TWONEWSPECIESOF THEGENUS GONEPLAX (DECAPODA, BRACHYURA,GONEPLACIDAE) FROM EAST ASIA

BY

HIRONORI KOMATSU andMASA TSUNE TAKEDA Departmentof Zoology, National Science Museum, 3-23-1 Hyakunincho, Shinjuku-ku, Tokyo169-0073, Japan

ABSTRACT

Twonew species of goneplacid crab, Goneplaxmarivenae and G. megalops,aredescribed from thePhilippines and Japan, respectively. Both new species are closely similar to G. sigsbei (A. Milne- Edwards,1880) from the western Atlantic Ocean in the form of the carapace, eyestalk, fourth ambulatoryleg, and male pleopods, but can be distinguished from G. sigsbei bythe form of the orbit.Both new species are also similar to G.maldivensis Rathbun,1902, but are distinguished from G.maldivensis bythe form of themale abdomen. Goneplaxmarivenae ismost closely similar to G. megalops,butcan be distinguishedfrom G. megalops bytheform of theextraorbital tooth and themale Ž rstpleopod.

RÉSUMÉ

Deuxespè ces nouvelles de crabes Goneplacidae, Goneplaxmarivenae et G. megalops, sont décrites respectivement, des Philippines et du Japon. Les deux espè ces nouvelles sont trè s prochesde G. sigsbei (A.Milne-Edwards, 1880) de l’océ an Atlantique occidental par la formede lacarapace, les pédoncules oculaires, la quatrième patte ambulatoire, et les plé opodes mâ les, mais s’ en distinguent parla forme de l’ orbite. Les deux nouvelles espè ces ressemblent é galementà G.maldivensis Rathbun,1902, mais s’ en distinguent par la forme de l’ abdomen mâ le. Goneplaxmarivenae est leplus proche de G. megalops,maisen diffè re par la forme des dents extraorbitales et le premier pléopode mâ le.

INTRODUCTION The genus Goneplax Leach,1814 currently consists oftwospecies fromthe At- lantic Ocean,viz., G.rhomboides (Linnaeus,1758) (the typespecies) and G. sigs- bei (A.Milne-Edwards,1880), and Ž vespecies fromthe Indo-PaciŽc (Serène & Umali, 1972),viz., G.sinuatifrons Miers, 1886, G.maldivensis Rathbun,1902, G.renoculis Rathbun,1914, G. wolŽ Serène, 1964, and G. serenei Zarenkov,1972. Rathbun(1918) synonymized Frevillea A.Milne-Edwards,1880 with Goneplax, © KoninklijkeBrill NV ,Leiden,2004 Crustaceana 76(10):1243-1256 Alsoavailable online: www.brill.nl 1244 HIRONORI KOMATSU&MASATSUNET AKEDA butGuinot (1969) clariŽ ed the identity of Frevillea and Goneplax,respectively. Serène & Soh(1976) established anewgenus, Singhaplax,to accommodatetwo species of Goneplax, G.ockelmanni Serène, 1971 (the typespecies) and G. nippo- nensis Yokoya,1933. InFebruary 2003, we hadan opportunity to take aŽeld trip to Balicasag Island, Bohol,Philippines, as apart ofaresearch project ofthe National Science Museum, Tokyo,entitled “Natural History ofthe West PaciŽc Archipelago”. Onthis, arrangedby Ms. MariveneR. Manuelof the National Museumof the Philippines, wecould obtain many brachyuran specimens fromthe local shell Žshermenof Balicasag Island.Some of these specimens provedto belongto a newspecies of the genus Goneplax.Inaddition to this material, some specimens collected from Japanese waters proveto belongto anothernew species of Goneplax. This new species is identical with the species recordedas Goneplax sp. aff. sigsbei by Nagai &Tsuchida(1996) from Miyake-jima I.,IzuIs., andoff Shiono-misaki Cape, Kii Peninsula,central Japan,and was misidentiŽed as G.renoculis byTakeda(1978) fromOshima I.,Izu Is., central Japan.In this paper,we clarify the differences betweenour two new species andtheir congeners,and describe them as newto science. Measurements, givenin millimeters (mm),are ofthe greatest carapacelength andwidth (including the anterolateral teeth), respectively.Pereiopodsare measured alongthe outermargin from ischium to dactylus.The specimens examined are depositedin the Departmentof Zoology, National Science Museum,T okyo (NSMT),the National Museumof the Philippines (NMCR),the Natural History Museumand Institute, Chiba(CBM), and the ShowaMemorial Institute, National Science Museum,T okyo(NSMT S).

TAXONOMY Goneplaxmarivenae n.sp.(Ž gs. 1-3, 7A)

Materialexamined. — Holotype:male (12 :3 18:2),NSMT -Cr15531, Balicasag I., Bohol, £ Philippines,tangle nets, coll. local shell Ž shermen,February 2003. Paratypes:2 males(11 :7 17:5, 11:4 18:0),1 ovig.female (10 :2 15:7),NSMT -Cr15532, £ £ £ samedata as holotype; 1 male(12 :2 19:1),NMCR, samedata as holotype. £ Description.— Carapace(Ž g. 1a) trapezoidal ingeneral outline, 1.5-1.6 times broaderthan long (holotype 1.5 times); uppersurface smoothand glabrous, convex dorsally; regionsill-deŽ ned, marked with H-shaped groove at center.Frontal region(Ž g. 1b) about0.3 times as broadas carapace,anteriorly slopingdownwards, transversely rectangular; marginsubtruncate, weakly concave medially ,withsmall mediantriangular tooth,forming almost right anglewith supraorbital margin, TWO NEW GONEPLAX 1245

Fig. 1. Goneplaxmarivenae n.sp.,holotype, male, Balicasag I., Bohol,Philippines, 12 :3 18:2 mm, £ NSMT-Cr15531: a, carapace,dorsal view; b, frontalregion, antero-dorsal view; c, same,frontal view ofleft half with  agellumof antennule retracted in fossa and  agellumof antenna cut at midway; d,e, right and left chelae (tip of immovableŽ ngerof left chela broken), respectively, external views; f,merus and carpus of cheliped,dorsal view; g, same, lateral view; h-k, left Ž rstto Ž fthambulatory legs,dorsal views. Scales: a, d-g, h-k 2 mm; b, c 1 mm. D D 1246 HIRONORI KOMATSU&MASATSUNET AKEDA markedwith weaksubmarginal line. Lateral marginof carapace moderately convergingposteriorly ,withlarge, triangular extraorbital toothwith blunt point oncorner with supraorbital margin; asmall, acute triangular toothis placedjust behindthe extraorbital tooth,both teeth directed antero-laterally.Posterior margin almost straight, double. Orbit (Žg. 1a, c): supraorbital marginsinuate, withoutsuture, slightly turned upwards,fringed with veryshort setae; infraorbital marginstraight in mesial half, arcuate in lateral half,slightly turnedupwards. Eyestalk (Žg. 1c)long, completely set in orbit whenretracted; dorsal extensiononto cornea narrowly rectangular, withrounded tip; corneamuch broader than stalk, 0.4times as longas total eyes- talk. Antennule(Ž g. 1c) transversely foldedinto fossa; basal segment occupying ventral 0.7,with weaktransverse ridgealong midline. Antenna(Ž g. 1c): Žrst segment transversely ovate; secondand third segments subcylindrical,occupying orbital hiatus; agellum long,slightly longerthan eyestalk. Epistome (Žg. 1c) triangular, separated frombuccal frame bytransverse Žssure. Mandible(Ž g. 2a): endopodwell calciŽed, with short suture at middle; palpthree- segmented; proximalsegment veryshort; secondsegment fringedwith plumose setae onoutermargin; distal segment entirely fringedwith simple setae. Maxillule (Žg. 2b): coxal endite elongatetongue-shaped, thin, fringed with short simple setae, withsome longplumose setae onproximal part oflower margin; basial endite triangular,fringed with thinand stout simple setae alongmesial margin; endopodtwo-segmented, fringed with plumosesetae alongmesial marginof proximalsegment, distal segment lobularand covering mandible in situ. Maxilla (Žg. 2c): coxalendite bilobed,both lobes elongatetongue-shaped, fringed with plumosesetae; basial endite distally bilobed,entirely fringedwith simple setae; endopodtriangular, narrowed at tip, fringedwith short plumosesetae excepttip; exopod(scaphognathite) longitudinally ovate, entirely fringedwith short plumose setae. First maxilliped (Žg. 2d):epipod present; coxalendite rounded,with dense simple setae; basial endite ovate,fringed with simple setae, with submarginalrow ofsimple setae alongmesial margin; endopodlobular, subsquare in distal half, fringedwith plumose setae alongproximal half ofmesial margin; exopodwith long agellum, agellum fringedwith longplumose setae. Secondmaxilliped (Žg. 2e): oneepipod and one podobranch present; ischium-merus withsubmarginal rowof simple setae alonglateral margin; propodusproduced outwards on lateral margin,with simple setae; dactylus with stout setae aroundtip; exopodelongate rectangular,fringed with short plumosesetae alongproximal 0.7 of lateral margin anddistal margin; agellum long,fringed with longplumose setae. Thirdmaxilliped (Žg. 2f): podobranchpresent; coxaand epipod covering af- ferent channel;basis triangular,fused with ischium butdivided by suture; is- TWO NEW GONEPLAX 1247

Fig. 2. Goneplaxmarivenae n.sp.,holotype, male, Balicasag I., Bohol,Philippines, 12 :3 18:2 mm, £ NSMT-Cr15531. Right mouthparts in external view ,setaeof each  agellumof exopod omitted: a,mandible; b, maxillule;c, maxilla; d, Žrstmaxilliped; e, second maxilliped; f, third maxilliped. Scales:a-e, f 1 mm. D chiumwith shallowoblique groove at center,dentate andfringed with short se- tae onmesial margin; merus triangularly projectingon mesial margin,dentate on proximalhalf ofmesial margin,produced on disto-lateral corner;palp exposed, fringedwith simple setae oninner margins ofpropodus and dactylus; exopodsub- rectangular,with long  agellum fringedwith long plumose setae, withtriangu- lar process ondistal 0.3of mesial margin,process concealedbeneath merus in situ. 1248 HIRONORI KOMATSU&MASATSUNET AKEDA

Cheliped(Ž g. 1d-g) stout, smooth,2.8-2.9 times as longas carapacein adult male (n 4;holotype2.9 times), 2.5times in adult female (n 1); coxalcondyle D D small, roundedin both sexes; merus subcylindrical,with small, obtusetriangular toothat proximal0.3 of outermargin of dorsal surface,covered with Žneattened granuleson innersurface; carpushemispherical, innerangle obtuse in adult male andacute inadult female. Chelae (Žg. 1d, e) dimorphicin both sexes, usually right chela slightly larger thanleft one;palm weaklyconvex, weakly dilated distally; Žngerstriangular, crossed at the tip, movableŽ ngerinside. Cuttingedges ofleft chela (Žg. 1e) irregularly dentate,entirely meeting,more strongly dentate onimmovableŽ nger.In adult male, cutting edgesof right chela (Žg. 1d)leaving gap;cutting edgeof movableŽ ngeralmost blunt;cutting edgeof immovable Ž nger armedwith large,obtuse triangular toothat middle,weakly dentate exceptmedian tooth.In subadult male andadult female, cuttingedges of right chela as in left chela. Ambulatorylegs (Žg. 1h-k) slender, long, sparsely fringedwith short setae, similar in shapeexcept fourth leg; Žrst tofourth leg about2.3-2.4, 2.5-2.6, 2.5,and 1.8 times as longas carapace(holotype 2.3, 2.5, 2.5, and 1.8 times), respectively; coxalcondyles small, roundedin bothsexes; merus,carpus, and propoduscompressed, sparsely fringedwith short setae alongmargins; dactylus slender,fringed with short setae alonginner margin; propodusand dactylus of fourthleg (Žg. 1k) strongly compressed and broadened, densely fringed with plumosesetae alonginner and outer margins. Male thoracic sternites (Žg. 3a) smooth; sternites 1and2 fused; sternite 3 dividedfrom sternite 2bytransverse suture; sternite 4fusedwith sternite 3,but leavingshort obliquesuture oneach side; sternite 5with small granularbutton; sternite 8slightly visible betweensecond and third somites ofabdomen on each side; sutures betweensternites 4and5, 5and6, and 7 and8 interruptedmedially; mediansuture reachingto suture betweensternites 6and7; abdominalcavity reachingto medial portionof sternite 4. Female thoracic sternites as in male in formula; sternite 5with ovategonopore; sutures as in male; abdominalcavity reachingto borderbetween sternites 3and4. Male abdomen(Ž g. 3b) smooth, almost at, dorsally exposedin Ž rst andsecond somites, all somites free; Žrst somite veryshort; secondsomite short,transversely rectangular; third to sixth somites subtrapezoidal,gradually decreasing in width distally; telson triangular with roundedtip. Female abdomensmooth, convex externally, ovate; eachsomite subrectangular, graduallygetting longerfrom Ž rst tosixth somite; lateral margins fringedwith short soft setae; telson transversely triangular with roundedtip, fringedwith short soft setae. TWO NEW GONEPLAX 1249

Fig. 3. Goneplaxmarivenae n.sp.,holotype, male, Balicasag I., Bohol,Philippines, 12 :3 18:2 mm, £ NSMT-Cr15531: a, sternum, ventral view; b, abdomen,external view; c, right male Ž rstpleopod, externalview; d, tip of same, external view; e, right male second pleopod, external view .Scales: a-b 2 mm; c, e 1 mm; d 0.2 mm. D D D Male Žrst pleopod(Ž g. 3c) weakly compressed, getting narrower distally, with some verysmall spinulesetae just belowapical aperture; medial shoulderwith some plumosesetae; mesial marginentirely fringedwith short plumosesetae; lateral marginfringed with plumosesetae alongproximal 0.3; apical aperture largely openon external surface; tip (Žg. 3d)triangular, directed laterally,roundly convexon mesial margin,with some veryshort setae alonglateral margin.Male secondpleopod (Ž g. 3e) elongate, slightly longerthan Ž rst one,two-segmented, fringedwith plumosesetae at base oflateral margin; tip curledexternally. Etymology.— This species is dedicatedto Ms. MariveneR. Manuelof the National Museumof the Philippines, whokindly arranged the Želd trip inthe Philippines. Thename is anounin the genitive singular. Distribution. —Philippines: Balicasag I.,Bohol(type locality). Exactdepth unknown. Remarks. — Goneplaxmarivenae n.sp.is most similar to G. megalops n. sp. in the formof the carapace,the fourthambulatory leg andthe male pleopod,but canbe distinguished from G. megalops bythat the extraorbital toothdoes not project beyondthe secondtooth, whereas it projects beyondthe secondtooth in G. megalops;the mesial marginof male Žrst pleopodis entirely fringedwith plumosesetae, whereasit is fringedwith short simple setae alongthe distal half in 1250 HIRONORI KOMATSU&MASATSUNET AKEDA

G. megalops;the tip ofthe male Žrst pleopodof G.marivenae is broaderthan that of G. megalops; and G.marivenae is abouttwice as large as G. megalops when consideringcarapace size. Thepresent twonew species canbe readily distinguishedfrom Goneplax renoculis Rathbun,1914 (type locality: offsouthern Luzon, 146-186 m) bythat the secondtooth of the lateral marginof the carapaceis situated just behindthe Žrst (extraorbital) tooth,whereas that of G.renoculis leaves ashort space with the Žrst one;the dorsal extensionof the eyestalk ontothe corneais narrow,whereas that of G.renoculis is broad;the fourthambulatory leg is fringedwith dense plumose setae, whereasthat of G.renoculis is simple andsimilar in shapeto the otherlegs; the formof the male Žrst pleopodis quite different fromthat of G.renoculis (see Takeda& Miyake,1968, Ž g.8c-e); andthe male secondpleopod is curledat the tip, whereasthat of G.renoculis is straight. Thepresent twonew species are similar to G.maldivensis Rathbun,1902 (type locality: Maldive Is., 36m) inthe formof the anterolateral teeth ofthe carapace andthe broadenedpropodus and dactylus ofthe fourthambulatory leg. But they canbe distinguished from G.maldivensis bythe fact that the frontal marginis subtruncate,whereas that of G.maldivensis has anotchat the outerangle in which the antennais lodged;the Žrst somite ofthe male abdomenis visible in dorsal view,whereas that of G.maldivensis is hiddenunder the carapace; andthe male abdomenis subtriangular,whereas that of G.maldivensis is muchnarrowed. Thepresent twonew species are also similar to G. sigsbei (A.Milne-Edwards, 1880)from the western Atlantic Ocean(type locality: Grenada,166 m) in the formof the anterolateral teeth ofthe carapace,the narrowdorsal extensionof the eyestalk ontothe cornea,the setose fourthambulatory leg, and the formof the male Žrst pleopod.But they can be distinguished from G. sigsbei by the fact that the infraorbital marginis almost straight in its mesial half,whereas that of G. sigsbei has asmall triangular notchjust nearthe mesial end(see Guinot,1969, Žg.68), and the carpusof the chelipedhas asmall orinconspicuous tooth on the innerangle, whereas that of G. sigsbei has alarge toothon the innerangle.

Goneplaxmegalops n.sp. (Ž gs. 4-6, 7B) Goneplaxrenoculis —Takeda,1978: 78 (inlist). [Not G.renoculis Rathbun,1914.] Goneplax sp. aff. Sigsibei [sic]— Nagai& Tsuchida,1996: 32, pl. 1 Žg.7. [Not G. sigsbei (A. Milne- Edwards,1880).]

Materialexamined. — Holotype:male (6 :2 9:5),CBM-ZC 7031,26 18:86 N 127 09:01 E, £ ± 0 ± 0 KeramaIs., Ryukyus, southwestern Japan, R V“Tansei-maru”cruise KT02-03, stn E5-2, dredge, 182-169m, coral rock and sand, coll. T .Komai,19 April 2002. Paratypes:1 youngmale (4 :8 7:3),NSMT -Cr5603, southeast of OshimaI., IzuIs., central Japan, £ dredge,23-65 m, coll. M. Takeda,12 July 1977; 1 ovig.female (7 :4 11:2),NSMT -Cr10015, off £ TWO NEW GONEPLAX 1251

Kushimoto,Kii Peninsula, central Japan, R V“Tansei-maru”KT84-12 cruise, stn12-1, dredge, 100- 101m,coll. E. Tsuchida,31 August1984; 3 youngfemales (3 :4 4:6-5:1 7:5;largest one infected £ £ by Thompsonia),NSMT-Cr5594, Oshima I., IzuIs., central Japan, dredge, 30 m, coll. M. Takeda, 12July 1977; 1 youngfemale (5 :0 7:3),NSMT -Cr6891, Omuro-dashi Bank, Izu Is., central £ Japan,dredge, 87-184 m, coll. T .Okutani,26 June 1972; 1 youngfemale (3 :5 4:1),NSMT -CrS3, £ 34±41:880N 139±20:350E,southof Oshima I., IzuIs., TR V“Shin’yo-maru” ,dredge, 106-103 m, coll. H.Komatsu,24 October 2002; 1 ovig.female (6 :2 9:3),CBM-ZC 5288,34 58:47 N 139 34:15 E, £ ± 0 ± 0 OkinoyamaBank, Sagami-nada, TR V“Shin’yo-maru” cruise SY96, stn 19, dredge, 121-129 m, coll. T.Komai,24 October1996.

Description.— This newspecies is closely similar to Goneplaxmarivenae n. sp.in most diagnostic characters, so that onlythe differences are describedin this section. Carapace(Ž g. 4a) 1.4-1.5 times broaderthan long (n 9; holotype1.5 D times); regionsill-deŽ ned, slightly depressedat center.Frontal region(Ž g. 4b) with small obtusemedian tooth. Lateral marginof carapace with large,acute triangular extraorbital toothon corner with supraorbital margin,with small acute triangular toothjust behindextraorbital tooth,both teeth directed antero-laterally. Eyestalk (Žg. 4c) long, slightly protrudingfrom orbit whenretracted; cornea muchbroader than stalk, 0.5times as longas total eyestalk. Mouthpartsshown in Ž g.5.First maxilliped (Žg. 5d): coxal endite withdense plumosesetae; basial endite fringedwith plumose setae; endopodfringed with veryshort setae alongdistal marginand with plumosesetae alongproximal half ofmesial margin.Second maxilliped (Žg. 5e): propoduswith longsimple setae aroundlateral margin; exopodelongate, fringed with short plumosesetae along proximalhalf oflateral marginand distal margin. Cheliped(Ž g. 4d-i)2.3, 2.0, 2.2, and 1.8-1.9 times as longas carapacein adult male (n 1; holotype),young male (n 1),adult female (n 2) and young D D D female (n 3),respectively; merus with small acute triangular toothat proximal D 0.4of outer margin of dorsal surface,tooth directed distally,Žnelygranulate on innersurface; carpuswith small triangular toothon inner angle. In adult male, cutting edgesof right chela (Žg. 4d) leaving gap; cutting edgeof movableŽ nger almost bluntexcept proximal rounded tooth. In young male andfemale, cutting edgesof right chela (Žg. 4f) entirely meeting,more strongly dentate thanthose of left chela,with roundedor roundlytriangular process onbase ofimmovableŽ nger androunded tooth on proximal-end of movableŽ nger. Ambulatorylegs (Žg. 4j, k): Žrst to fourthleg about2.1-2.3, 2.4-2.6, 2.3-2.5, and1.8-1.9 times as longas carapace(holotype 2.1, 2.5, 2.4, and 1.9 times), respectively. Thoracicsternites andabdomen (Ž g. 6a, b) of bothsexes as in G.marivenae . Male abdomen:Ž rst somite (Žg. 6b) longer than that of G.marivenae . 1252 HIRONORI KOMATSU&MASATSUNET AKEDA

Fig. 4. Goneplaxmegalops n.sp. a-e, h-k, holotype, male, Kerama Is., Ryukyus, southwestern Japan, 6:2 9:5mm, CBM-ZC 7031;f, g, paratype, young male, Izu Is., central Japan, 4 :8 7:3 mm, £ £ NSMT-Cr5603. a, Carapace, dorsal view; b, frontal region, antero-dorsal view; c, same, frontal viewof left half, with  agellumof antennule retracted in fossa; d, f,right chela, tip of immovable Žngerbroken in d, external view; e, g, left chela, external view; h, merusand carpus of cheliped, dorsalview; i, same,lateral view; j, k, leftthird and fourth ambulatory legs, dorsal views. Scales: a 2mm; b,c, d-e,f-g, h-i, j-k 1 mm. D D TWO NEW GONEPLAX 1253

Fig. 5. Goneplaxmegalops n.sp., paratype, young male, off Oshima I., IzuIs., central Japan, 4:8 7:3mm, NSMT-Cr5603. Right mouthparts in external view ,setaeof each  agellumof £ exopodomitted: a, mandible,medially broken; b, maxillule;c, maxilla;d, Ž rstmaxilliped; e, second maxilliped;f, third maxilliped. Scales: a-e, f 0.5 mm. D 1254 HIRONORI KOMATSU&MASATSUNET AKEDA

Fig. 6. Goneplaxmegalops n.sp., holotype, male, Kerama Is., Ryukyus, southwestern Japan, 6:2 9:5mm, CBM-ZC 7031:a, sternum and abdomen, ventral view; b, same, posterior view; £ c,rightmale Ž rstpleopod, external view; d, tip of same, external view; e, rightmale second pleopod, externalview .Scales:a-b 2 mm; c, e 0.5 mm; d 0.2 mm. D D D Male Žrst pleopod(Ž g. 6c) weakly compressed, getting narrower distally, with some verysmall spinule setae just belowapical aperture; mesial marginfringed with short simple setae alongdistal half; lateral marginfringed with plumosesetae atbasal part; apical aperturelargely openon external surface; tip (Žg. 6d)acutely triangular,directed laterally,with some veryshort setae alonglateral margin.Male secondpleopod (Ž g. 6e)as in G.marivenae . Colourof young female. —Dorsal surface ofcarapace,cheliped, and ambula- torylegs symmetrically speckledwith darkmagenta punctae; ventral surface ivory- white.Eyestalk darkmagenta; corneablack. Etymology.— ThespeciŽc nameis acombinationof the Greek,mega ( large) D and ops ( eye),in referenceto the characteristic, large stalked eye.It is anounin D appositionto the genericname. Distribution. —Japan: IzuIs., offKii Peninsula,and Ryukyus (type locality). Occurringat depthsof 23-184m. Remarks. — Goneplaxmegalops n.sp. can be distinguishedfrom its congeners as explainedin the ‘Remarks’section of G.marivenae n.sp.,above.

ACKNOWLEDGEMENTS

Theauthors wish to express their cordial thanksto Ms. MariveneR. Manuelof the National Museumof the Philippines andto local Žshermenof Balicasag Island fortheir kindhelp during the Želd trip inthe Philippines. Thanksare also dueto TWO NEW GONEPLAX 1255

Fig. 7. A, Goneplaxmarivenae n.sp., holotype, male, Balicasag I., Bohol,Philippines, 12 :3 £ 18:2mm, NSMT-Cr15531; B, Goneplaxmegalops n.sp., holotype, male, Kerama Is., Ryukyus, southwesternJapan, 6 :2 9:5mm, CBM-ZC 7031. £ Dr.T. Komai of the Natural History Museumand Institute, Chiba,for providing us with the valuablespecimens.

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Firstreceived 5 July2003. Finalversion accepted 19 August 2003.