Concepts Giving an Idea of Mineral Paragenesis Under One Set of Given
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Plant Fossils and Gondwana Flora
UNIT 12 PLANT FOSSILS AND GONDWANA FLORA Structure_____________________________________________________ 12.1 Introduction Vertebraria Expected Learning Outcomes Thinnfeldia 12.2 Plant Fossils Sigillaria Definition Nilssonia Classification Williamsonia Modes of Preservation Ptilophyllum Significance 12.5 Activity 12.3 Gondwana Flora of India 12.6 Summary 12.4 Descriptions of some Plant 12.7 Terminal Questions Fossils 12.8 References Glossopteris 12.9 Further/Suggested Readings Gangamopteris 12.10 Answers 12.1 INTRODUCTION The animals, plants and micro-organisms are the three main life forms surviving today. Even their fossilised remains are found in rocks that tell us about their past history. The animals comprise invertebrates and vertebrates. In Block 4, you will read about the invertebrates and their geological history that began in the latest Precambrian time. You also read about the microfossils in Unit 10 that too have a long geological record beginning from Precambrian onwards. In Unit 11, you read the evolutionary history of one of the vertebrate groups i.e., horse. In this unit, you will read the plant fossils and the Gondwana flora of India. Introduction to Palaeontology Block……………………………………………………………………………………………….….............….…........ 3 Like the kingdom Animalia, plants also form a separate kingdom known as the Plantae. It is thought that plants appeared first in the Precambrian, but their fossil record is poor. It is also proposed that earliest plants were aquatic and during the Ordovician period a transition from water to land took place that gave rise to non-vascular land plants. However, it was during the Silurian period, that the vascular plants appeared first on the land. The flowering plants emerged rather recently, during the Cretaceous period. -
Permophiles Issue #28 1996 1
Permophiles Issue #28 1996 1. SECRETARY’S NOTE I should like to thank all those who contributed to this issue of I have discussed the matter of financial support for Permophiles @Permophiles@. The next issue will be in November 1996 and with many of the members of the Permian Subcommission; the will be prepared by the new secretary, CIaude Spinosa. Please consensus is that we should request voluntary donations to offset send your contributions to him (see note below from Forthcoming processing, mailing and paper costs. We are asking for contribu- Secretary). tions of $10 to $25. However, Permophiles will be mailed to ev- I should like to express my gratitude to those of you who have eryone on the mailing list regardless of whether a contribution is submitted contributions during my eight year term of office. You made. have helped make =Permophiles@ a useful, informative, and timely We have established access to Permophiles via the Internet; the Newsletter of the Subcommission on Permian Stratigraphy. address will be: J. Utting http://earth.idbsu.edu/permian/permophiles Geological Survey of Canada (Calgary) 3303 - 33rd Street N.W. Our intention is to provide a version of Permophiles that is readily Calgary, Alberta, Canada T2L 2A7 available through the Internet. The Internet version of Permophiles Phone (404) 292-7093 FAX (403) 292-6014 will have multiple formats. Initially the format of the Internet ver- E-mail INTERNET address: [email protected] sion will be different from the official paper version. Because new taxonomic names can not be published in Permophiles, a hard copy, downloaded from the interned will suffice for many of us. -
Curriculum Vitae
CURRICULUM VITAE ORCID ID: 0000-0003-0186-6546 Gar W. Rothwell Edwin and Ruth Kennedy Distinguished Professor Emeritus Department of Environmental and Plant Biology Porter Hall 401E T: 740 593 1129 Ohio University F: 740 593 1130 Athens, OH 45701 E: [email protected] also Courtesy Professor Department of Botany and PlantPathology Oregon State University T: 541 737- 5252 Corvallis, OR 97331 E: [email protected] Education Ph.D.,1973 University of Alberta (Botany) M.S., 1969 University of Illinois, Chicago (Biology) B.A., 1966 Central Washington University (Biology) Academic Awards and Honors 2018 International Organisation of Palaeobotany lifetime Honorary Membership 2014 Fellow of the Paleontological Society 2009 Distinguished Fellow of the Botanical Society of America 2004 Ohio University Distinguished Professor 2002 Michael A. Cichan Award, Botanical Society of America 1999-2004 Ohio University Presidential Research Scholar in Biomedical and Life Sciences 1993 Edgar T. Wherry Award, Botanical Society of America 1991-1992 Outstanding Graduate Faculty Award, Ohio University 1982-1983 Chairman, Paleobotanical Section, Botanical Society of America 1972-1973 University of Alberta Dissertation Fellow 1971 Paleobotanical (Isabel Cookson) Award, Botanical Society of America Positions Held 2011-present Courtesy Professor of Botany and Plant Pathology, Oregon State University 2008-2009 Visiting Senior Researcher, University of Alberta 2004-present Edwin and Ruth Kennedy Distinguished Professor of Environmental and Plant Biology, Ohio -
The Seed Cone Eathiestrobus Gen. Nov.: Fossil Evidence for a Jurassic
American Journal of Botany 99(4): 708–720. 2012. T HE SEED CONE E ATHIESTROBUS GEN. NOV.: 1 F OSSIL EVIDENCE FOR A JURASSIC ORIGIN OF PINACEAE G AR W . R OTHWELL 2,3,6 , G ENE M APES 2 , R UTH A . S TOCKEY 3,4 AND J ASON H ILTON 5 2 Department of Environmental and Plant Biology, Ohio University, Athens, Ohio 45701 USA; 3 Department of Botany and Plant Pathology, Oregon State University, Corvallis, Oregon 97331 USA; 4 Department of Biological Sciences, University of Alberta, Edmonton, AB, T6G 2E9, Canada; and 5 School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, B15 2TT, UK • Premise of the study: Pinaceae and nonpinoid species are sister groups within the conifer clade as inferred from molecular systematic comparisons of living species and therefore should have comparable geological ages. However, the fossil record for the nonpinoid lineage of extant conifer families is Triassic, nearly 100 million years older than the oldest widely accepted Lower Cretaceous record for Pinaceae. An anatomically preserved fossil conifer seed cone described here extends the strati- graphic range of Pinaceae nearly 30 million years, thus reducing the apparent discrepancy between evidence from the fossil record and inferences from systematic studies of living species. • Methods: Material was prepared as serial thin sections by the cellulose acetate peel technique, mounted on microscope slides, and viewed and photographed using transmitted light. • Key results: A large cylindrical cone consisting of bract-scale complexes that diverge from the cone axis in a helical phyllotaxis has bracts and scales that separate from each other in the midregion and are of equal length and of nearly equal width. -
Early Permian Palaeofloras from Southern Brazilian Gondwana: a Palaeoclimatic Approach
Revista Brasileira de Geociências 30(3):486-490, setembro de 2000 EARLY PERMIAN PALAEOFLORAS FROM SOUTHERN BRAZILIAN GONDWANA: A PALAEOCLIMATIC APPROACH MARGOT GUERRA-SOMMER AND MIRIAM CAZZULO-KLEPZIG ABSTRACT In evaluating the parameters supplied by the taphofloras from different sedimentary facies in the Early Permian sedimentary sequences of the southern part of the Paraná basin, Brazil, it has become evident that the palaeofloristic evolution was related to palaeoecological and palaeoclimatic evolution. The homogeneous composition of Early Permian floral assemblages, which are characterized mainly by herbaceous to shrub-like plants considered to be relicts from the rigorous climate of an ice age (e.g. Botrychiopsis plantiana) suggest the persistence of the cold climate. The dominance of Rubidgea and Gangamopteris leaves with palmate venation associated with glossopterids with penate venation seems to indicate a gradual warming of climate. In roof-shales of coalbearing strata pinnate glossopterids related to Glossopteris are common, while Gangamopteris and Rubidgea (palmate forms) are poorly represented. The sudden enrichment of herbaceous articulates and filicoids fronds is characteristic of this stage and trunks of arborescents lycophytes become important elements. These antrocophilic paleofloras are characterized by typical elements of the "Glossopteris flora" associated to tree lycophytes and ferns communities. Therefore, the cool seasonal climate of Early Permian changed into the moist seasonal interval during the Artinskian-Kungurian. This climatic change was significant to the meso-hygrophitic to hygrophitic vegetation registered in roof-shale ferns of the Gondwana Southern Brazilian coalbearing strata. Keywords: roof-shale floras, Gondwana, Southern Brazil, Paraná basin, Glossopteris Flora INTRODUCTION The intracratonic Paraná basin, with a total area proposed by Milani et al. -
1058 Throughout the Evolutionary History Of
American Journal of Botany 99(6): 1058–1068. 2012. S EED CONE ANATOMY OF CHEIROLEPIDIACEAE (CONIFERALES): 1 REINTERPRETING P ARARAUCARIA PATAGONICA W IELAND I GNACIO H. ESCAPA 2,5,6 , G AR W . R OTHWELL 3,4 , R UTH A . S TOCKEY 3 , AND N. RUBÉN C ÚNEO 2 2 CONICET-Museo Paleontologico Egidio Feruglio, Av. Fontana 140, Trelew, Chubut 9100, Argentina; 3 Department of Botany and Plant Pathology, 2082 Cordley Hall, Oregon State University, Corvallis, Oregon 97331 USA; 4 Department of Environmental and Plant Biology, Ohio University, Athens, Ohio 45701 USA; and 5 Department of Biological Sciences, University of Alberta, Edmonton, Alberta AB T6G 2E9, Canada • Premise of the study: Seed cone morphology and anatomy refl ect some of the most important changes in the phylogeny and evolutionary biology of conifers. Reexamination of the enigmatic Jurassic seed cone Pararaucaria patagonica reveals previ- ously unknown systematically informative characters that demonstrate affi nities with the Cheirolepidiaceae. This paper docu- ments, for the fi rst time, internal anatomy for seed cones of this important extinct Mesozoic conifer family, which may represent the ghost lineage leading to modern Pinaceae. • Methods: Morphology and anatomy of cones from the Jurassic La Matilde Formation in Patagonia are described from a com- bination of polished wafers and thin section preparations. New photographic techniques are employed to reveal histological details of thin sections in which organic cell wall remains are not preserved. Specifi c terminology for conifer seed cones is proposed to help clarify hypotheses of homology for the various structures of the cones. • Key results: Specimens are demonstrated to have trilobed ovuliferous scale tips along with a seed enclosing pocket of ovulifer- ous scale tissue. -
Cryptic Diversity of a Glossopteris Forest: the Permian Prince Charles Mountains Floras, Antarctica
CRYPTIC DIVERSITY OF A GLOSSOPTERIS FOREST: THE PERMIAN PRINCE CHARLES MOUNTAINS FLORAS, ANTARCTICA by Ben James Slater A thesis submitted to the University of Birmingham for the degree of DOCTOR OF PHILOSOPHY School of Geography, Earth and Environmental Sciences College of Life and Environmental Sciences University of Birmingham September 2013 University of Birmingham Research Archive e-theses repository This unpublished thesis/dissertation is copyright of the author and/or third parties. The intellectual property rights of the author or third parties in respect of this work are as defined by The Copyright Designs and Patents Act 1988 or as modified by any successor legislation. Any use made of information contained in this thesis/dissertation must be in accordance with that legislation and must be properly acknowledged. Further distribution or reproduction in any format is prohibited without the permission of the copyright holder. ABSTRACT The Toploje Member chert is a Roadian to Wordian autochthonous– parautochthonous silicified peat preserved within the Lambert Graben, East Antarctica. It preserves a remarkable sample of terrestrial life from high-latitude central Gondwana prior to the Capitanian mass extinction event from both mega- and microfossil evidence that includes cryptic components rarely seen in other fossil assemblages. The peat layer is dominated by glossopterid and cordaitalean gymnosperms and contains sparse herbaceous lycophytes, together with a broad array of dispersed organs of ferns and other gymnosperms. The peat also hosts a wide range of fungal morphotypes, Peronosporomycetes, rare arthropod remains and a diverse coprolite assemblage. The fungal and invertebrate-plant interactions associated with various organs of the Glossopteris plant reveal the cryptic presence of a ‘component community’ of invertebrate herbivores and fungal saprotrophs centred around the Glossopteris organism, and demonstrate that a multitude of ecological interactions were well developed by the Middle Permian in high-latitude forest mires. -
Alien,· Elements in the Gondwana Flora of India
Alien,· elements in the Gondwana Flora of India A. K. Srivastava Srivastava AI< 1992. Alien elements in the Gondwana flora of India. Palaeobotanist 40 147·156. The occurrence of northern hemispheric taxa in Gondwana is considered as alien to the Glossopteris and Dicroidium floras of Southern Hemisphere. The morphology, evolutionary lineages, and stratigraphic distribution of Euramerian, Cathaysian and Angaran forms in the Gondwana flora of India are examined in view of their latest discoveries in the Permian and Triassic sequences. The study indicates that some of the elements of contemporaneous floras possess characteristic affiliation with the Gondwana flora, likewise some of the Gondwana elements exemplify the comparative characters of northern forms. Possible linkages, association, existence and ancestry of the Gondwana flora vis a vis Northern floras are discussed. " Key-words-Alien., elements, Evolutionary linkage, Gondwana flora, A K. Srivastava, Birbal Sahni Institute of Palaeobotany, 53 University Road, Lucknow 226007, India. ~T ~ ~ ~ ~;1' 'l'lWf<1 ..iil if 3Nlfll' '* ~~~ , ,l ~ I( I ~ ~~ ~ ~ ~ l<'1htTc~F<(1 ~ <:I~il~~ll'l 'l'1W~"11(11 ~ if <$ ott mroft <$ 1) 3RltiliTll' ~<$-.r'l'R ~ ~ ~ ~ ~ 'l'l14~"1ld ~, <t~ ~ ~ ~, 1Tt t I 'K4t if 'lIT«f <$ 1) ailm:'1' ott ~ ~ ~ ~ ~ ~ 'l'114~"1l<il ~~ ~3if Cf?lT 'iN 3fanr-r 11m tl 3fanr-r.r 0<f'fCf itcrr t f'q; <$ 3l'<fll'Cl' ~~t 3ltt>$~~~ 3l'<fll'Cl'~~<$~O<f'fCf~tl ~ 'l'114~"1iil Cf?lT~ 'l'1l4fc1"1ld.r w:IR"i\T ~ 'l'114Fd"1ldl <$ lltli W'llfC2l ~3if, ~'Cf?lT ~ ott ott 1Tt tl ruE Indian Gondwana -
Notes on the Evolution and Taxonomy of the Stachyospermic Gymnosperms
PR E S L I A 1959 31: 251- 272 Frantisek Ne me j c: Notes on the evolution and taxonomy of the stachyospermic gymnosperms Introduction Examining the ongm, phylogenetical evolution as well the taxonomy of the Gymnosperms, though most of the representants of this plant group be long at present to the rather remote geological past, we stand on a much safer and more concrete soil than in the case of the Angiosperms. The reason of that is no doubt the fact that this plant group left from its past much more numer ous fossils of such kind, which permits not only mere morphological studies of their external appearance but also studies of their finer anatomical struc tures. After the very numerous palacnotological researches and djscoveries of the last 25 years, it would seem at the first sight, that the Gymnosperms represent indeed one of the best recognized plant types as to its orjgin and evolution. But at a more detailed and thorough analysis of all our present knowledges we must however objectively admit, that by far this is not yet a true reality. Perhaps such problems and relations are much more evident in their phyllospermic branch (i. e. Pteridosperms and Cycads). Contrary in their stachyospermic branch (Cordaits, Ginkgos, Conifers) nearly all such principal relations disappear in the far past of the geological times, just as if we set to examine similar problems concerning the Angiosperms. And just several of these principal tasks concerning the origin and evolution of the stachyospermic branch of the Gymnosperms are subject of the following account. -
39. on SOME Nevv PLANT FOSSILS from the KARHARBARI BEDS, GIRIDIH COALFIELD, INDIA
STUDIES IN THE GLOSSOPTERIS FLORA OF INDIA- 39. ON SOME NEvV PLANT FOSSILS FROM THE KARHARBARI BEDS, GIRIDIH COALFIELD, INDIA P. fC MAfTHY Birbal Salmi Institut •• of Palaeobotany, Lllcknow ABSTRACT or spathulate-lanceolate in shape. The spa• thulate leaves have mostly broad obtuse Four new plant fossils Gang(unopleris oblanceoZ"la sp. nov., Euryphyllum obovalu1JI sp. nov., DoZian-ilia apex (PL. 4, FIG. 1) and the narrower leaves karharbarensis sp. novo and Buriadia jforinii sp. novo (spathulate-Ianceolate) tend to show some• are described from the Karharbari beds, Giridil1 Coalfield. what roundly obtuse apex (PL. 1, FIG. 2). Leaves have narrow tapering base. One INTRODUCTION of the margins of the leaf base shows more curvature than the other. Median sub• parallel veins are not prominent. Secon• beds, Giridih Coalfield have earlier PLA~T fossils from the Karharbari dary \'eins are gi\'en out from the median been described by Feistmantel (1879) veins at acute angle. The veins in the and Zeiller (1902). In recent years Maithy median portion are ± errect and show 1965a-e) has described the plant fossils radiating curvature towards the margin. from these beds. In a recent collection The course of the veins is ± straight. The some new plant fossils were encountered maximum width of the leaf is above the from the Karharbari beds, Giridih Coalfield. i length of the leaf. The densitv of the The\' ;Lr,', tl1"'reiore, described here in detail. veins is 12-16 per cm. - ComparisoN and Discussion-Feistmantel DESCRIPTION (1879) proposed Gangamopteris major for GA~GA:\IOPTERIS McCoy those leaves with a characteristic elliptical shape and ± pointed apex. -
On Vol Tziopsis, a Southern Conifer of Lower Triassic Age
PAPERS AND PROCEEDINGS OF THE ROYAL SOCIETY OF TASMANIA. VOLUME 101. ON VOL TZIOPSIS, A SOUTHERN CONIFER OF LOWER TRIASSIC AGE By JOHN A. TOWNROW Botany Department, University of Tasmania (With seven text figures and one plate.) ABSTRACT Diagnosis (emended). Conifers showing irregu Shoots ascribed to Voltziopsis africana Seward larly and sparingly branched shoots. Leaves dimor are described, with cuticular detail. These compare phic even on same shoot; smallest scale-like, about very closely with the shoot system, with an attached 3 mm. long and 2 mm. wide, more or less triangular, seed cone, caned V. wolganensis sp. nov., which is apex rounded or acute, about half as thick as associated with pollen of the Strotersporites Wilson broad. Largest leaves 2 or 3 cms. long and 3 to 5 sort. The shoot called, Brachyphyllum angustum mms. wide, directed forward, or forwards and out Walkom is redescribed though Walkom's speci wards, and if the latter, showing tendency to be mens cannot be found, and proves to be so close twisted into two rows up the shoot. Long leaves to the species referred to Voltziopsis that it is 2 or 3 mms. thick at base. All leaves having a reclassified V. angusta (Walkom) comb. nov. The thin but not scarious margin, lower (abaxial) material considered comes from the Lower Triassic surface rounded or angled, but lacking a distinct of New South Wales, and reasons are given for con keel, upper surface usually concave, or fiat, rarely sidering all Voltziopsis specimens Lower Triassic slightly raised over the midrib. All variants also. The classification of Voltziopsis in the Coni between extremes of leaf form found. -
The Botrychiopsis Genus and Its Biostratigraphic Implications in Southern Paraná Basin
Anais da Academia Brasileira de Ciências (2003) 75(4): 513-535 (Annals of the Brazilian Academy of Sciences) ISSN 0001-3765 www.scielo.br/aabc The Botrychiopsis genus and its biostratigraphic implications in Southern Paraná Basin ANDRÉ JASPER1, MARGOT GUERRA-SOMMER2, MIRIAM CAZZULO-KLEPZIG2 and RUALDO MENEGAT2 1Setor de Botânica e Paleobotânica, Museu de Ciências Naturais, Centro Universitário UNIVATES (SBP/MCN/UNIVATES),95900-000 Lajeado, RS, Brasil 2Instituto de Geociências, Universidade Federal do Rio Grande do Sul (IG/UFRGS) 91501-970 Porto Alegre, RS, Brasil Manuscript received on February 3, 2003; accepted for publication on July 23, 2003; presented by Alcides N. Sial ABSTRACT Botrychiopsis has been considered an important floristic element of Westphalian/Artinskian associations of the Paraná Basin. The occurrence of Botrychiopsis in roof-shales of the Rio Bonito Formation in Southern Paraná Basin (Quitéria area), supported by the identification of Botrychiopsis valida, enlarges the genus biochron. Consequently, the stratigraphic hierarchy for Botrychiopsis plantiana and Botrychiopsis valida was defined for the Paraná Basin. Although it is climatically controlled and related to a deglaciation icehouse stage, stratigraphic distribution of the genus presents a substantial climate tolerance, from cold/cool to warm/temperate conditions. A new phytostratigraphic zonation is proposed for the southern portion of the basin that includes the Botrychiopsis Zone (Asselian/Kungurian), which is subdivided into the Botrychiopsis plantiana (Asselian/Artinskian) and Botrychiopsis valida (Late Artinskian/Kungurian) subzones. Key words: Botrychiopsis, biostratigraphy, Paraná Basin, Permian, palaeoclimatology, Gondwana. INTRODUCTION sic – Early Cretaceous and Late Cretaceous) encom- pass continental strata and volcanic rocks. Each The intracratonic Paraná Basin covers 1.700.000 megasequence corresponds to previously proposed square kilometers of East and Central South lithostratigraphic units.