431

NOAA Technical Report NMFS Circular 431

Guide to Some Trawl-Caught Marine From to Cape Hatteras, North Carolina

Donald D. Flescher

March 1980

U.S . DEPARTMENT OF COMMERCE National Oceanic and Atmospheric Administration National Marine Fisheries Service NOAA TECHNICAL REPORTS

National Marine Fisheries Service, Circulars

The major responsibiilties III the "attllnol \Iann.' FI,hpnp, ",'rvllP I:-':\HSI ArI' t(, m(,ml"r snd delhI aIJundan(t and j(Pfll(raphlf dlStrihution IIf lisherv r~sources, til unlierslunli snd prelil' I flUCI'.I1t1 ",ns In t hi' 4U n',1\ and d, trJbut H'n (,I Ihe < re rune and til E' labli h leHI lor optImum use of the resour('es, ,\:\IF:-; IS also (harged wllh Ih., d,'v!'lopmenl and ,mT,lementst,,·n "I poil" , I'lf man gllll( nallonal II hlnl( grounds, development and pnfoflempnt 01 dome,t'l Ii,hen," rpgu~AtH·ns, "If\(,1;18nCI' (of f'·rell(n f, h nj( "ft I nl'('(l"l I" ('Ia till "dler ann thE' development and enlorcement "I IllternatillnallJsh.n ul(rf't'menls and pollCl"s .... :\1~" .,1,,· a" I Ihe" hlng Ind •• Ir thr"ul'h rnarhltnj( ef\ ce and e(,lInllmlt anal\", prClgnl'n,. and ""rtgage n, Han' e and \(,,,(,1 (·m trurtll" L,h"d'e It, (" e I anal\le lnd puh he \3f1I1US phases "f the Indust r\ Tht' ,\OAA Technical Hpp"rt '\IF:-. t'Jr(ular "'rJ!" r m',n Ie'" ,eflt th,,' hft heN, n ( le'He ,net'. II. The (,r- 11M publicallons "I general'l1lt'resl Inlennt'ri I" ~J(! (lI"pr\.lIH·O and manag('rr(nl Puhl" tl' In 'Ilat rt"~" .n ('Ill lripnhp dr'~" lechnl(alleHI cerlaln hn an arfd' It research ~T:)ear in 'hi ,tnt· I fllln 01 paper. r i! IOallr n C(ln' IT. I< tuelj( mel fr" '11 If gpmpnl n \'£>stlgatlnns appt'ar In the Clfl'tJ!ar senes "()AA Tf,hnlcal Reporl .... \IF" (lfCUH, an' ,J\d lahl.· Irep n lim,ted r,umber t 'j!()\t'rnm('r I 19pn(, , hoi h h'der I dnd al,,( a\~ilah e In t:'xchangt' lor ether Sl:t'rttlflc and 'PChOlJ1 ( enter, OA Iwck" Ie \10 LOh:>': Recent Clr· ('ulars .Ire

396 \\"hales. do,phlns, and porpo.se, of the we,tern ~(>r"1 A.t an'. A err m • t Pc n ng OHlce \\ a r n '''' U(, .l1J-IO~, tock L 003·01 gUide to the r Identlhcal1c n Hv Stephen Leathe", lod Ua\ Id h lalcJ lI<.-tO t well. and Hc\\'ard E. \\'lnn AL".1S· l'n, 1\ ~ l~6 P , l08 hg ., _ tat .. I~ app ligs. For sale b\ the "upcron'er.lent If Doc..l lents. l S (r.O\ Il~ \In~ ne 1 oru and f..lf of the n lrthes 'ern I IlIled t8 ernmer! Pnntlng OlflLe, \\ ashl1gt >n U( _(4u2 ~'cck () 1)0<·(2) I lphl zoa Hy Ibnald J Ld"'>J1 Au "ht 'l~t 11" I P 2.~ figs ~or sale by the :-'upennle'1dent 01 Docu nen's c.. l\fr'1 1 11' rthea tern llllted men' Pn'ltl'lg Ofl".,·e \\ dshl'1gt~n Dl _(4l2 COO, \\ood OO:lti, I cem"er .,~ I I )~ chart

19~ \Ian 1e lera and launa 01 the r Jrtheasterr l n ted Dte Hlghr .uS ll.('ctlc, I' JIl_ ba.tI, h papers 12 "aper;.) B} RIchard ~ Deuter~my,ete,. n) cete~ R lungl Ascc mneles. and Ha,ldlc 1:3) A t: lm..lrd edl' -I Decem"er !9~~ 11 ~ it ~ p l_val ere :'I Iarci' IJ7-, 11 + ~'1 p., 2~ Ilgs For ,a _ t) de ::-.upe r inlendent l Documents. l S (",vern 'len' Prlnt.ng Ollie. \\ a,hlJ1gtc:1, 41 \Iar ne I "3 ana fd .na the narthea tern l r..ted tates DC 2()~li:.. :-.tcck '\c Ol'l- ):!()·ltU26 _ l lpepod (h)PC ,J paras. tiC m hshe B\ Jl." he\ Ho Februa 19 .11" 12 P 1 IIg

399. :'Ilanne ;1ora and faun" 01 the northeastern L'n ted ,.,tates ra'" trip bl Tn m the. P\\ YJrk B,ght ('ause lopepoda Harpactlc( da B\ Bruce (' lOl•• 1 \1urct< 977 11'" 4 p., ",.,d ""(, iJt'4Le lee, B\ Thomd ( :\Ialcne \Ia} 19. , 1\ + 14 p., 1- 100 figs. for sale b} the Superintendent of Document>, l :'. Gcvern II,::" , 1 table ment Printing O;\ice \\ asbmgtun Dl 20~C2 :-.tock c lIl3 02U-ll(.I12.J· -l 411 'S"temlIlC, and blole-I:) 01 the lI.elbhe (Per~.fc.rmb Branchia­ 400 Fishery publication mdex. 19b5-74 B\' \Iar\, Ellen Engett and 'te~.Jtle ~nd \1,,:.l..tl'ltt< dtlel, \\ th dbcnptHm 01 two ne\\ .. pecle, B} Lee C Thorson :'Ilarch 19 •• , ill ~ 2:!O P For ,a.e b\ the :'uper Jilme, K Doole) Apn . • q-~ \ ~ -, P 44 flJ:s , :?6 table> mtendent of Documents, l ~. Government Prmtmg Office, \\ ashingt m. D(, 20-lJ:!. ,.,tock ;-"0 Jl):)·WO-OO.~7·1 112 ~\ ~op'" c! blO,"Jg, • .l1 -idta on the red porg:" Pagru. pa~rus ILIIl' 'laeu, H) ('harle- 'I \Ianooch III and \\ :!ham \\ Has>ler :'.Ia) 401 Fisheries and flshe" resources c I .... e\\ Y lrk Bight. By J L 197' I t 19 p .. l_ hgs , 7 tables. For sale b) the Superintendent 01 I\lcHugh :'Ilareh 19~-. \. + 5" p., 4.3 I'gs., -19 tables For sale l:>y the D('(Lmenl" l'.'1 (.overnment Pnntmg Oftice. \\"a~hlllgton. DC ~0402; Superintendent of Documents, L,.S Government Printmg Oflice, Wash­ :-.tock '\ l 00jl),-·OOH,,·(I" mgton, DC :20402, Stock;-"o 003-0:20·00129-~. H3 \Iarine nora and fauna ot tbe northeastern l'mted States. 402. Guide to the identilication of scorpIOn larvae (Family Seor­ ('n.,taeea· Hranlhiura Hy Ro!(er F Cre,.. e), :'Ila) 19-:~. Iii + 10 p., 15 paellldae) m the eastern PaCIfic wllh comparative notes on species of IIgs. For sale Dy the Superintendent of Document., L'.S. Government ::>eba,tes and Helicolenus from other oceans. By H. Geoffrey :'Iloser. PnntlJ1g Oltice, \\·a,hmgtan. DC 20~U~, ::;tock :-':0 OU3·017-lJ0419·8. Elbert H. Ahlstrom, and Elaine:\ol. Sandknop. April 1977, v + 71 p .. -IO figs .. 38 tables. For sale by the Superintendent of Documents. C S. Gov· ~14 ::;ynop,is of bIOlogical data for the wmter flounder, Pseudopleuro· ernment Printing Office, Washington, DC 20402; Stock :-':0. 003·020- neet .. " amencaf1W \\\albauml. By Grace KlelJ1·:'IlacPhee. :-':ovember 00128-9 ;9.8. Iii + ~3 p, 21 figs, 28 tables.

403. Marine flora and fauna of the northeastern United States Sipun· H5 A baSIS for c1asslfYlJ1g western Atlantic Sciaenidae (Teleostei: cula. By Edward B. Cutler. July 1977. Iii + 7 p., 6 figs. For sale by Perciformesl. By Labbisb :-':ing Chao. September 19,8, v + 64 p., 41 the Superintendent of Documents, U.S Government Pnnting Office, figs., 1 table. Washington, DC 20402; Stock No. 003-012-00404-0. 416. Ocean variability: Effects on C.S. marine fishery resources 404. Revision of the basses of the Diplectrum (Pisces: Ser­ 1975. (20 papers.) By Julien R. Goulet, Jr. and Elizabeth D. Haynes, rallldae). By Stephen A. Bortone. September 1977, v + 49 p., 15 figs., Editors. December 1978, iii + 350 p. 9 tables. 417. Guide to the identification of genera of the fish Order 405. Marine flora and fauna of the northeastern United States. Echino· with a tentative classification of the order. By Daniel M, dermata: Holothuroidea. By David L. Pawson. September 1 Co.hen...aruLJ0l"1l'en..G ieJR A""= .b..~ NOAA Technical Report NMFS Circular 431

Guide to Some Trawl-Caught Marine Fishes From Maine to Cape Hatteras, North Carolina

Donald D. Flescher

March 1980

U.S. DEPARTMENT OF COMMERCE Ph p M Klutznlc - Sec e a National Oceanic and Atmospheric Administration R chard A Fran Adr1" S ra''''' National Manne Fisheries Service Terry L Le tze Ass s'an Ad~ ua I" e es The ational ;\!a rine Fi henes en'ice (.'.1 F. ) do S not appro\' • r ec­ ommend or endorse any proprietary produc or proprietary mat rial mentioned in this publtcation. .'0 r fert'nCl, hall b mad to. 'Mr . • or to t his publication furni 'hed by • ',\1 F ~, in any ad\' r iSing or all' pro­ motion which would indicate or imply that. ·.\If appro\' • r comm nd or endorses any proprietary product or proprietary maLrial m nlion d herein, or which has as its purpose an intI nt to cau e dir etly or indir ctly the advertised product to ve used or purcha d b cau e of thi • '.1 pu blica tion. Introduction ...... Introductory key to fib he ...... 3-7 Shark. except angel hark ...... Skates, rays and angel bhark ...... , ray - skates ...... 10 Skate ,rays - rays ...... 11 kate ,rays - tin grays ...... 12 -shaped fi he ...... 13,14 Herring family ...... 1.5,1 ' Anchovy- haped fishes ...... 17,1 family - one or three dorsal fins ...... 1 Cod family - two dor al fins ...... 20 Bass-shaped fishes - one ...... 21 Bass-shaped fishes - two dorsal fins ...... 22 Searobins and sculpins - searobins ...... 23 Searobins and sculpins - sculpins ...... 24 - right-eyed flatfishes ...... 25 Flatfishes - left-eyed flatfishes ...... 26 Mackerel and - haped fishes ...... 27 Goosefish and butterfish ...... 2 Index of common names ...... 29-31 Index of scientific names ...... 32-:34

iii Guide to Some Trawl-Caught Marine Fishes From Maine to Cape Hatteras, North Carolina

DONALD D. FLESCHER1

ABSTRACT

Fishes covered are those regularly caught during trawling oper ations. S imila r shaped fhhc, arc grouped together. On each page the written keys are connected by lines to the fish illustration.: <:on,c­ quently, technical terms in the keys a re illustrated as they are used. ;\Iotes on the size and range or each fis h are included.

INTRODUCTION cation. The reader is referred to the following text for a more extensive coverage of the fishes: This guide is designed for the quick identification of trawl caught fishes under sometimes difficult field condi­ "Field Book of Marine Fishes of the Atlantic Coa -t" by tions. The species that are included are abundant in bot­ Charles M. Breder, Jr. 1948. G P. Putnam' Son~, tom trawl catches of National Marine Fisheries Service 332 p. This book also mcludes the e tuarine peCles a~ (NMFS) research cruises on the continental shelf. These well as those whose center of abundance IS south of Cape cruises cover the area slightly northeast of the Gulf of Hatteras. It is pocket-sized, which is helpful for in-the­ Maine to Cape Hatteras, N.C., from about 5 to 200 fiel d use. fathoms. Estuaries are not included. "Fishes of the Gulf of Maine" by Henry Bigelow and Fishes with similar characteristics are grouped William Schroeder. 1953. U.S. Fish and Wildlife 'er­ together even though they may not be related. When vice, Fishery Bulletin, vol. 53, 577 p. [Available as a identifying a fish, if you cannot decide on which page to reprint from the Museum of Comparative Zoology, Har begin after leafing through the guide, you can use the in­ vard University, Cambridge, MA 02138.) It include troductory key on pages 3 to 7. The keys are for identify­ not only the usual fi hes of the Gulf of 1\laine and ing adult fi hes. The body proportions of immature fishes Georges Bank but all that have ever strayed into that may be quite different, and some body parts may not area. Exten ive information i given on the bIOlogy and have developed yet. economics of each species. A geographical range is given for each species. This is "Fishes of Chesapeake Bay" by ~amuel Hildebrand the to~al area over which it has been found. It may be ex­ and William Schroeder. 192. Bulletin of the T pected to be abundant within a small area of this range. Bureau of Fisherie , 43(1): 1-366. [A 1972 repnnt i" Many species occur quite frequently in NMFS trawl available from T. F. H. Pubhcation , Inc ..• 'eptune, •.J catches but are not considered to be abundant. These 07753.J Although about 50 year old, thi publication species are omitted in order to keep the guide small. gives good coverage of the biology and economic impor­ Therefore any fi h that does not exactly fit the key char­ tance of each pecie . The T. F H PublicatIOn repnnt acteristics or that looks different from the majority of the bring the scientific name up to date. individuals can be preserved (10'0 Formalin or full "Fishes of the AtlantiC Coa t of Canada" by A. H strength alcohol works well) or frozen for later identifi- Leim and \\. B. 'cott. 1966. Fi hene Re earch Board of Canada, Bulletin 15.5. 4 5 p. It cO\er the

:\orthea t Fisheries Center Woods Hole Laboratory. :\'ational ~larine fi hes found between the Gulf of ~laine and Labrador out Fi heries Sen·ice. NOAA, Woods Hole. l\1A O~543. to 1,000 fathom .

1 Source of Drawings

Forty-eight of the drawings came from the files of the United States National Museum (Smithsonian Institution). Thirty-one are from the book "The Fishery Industries of the United States, Section I, History of Aquatic " by George B. Goode, 1884. Twenty-one are from the books "Fishes of the Western North Atlantic," Part 1, 1948; Part 2, 1953; Part 3, 1963; and Part 6, 1973 (Sears Foundation for Marine Research, Memoir 1). Four are from "Oceanic Ichthyology" by George B. Goode and Tarleton H. Bean, 1896. Six other government and museum publications were each the source of one or two drawings. Illustrators at the National Marine Fisheries Service, NOAA, Woods Hole, Mass., drew the undersides of the winter and little skates and the gill rakers of red and white hake.

Total fish length

Caudal fin Gill

Ventral fi n Lateral l ine Gill arch

Parts of a fish used for fish identification. UTRODUCTORY KEY lao Has five gill openings on each side. ~ o to 2. 1 lb. H ither one or no gill opening on each side. o to 3. cross-sectlon body in cross section ;s more or less rounded. • angel shark, p. o cross-sec-tlon b cross section is flattened from bell o bac". 1 sharI<. p. to 12.

• n in and no rnal eves.

n 4b. Mouth not enormous, most teeth in lower jaw not exposed when mouth closed. Go to 5.

5a. Body flattened in cross section; both eyes on the same side of the head.

See flatfishes, p. 25, 26.

5b. Body more or less rounded in cross section; one eye on each side of head. Go to 6. ..,.

6a. Body tapers to a whiplike tail ("rattail"). t

See grenadi er, P. 13.

6b. Tail not whiplike.

Go to 7. c ... ~

7a. Body long and slender: body's greatest height (not counting dorsal fin) less than or equal to 1/5 of total body length; has only one dorsal fin which is at least 2/3 as long as total body length. A See eel-shaped fishes, P. 13, 14. B < 1/5 A c > 2/3 A lb. Body shorter and stouter: body's greatest height (not counting dorsal fin) greater than or equal to 1/4 of total body length or the longest dorsal fin is less than 2/3 of the total body length. Go to 8.

8a. The belly in cross-section has a bottom edge that ~\@ o f ""o" ,,,, is sharp edged. )) ~ ~~ , See herri ngs, p. 15, 16 and butterfish, p. 28. - - . ~ \ \~ 8b. The belly in cross-section has a bottom edge that is more or less rounded. () Go to 9. CJl \ 9a. Numerous light-producing organs (photophores) along the ventral surface.

See pearlsides and lanternfish, ~ 17.

9b. No light-producing organs (photophores) along the ventral surface. Go to 10.

lOa. Four or more small fins between last dorsal fin an~ caudal fin and between anal fin and caudal fin.

See mackerel and tuna-shaped fishes, p. 27 . lOb. No small fins between last dorsal fin and caudal fin and between anal fin and caudal fin. Go to 11.

<4!• lla. Base of longest dorsal fin 1/7 or less of total ~(~ ~B ---~i~',~ body 1ength. , • ) ,f) ,-..;:: "" _ -:::.- ~ ~l ~ "'~ See anchovy-shaped fishes, ~ 17 , 18 . I .. ~I A llb. Base of longest dorsal fin 1/6 or more of total body length. B < 1/7 A Go to 12. CT>

12a. The front half of the first dorsal fin is supported entirely by segmented, fairly soft bones (called rays); start of ventral ~ fin is located directly beneath or forward of start of pectoral fin. ~ See cod-family, p. 19,20.

12b. The front half of the first dorsal fin is supported entirely by unsegmented, often very hard bones (called spines); or start of ventral fin is located behind Start of pectoral fin. Go to 13. ~ ~

13a. Two dorsal fins. Base of anal fin long, more than 1/5 of total body length. Pectoral fins large, usually more than 1/5 of total body B > 1/5 A length. usually C See searobins and sculpins, p. 23, 24.

13b. One or two dorsal fins. If two dGrsal fins are present, base of anal fin usually less than 1/5 of total body length. Pectoral fins small, usually less than 1/5 of total body length.

See bass-shaped fishes, p. 21,22. usually B < 1/5 A

usually C < 1/ 5 A -l .. .. A SHARKS EXCEPT ANGEL SHARK [-1

A chain-lik.e pattern of b'ack No d'l.l' r I he pattern of black stripes on back and ·,ides. ..tripes on back ~nL ~ides.

-I An ana fln. "1 ,;pine 1 ' No anal fin. A spine in frent t'~ e::ch dorsal fin. f"'Ont of" ea<:h dorsal fin. ~r---~~ 1- t CHAIN DOGFISH '" £ rh 81 ~. J~ ."1a:r:i-:1.Q1l ~ 2\ feet ,'T" . Range: Offshore (40 to 125 fathor.""'i, 're­ New York to North Caro 11 na ~ \

:'.J~ 'lY OCii.. " I(',ti

F rst at' tj lrsal (1'!,. r t I a ""Jcr about ~ u h __ n

First dorsal fin begins over ,"ear edte Of pectora~ a1 fir'! 00 fin. Teeth tiny, flat, and pavercnt-' ~_e nf 'leI-to .. a1 fin.

-'- . ., , ,,~~._~. - , :~.~ ."', .... -.J

SI'llOTH DOGFISH 1.f;:.nn;ut!f n.~ 5 feet Cape Cod to as far sou!.h as .Jrugua

.~e inne'" a.,~'e o""q);t f' ., 1a1"Qel'" ~ts~; ~ht is at leas1' t"; i'hir"lj ~P' openi'9

_ t /( ~~ .. I!{(I ---- ~~. ' -----. \..- -'" ~ - . _ ---,0' ~~~ ~ ~I SA.\D8AR SHAll/; =-<.,.".., n~ (8""... S~.r' offr"lf'l~ ..aters 7 2Jl , ••t I ..l-- n :m in inshO,.. and offsh:re wa t~rs f .... Cope Cod to FlorH. ,- SKATES, RAYS AND ANGEL SHARK Mouth located at very front of head. Distinct notches between Mouth located some distance back on underside head and front edges of "wings" (pectoral fins). of head. No notches between head and front edges of "wi ngs" (pectora 1 fi ns) . 1 There are two dorsal fins at tip of tail. There are no dorsal fins at tip of tail . / There ilre never any spines ("stingers") on tail. There may be one or mo re spi nes ("sti nge r s") on tail. SEE RAYS , PAGE 11 3 ...... , t·· ...... I I One or more rows of conspicuous thorns in middle of back No conspicuous thorns in middle of back, i n area A. ;'.(.: in area A. No black dots or dashes on lower surface. Lower surface has black dots or dashes. \ ANGEL SHARK Squatina dwnePi!i /\ Maximum siae: 3 feet Range: Cape Cod south to Jamaica. ,',,- Numerous thorns, a 11 very ti ny, One or more rows cover the top of rear 1/3 of tail. of moderate to large thorns on top of ~' , rear 1/3 of tail. il ~ Q; ~ -:. j: ~~{ .... ;/' _ 'i " ~ .,' "" .,,:. __ . /C / - Ie '" I... "- - /' ""-..qf(1:J -~ /~~IJ {~I SMOOTH SKATE Raja senta ., (Smooth-tailed skate) , , <:0 !rtzximum size: 24 inches total length. \:f ~ Range : 0 ff Newfound 1and to South Carol ina. Mostly in depths greater than about 40 to ':~!;!U;n\:" / ~ BARN DOOR SKATE Raja l.aevis '"'-.J 50 fathoms, ~\.v On the back there are no On the back there are conspi cuous Ma:r:irrrwn 8ize: 5 feet in length. \\ ,,' conspicuous ddrk rosettes. dark rosettes (a rosette is a group Rang" Off Newfoundland to North '•• ,~~ of 6 or more dark brown or black Carollna...... ~.".# ••• ~ / spots surrounding a central spot). I ~ The thorns of the mi tidl e row on No one row of thorns on the tal 1 "..,--. the tail are much larger and is much larger or more conspicuous ,,"' 'II ~ ~ conspicuous than any other thorns than the other thorns on the tai 1. ~ on the td 11. There are g or There are at least 15 thorns " !¥ f ' " \\ in each of the rows on the ta i 1 . ,:

~:~ ,.: ~~'. • :i< ,>,:::, ::~>:.',. ROSETTE SKATE Raja garmani ~ t~~~'~~;A~~~~ ';,-\4' ."... (leopard skate) ~J.i; 'k' Size Th l S one of the smaller skates '" >~:;:~·~/X:' ,» ,THORNY SKATE Raja radiata . ' and grows to a total lenqth of at Irtzxirm.mr size: 40 inches least 16 inches. Range: Both sides of the North Atlantic. Range: Off Nantucket to Florida, in In North America from west of Greenland depths of 30 to 300 fathoms, to South Carol; na. Restri cted in genera 1 to water deeper than 10 fathoms. (continued on next page) SKATES (continued from SKATES, RAYS - preceding page) , 1 Only one row of large thorns in the middle of the b~ck, There are at least three rows of thorns in middle of back in area B. Upper surface of body marked with roundlSh in area B. Upper surface of body marked with spots but spots and short bars. The first and second dorsal fins are has no short bars. The first and second dorsal fins are separated by a definite space or at least 1 or 2 thorns. not separated by d definite space or by a thorn or thorns. · I I I Generally less than 54 rows of teeth Generally roore than 80 rows of teeth ""'0:: in upper jaw. Rarely exceeds length of in upper jaw. Grows to a length of ~ , '\. ~ about 21 inches (54 centimeters) or about 43 inChe.s (l09 centimeters) and '_ ,.".Jfj I J.. \ weight of 2 pounds, often exceeds a weight of 2 pounds, ~ ~ ~ I ./',I ., ·\ J #. t ~ • \ i' .0.1.:_ i:: 8 't: 111 /'.' ". ~\ • ~ '"!. r- ">.. _,./);' . • \ ~,~. .~.. .. " ., ~ ., .... .,'.. :;.\ ".' • -----" ~. " . . .."!":"jl ,'..., ..~}; ,1 I -_ .... ,.., c ...... _ . ,.--/I /'. .. I"/. :.:...... ,...... :. ~ t:.:·' ... . j ~ -; ~ • J , ~ -...... \' I . "- I r;~:j/::=: '. ~", C>; I ,~ :. '\ . . '0. . ~ ::-0. ... ~, .:. ... ~. \... ./ " '\ "Z.,. .... ,' ~> /--'" CLEARNOSE SKATE Raja eg1An"" a ," .-.' I ..... (Brier skate) o Ma.rimwn size: 37 inches in total length Range: Massachusetts to Florida ,_ ...... LITTLE SKATE R .. ,u eM.>la" AkJ.z1.1!Mr' ria" Rarely ex{eeds. ci total length 0' about WI~TER SKATE ''''' 1 '"

Note In U S water$. fiSh longer than ~ :entllTl?ters (21 Inches: w1"l usually tum out to be winter skates, and s.peclmen~ longer than about 60 centc'X'ters (24 Inches) are allh)Sl certainly winter skates. There is a problem when separating winter skates that lire less than 54 centimeters from little s).ates by counting rows of teeth, sinle It is 'iO tite· :onsUftling as to be- hnpractical in the field. However. if each selt is conSidered separately. "hose ;n t·"!e size range of 35 centine~ers (14 inches) to S4 centllDeters can be readlly identified. This is the size range 1n which 1 ;ttle skates sMw matur' '\9 or mature e).ternal se~ chardt.:ters. and the winter skate shows tmnature external se. chuacters For fiSh under ab\lut 35 centimeters. though, both species are lar.ature and there is no easy characterist1C tor quickly separating the:;l In the field. The sues can be separated as follows l\ales have 2 cla~pers (copulatory organs). one at eden slde of the start of the tail: females lad claspers.

Fish between 35 and 54 cent1meters ---, On underside of body there are two patches of spines, one On undersl de of body there 1s no patch I on each side of the vent. (Note: The spi nes a re often of spines on each side of vent The claspers e).tend wel t. ''tI n The claspers e1 ther -.on t reach posterior difficult to see even in good light conditions but they no.terior ed9"1 of d'" ed~. of jl,C:'::.'"" ".",ly ~eyond it. can be fe 1t 1f you move your f1 ngerti pin the ta 11- toward·head direction). I

I" . .. I, Ifllf '1 '~J_~Ii" A~!) ' ''I ] -::.2 .;:::,. (~ , .~ J, \.\\.... ' /~ ,\\' ~' .... ,-~.t ~ ~ .

LITTLE SKATE . female. under.ide WINTER SKATE. f.male. under'ide LITTLE SMTE. ... le, unde"lde WINTER SKATE. ... 1•• underside SKATES, RAYS - RAYS

I I Outline of front edge of fish. from ~1ngtip to O~tline o~ front edge of fish, from wingtip to wingtip. interrupted by head protrudlng forward. wlngtip, 1S approximately v-shaped. That is, I head doesn I t protrude forwa.rd appreci~bly.

Underside of head

COWNOSE RAY Rhinoptera bOY/a.SU8 BULLNOSE RAY: MyliobatiB fl'eminvi7.lei ...... Maximum size: About 38 inches wingtip to M::z:cimum size: 34 inches wingtip to wingtip . wingtip ...... Range: Cape Cod to Brazi 1. Range: Vicinity of Cape Cod {Nantucket, Woods Hole} to Brazil.

Tail short, much shorter than distance from snout to start of tail. Wingtip to wingtip distance much Tail long, whiplike. much longer than distance from greater than distance from snout to end of tail. snout to start of tail. Wingtip to wingtip distance much less than distance from snout to tip of tail. F \ One or two spi nes No spines (stingers) (stingers) at base of tail. at base of tail.

il' / \ (j\ '1 f I if J r; I \ \- ; \ .. -' ( \ ,! j ('-~ \ I il J --,,,,-// \ !Ii ~-----~~-j, -~ .---,,'-- 'Il~l~ SPINY Gymnura aUave~a , Ma.x:imwn 8ize: In U.S. waters 6 feet 10 SMOOTH BUTTE RFl Y RAY Gymnura mioPUra inches wingtip to wingtip. Ma:r:imum size: 3 to 4 feet wingtip to Range: Both sides of Atlantic. In wingtip. Range : Cape Cod to Brazil western Atlantic Cape Cod to South America. (continued on next page) / /

)'. ) ./

l~~ ______EEL-SHAPED FISHES .----- Has no j aws. no pectoral f1 nand Has jaws . pectora 1 f1 nand no externa 1 eyes. external eyes. r -- Has one dorsal fin. Has two dorsa 1 fi ns but the second one ; 5 so much lower than the fi rs t tha t it may be diffi cu 1t to see. Body tapers to an e 1009a ted whi p 1 ike tail ("rattail") with no distinctive caudal fin .

~~~-~ . +" .... - f ATLANTIC HAGFISH Myxi"" glutinosa Maximwn siu: 31 inches Range: Both sides of Atlantic. In North America, Arctic to North Carol ina. GRENADIERS ("RATTAILS")

Note: There are many species of grenadiers and all live in deep water. In U.S. waters a few are occasionally caught in depths less than 100 fathoms. One, the Marlinspike, Three barbels above the mouth: one in front of each nostril and Nezwrria bail'di, is shown above. the third on the top of the snout. A fourth barbel hangs from the No barbels above the mouth. chin. r------Anal fin and caudal fin not Anal fin and caudal fin ...... separated by deep notch or gap. separated by deep notch or gap . W I

FDURBEARD RDCKLING gnahelyopus cimbrius No ventral fins or chin barbel. Ventral fins and chin barbel present. Maximum size: 12 inches in North America Range: Both sides of North Atlantic. In North America, Gulf o~ St. lawrence to North Carolina.

----e.: t CUS K 81'Osme bro8TTIe Ma:r:imwn size: 3~ feet. 27 pounds Range: Both sides of North Atlantic. In North America, Newfoundland to Virginia.

Snout rounded and large canine teeth present.

Snout pointed and no large canine teeth. --

ATLANTIC WOLFFISH Anarhicha8 lupus (Dcean ) SAND LANCE Ammodytes species (continued on Maximum size: 5 feet, 40 pounds Maximum size: 7 inches next page) Range: Both sides of North Atlantic. In Range: Labrador to North Carol ina. I North America, Greenland to . SKATES, RAYS - STINGRAYS

(continued from I preceding page) I ------.._------,

Tail is rough because of large number of Tail ;s smooth. lacks thorny tubercles. thorny tubercles on it. Tail has a Tail has a keel-like fold of skin on keel-like fold of skin only on underside. both':'ide and underSidj

~---~=~---.-:;:

Detail of tail showing thorny tubercles and single Detail of tail showing two keel-like keel-like skin fold located on underside of tail. skin folds just behind "stinger. n

1 >-' j Iv J / ~. \t_ iJ ~ j _ f J 'Ii :1I \. ( h.. -. \ '", ""- ~ /

ROUGHTAIL Daeyatie oentpo""a BLUNTNOSE STINGRAY Dasyatie eayi (Northern stingray) Maximwn size: One meter (about 39 inches) Ma:rimwn siae: Nearly 7 feet wingtip to wingtip to wingtip. wingtip. Range: Southern Massachusetts to Brazil Range: Cape Cod to Florida or farther south. EEL-SHAPED FISHES

Has no jaws. no pectoral fin and Has jaws. pectoral fin and no external eyes . external eyes.

/ ._-- ,\\ '. ?i Has one dorsal fin. Has two dorsal fins but the second one ;s so much lower than the first that it may be difficult to see. Body tapers to an elongated whiplike tail ("rattail ") with no distinctive caudal fin.

-...... f ATLANTIC HAGFISH Myxi"e glutinosa Maxirrrwn size: 31 inches Rallge: Both sides of Atlantic. In North America. Arctic to North Carol ina. GRENADIERS ("RATTAILS")

Note: There are many species of grenadiers and all live in deep water. In U.S. waters a few are occasionally caught in depths less than 100 fathoms. One, the Marlinspike, Three barbels above the mouth: one in front of each nostril and Nezumia bair>di, is shown above. the thi rd on the top of the snout. A fourth barbel hangs from the No barbels above the mouth. chin. / ... _-----, Anal fin and caudal fin not Anal fin and caudal fin ...... separated by deep notch or gap. separated by deep notch or gap . v:> I r FDURBEARD RDCKLING Enohe lyopus OimbriUB No ventral fins or chin barbel. Ventral fins and chin barbel present. Maximum size: 12 inches in North America Range: Both sides of North Atlantic. In North America, Gulf o~ St. Lawrence to North Carolina.

t CUS K B'1'Osme brosme Ma:cimwn size: 3~ feet, 27 pounds Range: Both sides of North Atlantic. In North America, Newfoundland to Virginia.

Snout rounded and large canine teeth present.

Snout pointed and no large canine teeth. -- ATLANTIC WDLFFISH AnarhichaB lupus (Dcean catfish) SAND LANCE Ammodytes species Maximwn size: 7 inches (continued on Maximwn size: 5 feet J 40 pounds Range: Labrador to North Carolina. next page) Range: Both sides of North Atlantic. In I North America. Greenland to New Jersey. (continued from preceding page) r. .... ---- EEL-SHAPED FISHES ( CONTINUED'

Dorsal fin seems separated from Dorsal. caudal, and anal f'ln;t caudal fin by a considerable gap form one cont inuou! fin

OCEAN POUT MaL'l'OBcar.- ~B "17/c}"1...:anus (Eel pout) Iohrinrwn eiBe: 3~~ feet, 12 pounds Range ~ Newfoundland to Delaware,

No barbel·like fins - f) .. he thr'Oat Barbel t.hrott

No "art (~lne on . 'lo ... t rtwlrt sha""P ~)ioe on the t..JP of snout no : o~re" wi h ). i ., is edsily (e t if not seen (for it i-, nearly hi,jden 1n Skin, l..pper sides tovered with pa'e round spots ...... \

~

Gape of mouth reache~ only about .!,'it tdr 4S red" th r"t't of eye; body th1c~, t1p of o.'Ii1 soft dod nlundt'>d r, ' r' "" .< ~ \

CONGER EEL Conger ocelmimdr 14lrif'11W1f Sill(; 7 feet, 22 pounds in North Amt'ricd &mgt': Continental shelf of eastern North _rica, ~4(h1n9 as far north as Novll Scotfa HERRING FAMILY

A ~ Belly (A) is sharp-edged B 0 Belly (B) is more or lfss , in cross section. rounded in cross section . ~------~I------~ \

No rows of distinctive scales along Two rows of distinctive scales along middle of back between head and dorsal middle of back between head and dorsal fin. Head (C) sma ller, about \; or less of (0). fin. Head (C) large, about 1/3 of body length to narrowest part of tail (0). ~~~ ~'----

Top view of fish to show scales

ROUND HERRING Et""'""us teres MaxirttUnl size: 10 inches Range: Bay of Fundy to Gul f of Mexico

c • 1/3 0

(E) much less than (F). Distance from tip of lower jaw to start of Belly very sawtoothed. dorsal fin (E) equals d1stance from start of dorsal ...... fin to last large scales on the middle of the <:n tail (F). Belly slightly sawtoothed.

E • F

ATlANTIC MENHADEN Br-evool'tia tY1'a1lnU8 (Pogy) Ma:cirTtWrI size: 20 inches, about 1 3/4 pounds Range: Nova Scotia to Florida.

'...... i:t...:/...... --.- i.l....'..:.W ....

ATLANTIC HERRING C!upea haJ'engus (S.. a herri ng) Maximum siae: 17 inches, about llz pounds Tip of lower jaw extends beyond Range: Both sides of the North Atlantic. In Tip of lower jaw does not extend much North America, Greenland to North Carolina. beyond upper jaw when mouth closed upper jaw when mouth is closed

(conti nued on next page) HICKORY SHAD AZosa mediocris Maxirmun size: 2 feet Range: Bay of Fundy to Florida. (continued from preted1 ng page) HERRING FAMILY (COHT! HUED)

Upper outl ine of forwdra pdrt Jf lower jdw nearly stra1q;}. Cheek he ...... bone muth higher (G) th,n long (,; thdn 0)

I > J

J I AMERICAN SHAD AL- ea C.l.P 4- 68 .-,- , "h._.lmum st4e Z., feet, Il'1 pounds R:rn(.1tl Newfound14nd to florida. and on the United States Pacific COdtt ~

'I"th rlro'.na ANCHOVY - SHAPED FISHES ------1

No light-producing organs (photophores) present. Light-produc;ng organs (photophores) present.

Mouth small, extends only to about front of eye. Mouth large, extends beyond eye.

PEARLS IDES Ma/AI'OtiC!UB TTTUE Lleri Maximwn size: 2~ inches LANTERNFISHES (MYCTOPH IDS) Range: The open Atlantic. Many species of lanternfishes exist. Identification depends mos t 1y on the pas it i on and number of the; r 1 i ~ht- praduc; ng organs (photophores). They are small fishes (most will be shorter than 3 or 4 inches) which are most abundant in the open ocean beyond the continental shelf. One fairly common species is the HORNED LANTERNFISH CerutoBoopelu8 nad£l'ensiB. shown above. which has a small spine ("horn") pOinting forward just above each eye. Start of ventral fins in front Start of ventral fins well behind of rear tip of pectoral fins. rear tip of pectoral fins .

...... -l

SHORTNOSE GREENEYE Ch~ol'ophthalmu8 aga88~"i Ma:r;irrrum size: About 6~ inches Range: South of Cape Cod to South America. in 81 to 400 fathoms. No sma 11 f1 eShy fi n (ad i pose A small fleshy fin (adipose fin) behind the dorsal fin. fin) behind the dorsal fin.

/~

ATLANT IC ARGENTINE Argentina sHus Maximum aize: 18 inches (continued on Range: Both sides of North Atlantic usually in next page) wa ter as deep as 80 to 300 fa thoms. I n North I America from Nova Scotia to off southern New England. ANCHOVY - SHAPED FISHES (CONTINUED) I (continued from preceding page)

r Ventral fins located far behind dorsal Ventral fins located in front of fin. Mouth doesn't extend beyond eye. dorsa 1 fi n. Mouth extends beyond eye.

/ Start of dorsal fin far'the' Start of dorsal fin di rectly ROUND HERRING EtrwneU8 te,,,. over start of anal fin. Marimum size: 10 inches forward than tar. 0)', anal fin Range: Bay of Fundy to Gul f of Mexico

--­"\ ...... @ \ 00 ~L~

BAY ANCHOVY ~""" ""..,"'1, Y.l..r'i.r4!1 Biu: About l's inches i43ine to Texas.

Bone (maxillary bone) forming the bottom edge Bone (,...1114ry bone) fanning bottom edge of upper jaw reaches about to g111 opening of upper jaw stops far in front of gill openlnq and is painted at its posterior tip. and 1s rounded at its posterior tip.

'"'= " ,....-'

STRIPED ANCHOVY Anchoa hOp •• 'UB S I LV ER ANCHOVY DIg_ L'" r""1/" Maximum sias: About 6 inches .~.l.l.t-imwnr sba -' Abou t 6 1nche!. Range: Nova Scot to Uruguay. 1a Iiim(l .. : In sUlmler It 1S cOIfrnOn offShore between Massachusetts and North Carolina COD FAMILY - ONE OR THREE DORSAL FINS

Til ree sepa ra te dorsa 1 fi ns One dorsa 1 fi n and two anal fins. and one anal fin. ------·-l

The ;s pale; "" The lateral 1 ioe is black; CUSK BT'Osme broBme a b 1 ad blotch on each shou 1der. there is no shoulder blotch. Maximum size: 3J.s; feet. about 27 pounds Range: Both sides of the North Atlantic. In North America. from Newfoundland to New Jersey in moderately deep water.

Although the cusk has the fin shape of the (that is, a Single, long dorsal fin and a single long anal fin) and is included in the eel section of this guide, it is also included here since the cusk is a relative of the cod-like fishes and has a stout body like them ...... CO

HADDOCK Me'Lanogrcurmus aeg7,efinu,~ Maximum size: 44 inches, about 37 pounds Range: Both sides of the Atlantic. In North America. from West Greenland to North Carolina.

The lower jaw projects beyond The upper jaw projects beyond the upper; the chin barbel is the lower; the chin barbel is large. very small, if there is one.

POLLOCK PoZlachiu8 virens Maximum size: 3".i feet. about 35 pounds ATLANTIC COO Cadus morhua Range: Both sides of the North Atlantic. In Maximwn size: More than 6 feet. about 210 pounds North America. from Gul f of St. Lawrence to Range: Both sides of the North Atlantic. In North North Carol ina. America, from West Greenland to North Carolina. COD FAMILY - TWO DORSAL FINS -, The ventral fins are short The ventral f1 ns are very and of ordinary fonn long and feeler-like

, -1

15-22 gill rakers on first gill arch (A). '9_11 gill rakers on first gill arch (A) • Scales larger (103 to 130 scales A Scales smaller (130 to 148 scales , along the lateral line). along the lateral line).

Lateral line ;s unbroken; Lateral line is broken up by front of first dorsal fin white spots; first dorsal fin projects as a thread doesn I t project as a thread

SILVER HAKE Met'~UCCiU8 biUnea1'is OFFSHORE HAKE Mer7..ucciu8 albidus (Whiting) Range: From Georges Bank southward, Ma:cimwn size: 23:! feet, 5 pounds 50 fathoms. Range: Off Newfoundland to South Carolina. otv

sponED HAKE IJrophyais regius Mazimwn size: About 16 inches, about l~ pounds Range: Nova Scotia to Florida. Occurs regularly from southern New England to Gulf of Mexico. The ventral fins long, reaching The ventral fi ns shorter. do not reach beyond the mi ddl e of the ana 1 fi n. back to the middle of the anal fin. I ,,<1177; -, r "M~}' Two gill rakers on the upper segment of the '.. "il~., ,·'.;~.j Three gill rakers on the upper segment of the "·~;'·· first gill arch (A). Scales smaller (120 to 149 !g~~~ first gill arch (A). Scales larger (95 to ~ ' scales a'?ng the lateral line). Grayish or A lififJ 117 scales.alonq the later:.al line). Brownish A I' purplish 1n color. Often heavier than 4 pounds. 'li-ifi~ to bronze ln color. Very rarely heavier Length often exceeds 20 inches (50 centimeters). -/';:' than 4 pounds. Length never over 20 inches ~i ~< (50 centimeters). and usua lly not over 16 inches 2~... , l. ~,.... (40 centimeters).

LONGFIN HAKE Phycis chesteri M7xirrrwn size: 15 inches Range: Off Newfound 1and to North Caro 1 ; na. In deep water, mainly between 100 and 500 fathoms.

WH 1TE HAKE U1"ophyoistenuis RED HAKE UT'Ophycis ohUBs (Ling) (Squirrel hake, ling) Maa:irrrum size:': 4 feet. 40 pounds frtz:ciT1TWTl size: 20; nches Range: to Florida. Range: Southern Nova Scotia to Nort~ Carolina. BASS - SHAPED FISHES - ONE DORSAL FIN Two long filaments on chin. No long filaments on chin. I r------~

A large fl eshy flap on top of head. No large fleshy flap on top of head.

BEARDFISH lowei / Maximwn size: About 7 1/2 ; nches No spines on cheeks. Body red or orange. Spi nes Range: New Jersey to South America. Deep water, Eyes smaller. on cheeks. Eyes larger. caught mostly between 100 and 300 fathoms .. TILEFISH Lophol.atilus chamael.eonticeps M:z:rimwn size: At least 42 inches. 35 pounds Range: Nova Scotia to Gulf of Mexico, in depths I of 45 to perhaps 200 fathoms. I I Black spot on dorsal fin. Dorsal fin No black spot on dorsal fin. Dorsal has 12 hard spines in front. followed fin has 14 or 15 hard spines in front. by fairly soft fin bones (rays). followed by fairly soft fin bones (rays). Tai 1 forked.

~ >-'

ROSEFISH Helicolenus dactylopterus Maximum size: 15 inches REDFISH Sebastes marinus Range: Both sides of the North Atlantic. In North (Ocean perch) America, from Georges Bank to Florida, in depths Maximum size: Off North America, 27 inches. 13 1/2 pounds of 68 to 373 fathoms. Range: Both sides of North Atlantic. In North America. west of Greenland to New Jersey.

SCUP Stenotomus chPysops (Porgy) Ma:rimwn size: 18 inches. 4 pounds Range: Gulf of Maine to North Carolina. Abundant along Start of pectoral fin ;s in front of start the mid Atlantic states up to southern r~assachusetts. but Start of pectoral fin is behind the start of of ventral fin. Pectoral fin shorter. end of never plentiful on Georges Bank or in the Gulf of Maine. ventral fin. Pectoral fin long. end of pectoral pectoral fin far ;n front of start of anal fin. fin often reaches to start of anal fin. I

Scales on rear part of gill cover. Snout No scales on rear part of gill cover. somewhat pointed. Profile of head rather flat. Snout rounded. Profile of head rounded

BLACK SEA BASS Centl"Opristis striata TAUTOG Tautoga oni tis l.f:Lximwn size: At least 2 feet. A weight of 7 1/2 pounds. CUNNER TautogoZab1"US adspel"sus Maximum size: 3 feet. about 22 pounds Range: Maine to Florida. Maximum size: 15 inches, 2 1/2 pounds Range: Nova Scotia to South Carolina. In shallow Range: Newfoundl and to Chesapeake Bay. depths in the immediate vicinity of the coast. , BASS-SHAPED FISHES TWO DORSAL FINS First dorsal fin ;s much First dorsal fin ;s as high lower than second dorsal fin. or higher than second dorsal fin. r Second dorsal fin about Second dorsal fin not much twice as long as anal. 1anger than ana 1 fi n.

BLUEFISH Pomatomus saltatrix \ (Young are called Snapper) Ma:cimum size: About 3 1/2 feet, 31 pounds Chin has one or more barbels. Chin has no barbels. Range: In several oceans. On east coast of the Americas, Nova Scotia to Argentina.

A single thick barbel A row of minute barbels STRIPED BASS /okll'm(. sa:ratiLis at tip of lower jaw. on each side of lower jaw Harimwn size: About 6 feet. 125 pounds Range: Atlantic and Pacific coasts of United States. On Atlantic coast. St. Lawrence River to Florida and the northern Gulf of Mexico.

t-:l t-:l

Oar~ spot on body just No dark spot on body just behind upper edge of gill cover behind upper edge of gi 11 cover ATLANTIC CROAKER I>t;,r ~ l' n ",,"u/.a ~ NORTHERN KINGFISH MentiJirrhuB sazat:i7':8 Ma.ri-Irrur.I oiae: A 1 ittle over d foot Maximum size: 17; nches, 3 pounds /I "11Jc New York to Texa~ Range: Maine to Florida. Most numerous from Chesapeake Bay to New York.

The northern kingfish is conmon only north of Chesapeake Bay. South of Chesapeake Bay " a close relative, the SOUTHERN KINGFISH

Mentioirrhug amerwanu83 wi 11 be cOlllTlOn. The two speci es can be di s ti ngui shed as follows . NORTHERN KINGFISH: A dark "V" shaped mark on the side of the body in the shaul der regi on. Front of first dorsal fin very high and when laid WEAKFISH )fn '.e'lc~~C" Nt" -"_s back it reaches well beyond start of second (Squet.ague , Sea trout) dorsal fin. Color is usually dark. "'h.rimk."'"! oi"! Rarely over 3 feet. Hay have reached 30 pounds in the past; 17 112 pounds was a recent record. SOUTHERN KINGFISH: No dark "V" shaped mark on Florida to "'assachusetts Say. may stray to Nova SCvtia the side of the body in the shoulder region. Front of first dorsal fin lower and when laid back; t barely reaches the start of second dorsal fin. Color usually light. SPOT " 08 tom!u.l .run thurus Going south from about Del aware. the SPODED SEATROUT l !<110~:·:·:n Mbuz.c.sws. Mlrirmtm oia. 14 inches. 1 pound 6 ounces a close relative of the weak.fish. will be encountered regularly. These two Ra"ll( Ma ssachuset ts Bay to Texas species can be distin9uished as follows. WEA"'fISH: Body covered wi th numerous irregular blotches. some of which fonn wavy lines runnin~ forward and downward. Soft seqmented bones of anal fio and second dorsa 1 fi n ha ve sea 1 es . SPOTIED SEATROUT: Body covered with round b lad spots. Soft se""",nted bones of anal fin and second dorsal fin scaleless SEAROBINS AND SCULPINS - SEAROBINS

Lower part of pectoral fins in the form of Lower part of pectoral fins not in the form of fee 1ers and sepa ra ted from rema i nder of fi n. feelers and not separated from remainder of fin.

l._1 /

(see SCU LPINS, next page)

--.:... I.\'.' ...... , \\

Snout seems double when seen from above. Snout not double when seen from above.

~ v:>

No stripe down side of body. Pectoral ARfo'DRED SEAROBIN Peristedion miniatwn fin shorter, reaching only 1/2 the way A prominent, dark-brown stripe down side of I>brimwn size: 14 inches to end of base of second dorsal fin . body. Pectoral fin longer, reaching about 3/4 of Range: Georges Bank to South Carol ina. In the way to end of base of second dorsal fin. deeper water. from 50 fathoms to over 200 fathoms. '#

,~

......

NORTHERN SEAROBIN rnonotuB carolinu8 Maximum size: 16 inches STRIPED SEARDBIN PPionotue evolane Range: Bay of Fundy to South Carolina, mainly west Maximum size: 18 inches and south from Cape Cod. Range: Gulf of Maine to South Carolina. SEAROBINS AND SCULPINS - SCULPINS r------·--- No fleshy flaps on lower jaw, lower jaw. top of head, and top edge top of head. or fi rs t dorsa 1 f1 n. of first dorsal fin have ragged fleshy flaps.

A ser; es of bony p 1a tes on each side of body. No bony plates on each si.:le of body.

SEA RAVEN Hemitripte:rus amel"icanuB M;u:imwn siae: 25 inches. at least 7 pounds MOUSTACHE SCULP IN Triglops """"",yi Range: Newfoundland to Chesapeake Bay (Mailed sculpin) Ma:cirmun siae: 8 inches Range: Both sides of the North Atlantic. In The long spine on the cheek is straight. The long spine on the cheek hooks upward. North America. north of Labrador to Cape Cod. ~ ~ / :"R y :J""'. ~-.;~ fi'~~ - '.~~~'-~ ~ ;' . ~ The longest (uppennost) cheek spine ;s four The longest (uppennost) cheek spine is not more ~- "I- - . ~ times as long as the spine below it and reaches than twice as long as the spine below it and back to the edge of the gi 11 cover. does not reach more than about 1/2 way to the edge of the gill cover. \

HOOKEAR SCULPIN ArteduH.iu8 species .'tl.ri'ffW'l SiB. 4 inches Ran.,.: Both sides of North Atlantic. In North Ameri ca, labrador to Cape Cod.

LON GHORN SCULPI N Myoxooepha~u8 ootodscsmspinosUB GRUBBY Myo.JXlt't6ph.J IUB o.l61klIilUB Maximum SiBB: 18 i nche s Ma.rtf1tW'f/ BiIlO: 8 inches Range: Newfoundland to Vi r gi nia Rangfl: Newfou ndland to New J ersey . FLATFISHES RIGHT- EYED FLATFISHES

,- ---

Large mouth 1 gap; n9 back Small mouth. not gaping ,- as far as the eye back as far as the eye Margin of tail rounded; lateral line Margin of tail has sharp corners and is slightly concave; nearly straight just behind gill opening lateral line arched just behind gill opening.

pl.atesBoides ATLANTIC HALIBUT Hippoglo88u8 hippog'108Bu8 1~~~~~~nP~:~:E D~~)Poglo8soide8 Maximum size: An 8-foot halibut weighs about 350 pounds. Maximum size: 32 1/2 inches, 14 pounds Known to reach about 700 pounds. Range: Both sides of the North Atlantic. Range: Both sides of the North Atlantic. In In North America, labrador to r~ontauk North America. from Virginia north to, or beyond, Labrador. Point, tlew York. Latera 1 1 i ne arched behi nd gill open; ng ~ Lateral line nearly straight ell behind gill opening. I Depth of ~base of tai 1 much more than Depth of base of tail about equal to distance between outer margins of eyes distance between outer margins of eyes

YELLOWTAIL FLOUNDER U=nda ferruginea Maximum size: 21 3/4 inches Range: Labrador to Chesapeake Bay.

WITCH FLOUNDER GZyptDaephaZus aynogZo88u8 (Gray sale) ~~~!~~b:~~~N~~~~8~~)Leuronecte8 cunericanu8 Maxirmon 8iae: 25 inches. about 4 pounds Range: Both sides of the North Atlantic. In North Maximwn size: 25 inches, 8 pounds Alrerica. north of Newfoundland to North Carolina. Range: Newfoundland to Georgia. Found in moderately ~eep water. FLATFISHES LEFT - EYED FLATFISHES

Lateral line arched over pectoral fin. Lateral line is nearly straight. Often much longer than 7 inches. Maximum is 7 inches or less. I ------1 Snout has a well developed bony No bony protuberance protuberance I extend; n9 hori lontal1y. extending horizontally on snout.

GULF STREAM FLOUNDER CithaPichthY8 arctifron8 SMALL MOUTH FLOUN DER EtropU8 micros~o"",s Ma:r:imwn size: 7 inches Ma:.cimwn oize: 6; nches Range: Georges Bank to Yucatan, Mexico. Occurs mainly at Range: New England to the Gulf of Mexico. Common at depths ~ depths of 25 to 200 fathoms. seldom as shallow as 12 fathoms. of 2-15 fathoms. seldom deeper than 25 fathoms. 0">

Body more round in outline. body about 1 1/2 Body more elongate in outlinej body about 2 times as long as broad. Front part of times as long as broad. Front part of dorsal dorsal fin ;s enlarged and fringe-like. fin ;s abl)ut like the rest of dorsal fin.

Four large~ spots on upper sit;le. Many small spots on upper side.

• ,,1.1/ f,U .... .f, •

~;'~~ ~ ... ~·'i..,~._. fl - !, ..,j. ~~:, ..:. (,,,jIf "1-.", ..

d" .~ rf:.:.f') ) /.'12/ ". WINDo\~PANE SaophthaZrrrus aquosu.s .~<~'~ (Sand dab. Sand flounder. Sundial) ... Maxirrrum size: 18 inches, 2 pounds '~:'-;--""'1 \\~~. , ... Range: Gulf of St. Lawrence to South Carol ina.

FOURS POT FLOUNDER HippogLo88U8 obLongus SUMMER FLOUNDER ParaUchthye dentatu8 Ma:cimum size: 16 inches (Fl uke) . Range: Georges Bank to South Carolina. Maximum size: A 37" fish has been caught which weighed 20 pounds. May reach 30 pounds. Range: Maine to South Carolina, maybe to Florida. MACKEREL AND TUNA-SHAPED FISHES -l

Snout short, not like a needle.

ATLANTIC SAURY Scombe!'eso:r; SQUPU8 The two dorsal fins touch each other or The two dorsal fins are separated (Needl efi sh) are separated by a space much shorter than by a space at least as long as the length of the first dorsal fin. the length of the first dorsal fin. Maximum si.a:e: 18 inches Range: Both s; des of the North Atl antic. In North AfTErica from Newfoundland to the Joiest Indies.

The sides below the midl ine The sides below the midline are spotted. are silvery, not spotted. - A group of similar look;n!) fishes, fast swimming and seldom caught in otter trawls. One spec; es. the BLUEFIN TUNA Thunnus thynnu8, shown above. attains a weight wel1 above 1000 pounds. t-:l -J .-, 'i?, ,,_.. ' -~, '~~ l~

ATLANTIC MACKEREL Soo,,",er 800,,",""8 CHUB MACKEREL ScomheT' jcrponicus Maximum size: A few reach 22 inches and about 4 pounds. Maximum size: 14 inches One weighing 7 1/2 pounds is mentioned in "Fishes of the Range: Gulf of St. Lawrence to Virginia. Gul f of Maine." Range: Both sides of the North Atlantic. In North America, Newfoundland to North Carolina. Another group of fishes. the jacks (also called carangids) contain several.fishes that look like the mackerels. The jacks can be separated from the mackerels by the following characters. MACKERELS: The length of the base of the second dorsal ;s about equal to or smaller than the base of the first dorsal fin. There are four or more little fins (called finlets) between the last dorsal fin and the caudal fin.

JACKS: When two dorsal fins are present some are present (some jacks have only one dorsal fin) the base of the second dorsal is much longer than the base of the first. There is one or no 1 ittle fins between the last dorsal fin and the caudal fin. One of the jacks, the MACKEREL SCAD Decapte:rus macareLZue is shown below.

MACKEREL SCAD DecapteruB macareLZus Maximum size : About 1 foot Range: Nova Scotia south to warm parts of the Atlantic. GOOSEFISH AND BUTTERFISH

Note These two fishes should be easily distinguishable from all other fishes in the guide They are placed to­ gether here for lack of a better place and not because they resemble each other

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BuPl..RF~S.H GOOSEFISH L)phiu8 ~.101' .r..... 12 .nche ... (Angler. Monkfish) nd Gul' f it lawren C:outh I an' tna ~ sias: 4 feet, 50 pounds t: Range: Near Newfoundland to North Cdrol inA Same or similar species off South Americd Another group of fishes. the jacks (also called c4r4n~~1s) contafn several ffshes thdt SOfM"What r"eSEi-;--b e the butterfish. They can be distinguished be<:".l~f all Ja("ks ha,"e ventra~ f1ns. Butter-'ish t .. k ypnt ral filS INDEX OF ------COMMON NAMES Page Page

Alewife. 16 Blueback herring. 16 Ameri can dab . 25 Bluefin tuna. 27 American plaice. 25 . 22 American shad. 16 Bluntnose stingray . 12 Anchovy . 18 Brier skate 10 Angel shark. 9 Brown shark 8 Angler . 28 Bullnose ray. 11 Argentine. 17 Butterfish. 28 Armored searobin 23 Butterfly ray 11 Atlantic argentine 17 Atlantic cod . 19 Catfish . 13 Atlantic croaker 22 Chain dogfish 8 Atlantic hagfish 13 Chub mackerel 27 t-:) (,0 Atlantic halibut 25 Clearnose skate 10 Atlantic herring 15 Cod. 19 Atlantic mackerel. 27 Common sea robin. 23 Atlantic menhaden. 15 Conger eel. 14 Atlantic saury . 27 Cownose ray 11 Atlantic wolffish. 13 Croaker 22 Cunner. 21 Barndoor skate 9 Cusk. 13, 20 Bass .21, 22 Cusk-eel. 14 Bay anchovy. 18 . 21 Dab 25 Big skate. 10 Dogfish . 8 Blackback. 25 Dusky shark 8 Blackbelly rosefish. 21 Black sea bass 21 Eel 13, 14 Blueback . 16 Eelpout 14 Fawn cusk-eel. 14 Longfin hake. 20 Flatfishes 25, 26 Longhorn sculpin . 24 Flounder . 25, 26 Fl uke. 26 Mackerel. 27 Fourbeard rockling 13 Mackerel scad. 27 Fourspot flounder. 26 MaileJ sculpin. 24 Freshwater herring 16 Menhaden . 15 Monkfish. 28 Goosefish . 28 Moustache sculpin 24 Gray sole. 25 Myctophid 17 Greeneye . 17 Grenadier. 13 Needlefish. 27 Grubby . 24 Northern kingfish 22 Gulf Stream flounder 26 Northern searobin 23 Northern stingray 12 . 19 ~ Hagfish. 13 Ocean catfish 13 Hake 20 Ocean perch . 21 Halibut. 25 Ocean pout. 14 Herring. 15, 16 Offshore hake 20 Herring family 15, 16 Hickory shad . 15 Pearlsides. 17 Hookear sculpin . 24 Perch 21 Homed lanternfish 17 Plaice. 25 Pogy. 15 Jack . 27, 28 19 Porgy 21 Kingfish 22 Pout. 14

Lance 13 Rattail 13 Lanternfishes. 17 Raven . 24 Lemon sole . 25 Rays. 11 Leopard skate. 9 Redfish 21 Ling. 20 Red hake. 20 Li ttle skate . 10 Rockling . 13 Rosefish. 21 Sole. 25 Rosette skate . 9 Southern kingfish . 22 Roughtail stingray. 12 11 Round herring 15, 18 Spiny dogfish 8 Spot. 22 Sandbar shark 8 Spotted hake. 20 Sand dab. 26 Spotted seatrout. 22 Sand flounder 26 Squeteague. 22 Sand lance. 13 Squi rrel hake 20 Sand tiger. 8 Stingrays 12 Saury . 27 Striped anchovy 18 Scad. 27 Striped bass. 22 Sculpin 24 Striped cusk-eel. 14 Scup. 21 Striped searobin. 23 Sea bass. 21 Sununer flounder 26 Sea herring 15 Sundial 26 Sea raven 24 w...... Searobin. 23 Tautog . 21 Seatrout. 22 Thorny skate. 9 Shad. 15, 16 Tilefish. 21 Shark . 8, 9 Tuna. 27 Shortnose greeneye. 17 Sil ver anchovy. 18 Weakfish. 22 Silver hake. 20 White hake. 20 Skate . 9, 10 Whiting. 20 Smallmouth flounder 26 Windowpane. 26 Smooth butterfly ray. 11 Winter flounder 25 Smooth dogfish. 8 Winter skate. 10 Smooth skate. 9 Wi tch flounder. 25 Smooth-tailed skate 9 Wolffish. 13 Snake eel 14 Snapper. 22 Yellowtail. 25 Yellowtail flounder 25 INDEX OF SCIENTIFIC NAMES

?..a.9.£ Page acan thias ~ Squalus . .. . 8 rarr ' hal'hirl1~ .~ )bs pu.s . 8 adspersus ~ Tautogo labr us. 1 aY'olinu.s , Prionotus . 23 aeglefinus ~ Me l anogrammus 19 Centm{ris tis .s tpiata. 21 aenaeus ~ MYoxocephalus . . 24 C'entrOUlYl ~ Das!}atis . 12 aestivalis ~ Alosa . . . . 16 Cepa ton, ' 7pP l us maderens1-s. 17 agassizi ~ Chloropht halmus 17 ,'ervinwft, Lep0l!hidium. 14 albidus ~ Merluccius {l0 ·hamcwk )}1f;cc'f's , [Q,h JZ:;.tiZus 21 Alosa aestivalis . .. 16 .:h(.'stel'i ~ Phyt ' ~'. s ... 20 Alosa mediocris . . . 15 t:hlm'vph t-halmus a.,j IS S ' ;o; i 17 Alosa pseudoharengus . 16 ChI'YStPS , '":tP}10t 1mt03 . 21 Alosa sapidissima . . 1 , ~ hU:; $ ~ [/n o altavela ~ Gymnura .. 11 c·imbl'iwl , 13 americanus ~ Hemitrip terus ~)4 ('7' tl' 26 americanus ~ Lophius . . . 'Jf, 15 americanus ~ Macrozoarces . 11 '.,)}lcjt I ' ol ' €',(nl> ~ li" 14 americanus ~ Menticil'rhus . 2 :~ L' l ' lIe l d i .t~) /', ot hidlthy;, 14 qr: americanus ~ Pseudopleuronec tes . < . ) La J l 0 ,; D fA:; , (;l ~/r tocC'pJw. Zw ; . 25 Ammodytes s pecies ].) 62 Anarhichas lupus . l.~ ::12 Anchoa hepsetus . 18 Anchoa mitchilli. lq t n7~:, aquo s us ~ Scophthalmus ~)C 12 arcti frons ~ Cithari~hthus 26 12 Argentina silus . . 17 27 " , Artediellus speci es 24 ~O 9 bair di ~ Nezumia . . 13 bilinearis ~ Merluccius . ?O 10 bonasus~ Rhinoptera 11 13 -, Brevoor tia tyrannus 15 c BY'osme br osme . 13 ~ 1.' o brosme ~ Brosme . l,~ , 1: 2a . 15 . 1S canis~ Mus teluf! q (;~ ., . ' /j ,-) Carangids ... I , Carchar hinus mi lberti tj f),", ., '-"r " 1 Gadus morhua. 19 Melanogrammus aeglefinus . 19 garmani .. Raja 9 Menticirrhus saxatilis . 22 glutinosa.. Mljxine 13 Menticirrhus americanus 22 Glyptocephalus cynoglossus . 25 Merluccius albidus . 20 Gymnura altavela. 11 Merluccius bilinearis 20 Gymnura micrura 11 Micropogon undulatus . 22 microstomus .. Etropus . 26 harengus.. Clupea. 15 micrura.. Gymnura . 11 Helicolenus dactylopterus 21 milberti .. Carcharhinus . 8 Hemitripterus americanus . 24 miniatum.. Peristedion 23 hepsetus .. Anchoa. . . 18 mitchilli .. Anchoa 18 Hippoglossoides platessoides. 25 morhua.. Gadus 19 Hippoglossus hippoglossus 25 Morone saxatilis . 22 hippogloscus .. Hippoglossus . 25 muelleri .. Maurolicus . 17 Hippoglossus oblongus 26 murrayi.. Triglops 24 Muste lus canis . 8 japonicus .. Scomber . 27 Mljliobatis freminvillei 11 w w Mljoxocephalus aenaeus 24 laevis.. Raja. 9 Mljoxocephalus octodecemspinosus 24 Leiostomus xanthurus. 22 Mljxine glutinosa. 13 cervinum. 14 Limanda ferruginea . 25 nebulosus .. Cynoscion . 22 Lophius americanus . 28 Nezumia bairdi . 13 Lopholatilus chamaeleonticeps 21 lowei .. Polymixia. 21 oblongus .. Hippoglossus . lupus .. Anarhichas 13 obscurus .. Carcharhinus . oceanus.. Conger macarellus .. Decapterus . 27 oce Uata.. Raja. Macrozoarces americanus 14 octodecemspinosus .. Mljoxocephalus . maderensis .. Ceratoscopelus . 17 Odontaspis taurus mm1ginata.. Risso La. 14 oni tis -' Tautoga marinus .. Sebastes 21 Ophichthus cruentifer Mau1'O l icus mue lleri 17 medi,ocY'is.. A losa. 15 Paralichthys dentatus. 26 Sc:yUophinus retifep. 8 Peprilus triacanthus 28 Febastes marinus . 21 Peristedion miniatum . 23 senta ~ Raja 9 Phycis chesteri . ... 20 siluR ~ Argentina. 17 platessoides~ Hi ppoglossoi des. 5 S'I,{c.1lu[; aC'~mthias 8 Pollachius virens .. 19 Squatinc.1 dumerili 9 Polymixia lowei. . . 21 Steno tomus r;hiY sopS 21 Pomatomus saltatrix. 2 striatc.1 ~ CentPOpi,istis. 21 ')7 Prionotus carolinus. u t..) Prionotus evolans .. 2 ,~ t 8 pseudoharengus~ Alosa . 1 21 Pseudopleuronectes ame r~ canus. ? 21 20 radiata ~ Raja. . 15~1'7 Raja eglanteria. 10 Raja erinacea. 10 6 7 Raja garmani . u8 Raja laevis . . 2'1 w.,. Raja ocellata. 10 15 Raja radiata . Raja senta .. 9 GG'''1 '1- regalis~ Cynoscion y 20 regius ~ Urophycis. " 0 20 retifer ~ Scyliorhinus. B 20 Rhinoptera bonasus 1 Rissola marginata. . 1 18

'1 .-, saltatrix~ Pomatomus :. sapidissima ~ Alosa . saurus ~ . saxatilis ~ Menticirrhu saxatilis ~ Morone . oj . -~ sayi ~ Dasyatis . . Scomber japonicus . Scomber scombrus . Scomberesox saurus scombrus ~ Scomber . Scophthalmus aquosu ;'; 6 ERRATA

NOAA TechnicaL Report NMFS Circular 428: Morphological Comparisons of North American Sea Bass Larvae (Pisces: Serranidae), by Arthur W. Kendall, Jr.

Page 7, Figure 6d is incorrect. See correct Figure 6d below.

d