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P" VENUS (Jap. Jour. Malac.) Vel. ,l7, No. 3 (]988):172--IS4

E rilioidea ma cD IJ = = 7" v' >if' -r $;F !x･ e[) Eil EEZI JM as 3 ;Yf mp a) EEI&

IJ . , 7n ･y 7' 7' Vx )-. e7 ;/ F'?L V L･. I-

Transfer of Exilioidea Grant & Gale, 1931 to ,

with Deseriptions of Three New Speeies ()

Philippe BoUCHET (Mus5um National d'Histoire Naturelle, Paris)

and

Anders WAREN

(Naturhistoriska Riksmuseet, Stoekholm)

A.bst,ract: Exilioidea Grant & Gale, 1931 is transferred from the family Buneini-

dae to the Turbinellidae based en its radmlar and shell niorphology, From an

examination of types and representative material of E. 7'ectirostr･is (Carpenter, 1864), E, ketseui (Dall, 1908), and Plicifusus obsoletus Talmadge, 1971, it is concluded that these names are synonyms and based on a northeast Pacifie species

with great, elinal, latitudinal and ba-thymetrieal variation, The East Pacific

abyssal Fttsi{s rufocattdatus Da]1, 1896 is transferred to Exilioidea and three new ･ind･ica, species are deseribed: E. Indian Oeean, 2900-4040m; E. costt.tlata, SE

Asia, le25 m; E. atlant･ica, Gulf ef Mexieo, 880-3365 m.

Introduction

The Exilioidea Grant & Gale, 1931 was described in the family

Neptuneidae, and currently includes two northeast Paeifie shelf and bathya!

speeies

been summarized by Bartsch (1945) who, despite finding no radula, ineluded

the genus in the Fusinidae, following Bentson (1940). Few reeent authors have

been coneerned with Exi･l･ioidea, whieh has rernained plaeed in the Buccinacea

(Palmer 1958, Abbett 1974, Hickman 1980).

Herein we transfer Exil･io･idea to the family Turbinellidae on the basis of

radular and shell morphology; we revise the northeast Pacific representatives of

the genus; we transfer to Emilioidea one additional east Pacific speeies described

as fiik'su.s and we deseribe three new species from the western Pacifie, the Indian 'the Oeean, and Caribbean, all from deep "rater.

Abbreviations used in the text:

RMNJ{--British Museum CNat, Hist.), London

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. Bouchet War6n; Transfer of Exitioidea to Turbinellidae 17/';

CAS-California Academy of Sciences, San Francisco

LACM - Los Angeles County Museum, Los Angeles

MNHN-Mus6um National d'Histoire Naturelle, Paris

USFC-United States Bureau of Fisheries

USNM-National Museum of Natural History, Washington, D.C.

ZMC - Universitetets Zoologiske Museum, Copenhagen - sh (s) shell (s) - spm (s) specimen (s)

St. - station

The radteta･ of Exilioidea: Bartseh (1945) disseeted five speeimens of the type

speeies of E'xilioidea and found no radula. The reason is probably that the radula is very small, its total length is less than 140 sim in the species we have

examined, It is rachiglossate, the central tooth has a broad, arched basa] plate,

with a strong arrow-like central cusp and two mueh smaller outer cusps. The

laterals are simp]e, roughly triangular, with a broad base and a more or less

projecting eusp. The radula has been examined in four of the five known species,

and was found to show little interspeeific differences (Figs. 4-15).

Such a radula is consistent with what is known in the family Turbinellidae

(see for instanee Bouehet & War6n 1985, figs. 389, 390, 393; Harasewyeh 1987,

figs. 17-22) and exeludes Exitioidea from the Buecinidae.

Systematics

Exilioidea･ Grant & Gale, 1931

Type species: ChTysodom'us rectiTostTi･s Carpenter, 1864, by original designa- tion,

･..,".A.-..,t./･''

Figs. 1-3. Opercula of Exilioidea. 1-2. E, costulata, holotype, outerAntonheight and inner views, height 3.2mm. 3. E indica,

Bruun specimen (USNM 717323), outer view, 3.5mm,

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174 vENuS: voL 47, No,3(1988)

Ex;itioidea rectirastris (Carpenter, 1864) (Figs. 4-9, 11, 16-22)

Ch.?'ysodo7n?ts rectirostris Carpenter, 1864: 664.

New synonyms:

Tr'ito7ie,;iLsus (Pliei,fztsus) kelseyi Dall, 1908b: 249.

Pl･ic･ifz{sus (Mic･roft{su.s> obsoletv,ts Talmadge, 1971: 43, figs. 3-4.

Type material: C, reetirostris, holotype USNM 4515. T. kelsegA, holetype USNM

110631. P, obsoletus, holotype CAS 13319.

Type loca･lities: C. rectirostris, Puget Sound, Washington State, USA. T.

kelseyi, off San Diego, California, 90m. P, obsoletbl.s, off northern California,

approximately 41005' to 41e25'N, 830m.

Material examined: The type material of C. TectdeostTis and T. kelseyi and

Alaska: Forrester Id,, 110m, leg. Willett, 3spms (USNM 216335); USFC St.

4230, Behm Canal off Indian Point, 32em, 1spm (USNM 222233).

British Columbia: USFC Sts. 4200, 4201, Queen Charlotte Sound, 200-260m, 5spms (USNM 222187, 222478, 226117: Fig, 18); USFC Sts. 4194, 419Z 4198,

Gu]f of Georgia, 110m, lspm (USNM 222463), 260-360rn, 7spms (USNM

222094, 222608, 223558) ; N end of Vancouver I., 51009'N, 127e56'W, 119 m, 1 spm

(LACM 68-176) ; N end of Vancouver I., 51008'N, 127e45'W, 75 m, 1 spm (LACM

71-219) ; Strait of Georgia, 50006'N, 125e18'W, 65 m, 2spms (LACM 71-218) ;

Strait of Georgia, 49050'N, 124050'W, 350m, leg. McLean & McCowan, 7spms

(LACM 63-17) ; W coast of Vaneouver I. 49043'N, 127e08'W, 90 m, 1 spm (LACM

67-35); W eoast of Vaneouver I,, 48054'N, 126e16'W, 130m, 4spms (USNM

71-221); Royal Roads, Vietoria, Vancouver I., 120m, leg. Riehardson, 1spm

(USNM 108994) ; Vietoria, Vancouver I., 130m, Ieg. Riehardson, 1 spm (USNM 220250).

Washington: San Juan Is,, Puget Sound, 48024'N, 122054'W, 90 m, leg. Shimek,

10 spms (LACM 74-87) ; R. V. Yaquina St. OTB-393, off Willapa Bay, 46a38'N, 124025'W, 85 m, 13 spms (LACM 71-360.3: Figs. 16-17).

Oregon: USFC St. 2882, 46009'N, 124023'W, 125m, 2spms (USNM 106865,

222493); R.V. Yaquina St. OTB-126, off Yaquina Head Lighthouse, 44039'N,

124e5YW, 800m, lspm (LACM 66-253.1); R.V, Yaquina St OTB-140, off

Yaquina Head, 44039'N, 124036'W, 200m, lspm (LACM 66-257.2); R.V.

Yaquina St. OTB-256, off Yaquina Head, 44036'N, 124"56'W, 800m, 3spms

(LACM 68-216.5: Fig. 22) ; R. V. Aeona St. OTB-37, off Yaquina Head, 44034'N,

124e56'W, 800 m, 3spms (LACM 64-105.2); USFC St. 2890, 43e46tN, 124057'W,

Figs. 4L9. Radulae of Exilioidea rectirostris. 4-5. leelseyi form, off California, 147m (LACM 71 120). 6 7. Typi'cal form, off Washington State, 85m (LACM 71-360.3). 8-9, Deep water obsoleta form, off Oregon, 800m (LACM 68-216.5) Scale lines: 10ptrn.

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Bouchet. War6n:Transfer ofExilioideato Turbinellidae 175

tut･t.l-.V/IU.titt-rf

l/.

tw/de

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510m, 2spms CUSNM 122662), California: off Ti'inidad Harbor, 41007'N, 124021'W, 165m, ]eg. J.avenberg &

Swift, 4 spms (I.ACM 71-121') ; off Trinidad Harbor, 41009'N, 124-20'W, 147 m, leg, Lavenberg & Swift, 5spms (LACM 71-120: Figs. 19-20>; off Trinidad

Harbor, 41004'N, 124021'W, 185m, leg, Swift & Lavenberg, lspm

71112); off Eureka, Humboldt Co,, 185m, ]eg. Talmadge, 2spms (LACM

127009); USFC St. 3172, off Bodega Head, 38024'N, 123014'W, 115m, 1sprn

(USNM 224346) ; USFC St. 3170, off Bodega Head, 38017'N, 123029'W, 307m, 1spm (USNM 206583: Fig. 21); USFC St. 4513, off Pt. Pinos, 389-456 m, 1spm

(USNM 210621) ; off Pt, Pinos, 36e38'N, 122002'W, 185 m, leg. McLean, 1 spm

(LACM 60-21) ; USFC St. 3193, off San Simeon Bay, 35026'N, 121C'09'W, 295m,

1 spm (USNM 206596'} ; USFC St. 3195, off San Luis Obispo, 35e14'N, 121007'W, 465 m, 8spms (USNM 122660, 205915, 206026); USFC St. 3199, Santa Barbara

Channel, 34n17'N, 120-26'VL', 430m, 1spm (USNM 213359); USFC St. 2936,

off San Diego, 32049'N, 117028'W, 660rn, 4spms

USFC St. 2935, off San Diego, 32045'N, 117023'W, 230m, lspm (USN"(

110632); off San Diego, 105m, leg. Paine, 1spm (USNM 186069); Pleistoeene

of San Pedro, 1 spm (USNM 193723).

DistTibution: From Alaska to California, in 60-800m.

Re･marks :

1. Nomenelature: The problems regarding the holotype of C. rectirostris have

been solved by Bentson, who presented the only existing illustration (1940: pl. 1

fig. 22). An extensive chresonymy of the name 7`ectirostris can be found in

Palmer (1958: 215),

Confusien coneerning the holotype of T. kelseyi was eaused by Dall who

did not figure T. kelseyi until l925 (plate 1, fig. 6). This figure does not

represent the ho]otype but a shel] with USNM eatalogue number 224346,

whereas the holotype is USNM ll0631. The situation was made st･ill more

confused by Bartseh (1945) whose aecount of the genus contains many mis-

Ieading or erroneous statements. Among others, his figure of E. Tectirostris

in fact depicted the holotype of T. keZseyi! It is not clear, ho-rever, how the

confusion arose because he eorreetly listed the type lot (USNM 110631) of

kelseyi under that name. The figure of E. kelseyi in Abbott (1974: 216, fig.

2388) is based on the holotype of keeseyi and is eorrectly assigned to that name.

2. Variation: Examination of the large suite of material present in USNM

and LACM has led us to eonelude that it represents a single species. In the .--.... -... .-.-...-... ..-...t...... r.

Figs. 10-15. Radulae of Exitioidea. 10. E. indica, Anton Bruun spec{men (USNM 717323), scale 1ine 30,um. 11. E. rectirostris, same speci- men as Figs. 6 7, scale line 10,um. 12-13, E. costttlata, holotype, scale lines 10 and 5"m, respectively. 14-15. E. rufocaudata, Gala- thea specimen, sca13 lines 3C and 10stm, re3pectively.

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northern part of its range (from Alaska to Puget Sound: Figs. 1648), specimens have a coarser spiral seulpture than further to the south (Oregon to California:

Figs. 19-21). In addition, there is a variation with depth: Shallow water speei-

mens (60-200 m, Figs. 16-17, 19-20) are more slender, while specimens from deep

water (500-800m, Fig. 22) have more convex whorls, with stronger axial ribs.

There exist all intermediates between these different forms (Figs. 18, 21) and it

should be noted that the transition to the deep water form appears to occur

more shallow in the north than in the south. The radulae of the different forms

(Figs. 4-9, 11) do not show signifieant differences.

The name ketseyi is based on the shallow water form from California, which

is very slender and weakly seulptured; the name obsoleta is based on the deep

water form, which shows little variation between northern and southern popula-

tions. Because the transttion is continuous between the different forms and

represents a twe dimensional eline, we eonsider that these names should not

be used to indieate subspecies.

Exilioidea rufocaucinta (Dall, 1896)

{Figs. 14-15, 25-26)

Fusus.? rufoeaztdatus Dall, 1896: 12; Dall 1908a: 302, pl.3, fig. 3.

Type materiat: Holotype USNM 123006.

Type locatity: Albatross St. 3415, 14046'N, 98'40'W, 3440m (off SW Mexico).

Material examined: The holotype and Galathea St. 716, 09e23'N, 89e32'W,

3590 m, 1spm (ZMC). Distribution: East Pacific off central America, 2325-3590m [listed from a few Albatross stations by Dall (1896) ; material not examined].

Exitioiclea indica, n. sp.

(Figs. 3, 10, 27-28)

Type material: Holotype in ZMC.

Type locality: Galathea St. 282, 05032'N, 78"21'E, 4040 m (SW of Sri Lanka).

Material examined: The type material and International Indian Ocean Expedi-

tion, Anton Bruun cruise 7, St. 363B, 23045'S, 43010'E, 2980 m (Mozambique

Figs. 16-22.Clinal variation in the NE Pacific species Exilioidea rectirostris. Upper row:Specimens from the northern part of the range;lower row: Specimens from the southern part of the range; specimen

to the right: Deepwater form. 16-17. Typical form, off Washington State, 85m (LACM 71-360.3), height 19.9 and 30.6mm, 18. Specimen from interrnediate depth, Queen Charlotte Sound, 260 m (USNM 226117), height 25.5mm. 19-20. kelseyi form, off N. Califbrnia, 147m (LACM 71-120), height 16.9 and 28.2mm. 2!. Specimen from intermediate depth, off central California, 307 m (USNM 206583), height 30,4mm. 22. obsoteta form, off Oregon, 800m (LACM 68-216.59), height 24.8mm.

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Channel), 1 spm (USNM 717323) ; John Murray Exp., St. 120, 05049'-05e52'S, 41028'-41"40'E, 2900 m (off Kenya), 1 sh (BMNH) ; Safari 2, St. CP2, 05042'N,

' 78056'E, 3625m (SW of Sri Lanka), 1spm (MNHN).

Distribution: Only known from the material examined, northern and western

Indian Ocean, 2900-4040m.

Description: Shell slender, fusiform, presumably consisting of at least 6 whorls;

3 corroded whorls remain, upper 3+ whorls and the Iarval shell dissolved, Whorls

convex with deep suture. Aperture evate with long, open siphonal eanal. Outer

lip thin, ehipped; columellar callus thin; eolumella straight, without fold, Axial

sculpture, fine incremental lines and strong axial, slightly episthoeyrt ribs; 19

on first intact and on penultimate whorl; becoming obsolete on body whorl. Spiral

sculpture of cords and grooves of about equal breadth; 12 on penultimate whorl,

but sutural zone partly decortieated and a few cords probably obliterated. Shell

yellowish brown; periostracum thin, yellowish.

Dimensions of shell: Height 36.6 mrri, breadth 11.0 mm; aperture with siphonal

eanal 18,5 rnm.

Radula and operculum: See Figs. 3 and 10.

Remarks: The 3 additional specimens known measure 16.4 × 5.5mm, 13 × 5mm

and 27 × 8,5 mm, respeetive]y. The 27 rnm high specimen has 16 spiral cords on

the penultimate whorl (suture not eorroded). The 16.4 mm high shell has 6 whorls;

the apex is corroded but probably not more than one whorl is missing. This

small shell has 15 axial ribs on the body whorl and it is white.

E. rufocaudata resembles E. i･ndica, but differs by its more oblique suture,

and by the whorls of more rapidly increasing diameter. The axial ribs are more

opisthocyrt and they do not become obsolete on the body whorl of a 50 mm high

specimen of E. rufocaudata.

Exitioidea castulata. n. sp･

Type material: Holotype and 2 paratypes, USNM 239408, Type Iocality: Albatross St. 5648, 05035'S, 122008'E, 1025m (Buton Strait, Sulawesi).

Materiat examined: Only known from the type material.

Description: Shell slender, selid, fusiform, eonsisting of 7 whorls. Upper 2 whorls corroded and apex totally dissolved. Whorls convex, suture shallow. Aperture ovate with long, open, siphonal canal; outer lip thin, slightly chipped,

Figs. 23-30. Genus Exilioidea, 23, E. costulata, holotype, helght 21,6mm, 24, E, sp., Galathea St. 299, height 29 mm. 25-26. E. rufocaudata, holotype, height 28,4mm and Galathea specimen, height 50.5mm. 27-28. E. indica. holotype. height 36.6mm and John Murray specimen, height 13.3mm. 29-30. E. atlantica, holotype, height 36.Imm and specimen from 878m, height 25.9mm.

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a little expanded. Columella straight, without fold and callus. Axial seulpture

of fine incremental lines and strong, broad ribs, slightly opisthocyrt; 23 ribs

on body-whorl, 21 on penultimate whorl. SpiTal sculpture of indistinctly defined

spiral furrows, best visible where they cross axial ribs and form nodulous

intersections. 5 rows of spiral undulations on penultimate whorl, 4 on upper

whorls. Spiral furrows and axial ribs obsolete in subsutural zone of apical

whorls. Secondary seulpture of a few mueh finer spiral threads. Shell whitish;

thin yellowish green periostracum lighter below the periphery and on the siphonal canal.Dimensions

of the shetl: Height 21.3mrn, breadth 5.7mm; aperture with

siphonal eanal 11 mm.

Raduta and operoulum: See Figs. 1-2 and 12-13,

Remarks: E. costulata differs from other species of Exilioidea by its sculpture

pattern. It resembles most closely Exilioidea sp. below, see remarks.

Exilioiclea sp. (Fig. 24)

MateTiat examined: Galathea St. 299, 17010'N, 84"30'E, 2860m (northern Indian Ocean), 1sh (ZMC). Description: Shell slender, fusiform, consisting of 4 whorls. Apical part of

the shell dissolved. Whorls convex. Outer lip thin, no columellar callus. Axial

sculpture of thin incremental lines and stronger axial ribs. 26 ribs on first

preserved apical whorl, 32 on body whorl. Ribs crossed by spiral threads, con-

siderably more narrow thall intervals. 2 spiral threads in subsutural zone of

apical whorls, gradually disappearing, obsolete on body whorl; 4, later 5, spiral

threads in middle and lower parts of whorls. Periostracum yellowish brown,

lighter on the siphenal eanal.

Dimensions of the shetl: Height 29 mm, breadth 8.5 mm; aperture with siphonal

eanal 17 mm.

Rem(zrks: This is probably a distinct, undescribed species. Its general propor-

tions, however, are the same as in E. costulata and it may be that the observed

differences in sculpture represent geographieal, rather than interspeeifie dif-

ferences. In E. costutata the axial ribs are fewer and broader, the spiral sculpture

consists of more indistinct spiral furrows and the intersections of spiral and

axial sculpture do not form wel! defined squares as in Exilioidea sp. Both have

weaker sculptuTe in the subsutural zone.

Exitioidea atlantica, n. sp.

(Figs. 29-30)

Type ?naterial: Holotype and one paratype in MNHN.

Type Zecatity: Gulf of Mexico, 25020'N, 86007'W, 3237m (L. Pequegnat St.

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Bouchet.Waren: Transfer of Exilioidea to Turbinellidae 183

68.A7.4A).

Materiat examined: The type material and Gulf of Mexieo, 25038'N, 96-07'W,

878m, 1sh (L. Pequegnat St. 68.A13.1); Gulf of Mexico, 25e14'N, 95e13'W,

3365m, 1sh (L Pequegnat St. 68.A13.9); Gulf of Mexico, 25030'N, 86009'W,

3230m, 1sh (L Pequegnat St. 69.A13.29) (all MNHN).

Distribution: Only known from the material examined, northern part of the

Gulf of Mexieo, 878-3365m.

Deseription: Shell very slender, fusiform, consisting of a bulbous, rounded protoconch of less than one whorl, and 10 teleoconeh whorls. The whole shell

is rather eorroded (as are all shells from below 3000m). Whorls convex with deep suture. Aperture ovate with rather shert siphonal canal (tip of the canal

may be broken but eanal short in all shells available). Outer lip thin, ehipped;

columellar callus thin; columella straight, without feld. Axial sculpture of strong

axial ribs, sharp and high on spire whorls, becoming obsolete on penultimate

and body whorls. About 12 ribs per spire whorl, more numerous towards base of

shell. Spiral sculpture not visible on holotype due to deeorticate condition; on

shell from 878m, 15-17 spiral groves present. Shell whitish, no trace of

periostracum left.

Dimensions of shelL: Height 36.1mm, breadth 8.8mm; aperture with siphonal

canal 11.5 mm.

Remarks: Despite the imperfect condition ef the holot･ype, E. atlantica can

be easily separated from other eongeneTs by its proportionally much higher

spire and short siphonal canal. It eannot presently be ascertained if the present

ExiZioidea from the Gulf of Mexieo is more related to the E. Pacifie species of

the genus, or to the Indo-West Pacifie ones. The paueispiral protoconch indicates

non planktotrophie larval development; if this character has been present for a

long time in Exilioutea, cornmunication across shallow water straits of Mioeene

and Pliocene central America would not have been possible. Acknowledyements:-We thank R. Houbriek, J. Rosewater, D. Bohmhauer (USNM), J.

Knudsen (ZMC), J. McLean (LACM), K. Way (BMNH> for access to the material

under their care. We also thank C. Monniet and L. Pequegnat for plaeing collections

of deep sea molluscs colleeted under their supervision at our disposaL J, McLean

generously shared with us unpublished information on the northeast Paeifie speeies.

C. Lamb revised the language.

ff ts -C "(i ttX:= V'.;f Fkl: Sdihii'L L i)Z Exitioidea tsVt caE' ee op meghi 5t = = 7" t･' trd $" fo 4 tl 8 rb:\U -)k. ffiStec7tsOasfiOmeM, At)&:Vt"tieD Tritonofusus ketseyiDall, 1908 'P PIici.fusus obsoletus

Talmadge, 1971 C'XLx-9=a'L ig E, rectirostris (Carpenter, 1864) OIWfiIQXv(fa4t lrb:\goko y-: tcl'(IC･:tLfrLkvCiOkh:OCMJt(SiZt\recD E. rufocaudata (Dall, 1896) Lblicalb2'L`CLhtshi-)7k:ioR, Ukla

4pth:kJtciP?2fi,4eF¢ eethhi6re.n$in.fo.

E. indica, n. sp. JK V 9VhMru, lt

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E. costulata. n. sp. XiV.V, i-hVtal[pt, zkme 1,025m, E, sp. Iyr#"ices, zkee 2,860rn,

E. atiantica n, sp. I S'v=n, zkva 3,237m.

References

Abbott, R,T. 1974. Amerioan Seashells, 2nd edition, New York, Van Nostrand Rein-

hold Ce. 663pp. Bartsch, P. 1945. Some notes on the genus Exil･ioidea. Na"titzts, 59: 61-63. Bentson, H. 1940. A systematie study of the fossil gastropod Exilia. Univ. Califor- nia Publ., BulL Dep. Geol. Sci., 25(5): 199-238.

Bouchet, P. and A. Waren, 1985. Revision ef the Northeast Atlantie bathyal and

abyssal Neogastropoda exeluding Turridae, Boll. Mal-eol. suppl. 1: 121-296, Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the of North Ameriea. Rep. BTit. Assoc. Adv. Sci. (1863) : 517-686.

Dall, W.H. 1896. Diagnoses of new species of mollusks from the west coast ef

Ameriea. Proc. US. Nat. M?ts., 18: 7-20. Dall, W.H. 1908a. Reports on the dTedging operations off the west coast of Central

America to the Galapagos, the west coast of Mexieo and in the Gulf of CalifoTnia, 14: the Mollusca and Braehiopoda. B2tlt. Mus. Comp. Zool., 43(6) : 205-487. Dall, W.H. 1908b. Deseriptions of new species of mollusks from the Pacific coast of the United States, with notes on other mollusks from the same region. Proc. U.S. Nat. Mus., 34: 245-257.

Dall, W.H. 1925. Illustrations of unfi.aured types of shells in the eollection of the

United States National Museum. Proc. U.S. Nat. Mus., 66: 1-41.

GTant, U.S. and H.R. Gale 1931. Catalogue of the rnarine Plioeene and Pleistocene

Mollusca of California and adjacent regions, Mem. Sdin Diege Soe. Nat. Hist., 1: 1-1036,

Harasewych, M.G. 1987. A revision of the genus Benthovolieta with notes on the evolutien of the subfarnily Ptychatraetinae (Prosobranchia: Turbinellidae). Nautilus, 101(4): 166-181.

Hickman, C, 1980. Paleogene marine gastropods of the Keasey formation in Oregon. Bull. A7ner. Paleont., 78(310) : 1-112.

Plalrner, K. 1958. Type specimens of marine Mollusca deseribed by P. P. Carpenter from the West Coast. Geot. Soc. Ameriea･, Mem,, 76: 1-376. Talmadge, R.R. 1971. The benthie Mollusca, Plicifuszes, in California, Veliger, 14: 42.44.

[Received: April 14, 1988]

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