Aquatic Beetles of the Families Dryopidae and Elmidae (Insecta: Coleoptera: Byrrhoidea) of Thailand: Annotated List and Illustrated Key to Genera
NAT. HIST. BULL. SIAM SOC. 61(2): 89–126, 2016 AQUATIC BEETLES OF THE FAMILIES DRYOPIDAE AND ELMIDAE (INSECTA: COLEOPTERA: BYRRHOIDEA) OF THAILAND: ANNOTATED LIST AND ILLUSTRATED KEY TO GENERA William D. Shepard1 and Robert W. Sites2* ABSTRACT Over the past two decades, Thailand has been surveyed extensively for its fauna of aquatic insects, including the beetle families Dryopidae and Elmidae, which are abundant in stream systems. Previously, nine genera were known from the country, all but one of which were re- collected: Parahelichus, Stenomystax (Dryopidae); Ancyronyx, Aulacosolus, Dryopomorphus, Graphelmis, Macronevia, Macronychus, Pseudamophilus (Elmidae). Here we add 13 genera not previously recorded from Thailand (Dryops, Elmomorphus, Helichus, Pachyparnus, [Dryopidae]; Grouvellinus, Indosolus, Leptelmis, Ordobrevia, Potamophilinus, Stenelmis, Urumaelmis, Zaitzevia, Zaitzeviaria [Elmidae]) and provide an annotated list and illustrated key to the genera. Additional information is provided to guide new work with Dryopidae and Elmidae in Thailand. Keywords: Coleoptera, Dryopidae, Elmidae, Thailand, aquatic, taxonomic keys, checklist INTRODUCTION Thailand is a topographically and ecologically diverse country. Its many mountain ranges and high rainfall average of 240 cm/yr (ARBHABHIRAMA ET AL., 1988) give rise to a multitude of streams and rivers. These streams and rivers support diverse lotic communities, especially of insects. Thai aquatic insects have been studied taxonomically across most orders includ- ing Ephemeroptera (UÉNO,
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