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Jacob Gardner Curriculum Vitae jacobdgardner.com March 2021 Department of Earth Sciences State University, Bozeman, MT 59717 [email protected] | +1 805 801 9197

Education & Training 2016– Ph.D. in Earth Sciences, Graduate Certificate in Applied Statistics, Department of Mathematical Sciences Dissertation: How Sexual Selection Shapes Macroevolution

2012–15 B.Sc. in Earth Sciences: Paleontology Option, Montana State University Geology field school: Indiana University Judson Mead Geologic Field Station, G429: Subsurface Stratigraphy Concentration

Appointments 2016– Graduate student researcher MSU Macroevolution Lab (PI: Chris Organ), Montana State University

2020– Graduate teaching assistant Department of Microbiology and Immunology, MSU

2019–20 Visiting research assistant Laboratory on Molecular Paleontology (PI: Qiaomei Fu) Voted top 10 in China’s scientific progress of 2020 Institute of Vertebrate Paleontology and Paleoanthropology, CAS

Freelance scientific editing Earth, Atmospheric, and Planetary Sciences, Cactus Communications

2016–19 Graduate teaching assistant Department of Earth Sciences, MSU Department of Cell Biology & Neuroscience, MSU

2014–15 Lab technician Gabriel Laboratory for Molecular Paleontology, Museum of the Rockies

2012–13 Lab technician Whitlock Paleoecology Laboratory, MSU

Fellowships & Internships 2017 National Science Foundation, East Asia & Pacific Summer Institutes Fellowship Host: IVPP, CAS

2015 Paleontological Society, Student Ambassador

2014 Smithsonian National Museum of Natural History, Research Internship

Teaching & Student Engagement Courses Taught Sp. 2018 PSPP 594 – Graduate Seminar on Phylogenetics, MSU Co-taught by Dr. Matt Lavin, Dr. Chris Organ, Frank Etzler, and myself

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Teaching Assistantships (Avg. Instructor Rating = 4.84 / 5.0, n = 176 students) 2021 GEO 413 – Macroevolution & The Fossil Record, STA*, MSU

2020– BIOH 201 – Human Anatomy and Physiology I, GTA*, MSU (4.86, 150)

2016–19 –

2016 ERTH 101 – Earth System Science, GTA, MSU (4.82, 26) GEO 310 – Invertebrate Paleontology, STA, MSU

2015 BIOO 310 – Comparative Vertebrate Anatomy, STA, MSU * STA: Student teaching assistant (unpaid); GTA: Graduate teaching assistant (paid)

Undergraduate Mentorship Carolyn Kocken (2018–19), Rudy Hummel and Thomas LaBarge (‘17–19), Gia Fisher (‘16–19), Kevin Surya (‘16–19), Kevin Jones (‘16–18), Mikayla Struble (‘16–17)

Publications *contributed equally

In Review • Gardner, Jacob D., and Mark V.H. Wilson. A Mostly Complete Bowfin (Amiidae: Amia sp.) from the Coal Creek Member of the Eocene Kishenehn Formation, Northwestern Montana. In review for Acta Palaeontologica Polonica

• Krumenacker, L.J., David J. Varricchio, Chris L. Organ, Jacob D. Gardner, Clint Boyd, and Brooks Britt. Osteology and Phylogeny of the Neornithischian Oryctodromeus cubicularis from the Middle Cretaceous (Albian-Cenomanian) of Montana and Idaho. In review for the Journal of Vertebrate Paleontology

• Varricchio, David J., Jason D. Hogan, and Jacob D. Gardner. Troodontid specimens from the Cretaceous Two Medicine Formation of Montana (USA) and the validity of Troodon formosus. In review for the Journal of Paleontology

Peer-Reviewed Articles 5. Wang, Wenjun*, Manyu Ding*, Jacob D. Gardner*, Yongqiang Wang*, … [25 authors] …, Vikas Kumar, and Qiaomei Fu. 2021. Ancient Xinjiang Mitogenomes Reveal Intense Admixture with High Genetic Diversity. Science Advances. DOI: 10.1126/sciadv.abd6690

4. Gardner, Jacob D., and Chris L. Organ. 2021. Evolutionary Sample Size and Phylogenetic Models of Correlated Evolution. Systematic Biology. DOI: 10.1093/sysbio/syab017

3. Gardner, Jacob D., Michel Laurin, and Chris L. Organ. 2020. The Relationship Between Metabolic Rate and Genome Size in Extant Vertebrates. Philosophical Transactions of the Royal Society B, 375(1793): 20190146. DOI: 10.1098/rstb.2019.0146 bioRxiv DOI: 10.1101/659094

2. Gardner, Jacob D.*, Kevin Surya*, and Chris L. Organ. 2019. Early Tetrapodomorph Biogeography: Controlling for Fossil Record Bias in Macroevolutionary Analyses. Comptes Rendus Palevol, 18(7): 609-709. DOI: 10.1016/j.crpv.2019.10.008 bioRxiv DOI: 10.1101/726786 Nature Ecology & Evolution Behind the Paper: go.nature.com/2TZDiLA Jacob Gardner, CV | 2

1. Wilson, John P., D. Cary Woodruff, Jacob D. Gardner, Holley M. Flora, John R. Horner, Chris L. Organ. 2016. Vertebral Adaptations to Large Body Size in Theropod . PLoS ONE, 11(7): e0158962. DOI: 10.1371/journal.pone.0158962

Conference Abstracts 19. Struble, Mikayla K., Jacob D. Gardner, and Alice Gibb. 2020. Grasping Behavior in Birds Drives Pedal Adaptations. Society for Integrative and Comparative Biology. Oral

18. Hummel, Rudy R., Jacob D. Gardner, and Chris L. Organ. 2018. Origin and Dispersal of Gorgonopsia: A Phylogenetic Approach to Permian Therapsid Biogeography. Poster

17. Wilson, John P., Ashley Ferguson, and Jacob D. Gardner. 2018. ‘Pendular’ Swings in the Evolution of Centrosaurine Cranial Ornamentation. Society of Vertebrate Paleontology. Poster

16. Struble, Mikayla K., Jacob D. Gardner, and Alice Gibb. 2018. The Evolution of Grasping Behavior in Birds and Associated Pedal Adaptations. Society for Integrative and Comparative Biology. Oral

15. Gardner, Jacob D., and Chris L. Organ. 2017. The Macroevolutionary Dynamics of Vertebrate Genome Size. Geological Society of America Annual Meeting. Oral

14. Surya, Kevin, Isabelle M. Brenes, Jacob D. Gardner, Lázaro W. Viñola López, Chris L. Organ, and David J. Varricchio. 2017. Pelvic Coevolution with Egg Size and Shape in Modern Birds: Implication for Reproductive Biology of Extinct Dinosaurs. Geological Society of America Annual Meeting. Oral

13. Gardner, Jacob D. and Mark V. H. Wilson. 2017. A Mostly Complete Bowfin (Amiidae: Amia sp.) from the Coal Creek Member of the Eocene Kishenehn Formation, Northwestern Montana. Society of Vertebrate Paleontology. Poster

12. Struble, Mikayla, Jacob D. Gardner, and Chris L. Organ. 2017. Biomechanical Stresses of Pedal Grasping Behavior within Modern Aves: Morphological Adaptations and Mesozoic Implications. Society of Vertebrate Paleontology. Oral

11. Surya, Kevin, Isabelle M. Brenes, Jacob D. Gardner, Lázaro W. Viñola López, Chris L. Organ, and David J. Varricchio. 2017. Pelvic Sexual Dimorphism in Modern Birds (Aves: Neornithes) and Its Evolutionary Relationship with Relative Egg Size. National Conference for Undergraduate Research. Poster

10. Jones, Kevin, Jacob D. Gardner, and Chris Organ. 2017. The Evolution of Human Pregnancy. National Conference for Undergraduate Research. Poster

9. Gardner, Jacob D., Chris L. Organ. 2016. Discrete Models of Correlated Evolution are Prone to False Positive Results. Society of Vertebrate Paleontology. Oral

8. Struble, Mikayla, Jacob D. Gardner, David J. Varricchio, and Chris L. Organ. 2016. Convergent Evolution and Biomechanics of the Raptorial Foot. Society of Vertebrate Paleontology. Oral

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7. Gardner, Jacob D. and James G. Schmitt. 2016. Mineralogic Evidence of Preservational Conditions in the Oligocene Canyon Ferry Insect Lagerstatte, Southwest Montana. National Conference for Undergraduate Research. Poster

6. Gardner, Jacob D. and James G. Schmitt. 2015. Mineralogic Evidence of Preservational Conditions in the Oligocene Canyon Ferry Insect Lagerstatte, Southwest Montana. Geological Society of America Annual Meeting. Poster

5. Gardner, Jacob D., D. Cary Woodruff, John P. Wilson, Holley M. Flora, John R. Horner, and Chris L. Organ. 2015. Biomechanical Adaptations to Increased Body Size in the Neural Spines of Theropod Dinosaurs. Society of Vertebrate Paleontology. Poster

4. Flora, Holley M., John P. Wilson, Jacob D. Gardner, and Denver W. Fowler. 2015. A Probable Oviraptorosaur from the of Montana. Society of Vertebrate Paleontology. Poster

3. Gardner, Jacob D., John P. Wilson, Holley M. Flora, and Chris L. Organ. 2015. Dinosaur Running Evolved in Episodic Bursts. National Conference for Undergraduate Research. Oral

2. Gardner, Jacob D., John P. Wilson, Holley M. Flora, and Chris L. Organ. 2014. Dinosaur Running Evolved in Episodic Bursts. Geological Society of America Annual Meeting. Oral

1. Gardner, Jacob D., John P. Wilson, Holley M. Flora, and Chris L. Organ. 2014. Accelerated Biomechanical Evolution Precedes Shifts in Locomotion. Geological Society of America Joint Rocky Mountain and Cordilleran Section Meeting. Oral

Honors, Awards, and Grants (Total Awarded Amount = $19,906) • University of CAS Partial Scholarship for International Students (2019-20) • Student Club Match Funding (MSU) for the Earth Science Student Colloquium ($750, 2019) o Awarded after raising $2,000 in donations • Student Club Mass Funding (MSU) for the Earth Science Student Colloquium ($3,389, 2018-19) • Dinosaur Research Institute Student Project Grant ($1,600 CAD, Fall 2017) • Annual Geological Society of America (GSA) Conference Travel Grants (Fall 2017) o MSU Graduate School Travel Grant ($185) o GSA Travel Grant ($90) o College of Letters and Science (CLS) Student Research Travel Grant ($375) • Student Club Mass Funding for the Dead Lizards Society ($1,600, 2017–18) o Organized Society of Vertebrate Paleontology Conference Trip in Calgary, Alberta • Student Club Mass Funding for the Dead Lizards Society ($2,500, 2016–17) o Organized Society of Vertebrate Paleontology Conference Trip in Salt Lake City, Utah o Invited Speaker: Dr. Anthony Martin (Emory University) • Earth Sciences (MSU) Donald L. Smith Memorial Graduate Scholarship ($2,700, 2016) • Undergraduate Scholars Program (USP) Research Grant (Fall 2015) o Description of a Nearly Complete Bowfin (Family: Amiidae) Fossil from the Eocene Kishenehn Formation, Northwestern Montana ($900) o Advisors: James G. Schmitt and Mark Wilson • Annual GSA Conference Travel Grants (Fall 2015) o GSA Travel Grant ($110, Fall 2015) o USP Travel Grant ($500, Fall 2015) • USP Research Grant (Summer 2015) Jacob Gardner, CV | 4

o Comparative Deposition and Fossilization of the Cenozoic Insect Lagerstatten of North America ($616.17) o Advisor: James G. Schmitt • ConocoPhillips Indiana University Field Camp Scholarship ($1,500, Su 2015) • GSA Rocky Mountain Section Undergraduate Research Grant (Sp 2015) o Deposition & Fossilization of the Cenozoic Insect Lagerstatten of North America ($750) • Annual GSA Conference Travel Grants (Fall 2014) o GSA Travel Grant ($150) o GSA On to the Future Travel Grant ($375) o USP Travel Grant ($500) o CLS Student Research Travel Grant ($500) • USP Research Grant (Summer 2014) o Taphonomy and Paleoecology of an Oligocene Lacustrine Insect Lagerstatte, Canyon Ferry Reservoir, Montana ($1,360) o Advisor: James G. Schmitt • Montana State University Letters and Science Dean’s List (2013–15)

Skills • Programming – Amazon Web Services, Bash, C++, Python, R, Unix/Linux, • Phylogenetics – BayesTraits, BayesTrees, BEAST, CIPRES, FigTree, Mesquite, MrBayes, PAUP*, RaxML, RevBayes, Tracer, TreeView, TNT; ape, geiger, MCMCglmm, phangorn, and phytools in R • Population genomics – AdmixTools, ADMIXTURE, admixr, fastsimcoal2, PCA, PLINK, READ • Parametric, nonparametric, and multivariate statistics – R, SAS • Geological and paleontological fieldwork – excavation, mapping, prospection, sedimentological description • Geological and paleontological lab work – fossil preparation, molding/casting, paleohistological techniques, rock and lake sediment core description and processing • Microscopy – histology and paleohistology, petrology, scanning electron microscopy, x-ray diffraction • Dissection – human and non-human vertebrates • ArcGIS

Professional Service • Journal Article Peer-Review (n = 7) o Journals: PNAS, Frontiers, Nature, Nature: Ecology and Evolution, Biology Letters, Integrative Organismal Biology, Philosophical Transactions of the Royal Society B

• Book Chapter Peer-Review (n = 2) o How Do We Know What Dinosaurs Looked Like? (Chapter 15) in: How Do We Know What We Know About Dinosaurs? (1 ed.) by M.H. Schweitzer, E.R. Schroeter, and C.D. Czajka. 2020. o Macroevolution (Chapter 20) in: Evolution (4 ed.) by D. Futuyma and M. Kirkpatrick. 2017.

• Society Memberships: o Geological Society of America o Paleontological Society o Society for the Study of Evolution o Society of Systematic Biologists o Society of Vertebrate Paleontology

• Institutional: o MSU Earth Sciences Annual Colloquium committee (2017–21; Co-Chair: ‘17–18; Chair: ‘18–19) o MSU Earth Sciences Weekly Seminar committee (2016–19; Co-Chair: ‘17–18) o Co-organized talk session on interdisciplinary research on macroevolution for the Montana State University Student Research Celebration. “Macroevolution: The Fellowship of the Tree” (2018) o Visiting researcher visits organized: § Dr. Sara Wylie, Northeastern University (2019) § Dr. Dough Emlen, University of Montana (2019) Jacob Gardner, CV | 5

§ Dr. Tracy Heath, Iowa State University (2018) § Dr. Tony Martin, Emory University (2017) o MSU Dead Lizards Society (2012–Current; Vice President: ‘14–15; President: ‘15–17) o Tennis Club at Montana State (2012–13, ‘16–19; Advisor: ‘18–19)

Invited Presentations/University Events 33. GEO 211 – Earth History and Evolution (Montana State University): Archean Life (Sp 2021). Guest lecture 32. Gardner, Jacob D., Kevin Surya, and Chris L. Organ. 2019. Phylogeography of the Tetrapod Water-Land Transition. Montana State University Fourteenth Annual Earth Sciences Student Colloquium. 31. LaBarge, Thomas, Jacob D. Gardner, and Chris L. Organ. 2019. Phylogeography of the Tetrapod Water- Land Transition. Montana State University Fourteenth Annual Earth Sciences Student Colloquium. 30. GEO 211 – Earth History and Evolution (Montana State University): The Cretaceous Period (Sp 2019). Guest lecture 29. GPHY 411 – Biogeography (Montana State University): Biogeography in Deep Time (Sp 2019). Guest lecture 28. GEO 211 – Earth History and Evolution (Montana State University): Sedimentary Environments (Sp 2019). Guest lecture 27. Gardner, Jacob D., and Chris L. Organ. 2018. The Evolution of Culture and the Rise of Civilization. Montana State University Earth Sciences Graduate Student Rough Cut. 26. Gardner, Jacob D., and Chris L. Organ. 2018. The Evolutionary Dynamics of Vertebrate Genome Size. Montana State University Thirteenth Annual Earth Sciences Colloquium. 25. Hummel, Rudy, and Jacob D. Gardner. 2018. Origin and Dispersal of Gorgonopsia: A Phylogenetic Approach to Permian Therapsid Biogeography. Montana State University Thirteenth Annual Earth Sciences Student Colloquium. 24. Gardner, Jacob D., and Chris L. Organ. 2018. The Evolution of Culture and the Rise of Civilization. Montana State University Student Research Celebration. Topical Session – Macroevolution: The Fellowship of the Tree. 23. Gardner, Jacob D., and Chris L. Organ. 2018. The Evolutionary Dynamics of Vertebrate Genome Size. Montana State University Student Research Celebration. Topical Session – Macroevolution: The Fellowship of the Tree. 22. Hummel, Rudy, and Jacob D. Gardner. 2018. The Evolution and Biogeography of the Carnivorous Mammal Predecessors, Gorgonopsia. Montana State University Student Research Celebration. Topical Session – Macroevolution: The Fellowship of the Tree. 21. Montana State University’s Department of Earth Sciences Seminar – Fossils and Diplomacy: Research and Collaboration in China (Fall 2017). Invited talk 20. ERTH 101 – Earth System Science (Montana State University): Consilience: An Approach to Studying Deep Time (Fall 2017). Guest Lecture 19. ERTH 101 – Earth System Science (Montana State University): Life in the Rocks – How to Study and Interpret the Fossil Record (Fall 2017). Guest Lecture 18. GEO 211 – Earth History and Evolution (Montana State University): Neogene & Holocene (Sp 2017). Guest Lecture 17. GEO 211 – Earth History and Evolution (Montana State University): The Paleogene World (Sp 2017). Guest Lecture 16. Jones, Kevin, Jacob D. Gardner, and Chris L. Organ. 2017. The Evolution of Human Pregnancy. Montana State University Student Research Celebration. 15. Surya, Kevin, Lázaro W. Viñola-López, Isabelle Brenes, Jacob D. Gardner, Chris L. Organ, David J. Varricchio. 2017. Pelvic Sexual Dimorphism in Modern Birds (Aves: Neornithes) and Its Evolutionary Relationship with Relative Egg Size. Montana State University Student Research Celebration. 14. ERTH 101 – Earth System Science (Montana State University): Life in the Rocks – How to Study and Interpret the Fossil Record (Sp 2017). Guest Lecture 13. Gardner, Jacob D., and Mark V. H. Wilson. 2017. The Big Bowfin of Glacier: Biodiversity and Preservation of Eocene Fish in Northwestern Montana. Montana State University Twelfth Annual Earth Sciences Student Colloquium. 12. Surya, Kevin, Lázaro W. Viñola-López, Isabelle Brenes, Jacob D. Gardner, Chris L. Organ, David J. Varricchio. 2017. Pelvic Sexual Dimorphism in Modern Birds (Aves: Neornithes) and Its Evolutionary Relationship with Relative Egg Size. Montana State University Twelfth Annual Earth Sciences Student Colloquium.

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11. GEO 211 – Earth History and Evolution (Montana State University): Sedimentary Environments (Spring 2016, ‘17). Guest lecture 10. ERTH 101 – Earth System Science (Montana State University): Consilience and Deep Time (Sp, Fall 2016). Guest lecture 9. ERTH 101 – Earth System Science (Montana State University): Life, Death, and Evolution (Sp, Fall 2016). Guest lecture 8. Gardner, Jacob D., and James G. Schmitt. 2016. Mineralogic Evidence of Preservational Conditions in the Oligocene Canyon Ferry Insect Lagerstatte, Southwest Montana. Montana State University Eleventh Annual Earth Sciences Student Colloquium. 7. Honors Presents: The Undergraduate Research Experience: A Paleontology Student’s Perspective (Spring 2016). Invited talk 6. FYRE: First Year Research Experience Program talk and Q&A: How to get started on research (Fall 2015). Invited talk 5. Museum of the Rockies Volunteers: An Overview of Research, Fieldwork, and Lab Work Experience (Fall 2015). Invited talk 4. Gardner, Jacob D., D. Cary Woodruff, John P. Wilson, Holley M. Flora, John R. Horner, and Chris L. Organ. 2015. Extreme Rugosities in the Neural Spines of Theropod Dinosaurs Were Biomechanical Responses to an Increased Body Size. Montana State University Tenth Annual Earth Sciences Student Colloquium. 3. Gardner, Jacob D., and Helen Dailey. 2015. Smithsonian Paleobiology Internships: Fossil Insects and Fish from the Kishenehn Formation, Montana. Montana State University Student Research Celebration. 2. Gardner, Jacob D., John P. Wilson, Holley M. Flora, and Chris L. Organ. 2014. Rates of Biomechanical Evolution Preceding Transitions in Dinosaur Locomotion. Montana State University Student Research Celebration. 1. Gardner, Jacob D., John P. Wilson, Holley M. Flora, and Chris L. Organ. 2014. Rates of Biomechanical Evolution Preceding Transitions in Dinosaur Locomotion. Montana State University Ninth Annual Earth Sciences Student Colloquium.

Volunteer/Outreach • Organizer, Fossils Competition, Montana Science Olympiad, Montana State University (2021) • Behind the Paper blog in Nature Ecology & Evolution Community o Geographic Dispersal across the Tetrapod Water-Land Transition (2020) https://go.nature.com/2TZDiLA • Interviewee, graduate film semester project by Harrison Bach, Montana State University (2019) o https://www.youtube.com/watch?v=2LWwvNV56VM • Co-organizer, Fossils Competition, Montana Science Olympiad, Montana State University (2018) • National Fossil Day volunteer, Museum of the Rockies, MT (2018) • Interviewee, Graduate School promotional video, Montana State University (2017) o https://www.youtube.com/watch?v=Y1wAXXAZQvA • Scouts Day volunteer, 5 Dinosaur Families activity, Museum of the Rockies, MT (2017) • Fossil preparation, field work, and exhibits (1,300+ hours), Museum of the Rockies, MT (2012–16) • Organizer of the Annual Paleontology Symposium at Montana State University (2015, ‘16) • Scouts Day volunteer, Make Your Own Dinosaur Dig activity, Museum of the Rockies, MT (2016) • Volunteer Assistant AV-Technician (10 hrs) at the Geological Society of America annual meeting (2015) • Montana State Renne Library Paleontology Exhibit (2015) • SCOPE (Student Community Outreach ProjEcts) (2013–14) o Preparation, field work, exhibits, and curation (200+ hours), Carter County Museum, MT • Big Sky Breaks Away student volunteer assistant leader, Carter Country Museum, MT (Sp 2014) • Destination Imagination volunteer official, Manhattan Elementary School, MT (2014) • Volunteer program assistant (8 hrs) at the GSA Rocky Mountain/Cordilleran Section meeting (2014) • Galapagos Islands/Ecuador Travel project (2013) o Were there dinosaurs in the Galapagos Islands? https://www.youtube.com/watch?v=Ow7ATpqQM1s • Galapagos Islands/Ecuador Travel Website for Montana Elementary Schools (2013) o http://msugalapagos.wordpress.com/ • AmpSurf volunteer: sharing the experience of surfing with veterans and citizens with disabilities (2012)

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