Zootaxa 2842: 1–188 (2011) ISSN 1175-5326 (Print Edition) Monograph ZOOTAXA Copyright © 2011 · Magnolia Press ISSN 1175-5334 (Online Edition)

Home , Dorsum (moth), Flax (moth), Medius, Paradoxica, Parens (moth)

ZOOTAXA

2842

Revision of the Micronoctuidae (Lepidoptera: Noctuoidea). Part 4, Taxonomy of the subfamilies Tentaxinae and Micronoctuinae

MICHAEL FIBIGER* Molbechs Allé 49, DK-4180 Sorø, Denmark *Michael Fibiger passed away prior to publication; inquiries should be addressed to Ole Karsholt at the Natural History Museum of Denmark (Copenhagen), where Fibiger's material has been sent.

Magnolia Press Auckland, New Zealand

Accepted by L. Gall: 20 Jan. 2011; published: 29 Apr. 2011 MICHAEL FIBIGER Revision of the Micronoctuidae (Lepidoptera: Noctuoidea). Part 4, Taxonomy of the subfamilies Tentaxinae and Micronoctuinae (Zootaxa 2842) 188 pp.; 30 cm. 29 Apr. 2011 ISBN 978-1-86977-663-3 (paperback) ISBN 978-1-86977-664-0 (Online edition)

FIRST PUBLISHED IN 2011 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/

© 2011 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use.

ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition)

Abstract ...... 5 Introduction, material, and methods ...... 6 Systematic part, taxonomy, Part 4 ...... 8 FAMILY MICRONOCTUIDAE Fibiger, 2005 ...... 8 SUBFAMILY TENTAXINAE Fibiger, new subfamily ...... 11 Genus Acusa Fibiger, new genus ...... 13 Checklist of species of genus Acusa Fibiger ...... 13 Acusa acus Fibiger, new species ...... 13 Genus Tentasetae Fibiger, new genus ...... 14 Checklist of species of genus Tentasetae Fibiger ...... 15 Tentasetae cambodiana Fibiger, new species ...... 15 Genus Pseudobscura Fibiger, new genus ...... 16 Checklist of species of genus Pseudobscura Fibiger...... 17 Pseudobscura karwaria Fibiger, new species ...... 17 Genus Parens Fibiger, new genus ...... 18 Checklist of species of genus Parens Fibiger ...... 19 Parens occi Fibiger & Kononenko, 2008, new combination...... 19 Parens paraocci Fibiger, new species ...... 21 Parens melli Fibiger, new species ...... 22 Parens chekiangi Fibiger, new species ...... 23 Genus Alienia Fibiger, new genus ...... 24 Checklist of species of genus Alienia Fibiger ...... 26 Alienia flavofascia Fibiger, new species ...... 26 Genus Tentaspina Fibiger, new genus ...... 27 Checklist of species of genus Tentaspina Fibiger ...... 28 Tentaspina venus Fibiger, new species ...... 28 Tentaspina duospina Fibiger, new species ...... 29 Tentaspina feriae Fibiger, new species ...... 30 Tentaspina balii Fibiger, new species ...... 31 Tentaspina sinister Fibiger, new species ...... 32 Tentaspina orienta Fibiger, new species ...... 33 Tentaspina paraorienta Fibiger, new species ...... 34 Genus Tentax Fibiger, new genus ...... 35 Checklist of species of genus Tentax Fibiger ...... 36 Tentax argentescens (Hampson, 1912), new combination ...... 36 Tentax mini Fibiger, new species ...... 38 Tentax vetus Fibiger, new species ...... 39 Tentax minima (Hampson, 1926), new combination ...... 40 Tentax makasi Fibiger, new species ...... 41 Tentax brunnea Fibiger, new species ...... 42 Tentax scoblei Fibiger, new species ...... 43 Tentax penicilla Fibiger, new species ...... 45 Tentax ferax Fibiger, new species ...... 46 Tentax malleus Fibiger, new species...... 47 Tentax tentaxia Fibiger, new species ...... 48 Tentax badasi Fibiger, new species ...... 50 Tentax bruneii Fibiger, new species ...... 51 Tentax musculus Fibiger, new species ...... 52 Genus Flax Fibiger, new genus ...... 53 Checklist of species of genus Flax Fibiger ...... 54 Flax basis Fibiger, new species...... 55 Flax amita Fibiger, new species ...... 56 Flax clavus Fibiger, new species...... 57

MICRONOCTUIDAE IV Zootaxa 2842 © 2011 Magnolia Press · 3 Flax poseidon Fibiger, new species ...... 59 Flax tempuensis Fibiger, new species ...... 60 Flax neptun Fibiger, new species ...... 62 Flax brevipennis Fibiger, new species ...... 63 Flax bibrevipennis Fibiger, new species ...... 64 Flax serami Fibiger, new species ...... 65 Flax tamborai Fibiger, new species ...... 67 Flax kuchingi Fibiger, new species ...... 68 Flax sulawesii Fibiger, new species ...... 69 Flax kalliesi Fibiger, new species ...... 70 Flax longus Fibiger, new species ...... 72 Flax semilongus Fibiger, new species...... 73 Flax micronesia Fibiger, new species ...... 75 Flax palaui Fibiger, new species...... 76 Flax biaki Fibiger, new species ...... 77 Flax newirlandi Fibiger, new species ...... 78 Flax lueneborgi Fibiger, new species ...... 79 Flax newbritaini Fibiger, new species ...... 81 Flax honeyi Fibiger, new species ...... 82 Flax fulturai Fibiger, new species...... 83 Flax fletcheri Fibiger, new species ...... 85 Flax solomoni Fibiger, new species ...... 86 Flax sanchristobali Fibiger, new species ...... 87 Flax elachista (Fletcher, 1957), new combination...... 89 Flax rennelli Fibiger, new species ...... 90 Flax vanuatui Fibiger, new species...... 91 SUBFAMILY MICRONOCTUINAE Fibiger...... 93 Genus Medius Fibiger, new genus ...... 94 Checklist of species of genus Medius Fibiger ...... 95 Medius calceus Fibiger, new species ...... 95 Medius khasisiodes Fibiger, new species ...... 97 Medius brassi Fibiger, new species...... 98 Medius nepali Fibiger, new species ...... 98 Medius unguisi Fibiger, new species...... 100 Medius kalimantani Fibiger, new species ...... 101 Genus Sternitta Fibiger, new genus ...... 102 Checklist of species of genus Sternitta Fibiger ...... 103 Sternitta parasuffuscalis Fibiger, new species ...... 103 Sternitta suffuscalis (Swinhoe, 1886), new combination...... 104 Sternitta hackeri Fibiger, new species ...... 106 Sternitta gregerseni Fibiger, new species ...... 107 Sternitta goateri Fibiger, new species...... 109 Sternitta gabori Fibiger, new species ...... 110 Sternitta magna Fibiger, new species ...... 112 Genus Micronola Amsel, 1935 ...... 113 Checklist of species of genus Micronola Amsel, 1935 ...... 114 Micronola wadicola Amsel, 1935 ...... 114 Micronola yemeni Fibiger, new species ...... 117 Micronola yemeni occidentalis Fibiger, new subspecies ...... 118 Micronola zahirii Fibiger, new species ...... 119 Micronola irani Fibiger, new species ...... 120 Genus Micronocta Fibiger, 1997 ...... 122 Checklist of species of genus Micronoctua ...... 123 Micronoctua karsholti Fibiger, 1997...... 123 ADDITIONS and CORRECTIONS ...... 126 Additions to Fibiger 2007, subfamily Pollexinae ...... 126

4 · Zootaxa 2842 © 2011 Magnolia Press FIBIGER Tolpia kalimantania Fibiger, new species ...... 126 Additions to Fibiger 2008a, subfamily Belluliinae ...... 127 Bellulia bibella Fibiger, 2011 ...... 127 Subfamily Parachrostiinae ...... 128 Parachrostia pura Fibiger, 2011 ...... 128 Mimachrostia fasciata Sugi, 1982...... 129 Additions to Fibiger 2010, subfamily Tactusinae ...... 129 Tactusa ostium Fibiger, 2010 ...... 129 Tactusa virga Fibiger, 2011...... 130 Tactusa flexus Fibiger, 2011 ...... 131 Tactusa brevis Fibiger, 2011 ...... 131 Tactusa discrepans yunnanensis Fibiger, 2011 ...... 132 Tactusa pars Fibiger, 2010 ...... 133 Description of three additional genera of Tactusinae Fibiger, 2010 ...... 134 Genus Dextella Fibiger, new genus ...... 134 Checklist of species of genus Dextella Fibiger ...... 135 Dextella alleni Fibiger, new species ...... 135 Dextella khaoyaiana Fibiger, new species ...... 136 Dextella flavus Fibiger, new species ...... 137 Genus Paradoxica Fibiger, new genus ...... 138 Checklist of species of genus Paradoxica Fibiger...... 139 Paradoxica parki Fibiger, new species ...... 139 Paradoxica asymmetrica Fibiger, new species ...... 140 Paradoxica proxima Fibiger, new species ...... 141 Genus Dorsum Fibiger, new genus ...... 142 Checklist of species of genus Doesum Fibiger ...... 144 Dorsum brunescens Fibiger, new species ...... 144 Dorsum kwaii Fibiger, new species ...... 145 Dorsum teraii Fibiger, new species ...... 146 Dorsum bengali Fibiger, new species ...... 147 Dorsum atlas Fibiger, new species ...... 148 Acknowledgements ...... 149 References ...... 149 Illustrations ...... 151–188 Imagines plates ...... 151–165 Genitalia plates males ...... 166–180 Genitalia plates females ...... 181–187

Abstract

This is the fourth part in a series of papers on the recently described family Micronoctuidae Fibiger, 2005. Part 4 includes the description of Tantaxinae, new subfamily, and Micronoctuinae, and 15 genera (13 new): Acusa, Tentasetae, Pseudob- scura, Parens, Alienia, Tentaspina, Tentax, Flax, Medius, Sternitta, Dextella, Paradoxica, and Dorsum (all new). Two genera are redescribed: Micronola Amsel, 1935 and Micronoctua Fibiger, 1997. A total of 93 species are treated (81 new, one new subspecies; 12 species are redescribed). Six new combinations are given. Additions are given to the previously published parts of the revision: Pollexinae Fibiger, 2007 (one new species), Belluliinae Fibiger, 2008a (one species from Fibiger et al. 2011), Parachrostiinae Fibiger, 2011 (one species from Fibiger et al. 2011), Tactusinae Fibiger, 2010 (three species and one subspecies from Fibiger et al. 2011, and 11 new species in three new genera). Taxa inhabit the Arabian Peninsula, and Asian, Oriental, and Oceania regions in tropical, subtropical, and temperate zones both north and south of the equator. Three preoccupied generic names, Dignus Fibiger, Fustis Fibiger, and Obscura Fibiger, proposed as new in Fibiger 2010, are replaced by Dignius Fibiger, 2011, Fustius Fibiger, 2011, and Obscurior Fibiger, 2011, resulting in 15 new combinations.

Key words: Micronoctuidae, taxonomy, new subfamilies, new genera, new species, new combinations, Arabian Peninsu- la, South-West, South, East, and South-East Asia, Oriental, and Oceania regions