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Kenai National Wildlife Refuge Species List, Version 2018-07-24

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Kenai National Wildlife Refuge Species List, Version 2018-07-24 Kenai National Wildlife Refuge Species List, version 2018-07-24 Kenai National Wildlife Refuge biology staff July 24, 2018 2 Cover image: map of 16,213 georeferenced occurrence records included in the checklist. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Invertebrates ......................................................... 55 Vascular Plants........................................................ 91 Bryophytes ..........................................................164 Other Plants .........................................................171 Chromista...........................................................171 Fungi .............................................................173 Protozoans ..........................................................186 Non-native species 187 Vertebrates ..........................................................187 Invertebrates .........................................................187 Vascular Plants........................................................190 Extirpated species 207 Vertebrates ..........................................................207 Vascular Plants........................................................207 Change log 211 References 213 Index 215 3 Introduction Purpose to avoid implying that any of the provisional names we use are formally accepted by formatting them as proper scien- A primary purpose for which the Kenai National Wildlife tific names. Also, note that our database enforces lower case Refuge was established in the Alaska National Interest letters in our specific epithets so that, for example, \Rab- Lands Conservation Act of 1980 was, \to conserve fish and dophaga sp. BOLD:ADA6450" becomes \Rabdophaga sp. wildlife populations and habitats in their natural diver- bold:ada6450." sity. ," where the term “fish and wildlife" was defined Synonymies we included do not necessarily reflect our as \any member of the animal kingdom, including without opinions and are not intended as statements on correct tax- limitation any mammal, fish, bird. , amphibian, reptile, onomy (we generally follow ITIS); rather, synonymns are mollusk, crustacean, arthropod or other invertebrate." An included in the index as an aid to the reader to help find obvious first step toward fulfilling this purpose is to know taxa for which names may have changed. what fish and wildlife, habitats, and natural diversity are to In the checklist species represented by both native and be conserved. This checklist is intended to be a frequently- non-native populations are indicated with a double dagger updated document reflecting our current knowledge of which symbol (z). living things call the Kenai National Wildlife Refuge home. This list is dynamic and far from perfect with many changes and corrections yet to be made. Please send comments and corrections to Matt Bowser at About the list matt [email protected]. The present list includes a total of 2,183 species. Of these, 2,073 are considered to be native, 105 species are con- Acknowledgments sidered to be non-native, and five species are represented by both native and non-native populations. Nineteen non- Thanks and credit are due to many people who have re- native species are believed to have been extirpated from the viewed this list, provided identifications, and helped in other Refuge. ways. The list below is by no means exhaustive. The current list includes 208 vertebrates, 848 inverte- brates, 182 bryophytes, 531 vascular plants, 372 fungi, 27 James Bergdahl (Conservation Biology Center, Spokane, protozoans, and 15 algae. Washington) reviewed and made helpful comments on Since the last version of this list (Kenai National Wildlife the Refuge's list of Carabidae. Refuge biology staff, 2017) where 2,139 species were listed, there has been a net increase of 44 species added. See the Peter Hovingh provided a list of leeches (Hirudinea) col- change log (page 211) for details. lected on the Refuge. Although accepted scientific names are used wherever these are known, we have included provisional identifications John Hudson (USFWS, Juneau, Alaska) provided lists of (for example, \Typhula sp.") where these represent the best Odonata observed on the Refuge. information available. Species recognized only as molecu- Richard Payne (University of York, Heslington, UK) pro- lar operational taxonomic units are also included, mainly vided a list of protists from Jigsaw Lake from Payne entities recognized by Barcode Index Numbers (BINs, Rat- et al.(2006). nasingham and Hebert, 2013). We have chosen not to fol- low the usual convention of italicizing genus and species David Wartinbee (retired from Kenai Peninsula College, names because our current system does not enable us to Soldotna, Alaska) shared a list of Chironomidae he had handle accepted and provisional names separately. We want collected on the Refuge. 5 Native species occurring on the Kenai National Wildlife Refuge Vertebrates Pungitius pungitius Common names: Tenspined stickleback Phylum Chordata Order Salmoniformes Class Actinopterygii Family Salmonidae Order Cypriniformes Oncorhynchus gorbuscha Family Catostomidae Common names: Humpbacked salmon Catostomus catostomus Oncorhynchus keta Common names: Longnose sucker Common names: Chum salmon Oncorhynchus kisutch Common names: Coho salmon or silver salmon Female Catostomus catostomus from Kelly Lake. Image CC0 Dean and Rickabough(2005). Taxonomic notes: Two forms have been documented from the Refuge: the typical form and a semi-dwarf form known from Wolf and Finger Lakes (Dean and Rickabough, 2005). Oncorhynchus kisutch caught at Swanson River Landing, September Distribution and trends: Widespread across Alaska, northern 17, 2008. Image CC BY Matt Bowser, http:// www:inaturalist:org/ North America, and into Arctic Siberia (Page and Burr, 1991), this observations/ 4891214. species is present in the Kenai River and Swanson River watersheds : Friedersdorff and Jakubas(1984 a) on the Refuge (Dean and Rickabough, 2005). Literature records iNatu- ralist observations: 4891214 Biology: Catostomus catostomus is a bottom-feeder, consuming mainly invertebrates (Mansfield, 2004). Oncorhynchus mykiss Literature records: Dean and Rickabough(2005), Friedersdorff Common names: Steelhead (1986), Friedersdorff and Jakubas(1984 a), Friedersdorff and Jakubas Literature records: Kyle(1992): iNaturalist observations: (1984b) 811982, 1814201 Oncorhynchus nerka Order Gasterosteiformes Common names: Sockeye salmon or kokanee : Kyle(1992) : Family Gasterosteidae Literature records iNaturalist observations 1021584 Gasterosteus aculeatus Oncorhynchus tshawytscha Common names: Threespine stickleback Common names: Chinook salmon or king salmon Prosopium cylindraceum Common names: Round whitefish Literature records: Kyle(1992) Salvelinus alpinus Common names: Arctic char Gasterosteus aculeatus at Headquarters Lake, October 15, Salvelinus malma 2015. Image CC0 Matt Bowser, http:// www:inaturalist:org/ Common names: Dolly varden observations/ 2138344. Literature records: Kyle(1992) iNaturalist observations: 811997 Literature records: Friedersdorff and Jakubas(1984 a), Kyle (1992), Sprehn et al.(2015) iNaturalist observations: 2138344 Salvelinus namaycush 7 8 NATIVE SPECIES Common names: Lake trout Literature records: Kyle(1992) Order Scorpaeniformes Family Cottidae Cottus aleuticus Common names: Coastrange sculpin Literature records: Friedersdorff and Jakubas(1984 a), Kyle (1992) Cottus cognatus Common names: Slimy sculpin Literature records: Friedersdorff and Jakubas(1984 a), Kyle (1992) Class Amphibia Order Anura Family Ranidae Lithobates sylvaticus Synonyms: Rana sylvatica Map of records of Accipiter striatus. Common names: Wood frog iNaturalist observations: 115158, 115166, 117220, 1816454 eBird checklists: S25492838 Class Aves Order Accipitriformes Aquila chrysaetos Family Accipitridae Accipiter gentilis Common names: Golden eagle Common names: Northern goshawk Map of records of Accipiter gentilis. Map of records of Aquila chrysaetos. eBird checklists: S8749697, S11505349, S3491054, S5866265, S5866269, S5866305, S5866306, S5866309, S5866310, S5870989, S5870990, S5870993, S5870994, S5878395, S6551001, S6980323, eBird checklists: S2627102, S2960113, S6526683, S8519986, S6551002, S6980324, S6550987, S6980344, S6709117, S6873993, S8542506 S7333547, S7372812, S8309514, S8365991, S8366301, S8388735, S8458336, S8414468, S8414915, S25492838 iNaturalist observations: 114273 Buteo jamaicensis Accipiter striatus Common names: Sharp-shinned hawk Common names: Red-tailed hawk VERTEBRATES 9 Map of records of Buteo jamaicensis. Map of records of Circus cyaneus. Literature records: Morton et al.(2017) eBird checklists: S2620561, S4878668, S5866263, S5866264, S5866266, S5866268, S5866269, S5866277, S5866278, S5866280, S5866237, S5866281, S5866282, S5866241, S5866243, S5866288, S5866290, S5866303, S5866310, S5866284, S5870974, S5870987, S5870994, S5870995, S6312021, S6551000, S6980322, S6446826, S6447196, S6723932, Literature records: Morton et al.(2017) eBird checklists: S6470158, S6551032, S6980247, S6980254, S6980279, S6980233, S5882060, S4342574, S5866263, S5866236, S5866267, S5866268, S6980283, S6980284, S6980308, S8191831, S8343529, S8565098, S5866272, S5866275, S5866276, S5866277, S5866241, S5866245, S8521206, S8542654,
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