Heimatjahrbuch für den ehemaligen Landkreis tology 2-10 August 1997 Prague, Czech Republic, Alzenau: 68-76. Prague: 126. 32. LENK, P., U. JOGER & M. WINK (2001): Phylo-

195 ULRICH JOGER genetic relationships among European ratsnakes by the Iron Curtain. – Biologia, 59 (Suppl. 14): of the genus Elaphe FITZINGER based on mitochon- 19-25. drial DNA sequence comparisons. – Amphibia- 38. KALYABINA-HAUF, S., N.B. ANANJEVA, U. JOGER, Reptilia, 22: 329-339. P. LENK, R.W. MURPHY, B.L. STUART, N.L. ORLOV, 33. LENK, P., S. KALYABINA, M. WINK & U. JOGER C.T. HO & M. WINK (2004): Molecular phylogeny (2001): Evolutionary relationships among the true of the genus Acanthosaura (Agamidae). – Current vipers (Viperinae) suggested by mitochondrial Herpetology, 23: 7-16. DNA sequences. – Molecular Phylogenetics and 39. FRITZ, U., A. CADI, M. CHEYLAN, C. COÏC, M. Evolution, 19: 94-104. DÉTAINT, A. OLIVIER, E. ROSECCHI, D. GUICKING, P. 34. LENK, P. (2002): Verdriftung Maurischer Bach- LENK, U. JOGER & M. WINK (2005): Distribution of schildkröten (Mauremys leprosa) und Vipernattern mtDNA haplotypes (cyt b) of Emys orbicularis in (Natrix maura) an der südportugiesischen Küste. France and implications for postglacial recoloni- – Zeitschrift für Feldherpetologie, 9: 221-223. zation. – Amphibia-Reptilia, 26: 231-238. 35. JOGER, U., D. GUICKING, S. KALYABINA, P. 40. FRITZ, U., T. FATTIZZO, D. GUICKING, S. TRIPE- LENK, Z.T. NAGY & M. WINK (2003): Pleistocene PI, M.G. PENNISI, P. LENK, U. JOGER & M. WINK speciation and patterns of Holocene dispersal of (2005): A new cryptic species of pond turtle from European reptiles from Mediterranean refugia. South Italy, the hottest spot in the range of the – Verhandlungen der Gesellschaft für Ökologie, genus Emys (Reptilia: Testudines: Emydidae). 33: 214. – Zoologica Scripta, 34: 1-21. 36. LENK, P., U. FRITZ, D. GUICKING, U. JOGER & 41. LENK, P., B. EIDENMUELLER, H. STAUDTER, R. M. WINK (2003): Dispersal of the European Pond WICKER & M. WINK (2005): A parthenogenetic Turtle from Pleistocene refugia. – Verhandlungen Varanus. – Amphibia-Reptilia, 26: 507-514. der Gesellschaft für Ökologie, 33: 231. 42. JOGER, U., D. GUICKING, S. KALYABINA-HAUF, 37. FRITZ, U., D. GUICKING, P. LENK, U. JOGER & P. LENK, Z.T. NAGY & M. WINK (2006): Phylogeo- M. WINK (2004): When turtle distribution tells graphie, Artbildung und postpleistozäne Einwan- European history: mtDNA haplotypes of Emys derung mitteleuropäischer Reptilien. – Zeitschrift orbicularis reflect in Germany former division für Feldherpetologie, Suppl. 10: 29-59.

Author‘s address: ULRICH JOGER, Staatliches Naturhistorisches Museum, Pockelsstraße 10, D-38106 Braunschweig, Germany, E-Mail: [email protected].

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