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1 C M 2 B H 3 B M 4 C M 5 B L 6 B M 7 B M 8 SPECIES EVALUATION Haplopappus pygmaeus, Priority 1. Tonestus pygmaeus (Torrey & Gray) A. Nelson (TOPY). pygmy goldenweed. CNHP G4 / SR, Track N G4 N?. CO SR, WY S1. WY Peripheral 2 MBNF Confi- Criteria Rank dence Rationale Sources of Information Distribution (see map below) is “nearly continuous,” so rating C was chosen, but Specimens at COLO and RM, Dorn 2001, 1 none of the pictures or descriptions really fit this species. Tracked by WY, but not Weber & Wittmann 2001ab, PLANTS 2002, Distribution C M tracked by CO or NM. WYNDD 2002. within R2 Tonestus pygmaeus is known principally from Colorado, but also from adjacent Specimens at COLO and RM, Harrington 2 southern Wyoming and northern New Mexico, and disjunct in southwestern Montana. 1954, Dorn 2001, Weber & Wittmann Distribution B H 2001ab, PLANTS 2002, WYNDD 2002, outside R2 MTNHP 2002. Seeds are lightly hairy and pappus is deciduous, so seeds probably cannot disperse Harrington 1954. 3 long distances; viability unknown. Pollen viability and dispersal unknown. Dispersal B M Capability Tonestus pygmaeus is found moderately frequently in the Alpine zone, but is not Fertig 2000, specimens at COLO and RM, really common in the normal sense. “Demographic stochasticity” is probably irrelevant Harrington 1954, Dorn 2001, Weber & 4 to this species. About 50 recorded occurrences in Colorado, three in Wyoming, two or Wittmann 2001ab, PLANTS 2002, WYNDD Abundance in C M three in New Mexico. “Abundance not known. Population censuses have not been 2002, MTNHP 2002. R2 undertaken, but the species is believed to be at least moderately common within its restricted range” (Fertig 2000). “Presumed to be stable, although long-term trend data are not available” (Fertig Fertig 2000. 5 2000). Population B L Trend in R2 “Common on rocky tundra” (Weber and Wittmann 2001a). Rocky tundra is not My observations. 6 undergoing any significant decline at this time. “Cushion plant communities Habitat Trend B M interspersed among Krummholz in gravelly fellfields, toe slopes, and ridge crests at or in R2 above treeline” (Fertig 2000). “Threats are relatively low, and there are not reports of trampling due to recreation My observations, Handley and Laursen or grazing. [In Wyoming,] pygmy goldenweed occurs in the Medicine Peak Special 2002. 7 Botanical Area and on lands managed for multiple use by the Medicine Bow National Habitat B M Forest” (Handley and Laursen 2002). Vulnerability or Rocky tundra is somewhat vulnerable to some recreation activities (trails and Modification routes), and these habitats would be difficult to revegetate. But the habitat is also somewhat resilient to disturbance. Effects on the species itself are not known. Tonestus pygmaeus is probably not palatable to wild or domestic herbivores. No observations on life history or demographics. 8 Life History and D H Demographics 1 SPECIES EVALUATION National Forests in the Rocky Mountain Region where species is KNOWN (K) or LIKELY (L)* to occur: *. Likely is defined as more likely to occur than not occur on the National Forest or Grassland. This generally can be thought of as having a 50% chance or greater of appearing on NFS lands. COLORADO NF/NG K L K L K L Arapaho-Roosevelt NF K NEBRASKA NF/NG WYOMING NF/NG White River NF K Samuel R. McKelvie NF Shoshone NF Routt NF K Halsey NF Bighorn NF Grand Mesa Uncompahgre Gunnison NF K Nebraska NF Black Hills NF San Juan NF K Ogalala NG Medicine Bow NF K Rio Grande NF K SOUTH DAKOTA NF/NG Thunder Basin NG Pike-San Isabel NF L Black Hills NF KANSAS NF/NG Comanche NG Buffalo Gap NG Cimarron NG Pawnee NG Ft. Pierre NG Taxonomy. Should be called Tonestus pygmaeus, rather than Haplopappus pygmaeus. This is considered to be a valid species by all botanists, endemic to Colorado, southern Wyoming, and northern New Mexico, with disjunct populations in southwestern Montana. Discussion. Based on distribution and abundance, and its habitat in rocky sites, this species probably does not have great concerns for viability. Any concerns would probably better be handled by improved management of Alpine zone. References Albee, Beverly J.; Leila M. Shultz; and Sherel Goodrich. 1988. Atlas of the Vascular Plants of Utah. Utah Museum of Natural History. http://www.nr.usu.edu/Geography-Department/utgeog/utvatlas/ut-vascatlas.html, accessed September, 2002. Fertig, Walter. 2000. State species abstract: Tonestus pygmaeus. Laramie, WY: Wyoming Natural Diversity Database, 2 pp. http://uwadmnweb.uwyo.edu/WYNDD. Handley, Joy; and Scott Laursen. 2002. Sensitive species evaluation: Haplopappus pygmaeus. Attached below. Harrington, Harold D. 1954. Manual of the plants of Colorado: For the identification of the ferns and flowering plants of the state. Denver, CO: Sage Books. 666 pp. Montana Natural Heritage Program. 2002. Field Guide: Haplopappus pygmaeus. http://nhp.nris.state.mt.us/plants. Rocky Mountain Herbarium. 1998. Atlas of the vascular plants of Wyoming. http://www.esb.utexas.edu/tchumley/wyomap/atlas.htm, accessed September, 2002. Spackman, Susan; Bill Jennings; Janet Coles; Carol Dawson; Mark Minton; Andrew Kratz; Carol Spurrier; Christopher S. Johnson; and Mike Barry. 1999. Colorado rare plant field guide. http://ndis.nrel.colostate.edu/ndis/rareplants/cover.html, last updated March 1, 1999. Weber, William A.; and Ronald C. Wittmann. 2000. Catalog of the Colorado flora: A biodiversity baseline. Boulder, CO: University of Colorado Museum. Revised March 11, 2000. http://www.colorado.edu/CUMUSEUM/research/botany/Catalog/Catalog.htm, downloaded September, 2002. Weber, William A.; and Ronald C. Wittmann. 2001a. Colorado flora: Western slope, Third Edition. Niwot, CO: Colorado Associated University Press. 488 pp. Weber, William A.; and Ronald C. Wittmann. 2001b. Colorado flora: Eastern Slope, Third Edition. Boulder, CO: University Press of Colorado. 521 pp. Author and date: Barry C. Johnston, Botanist, Grand Mesa-Uncompahgre-Gunnison National Forests, 216 N. Colorado St., Gunnison, CO 81230-2197. (970) 642-4467. [email protected]. Last modified September 23, 2002. 2 SPECIES EVALUATION REGION 2 SENSITIVE SPECIES EVALUATION FORM Species: Haplopappus pygmaeus / Pygmy Goldenweed (Tonestus pygmaeus) Criteria Rank Rationale Literature Citations Pygmy Goldenweed occurs in the Snowy Range in Albany and Carbon counties on the • Dorn 2001 1 AD Medicine Bow National Forest in Wyoming. It also occurs in Colorado, therefore, additional Distribution • Fertig 2000 within R2 information is necessary to determine its status in Region 2. • Harrington 1954 This species is found in cushion plant communities interspersed among krummholz in gravelly • Nelson 1904 fellfields, on toe slopes, and ridgecrests at or above treeline. The elevational range is from • Nelson 1984 10770 to 11600 feet. Its alpine habitat is very limited in this area of the state. • University of Wyoming 1998 Confidence in Rank High Pygmy Goldenweed is known from southeastern Wyoming to northern New Mexico, with a • Cronquist 1955 2 B disjunct population in the Bridger Mountains of southwestern Montana. Distribution • Duft and Moseley 1989 outside R2 • Lesica and Shelly 1991 Confidence in Rank High • Scott 1997 It may be inferred that the bristly pappus is appropriate for wind dispersal across unsuitable • Cronquist 1955 3 CD habitat. Dispersal • Fertig 2000 Capability Confidence in Rank High Pygmy Goldenweed is known from 2 extant occurrences in Wyoming, both observed since 1993 • Fertig 2000 4 AD (most recently in 2000). Both occurrences are on the Medicine Bow National Forest. Population Abundance in • Harrington 1954 R2 censuses have not been undertaken, but the species is believed to be at least moderately to • NatureServe 2002 locally common within its restricted range. This species is ranked “S1” in Wyoming and “SR” in • Nelson 1984 Colorado. • University of Wyoming 1998 Confidence in Rank High • WYNDD 2002 Presumed to be stable, although long-term trend data are not available. • - 5 D Population Trend in R2 Confidence in Rank High The availability of its rocky, alpine habitat is presumed stable. • - 6 BD Habitat Trend in R2 Confidence in Rank High Threats are relatively low, and there are not reports of trampling due to recreation or grazing. • Fertig 2000 7 BD Pygmy Goldenweed occurs in the Medicine Peak Special Botanical Area and on lands managed Habitat • Nelson 1984 Vulnerability or for multiple use by the Medicine Bow National Forest. • University of Wyoming 1998 Modification Confidence in Rank High • WYNDD 2002 Pygmy Goldenweed is a mat-forming, perennial forb. Flowering and fruiting occur from July • Cronquist 1955 8 D through August. Life History and • Fertig 2000 Demographics Additional information on the species, including life history stages, population structure, longevity, mortality, and seed biology, are not available. Confidence in Rank High Initial Evaluator(s): Date: March 26, 2002 Joy Handley and Scott Laursen 3 SPECIES EVALUATION National Forests in the Rocky Mountain Region where species is KNOWN (K) or LIKELY(L)1 to occur: Colorado NF/NG Kansas NF/NG Nebraska NF/NG South Dakota Wyoming NF/NG NF/NG Likely Likely Known Known Known Likely Known Likely Known Likely Arapaho-Roosevelt NF Cimmaron NG Samuel R.McKelvie NF Black Hills NF Shoshone NF White River NF Halsey NF Buffalo Gap NG Bighorn NF Routt NF Nebraska NF Ft. Pierre NG Black Hills NF Grand Mesa, Ogalala NG Medicine Bow NF X Uncompahgre, Gunnison NF San Juan NF Thunder Basin NG Rio Grande NF Pike-San Isabel NF Comanche NG Literature cited Cronquist, A. 1955. Pt. 5. Compositae. In: Hitchcock, C.L., A. Cronquist, M. Ownbey, and J.W. Thompson (eds). Vascular Plants of the Pacific Northwest. Univ. Washington Publ. Biol. 17(5): 1-343. Dorn, R.D. 2001. Vascular Plants of Wyoming, third edition. Mountain West Publishing, Cheyenne, WY. Duft, J.E. and R.K. Moseley. 1989. Alpine Wildflowers of the Rocky Mountains. Mountain Press Publ. Co., Missoula, MT. Fertig, W. 2000. State Species Abstract: Haplopappus pygmaeus. Wyoming Natural Diversity Database. Available on the internet at www.uwyo.edu/wyndd Hall, H.M.
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