P.. I.... nl .. I(J~;• .ul l",,,,,,.,. IM. J9. No, 5. 2005. pp. 4/17-497. Tro m.lu't rl/,,,," /'ulro ""'lo~; , - hr,'t; i Zilurn Genus Delphinella (Ammonoidea) from the Berriasian of the Crimean Mountains V. v. Arkadiev* and T. N. Bogdanova** *St. Petersburg State University, Universitetskaya nab. 719. St. Petersburg, 199034 Russia e-mail: [email protected] **Karpinsky All-Russia Research Institute of Geology. Srednii pr. 74, St. Petersburg, 199026 Russia Received April 28. 2004 Abstract-Seven species of the genus Delphinella, i.e., D. subehaperi (Retowski), D. erimensis (B urckhardt), D. obtusenodosa (Retowski), D. tresanensis Le Hegarat, D. delphinensis (Ki lian). D. janus (Retows ki ), and D. peetinata sp. nov., and the species Berriasella berthei (Toucas) from the upper grandis Subzone ofthejaeobi Zone of the Berriasian of the Crimean Mountains are described. Their stratigraphic distribution is different from that in the stratoty pe area, Key words: new species, Delphinella, Ammonoidea, Berriasian, Crimean Mountains. INTRODUCTION vi llages of Nanikovo and Sultanovka also belong to the jacobi Zone (Fig. 2) (Bogdanova et af., 1984). Species of the genus Delphinella frequently occur in the Berriasian of the Crimea and are very important for In the Tonas River basin, infrequent Delphinella sp. stratigraphy. Along with other Berriasian genera (Pseu were found together with Pseudosubplanites pontieus dosubplanites. Berriasella, Fauriella, Tirnovella, etc.) and P. lorioli (Bogdanova et al., 198 1). they form stable zonal assemblages easil y traceable in The collection contains molds completely rep laced the section s. Delphinella have been known from the by rock matrix, so detailed study of the shell ontogeny Berriasian of the Crimea for a long time, although they is impossible .. The collection is housed in TsNIOR received little monographic study except for the works Museum (St. Petersburg) (coil. no. 13055). We also of Retowski (1893) and Druschits (1960). D. delphin studied the collection of Retowski from the Feodosiya ensis, D. ex gr. obtusenodosa, D. subchaperi, D. cri Section (TsNIGR Museum, coIl. no. 10916). mens is. and D. benhei were recorded but not descri bed from the vicinity of Feodosiya (Bogdanova et al., 1984). HISTORY OF STUDIES . The authors of thi s paper have a large collection OF DELPHINELLA SPECIES (several tens of specimens) of Delphinella shells, col lected over years in the Crimea by v.v. Druschits, Retowski (1893) described an usual grou p of BT. Yanin, T.N. Bogdanova, S.v. Lobatscheva, v.A. Pro ammonoids characterized by ornamentation that zorovskii, T.A. Favorskaya, V.V. Arkadiev, and Yu.N. Savelieva. These shells were found predomi nantly in the eastern Crimean Mountains (vicinity of Feodosiya) and, to a lesser extent, in the central Crimean Mountains (basin of the Tonas River), where the clayey carbonate beds of the jacobi Zone are wide spread (Fig . I). In the section on Cape St. lIya (Fig. 2), the assem blage of Delphinella (D. subchaperi, D. crimensis. D. obtusenodosa, D. tresanensis. D. delphillensis, D. janus, and a new species, D. pectinala) was found together with Pseudosubplallites ponticus (Re!.), BLACK SEA P gralldis (Maz.), P. lorioli (Zill.), and P. combesi o 25 Le Hegarat in the 13-m-thick "Feodosiya Marl" Mem ber, wh ich is presently dated the grandis Subzone of the Fig. l. Locati on of the sections studied: (1) Cape SI. l1ya. jacoiJi Zone (Arkadiev and Savelieva, 2002; Arkadiev, (2) village of Suttanovka. and (3 ) village of Nanikovo. Bar· 2(03). Delphillella that were found in the vicinity of the ak ol Valley. 487 A ~ ~ Sultanovka Nanikovo ; ; E '" E Cape St. Il ya ~ Litho)og,cal Litholog,cal " ,"' column column g, E .:; ~ j .E .\< ; Lithological i ~ ~ 1 • PrrutWsubplunilrs luri(J1i ~ .: ~ column ~~i"~~z:::"'::~ i'-flZ==Z==2j Al Puudo.ubplolli't J lorioli. P. ponli· " '-' £ .!! .... : : : : : : :: .. CUJ. Btrrio"l/a opprli ~ g ~ !i ,!< PrrwJoJubp!Qn;ttJ grandis. P. 8 11 : : : : : : : : Vl N Vl ::;: ~ pom;(uJ, P./orloli. P. combni. BrrriaJtl/o mMsicD, 8. J"beat· z OrlphintIlOlrt Jalltl1JiJ.D. jollus. ______8uriastllu IJ.(rthti. Dtlphintllo "" jisto, Dtlphinrllo subcbop PALEONTOLOGtCAL JOURNAL Vol. 39 No. 5 200S 41)(l ARKADIEV, BOGDANOVJ\ Plalc 5 1',\J.EON I{lI.Otjlt 'Al, JOUk NI\1. Vu!. III Nil,.' 2011.' GENUS fJELPI-IINELLA (AMMONOIDEA) r:1~OM THE BERRIASIAN 4lJ I f)e/phinella )'"bclraperi (I{clo.. '.~ ki , 11I9J) ornamentation in early whorls, more pronounced PI~le .'\. figs. )\ ilnd 9 umbilical ridges, and by the lateral nodes in adults. 1foJllile.t subdwperi: Retowski. 1893. p. 64, pI. 4, figs. 3 and 4; R e m ark s. Follow ing Grigorieva (1938) we Simioncseu, 1899, p_ 5. pI. 1. fig_ 2. include the specimen figured by Si mionescu, 1899, Olllmasiam:; ? suhchaperi: Grigorieva. 1938. p. 118. pI. 7. pI. I, fig. 2 in the synonymy list of this species. figs. 2a and 2b. although it differs somewhat from the typ ica l subcha Berriaselfa Subc1wpf'ri: Luppov el al.. 1949. p. 220. pI. 63. peri in the longer (reaching the umbilicus) intercalating fig. I; non Berriaselfll $uhchaperi: Drusehils. 1960, p_ 276, pi. 21, ribs, we th ink that thi s is just variability. The specimen fig. I (= Relowskiceras tlntirus.wwi (Ret. ). figured and desc ri bed by Nikolov (1982, p. 86, pI. 20, Oelphinelfa subcJllIperi: Le Hcgarat, 1971. p. I 12, pI. 13, fig. 12. pI. 42. fig. 10: Bogdanova el al.. 1984. pI. 4. figs. 4 and 5: fig . 5) as D. obtusenodosa has st rongly pronounced Khimshiashvili. 1989, p. 17, pI. 4. fig. 1 umbilical nodes. We assign thi s specimen to D. subcha Oelphinella oblusellOdosa: Nikolov. 1982. p. 86. pI. 20, fig. 5. peri. The specimen fi gured by Druschits (1960, pI. 21. Jabrollellll subchaperi: Tavera. 1985 . p. 304. pI. 46. fig. I. text~ fig . 1) has coarse ribs (wit hout smoothening) and fig.2]/A. sharpl y pronounced lateral nodes and most likely S h e ll s ha pe. She ll s are large, discoidal, wi th should be assigned to Relowskiceras andrussovi. slowly expanding whorls. Flan ks are wide, almost flat. o c cur r e n C e. Berriasian. jacobi Zone of the The venler is ftauened. The umbilicus is shallow, mod Crimea (graf!di.~ Subzone), Caucasus, Bulgari a, south erately wide, with a narrow, sloping umbilical wall. The eastern France, Spain; Berriasian of Switzerland. umbilical shoulder is gently sloping. The whorl cross M a t e r i a I. Three specimens (nos. 1113055-3113055) section is rectangul ar to oval, compressed. from Cape St. lIya and the vi ll age of Sultanovka; coil. Orname nt ation. The whorls less than 50- by T. N. Bogdanova. 60 mm in diameter have very th in and dense ribbing. Pai red ribs dichotomi zing at mid¥f1 an k or slight ly Delphinella crimt niis (Burckhardl, 1912) above predominate. Very rarely the bi furcation occurs Plate 6, figs. 7-9 near the umbilical shoulder with fu rther deviati ng of Hopliles calislO d'Orbigny var. de/phillensis: Retowski. 1893. both ribs. Single intercalating ribs are present. In the p. 57. pl.]. fig. 4. adul t whorl two pairs of thickenin gs appear on the ribs: HopUles occiJaniclIs: Retowski. 1893, p. 60, pI. 3. figs. 7 and 8. umb ilical and lateral (at the point of rib bifu rcation). non fig. 9. Umbilical swellings virtually do not develop into nodes Sleueroceras cr;mense: Burckhardt, 1912. p. 165. bu t remain as elongated sharp ridges. These ridges give De/phinefla crim€me: Le Hcgarat. 1971. p. 10]. pI. 42. figs. 6 rise to indi sti nct dichotomously branching ri bs. and 8: l3 ogdanova el al .. 1984. pI. 2. fig. 6; pI. ]. fig. 6. Approximately in the middle of the whorl, these S h e II s h ape. Shell s are medium-s ized. wi th dis branches are connected by nodes, which in some spec coidal, moderately low and weakly expand ing whorls. imens are elongated and poimed, and are oriented pe r The flanks are strongly flattened . The venter is narrow, pendicular to the ribs. Aft er the lateral node in the upper very weakly convex. The whorl cross section is high part of the whorl wi th in the pair of ribs, a th ird ri b and rectangular to ova1. The umbilicus is moderately appears. Two or three sing le intercalating ribs are wide and shallow. The umbi lical shoulder is smooth. present between the bundles. These ribs begin from the The umbilical wall is narrow and gently sloping. mid-whorl or slightly lower. Near the lateral nodes the Or n a m enta ti on. On the inner whorls (up to ribs are smoothened and indistinct. Near the venter the Dm = 30-35 mm), the ornamentation consists of very ri bs are distinct, wide, and ri bbon-like. They are inter th in , densely spaced, weakly curved ribs. Tri ply bun rupted on the ve nter by a smooth band. The rib tennina dled ri bs with low bifurcation points at the umbilical tions on the ventrolateral shoulder are directed slight ly shoulder and high bifurcation poi nt s of one of the forward. branches in the mid-flank dominate. In addition, doubly Co m pa r i son. This species is distinguished from bundled ribs wit h bifurcation al the umbilical shoulder the si milar species D. obwsellodosa by the less robust and paired ri bs dichotomizi ng at the mid-whorl are Explanation of Plale 5 In all cases Ihe size is natural. figs. 1-3. OelphilreflajanllS (Relowski): (I) specimen no. 35/ 1305: (I a) laler,1I view. (I b) ventral view: village of Sultanovka; gratl d;s Subwne: (2) specimen no. 3311 3055. later::t! view: village of Nanikovo: grandis Subwne: (3) specimen no. 32/ 13055: (3a) lal eral view. Ob) ventral view: Cape SI. Ilya: gramlis Subzone. Figs. 4 and 5. Del/,/rinel/(I /reswrmsis Le Hcgarnt: (4) specimen no. 20113055. l,lIer;!1 view: village of Nanikovo; gmntlis Subzolle; (5) specimen no. 19/13055. lateral view: l3:lrakol Valley: gml1dis Subzone. Fi~~ . 6 and 7. Dell,/riwlla O/JllI.\·/'rr"dll,wr (Retowski): (6) specimen no. 16/U055. lateral view: Cape St. Ily:l: .~m/l{lis Suhzone: (7) ~pecitnen no. 1.'1/13055,laler,lI view: the same ,lge ;uullocality. F il!-~' Hand 9. Ild"iJilldlu .m/x:/IUI",ri (RelOws ki ): (8) spcdl11en no. 1/13055. 1:I(cr;11 vie\\': village of Sultamwb: .~rmrd;J Subz(lllc: (')1 sll\;(imen no . .V I 3055: (9a) lillcr:11 view. (9b) VC1llr,rl vie\\'; thc .~amc age and lo ..:alil'! . 1';\I.I:(),'.:- rOI.OlilCAl. JOUI{ 01 AI. Vpl. 1') Nu. 5 20()5 ARKADIEV, BOGDANOVA Plalc6 I'i\LEON TOLO(;ICi\\, JOURNi\L Vul. ,I'J Nu, ~ 200~ GENUS IJELPNlNELLII (AMMONOIDEA) fROM THE BERRIASIAN 493 present. Rarely, bidichotomou s ribs and. very rarely. M at e r i a I. Nine specimens (nos. 4/1 3055- 121 13055) si ngle ribs arc present. No pattern in alternation of the from Cape SI. l1 ya, Barakol Valley. and the village of rib types is present. In the body chamber at WH = 23- Sultanovka: coli. by Y.Y. Drusc hits, BT. Yanin . 25 mm , the whorl s increase more slowly in height. T.N . Bogdanova, and vv. Arkadiev. while the ribbin g sll1 00thens in the mid-flank. At thi s stage. thin single crestlike ribs are seen running from Delphinella obluu nodosa (Rcl(lwsk i. 11193) the umbilical shoulder to one-third of the who rl height. Further, the ri bs smoothen to the extent that the pl aces Plale S. figs. 6 and 1: Plate 6. fig. 6 of their bifurcation become very indi stinct. After Hoplites obtusenodosa: Retowski. 1893. p. 62. pI. 3, figs. 10 and 1 \. branching, the ornamentation consists of bundles of IJerriasel/u obtu.renodosa: Ma7cnol. 1939. p. 73, pI. 8. figs. 3 two or three ribs, which are sharpl y pronounced near and4; Druschits. 1960, p. 275. pI. 22. fig. l. the ventrolateral shoulder. In the inner whorls, the ribs lJerriasd/a n. sp. ind. (gr. n. delphinellsis ): ,\i3z.cnot. 1939. are separated on the venter by a smooth band that dis p. 71. pI. 6. fig. 19. appears in the body chamber because ribs continue onto non Berriasella aff. oblusellodosa: Maz.cnot. 1939. p. 72, pI. 7. figs . 6 and 7 (= De/phinel/a el/enica (Nikolov ). the venter. non Benimel/a oblUsenodosa: Amould-Sagct, 1953. p. 46. Dim e n s i o n s in mm, ratio s, a nd num pI. 5. fig. I (=? Berriasella). ? Berriasella aff. obltlsenodosa: Lc Hcgarat and Rcmane. 1968. b er of whorl s p e r half whorl: pI. 4. fi g. 2. non Oelphinella oblusenodosa: Khimshiashvili, 1976. p. lOO. Number of ri bs E pI. 16. fig. 6. Speci- E E per half whorl non Berriruelia (De/phinel/a) obtusenodosa: Sapunov. 1979. men no. e p. 178. pI. 56. fig. 8 (= Delphinellajanus (Rctowski ). E ~ ~ ~ inner outer Cl '"~ ~ ::> '"~ ~ ::> De/phinel/a obtusenodosa: Le Hegarat. 1971. p. 109. pI. )3. fi gs. 10. 11 , 13. and 14; pI. 41. figs. 6 and 7 (non fig. I): Nikolov. 4/13055 68 24 10 23" 0.35 0.15 0.34 34 64 1982. p. 86. pi. 21. fig. 1 (non pi. 20, fig. 5): Bogdanova et al .. 1984 . 30 25 pI. 4. fig. 2. 5/ 13055 50 23 14 0.46 0.28 Sh e II s h ap e. Shell s are medium-sized to large, 28 20 4S wi th di scoidal, low, moderately expanding whorls. The 6/ 13055 71 32 18 0,45 0.25 whorl cross secti on is high and rectangular. The umbi licus is shallow and moderately wide. The umbilical 7113055 46.5 19 12.5 0.41 0.27 24 49 wall is narrow, poorly delineated, and gently sloping. Co m par i so n. This species differs from other Or n a men tat i on. Flanks on the inner whorls up species in the di versity of ribs and their considerable to Om = 25- 30 mm are covered by thin bifurcating smoothening in the body chamber. strai ght ribs. They begin on the umbilical wa ll , lean weakly backwards on the umbilical shoulder, and after R e mark s. Burckhardt (1912) only named the that sli ghtly slanting orad run across the flank. Some species crimeflsis, without describing or fig uring it. what above the mid-flank , the ribs branch into two sim Le Hegarat ( 1971) was the first to descri be the species. ilarly pronounced ribs, with the posterior leaning This author placed th is species in the synonymy list of slightly backwards. On the venter, the ribs are inter Rop/ires calisto d 'Orbigny var. de/phinensis, described rupted by a smooth narrow band. The ornamentation of by Retowski ( 1893). We studied Retowskii 's co llecti on ad ult whorls at Dm = 50 mm consists of paired and. and concl uded that the variety delphifleflsis, similar to more rarel y, single ribs. Weak crests develop on the specimens described by Le Hegarat, is characterized by umbilical shoulders of the body chamber. Swel lings the development of various ribs: triple (bundled), bidi develop in the bifurcation points. These swellings chotomous, paired, and single. sometimes become indisti nct elongated nodes. Imme o c c ur r e n c e. Berriasian, jacobi Zone of the diately after the swellings, ribs smoothen. This effect Crimea (gralldis Subzone), southeastern France, Spain. disappears near the venter. The ribs on the venter of the E xplanati o n of P la te 6 In all cases the size is natural. Figs. I and 2. Delphillel/a peetillata sp. nov.: (t) holotypc no. 3911 )05.'i . lateral view : Cape SI. tly a: gnmdis Subzonc; (2) sp..:ci men no. 40/1 3055. lateral v\cw: th e same age and locality. Figs. J and 4. Berriaulla bertlrei (Toucas): (3) specimen no. 421 13055: (3a) ve nt ral view. (3b) lateral view; village of Sultanovka: grmrdis Subzone: (4) .~pccime n no. 41/13055. lateral view: Barakol Valley: /i,mrdis Subzonc. Fig. 5. De/phincl/a dcll'iri.rl'lrsis ( Ki lian): specimen no. 28/ 13055. lateral vi ew; village of Nanikovo; gramlis Subzone. Fig. 6. Delplrind!a ohtUSI'/I(){!OSll (Retowskil; specimen no. 1311)05): (6a) ventral view. (6h) aper1ural view. (6c) lateral view: Cape SI. Ilya: gmndis Subznl1e. Figs. 7- 9. Ol"philrel/a uil/J/'lIsix (Bu rc khardt ): (7) specimen no . .~I t 3(5). lateral view: Cap..: SI. Ilya: gmm/iI SUblOlle; 8) .~pccimen 110. 4/1305.'i: (ga) ventral view. (Xh) brteral vi ew ; Barakol Valley: ,~mlldi.{ Sub/.one; (9) specimen no. 6/ 130.'i). lateral view: village of Suhanovka: g/"lllllfi.\· Suh/.onc . •' il-: ' tU . {)d/J llindlajllllus (Retowskil; spec im en no. 341130.'i5. !:ttcral view: Cape SI. lI ya: gmlldix Suhl.llne. !'A1.EONTOl.(l(iICM. J(ll IRNA I. V"I. .N Np . .'i ::! 1I0~ 494 ARKAD1 EV. 130GDANOVA body chamber arc separated by a smooth band. Rib ter Dim e n s io n s in m m. rat i 0 s, and n u m - minat ions are curved antcriorly. b e r o f whorls p e r half wh o rl: Dim e n s ion s in mm, ratio s, and num Number of ribs ber o f who rl s p e r half wh o rl: E Speci men E E per hal r whorl 00. Number of rihs E 3 3 "'i' ii" ii" inner ouler Specimcn E E E per half whorl '" => 3 3 => 00. " t9113055 63" 28 18.5 0.44 0.29 27 60 E "- ii ii" inner outer " " '" => '" => 201 13055 62 J2 13 10.5 0.52 0.2 1 0.17 24 55 13/1305551" 20.5 9.5" 16.50.400.19" 0.32 22 42 " " " " 21113055 39 18 11 0.46 0.28 24 48 141!3055 26.5 I I 7 0.42 - 0.26 25 48 22113055 27 14 8 0.52 0.30 2J 47 Co m par i so n. This species differs from D. sub chaperi in the less strongly pronounced umbilical Co m par i s 0 n. This specie s differs from the sim ridges and weakly developed lateral nodes. ilar, densely ribbed spaces D. crimensis in the sparser and straighter ribs in the inner whorls (up to Om = 25- R e m ark s. Khimshiashvili (1976) described 30 mm) and also, to a lesser extent, in the smoothening D.obruSelZodosa from the Northern Caucasus. The of the ribbing on the body chamber. attentive study of hi s paper shows that the only speci men from the Caucasus possesses tripl ed ribs and has o c c ur r e n c e. Berriasian, jacobi Zone of the no smoothening of the ornamentation, femures that are Crimea (gralldis Subzone), southeastern France, and not characteristic of D. obtuseflodosa. Because of this, Spain. the authors of this paper do not place th e spec imen from M a t e r i a I. Nine specimens (nos. 19/ 13055- the Northern Caucasus in the synonymy list. 27113055) from Cape St. Il ya, Barakol Valley, and the O cc urre nce. Berriasian, jacobi Zone of th e villages of Nanikovo and Sultanovka; coil. by Crimea (gralldis Subzone); Bulgaria, southeastern y.y. Druschits, T.N. Bogdanova, and VY. Arkadiev. France, and Spain. M a t e r i a I. Six specimens (nos. 13/13055- 18/1 3055) Delphinella delphinensiJ (Kilia n, 1889) from the Cape St. Il ya and from the vi llage of Sul tanovka; col I. by T.N. Bogdanova. Plate 6. fig. 5 Hop/iles de/phinensis: Kilian. 1889. p. 662. fig. !. Hop/iles ca/islo: Rctowski. 1893, p. SS. pI. 3, fig. 1, non Hop D<.>lphinellll IrtSllnt nsis Le Hcgaral, 1911 lires calislo var. de/phinensis: Rctowski. 1893. p. 57, pI. 3. fig. 4 Plate 5. figs . 4 and 5 (= D. crime/uis). f/op/iles occilanicus: Relowski. 1893. p. 60. pI. 3. fig. 9. Berriasella de/phinensis: Ma:tenot. 1939. p. 67. pI. 6. figs. 14 non figs. 7 and 8. and 15; Amould -Sagel. 1953. p. 45. pI. 4. figs. JO and Il.!cx[-fig. 14. De/phinel/a lresanensis: Le Hegarat. 1971. p. 113. pI. 13. De/phi/Jella de/plrinensis: Le Hl!garal. 1971. p. 104, pt. 13. fig. 15; pI. 42. figs. 1 and 2. figs. 7 and 8; pI. 42. figs. 3 and 9: Nikolov. 1982, p. 86. pI. 20. Berriasella (Deiphinel/a) lresanensis: Hocdemaeker. 1982. figs. 2-4. non Berriasella (DelphinellaJ cf. de/phinensis: Sapunov. pI. 1. fig. 4. 1977, pI. 6, fig. 5; 1979. p. 177. pI. 56. fig. 7 (= De/phil/el/a mol/ovi , S h e 11 s h ape. Shell s are medium-sized, flat Nikolov). tened, wi th high moderately expanding whorls. The She 11 s h a p c. She ll s are small (mi croconches). flanks are weakly convex. slightly diverging toward the with slowly ex panding whorl s. The flan ks are flattened . umbilicus. The venter is narrow, flat , and smooth. The The venter is flat. The umbilicus is shallow and wide. umbilicus is shallow, moderatel y narrow, with low The umbilical wall is narrow. indistinct, and gently slop sloping wall s. The umbilical shoulder is very rounded . ing. The whorl cross secti on is high, rectangular to oval. Or n a m e n tat i o n. The ribbing is thin and den se. o rn a men ta t i on. The flanks are covered by The ribs are weakly curved, paired. tripled (bundled), straight or weakly curved bifurcating or. less com and occasionally single. In the tripled rib. the bottom monly, single ribs. The whorls 20-30 mm in diameter branching occurs somewhat above the umbilical shoul + possess weak swelling and flatt ening of ribs. A general der. One of the branches again fork into two branches smoothening of the ribbi ng is developed above the mid slightly below the middle of the flank. Paired ribs, flank . The ribs do not cross the venter, on which a wide branching on the umbilical shoulder or subdividing smooth band is devel oped. approximately in the mid-flank are present. Sparse sin Dimen s i o n s in mm. ratio s, and num gle ribs, appearing near the umbilical shoulder, arc b e r of whor l s p e r half whorl: present. Whorls about 60 nUll in diameter show weak ening of the ribbing in the mid-flank. which is clearly Number of rihs E vis ible in the specimen fi gurcd by Retowski ( 1893. Specimen E E pe r half whorl pI. 3. fi g. 9). All ribs arc flatt ened toward the pcriphery no. E ~ "'i' ii" ii" imlcr ()u[cr and arc interrupted 011 the vcntrolateral shoulder. It.::av "~ " => ~ => ing thc vcntcr smoot h. 1Xf1 .~05 .~ "41 15.5 " 11 .~ (Ut-: " 0,:\0 25 41i GENUS OELI)/IINEtLA (AMMONOIDEA) FROM THE 3ERRIAS IAN 49, Co III par i so n. This species differs from most D ime n s i o n s in mm. r at io s, and n um species in the smaller size (microconches) and early ber o f w horl s per h a Ir whorl: srnootheni ng of the ribbing. The latter character is very weakl y developed. Numhcr ofribs Specimen E E R em a rk s. The study of Retowski 's collection ~ 0 0 per half whorl '0. ~ ~ showed that the specimen of Hoplifes caliSlO figured by E ~ :> -~ inner oUler 0 ~ ~ ::> ~ ~ ::> him (RelOwski . 1893. pI. 3. fig. I) has the ribbi ng '" 3 1113055 66 25.5 ]I 21.5 0.39 0. 17 0033 28 60 smoothened on the body chamber and. hence. should be 321 13055 18.5 9 0.36 0. 17 assigned to D. deipliillensis. Previously Arkadiev and - 25 Bogdanova (2004) identifi ed thi s specimen as Berria )3113055 " 19 9.218 0.36 0. 17 0.34 25 53" sella ca/isto. The variety delphillellsis. described by " RelOwski. 1893. shows va rious types of ribs (t riple. bid Co m par i so n. This species differs from D. del ichOlomous, etc.). which does not agree with the diag phill ensis in the larger size and more intense ribbing, nosis of the species deiphillellsis given by Kil ian wh ich become smoother later in ontogeny. It is distin (1889). Triple ribs are present in B. (D.) cL delphillell guished from the closely related D. crimensis by the sis. described by Sapunov ( 1977. 1979). which appar less dense and more crestlike ribs on the body chamber. entl y instigated Nikolov ( 1982) to recognize it as a new o c c ur r e n c e. Berriasian, jacobi Zone of the species. D. moflovi. Crimea (g randis Subzone). Bu lgaria , and southeastern o c c ur r e n c e. Berriasian. jocohi Zone of the France. Crimea (gral/dis Subzone). Bulgaria. southeastern M ate r i a I. Eight specimens (nos. 3111 3055- France. Spain, and Tunisia. 38113055) from Cape SI. lI ya, Barakol Va lley. and the M ate r i a I. Three specimens (nos. 28/13055- vi llages of Sultanovka and Nan ikovo; colI. by 3011 3055) from the Cape SI. lI ya, Barakol Va lley, and T. N. Bogdanova, Y.Y. Dru sch its, and V.V. Arkadiev. lhe village of Sultanovka; coli . by V. V. Druschits and T.N. Bogdanova. Ihlphilltlla ptclinata Arkadie .. et 8ogdnn D ~· II. sp. no ... Platt 6. figs. 1 and 2 Dtlphintllajanus (Relowskl . 1893) E l y m 0 l og y. From the Latin pectinarus (pecti Pl ate S. figs. 1-3; Plme 6. fig . 10 nate). Hoplittsjmuu: RClowski, 1893, p. 59, pI. 3, figs. 5 and 6. H o l o t Y p e. TsNIGR Museum, no. 39113055; Berriastlla aff.jmUls: MazenOl, 1939. p. 70, pi . 6. figs. 20 and 21. Feodosiya, Cape SI. lIya. Berriasian, jllcobi Zone, Delpilinellajamu: Nikolov. 1982. p. 88. plo 21. fig. 5. gramlis Subzone. Delphinella dd(llzinensis: Bogdanova et al., 1984. pI . 2. fig. 3. She I I s h a p e. Judg ing from two fragments of S h e II s h a p e. The she ll s are discoidal and adult whorls. Ihe shells are discoida l, with wide flal medium-sized. The whorls are high. weakl y overlap tened fl anks and narrow ven ter. pin g. and slowly ex pandi ng. The fia nks are very weakl y Or n a III e n t at i on. Two fragments of a quarter of c-onvex. The venter is narrow and weakl y convex. The a whorl possess a distinctive ornamen tation. Six inner umbilicus is wide and shallow. The umbilical shoulder ri bs arc present. They begi n on the umbilical shoulder is rounded. The umbilical wall is narrow and steep. wi th small spiky nodes and slan t orad. Somewhat above Orna m e nt a ti o n. In juvenile whorl s (up to one-third of the height of the whorl a crestlike swelling Dm = 40 mm), the fiank s possess va ri ous ribs. mai nly on the fl anks is present. Ribs near nodes are naITOW and paired and tripled (bundled) and. less commonl y. bidi sharp and, after entering the swelling. become fiat chotomous and single. The ribs begin in the middle of tened. wide. and indistinctly bifu rcate. The swelling the umbilical wall. Their terminations are directed subdivides the fl ar:k in to two unequal pans. Before the sharply forward. On the umbilical shoulder. they form swell ing. the thickness of the shell increases to become an arc di rected backward, whereas on the fi anks again slightly reduced later. After the swelli ng. a wide depres slant forward. The ribs run straight across the flanks. sion is present on the shel l. Hardly no ticeable ribs run The first branching occurs on the umbilical shoulder; across the depress ion, slanting backwards. Near the the upper branching is in mid-fl ank . In the upper pall of vent rolateral shoulder, the ends or the ribs become the whorl . a ll ribs become flatt ened. and soon after sharp. distinct. and thin. They are then turned fo rward crossing the vent rolateral shoulder di scontinue to leave again . An additional two or three ri bs appear belween a smoot h band on the venter. The shape of the ribbing the bundles of ribs. The number of exte rnal ribs on a on the body chamber at Om = 50- 60 mm changes. quarter of the whorl reac hes 25. On the ve nter, the ribs Paired ri bs prevail. and single ribs arc less common. In are intermpted by a narrow smooth band . the Illiddle part. the ri bs smooth CIl: hence. the tipper Co III par i so n. Th is spccie$ di ffers from DeI br.mching is indistinct. The frequency of ribbings com phillel/a in the very distincti ve orn amen tation. pared to younge r Slages is reduced. The ribs become o c c tI r r e n c e. Berrias ian . jac:o/Ji Zone, Kramlis th icker and sparser. Suhzont! of the Crimea. 496 ARKADIEV, BOGDANOVA M ate r i a I. Two speci mens (nos. 39113055 and Bcrriasiiln , jacoh; Zone of the Crime;] (grandiJ Sub 40113055) fro m Cape SI. lIya: coli. by v.Y. Arkildiev. zone), Spain, and Tuni sia. Materia l. Two specimens (nos. 41 / 13055- Genus Herriaselfa Uh lig, 1905 42/ 13055) from Cape St. llya and from the Barakol Val Rtrri.o.stlla btrtht'; (Toucas.. 1890) Icy; coil. by T N. Bogdanova. Plate 6. fi gs. 3 and 4 lIoplites callislO ... ar. her/hei: Toucas. 1890, p. 601, pI. 17. CONCLUS IONS figs. 6 arid 7: Rctowsld. 1893. p. 55, pI. 3, fi g. 3. Berriasdla /x!rtirei: Mazcnot, 1939. p. 48. pI. 2. figs. 9 and 10. As a resuh of th is study, a rich assemblage of Del· non figs . 11 and 12 (= D. sevtl1ieri): Amould-$aget. 1953. p. 26. phinella species (seven spec ies, including D. subcha· pI. 2. I1gs. 12 and 13; pI. 3. fi g. 3: Le HcgarJl and Remanc. 1968. pI. 5. fig . 5. peri, D. crimensis, D. obtll senodosa, D. lres(lnensis, De/p /rinella bertllt!i: Le Hcgarat. 1971. p. 99. pI. 13. figs. 3-5: D. deipliinensis, D. janus. and D. pecliflOta) has been pI. 42. figs . I1 and 12: Nik olo.... 1982. p. 87. pI. 21. figs . 2-4. described for the fi rst time from the Berriasian of the Berriasdla (BerriasellaJ herthe;: Ta ...era, 1985, p. 248. pt. 34, Crimean Mountains. This assemblage a ll ows the recog fi gs. I arid 2. text.fig. I9IH. nition of the standard jacobi Zone in the section and 5 h e ll s h a p e. The shells are small (micro- correlation with southwestern France (Le Hegara t, conches), fl attened, wit h high, moderately ex pandi ng 197 1), Spain (Ta vera, 1985), Bulgaria (Nikolov, 1982), whorls. Flanks are weakl y convex . The venler is fl at CaucaslI s (Khimshi ashvili, 1976, 1989), and other tened. The umbi licus is wide, with low, gently or areas of the Mediterranean region. steeply sloping wa ll. The cllOmeracellsis Subzone, recognized by Ark Or n a me n ta t ion. The flanks possess relati ve ly adiev in the Feodosiya section in the lower part of the coarse, straight. mainly bifurcating ribs. Branc hing jacobi (Arkadiev and Savelieva, 2002; Arkadiev, 2003), occurs in the middle part of the flan ks. Sparse single ribs is presently characteri zed only by members of the gen beg inning from the umbilicus are present. On the body era Berriasella and Fauriel/a. De/phinella have not chamber, al the place where ribs are branching, crestl ike been fou nd in thi s subzone. In the Crimea all records of swellings are developed. The ribs become coarser, fl atter, thi s ge nu s are restricted to the grandis Subzone, where and sparser. The rib tenninations on the ventrolateral they have been found together with Pselldosubp/anites, shoulder are weakly curved forward. On the venter the Berriasella, and RelolVskiceras. The study by Arkadiev ri bs arc interrupted by a narrow smoOlh band. of the section of the jacobi Zone in the bas in of the Dime n s i o n s i n mm, rati os, and num Tonas Ri ver in 2003 showed that very few Delphiflella b e r of who rl s p e r ha lf w h o rl: occur with Pseudosllbp/aflires grondis and other Pse/ldoplmllllites, i.e., they are also characteri stic onl y Number of ribs of the grandis Subzone. Furthermore, in the lower part E Specimen E E per half whorl of the Tonas section, below the levels wi th Pse/ldosllb "0. ,. e " p/anites grandis, the species Berriasella jacobi was E ,. ~ "~ ,.'" => ,.'" ,. => inner outer found. This allowed the subdivision of thejacobi Zone 4 tll 3055 "45 17.5 '"- 16 0.39 0.36 into two subzones. jacobi and grandis, rather than 421 13055 Jl 12 6.5 9 0.39 0.21 0.29 20 39 chomeracellsis and gram/is, as was suggested prev i ously (A rkadiev, 2003). Co rn par i so n. This species is dist in gui shed from ot her species by the type of ri bbing near the venter and For southeastern France Le Hegarat ( 197 1) noted slight swell ings at the points of rib bifurCal ion. that D. crimensis, D. delphillensis, D. oblllsenodostl, D. subchaperi, and D. Iresanensis are characteri stic R e in ark s. This species has a conflicting set of only of the jacobi Zone in Le Hegarat's terminology. characters. On the one hand. it shows a weak thickening i.e., fo r the lower part of the jacobi Zone in its modem and flatteni ng of the ribs on the body chamber, which is understanding (Hoedemaeker and Bulot. 1990). No characteristic of the genu s Delphinella; on the other separation of the find s of De/phinel/o and Pseudosub hand , the ribbing (pai red and sing le ribs) and the p/anites have been so far found in the Crimea, perhaps, absence of smoothening of the ri bbing suggest it s affin because a ll the sections stud ied (Cape SI. lIya, Sul ity 10 the genus Berriasella. Such a combination of tanovka, and Nan ikovo) are incom plete. They do nOl characters certainly indicates a close connection ex pose deposits covering thejacobi Zone in a complete between these genera. Wc decided to leave the species section. Hence, wc can correlate the Crimean gram/is berrhei in the genus Berriasella. However, its precise Subzone onl y with the en tirejacobi Zone. However, the diagnost ics will be possible onl y when new dala on the analysis of ammonites from Retowski 's co llection morphogenesis of the shell s and the sUlura l out line a llows the recognition of the overlyi ng occitallica Zone become ava ilable. in th l! Feodosiya Secti on (Bogdanova er (11., 1999). o c cu r r e n cc. Bcrri'l" ian .jllcohi- occ:ifllllic{I Zones Most likel y. deposits of th is zone arc presentl y in the (.~ III}{/lpi"(I Subzone) soutllc;tstcrn Franc\!, Bu lgaria; bu il! up part of the town and are not exposed. I'ALt:ON']'OI.(XiICAL JOURN AL vU!. W Nu. S 200S GENUS fJ ELPIIINEU..II (AMMONOlDEA) FROM THE BE RRIAS IAN 497 We emended the di agnosis of the genus De/phille/lo, ian along the Ri o Argos (Caravaca, SE Spain)," Scripta according to w hich the s pecies bertltei is assigned to the Geo!. 65,1--8 1 (1982). genus Berriasellll. 12. P. J. H04.:demacker and L. Bulm, "'Preliminary Ammonite A close connection of the Berriasian marine basin of Zonation for the Lower Cretaceous of the Mediterranean Region," Gcol. Alp. 66, 123-127 ( 1990). the C rimean Mountains ex.isted with the western Euro pean basins. Of 15 s pecies of the genus De/phinel/o 13 . M. K. Howarth, ''Tithonian and Berriasian Ammoni tes fro m the Chia Gara Formation in Northern Iraq," Palae known now, seven are recorded from the C rimea, of ontology 35 Part 3, 597- 655 ( 1992 ). whic h five arc shared with France. 14. N. G. Khimshiashvili. AmmOlloidea of the Tifhoniall and Berri(lsi{/II of the Ca ucasus (Metsniereba, Tbilisi , 1976) ACKNOWLEDGM ENTS (in Russian ]. 15. N. G. Khimshiashvi li , Berriasellids of the Caucasus The study was supported by grant E02-9.0- 111 of (Metsni crcba, Tbi lisi, 1989) (in Russian ). the M inistry of Education o f R ussia in the Field of Nat 16. W. Kilian, "Mission d'Andalousie. 2. Etudes paleon ural Sciences. tologiques sur lcs terrai ns sc<:ondaire et tertiaires de l' Andalousie," Mem. pres. par divers savants Acad. Sci. Inst. Francc 30, 601 - 75 1 ( 1889). REFERENCES 17. G. Le Hegarat, "Perisphi nctidae et Beniasellidae de la I . V. V. Ar kadicv, "The Berriasellajacobi- Pseudosubp/all lim ite Jurassique Crctace. Genres nouveaux et rcvision ires grandis Zone of the Berriasian in the Crimean critiquc de quelques defi nitions taxonomiques Mountains," Byull. Mosk. O- va Ispyl. Prir., Otd. Geo!. anterieures," C.R. Acad. Se i. Pari s. Scr. D 273 (10), 850- 78 (6). 29- 35 (2003). 853 (197Ia). 2. V. V. Arkadiev and Vu. N. Savelieva. '''rnejacobi-grall 18. G. Le Hcgarat, "Le Berriasien du Sud-East de la Francc." dis Zone of the Berriasian in the Crimcan Mountai ns" in Doc. Lab. Gcol. Fac. Sci. Lyon, No. 43/1. 1-309 Abstracts of Papers of the 48th Session of the Pa/eol!1o ( 1971b). logical Society 0 11 Problems of BiochrrJllology ill Pale 19. G. Le Hegarat and 1. Remane. "Tithonique superieur el ontology and Geology, St. Peursburg. April 8-12. 2002 Berriasien de I' Atdeche et I' Heraull. Correlation des (SI. Pctersburg, 2002), pp. 11-13. ammonites et des calpionell cs," Gcobios, No. I, 7--69 3. V. V. Arkadiev and T. N. Bogdanova, "Genus Berriasella ( 1968). (Ammonoidea) and Ammonoid Zonation in the Benia sian of the Crimean Mountai ns," Stratigr. Geo!. Korrc 20. N. P. Luppov, V. I. Bodylevskii, and A. E. Glazunova. Iyatsiya 12 (4),54--67 (2004) (Strati gr. Gcol. Corrc lation "Ammonites," in Atias of /IJdex Forms of/he Fossil Fau 12 (4). 367-379 (2004)). IIQS of th e USSR, Vo!. 10: Lower Cretaceous Series 4. S. Arnould-Saget, "us ammonites pyritcuses du Titho (Gostoptekhizdat, Moscow. 1949), pp. 183- 253 [i n Rus nique superieur et du Berriasien de Tunisie centralc:' sian ]. Ann. Min. Gcol. Publ. Serv. Geo!. Tunisie, No. 10. 132 21. G. Malenot, "Les Palaeohoplitidae Tithon;queset Berri (1953). asiens du Sud-Est de la France,'" Mem. Soc. Geol. 5. T. N. Bogdanova. E. D. Kalacheva. and I. I. Sei, "On the France, N. Scr. 18 (Fasc. 1-4), 1-303 ( 1939). Presence of the 7imovelfa ocdranica Zone (Lower Cre 22. T. G. Nikolov, us ammonites de la fa mille Berriasel taceous. Berriasian) in the Feodosiya Section of Eastern lit/ae Spalh, 1922. Tithonique sJlperieur-Berriasien Crimea," Reg. Geol. Metallog., No. 9, 27-32 ( 1999). (Sofia, 1982). 6: T. N. Bogd,"o,". S. V Lob"" h",. V. A. Pmzomd;;. 23. O. Retowski, "Die tithoni se hcn Ablagerungen von The and T. A. Favorskaya, "On the Subdivision of the Berri odosia," Bull. Soc. Natur. Moscou, N. Scr. ' (2-3),1-95 asian Stage in the Cri mean Mountains," Ve stn. Lcningr. ( 1893). Univ., No. 1,5-14 (198 1). 24. I. G. Sapunov, "Ammon ite Stratigraphy of the Upper 7. T. N. Bogdanova, S. V. Lobatscheva, V. A. Pr07.0 rovskii, Jurassic in Bulgaria. 4. Tithonian: Substages, Zones, and and T. A. Favorskaya, "The Berriasian of Eastern Crimea Subzones," Geol. Balcan. 7 (2), 43-64 (1977). and the Jurassic-Crctaceous Boundary," in BOllndary 25. I. G. Sapunov, Lesfossiles de Bu/garie. Ill. J. Jurassiqlle Stages of Ihe Jurassic-CreraceOllS Systems (Nauka. sllpirieur. AmllJolJ oidea (Acad . Bulg. Sci., Sofia. 1979). Moscow, 1984), pp. 28-35 1in Russian]. 26. Simionescu, "Note sur quelques ammonites du Nco 8. C. Burckhardt, " Faunesjurassiqucs et crctaciques de San 1. Pedro dd Gello," Bol. Inst. Gco!. Mexico. No. 29 comien fram,a is:·Trav. Lab. Gcol. Fac. Sci. Uni v. Grcno ( 191 2). bIc 5 (Fasc. I). 1- 17(1899-1900). 9. v. V. Druschits, "Cephalopod Moll usks: Ammonites, 27. 1. M. Tavera, Los ammonites del fith 6nico superior-ber Part I ," in A//as of the Lower CreltlCeOllS F(/fl//{/ of fhe ria.rell.re de la ZO/Ja SJ/ bbitica (Comilferas Biticas) Nor/hem Caucasus and Crimea (Gostoptckhi zdat. Mos (U niv. Granada. Granada, 1985). cow, 1960), pp. 249- 308 [in Russianl. 28. A. Toucas, "Etude de la faune des couches tithoniques de 10. O. K. Grigorieva, "Lower Valanginian Ammonite Fauna I' Ardcche," Bull. Soc. Gco!. France. Ser. 18. 560-630 from the Ddaya River Basin on the Nort hern Slope of ( 1890). the Caucasus Mountains," in Data 011 Geology ali(I Mill 29. C. W. Wright. J. H. Cnlloman. and M. K. Howart h. Trea· eral R es() ll rce.~ (A1.Ovo -Chernomorskii Geo!. Trest. ti.fC OIl IlJ l'ertebrme Pafeollfology. Pt L M (llil l.~ca 4. 193R), Issue I, pp. R~ - I 22 lin Russi:ml . Re \ ~. Vo!. 4: C retaceo ll .~ AIIIIIWlloi(I('(l (Geol. Soc. Am. 11 . P. J. HocdemOleker. "Amma nile Biostfatigraphy of the i\J\d Univ. Kansas Press, Bou lder, Bou lder- Lawrence. Uppermost Tithonian, Berriasian, and Lower Val:lIl~in~ 1".1".16) . I 'AIH ' ~ ruLll(j[CAI. J{)UI~0..\L \"01. .t" :0.,, ).:; .! O(J.'i