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An Annotated Checklist of the Marine Macroinvertebrates of Alaska David T

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An Annotated Checklist of the Marine Macroinvertebrates of Alaska David T NOAA Professional Paper NMFS 19 An annotated checklist of the marine macroinvertebrates of Alaska David T. Drumm • Katherine P. Maslenikov Robert Van Syoc • James W. Orr • Robert R. Lauth Duane E. Stevenson • Theodore W. Pietsch November 2016 U.S. Department of Commerce NOAA Professional Penny Pritzker Secretary of Commerce National Oceanic Papers NMFS and Atmospheric Administration Kathryn D. Sullivan Scientific Editor* Administrator Richard Langton National Marine National Marine Fisheries Service Fisheries Service Northeast Fisheries Science Center Maine Field Station Eileen Sobeck 17 Godfrey Drive, Suite 1 Assistant Administrator Orono, Maine 04473 for Fisheries Associate Editor Kathryn Dennis National Marine Fisheries Service Office of Science and Technology Economics and Social Analysis Division 1845 Wasp Blvd., Bldg. 178 Honolulu, Hawaii 96818 Managing Editor Shelley Arenas National Marine Fisheries Service Scientific Publications Office 7600 Sand Point Way NE Seattle, Washington 98115 Editorial Committee Ann C. Matarese National Marine Fisheries Service James W. Orr National Marine Fisheries Service The NOAA Professional Paper NMFS (ISSN 1931-4590) series is pub- lished by the Scientific Publications Of- *Bruce Mundy (PIFSC) was Scientific Editor during the fice, National Marine Fisheries Service, scientific editing and preparation of this report. NOAA, 7600 Sand Point Way NE, Seattle, WA 98115. The Secretary of Commerce has The NOAA Professional Paper NMFS series carries peer-reviewed, lengthy original determined that the publication of research reports, taxonomic keys, species synopses, flora and fauna studies, and data- this series is necessary in the transac- intensive reports on investigations in fishery science, engineering, and economics. tion of the public business required by law of this Department. Use of funds Copies of the NOAA Professional Paper NMFS series are available free in limited for printing of this series has been ap- numbers to government agencies, both federal and state. They are also available in proved by the Director of the Office exchange for other scientific and technical publications in the marine sciences. Profes- of Management and Budget. sional Papers are published online in PDF format at http://spo.nmfs.noaa.gov NOAA Professional Paper NMFS 19 An annotated checklist of the marine macroinvertebrates of Alaska David T. Drumm Katherine P. Maslenikov Robert Van Syoc James W. Orr Robert R. Lauth Duane E. Stevenson Theodore W. Pietsch November 2016 U.S. Department of Commerce Seattle, Washington Cover figure (top left to bottom right): Halichondria oblonga (Porifera); Cheilonereis cyclurus (Polychaeta); Tonicella insignis (Polyplacophora); Neptunea lyrata (Gastropoda); Pandalus tridens (Decapoda: Caridea); Oregonia gracilis (Decapoda: Brachyura); brachiopod unidentified (Brachiopoda); Pagurus ochotensis (Decapoda: Anomura); Ophiura sarsii (Ophiuroidea); Pteraster octaster (Asteroidea). Suggested reference Drumm, David T., Katherine P. Maslenikov, Robert Van Syoc, James W. Orr, Rob- ert R. Lauth, Duane E. Stevenson, and Theodore W. Pietsch. 2016. An annotated checklist of the marine macroinvertebrates of Alaska. NOAA Professional Paper NMFS 19, 289 p. doi:10.7755/PP.19 Online dissemination This report is posted online in PDF format at http://spo.nmfs.noaa.gov (click on Professional Papers link). Copyright law Although the contents of the Professional Papers have not been copyrighted and may be reprinted entirely, reference to source is appreciated. Proprietary products The National Marine Fisheries Service (NMFS) does not approve, recommend, or endorse any proprietary product or proprietary material mentioned in this publi- cation. No reference shall be made to NMFS, or to this publication furnished by NMFS, in any advertising or sales promotion which would indicate or imply that NMFS approves, recommends, or endorses any proprietary product or proprietary material mentioned herein, or which has as its purpose an intent to cause directly or indirectly the advertised product to be used or purchased because of this NMFS publication. Disclaimer The findings and conclusions in this report are those of the author(s) and do not necessarily represent the views of the National Marine Fisheries Service (NOAA). CONTENTS Introduction ................................................................1 Order Astrophorida ........................................16 Methods ......................................................................2 Family Ancorinidae ...................................16 Data sources ................................................................3 Family Geodiidae ......................................17 Marine geography of Alaska .......................................4 Family Vulcanellidae .................................17 Current status and future directions ............................7 Order Clionaida .............................................17 Checklist ...................................................................12 Family Clionaidae .....................................17 Phylum Porifera—The Sponges .................................12 Order Polymastiida ........................................17 Class Calcarea ......................................................12 Family Polymastiidae ................................17 Subclass Calcaronea ...........................................12 Order Tethyida ...............................................18 Order Baerida .................................................12 Family Tethyidae ......................................18 Family Baeriidae .......................................12 Order Suberitida .............................................18 Order Leucosolenida ......................................12 Family Suberitidae ....................................18 Family Amphoriscidae ..............................12 Family Halichondriidae ............................18 Family Grantiidae .....................................12 Family Stylocordylidae .............................19 Family Leucosoleniidae .............................13 Order Haplosclerida .......................................19 Family Sycettidae ......................................13 Family Callyspongiidae .............................19 Subclass Calcinea ................................................13 Family Chalinidae .....................................19 Order Clathrinidae .........................................13 Family Niphatidae ....................................20 Family Clathrinidae ..................................13 Family Petrosiidae ....................................20 Class Hexactinellida .............................................13 Order Poecilosclerida .....................................21 Subclass Amphidiscophora ................................13 Family Acarnidae ......................................21 Order Amphidiscosida ....................................13 Family Microcionidae ...............................21 Family Hyalonematidae ............................13 Family Coelosphaeridae ............................22 Subclass Hexasterophora ....................................13 Family Crambeidae ...................................23 Order Hexactinosida ......................................13 Family Crellidae .......................................23 Family Farreidae .......................................13 Family Hymedesmiidae .............................23 Family Euretidae .......................................13 Family Iotrochotidae ................................23 Family Aphrocallistidae ............................14 Family Myxillidae .....................................23 Family Tretodictyidae ...............................14 Family Phellodermidae .............................24 Order Lyssacinosida .......................................14 Family Tedaniidae .....................................24 Family Euplectellidae ................................14 Family Cladorhizidae ................................24 Family Rossellidae ....................................14 Family Guitarridae ...................................24 Class Homoscleromorpha .....................................16 Family Esperiopsidae ................................25 Order Homosclermorphida ............................16 Family Mycalidae .....................................25 Family Plakinidae .....................................16 Family Isodictyidae ...................................26 Class Demospongiae .............................................16 Family Latrunculiidae ...............................26 Subclass Keratosa ...............................................16 Order Axinellida ............................................26 Order Dendroceratida ....................................16 Family Axinellidae ....................................26 Family Darwinellidae ................................16 Family Raspaillidae ..................................26 Order Dictyoceratida ......................................16 Order Bubarida ..............................................26 Family Dysideidae ....................................16 Family Bubaridae ......................................26 Subclass Heteroscleromorpha .............................16 Order Biemnida ..............................................26 Order Tetractinellida ......................................16 Family Biemnidae .....................................26 Suborder Spirophorina .................................16 Phylum Cnidaria .......................................................27
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