SBORNÍK NÁRODNÍHO MUZEA V PRAZE ACTA MUSEI NATIONALIS PRAGAE Řada B – Přírodní vědy • sv. 64 • 2008 • čís. 1 • s. 5–37 Series B – Historia Naturalis • vol. 64 • 2008 • no. 1 • pp. 5–37

WALLROTH´S COLLECTION OF VASCULAR PLANTS IN THE HERBARIUM OF THE NATIONAL MUSEUM, PRAGUE

BLANKA SKOČDOPOLOVÁ National Museum, Prague, Department of Botany, CZ - 252 43 Průhonice, Czech Republic, e-mail: [email protected]

JINDŘICH CHRTEK sen. Mimoňská 639, CZ - 190 00 Praha 9, Czech Republic

Skočdopolová B. and Chrtek J. (2008): Wallroth´s collection of vascular plants in the herbarium of the National Museum in Prague. – Acta Mus. Nat. Prague, Ser. B, Hist. Nat., 64(1): 9-42, Praha. ISSN 0036-5343.

Abstract. The National Museum in Prague in the past bought a large part of the botanical inheritance of German botanist K. F. W. Wallroth (1792-1857) - about 8 000 specimens of phanerogams including Wallroth´s own type material. In this present article sixty three new names of taxa are discussed including some remarks relating to the original diagnoses, descriptions of herbarium specimens and typification. Wallroth´s life and the history of his phanerogamic collection, preserved in Prague, are also briefly considered.

n Botany, Wallroth´s herbarium, phanerogams, type collection, lectotypification

Received December 20, 2007 Issued September, 2008

Introduction Heringen, later as district doctor in Nordhausen (1825- 1855). He died on March 22 1857 in Nordhausen. The preservation of Wallroth´s collection in the herbar- His renowned work “Schedulae Criticae de Plantis Flo- ium of the Botany Department of the National Museum in rae Halensis selectis” appeared in 1822. This publication Prague (PR) is a fact not very well-known in botanical cir- was intended as an addition to Sprengel´s Florae Halensis cles. The collection includes especially the material from (1806). In his interesting floristic and taxonomic work, Central Germany, which was mainly used in the elaboration Wallroth listed 419 species of phanerogams with more or of his two main works: Schedulae criticae (1822) and Erster less detailed descriptions and in addition described 292 new Beitrag zur Flora Hercynica (1840). From his herbarium taxa. Further new taxa of phanerogams he described in collection sixty three type specimens, mostly at species Erster Beitrag zur Flora Hercynica (1840), which was pos- level, were selected and reported on. The article may also sibly preprinted or reprinted from Schlechtendal´s Linnaea serve as a short survey of Wallroth´s life and work, includ- ing the history of his collection deposited in Prague. in 1840 (Stafleau et Cowan 1988:45). In this paper we The aim of the present paper is to contribute to the fund quote from Linnaea (Linnaea 14:1-158, 529-704, 1840), of knowledge on the museum´s herbarium and to point out where Wallroth´s text appeared with the title „Scholion zu the value of Wallroth´s materials. Hampe´s Prodromus Florae Hercyniae” and the subtitle “Ein Sendschreiben an den Apotheker Herrn Ernst Hampe Life and work of K. F. W. Wallroth zu Blankenburg“. Originally a mutual work on the flora of the Harz Region with a pharmacist Ernst Hampe was pro- German botanist Karl Friedrich Wilhelm Wallroth was posed, but Hampe already published Prodromus Florae born on March 13 1792 in Breitenstein (South Harz Reg- Hercyniae in 1836. This publication annoyed Wallroth, who ion), as the son of a vicar. From youth he was interested in responded with his own detailed revision of Hampe´s previ- botany. During his study at the monastery school in ously mentioned work. Rossleben (1805-1810) he investigated the flora in the Wallroth´s best work was said to be Flora cryptogamica vicinity of the town. In 1810 Wallroth started to study med- Germaniae, in which he described Filices, Lichenes and icine under the influence of Kurt Sprengel, Chief of Musci (1831), Algae and Fungi (1833). In his last published Medicine and Botany at the University of Halle. He fin- work Beiträge zur Botanik (1842-1844) eight complicated ished his medical study in Göttingen in 1815. In the same taxa of phanerogams were discussed. year he began serving as a military surgeon in the army. Wallroth became a well-known European botanist, spe- Wallroth worked one year, 1816, as a practicing physician cialized in floristic research in the South Harz Region and and from 1817 he was employed as a regional physician in he combined this work with taxonomic and nomenclatural

5 studies. He proposed many new names, but there were not National Museum in Prague. The main part consisted of his many taxa which remain accepted nowadays. literature searches e. g. Koch´s Additiones Kochianae zur Flora Teutschland´s (Synopsis florae germanicae et helveti- History of Wallroth´s estate cae) and excerpts concerning descriptions of various taxa. Also the manuscripts with descriptions of Wallroth´s either After Wallroth´s death, his numerous collections were published and unpublished taxa were present (Text-fig. 1). offered for sale (Anonymus 1858): herbarium of phanero- Further there is the manuscript of a monograph, dated 1822, gams, collections of lichens and other cryptogams, a collec- on roses – Symbolae de Rosis germanicis (Text-fig. 2). The tion of the genus Rosa and a collection of native woody second part of this manuscript contained an incomplete text, plants. Many of the plants which were for sale originated published in 1828 as Rosae plantarum generis historia suc- from the T. P. Ekart collection which Wallroth bought in cincta. There is also a small number of papers dealing with 1846. The cryptogams were bought by Swiss botanist J. E. lichens and fungi. Duby (1798–1885), lichens by G. L. Rabenhorst (1806– 1881) from Dresden and Ekart´s material was bought by Wallroth´s herbarium in Prague another German botanist J. F. T. Irmisch (1816-1879) from Sondershausen. In the above mentioned sale details from In the second half of the 19th century, during the cura- July 1858 Wallroth´s collection of phanerogams including torship of Čelakovský, all herbarium material was gradual- ferns and the fern allies (Lycopodium, Equisetum) were still ly classified according to Endlicher´s system including offered for sale. According to this reference Wallroth´s Wallroth´s collection. His herbarium was in those days sort- material was placed in blue thick folders and packed in 157 ed into individual species (exceptionally genera) and placed packets (originally fascicles), with many specimens in each in the common herbarium cover at the end of each taxon, taxon. where it has from that time always been kept in the general A further article published in Oesterreichische Botanis- herbarium of PR. Simultaneously the majority of Wallroth´s che Zeitschrift (Anonymus 1859) already detailed the pur- specimens were identified either by handwritten labels: chase of Wallroth´s phanerogams herbarium by the Muse- Wallr., Wallroth, Herbar. Wallroth or with oval-shaped um in Prague in April 1859. But in our acquisitions cata- stamps with an embossed inscription (without ink): MUS. logue of the Botany Department about 8 000 specimens BOH. COLL. WALLROTH. But also a rather large number from Wallroth´s collection were first recorded in 1861. of unmarked sheets existed, which it was not difficult to Obviously the new custodian, Ladislav Čelakovský, who recognize either by Wallroth´s typical handwriting or by the served in the museum from 1861 till 1902, was the first to reg- special common cover placed at the back of each taxon. ister Wallroth´s collection, with some inevitable delay. The Wallroth´s herbarium in the National Museum in Prague price of the herbarium was 80 thalers. According to the previ- contained material mostly from Thuringia, with many spec- ously mentioned article from 1859, the heiress handed over imens from the environs of Halle and from Harz. A fair Wallroth´s manuscripts in addition to the herbarium material. number of plants came from collections of other German The largest part of the phanerogamic herbarium was botanists, e. g. of his contemporaries E. G. Hornung, F. T. sold partly to Prague and partly to St. Petersburg (LE). Kützing, P. Salzmann, A. Spitzel and L. E. Schaerer from According to information from J. Štěpánek, Wallroth´s very Switzerland. Some specimens originated also from abroad, large collection of vascular plants preserved in LE originat- because Wallroth received or exchanged them with ed from the herbarium of his contemporary, German renowned European botanists. It is still possible to find botanist F. C. Mertens (1764-1831). a few of his original thick bluish folders in the herbarium PR. Stafleu et Cowan in Taxonomic Literature (1988) stated Wallroth´s herbarium had one special peculiarity – the that more of Wallroth´s material is preserved in the follow- labels. Some of them were inaccurate and often without any ing European herbaria: Bruxelles, Genève – also in Herb- localities. Date, rather the year or month of collecting was arium de Candolle, Göttingen, Halle, Jena, Kiel, Moscow published occasionally. The labels were mostly written in and Warsaw. The cryptogams are deposited in Strassbourg. Wallroth´s characteristic somewhat illegibile handwriting in In Berlin and Leipzig the material was mainly destroyed pen, less frequently in pencil. According to the above men- during the war. tioned botanist Kützing (1857), Wallroth kept the localities One of us (B. S.) has seen Wallroth´s material in the especially of his new and rare species secret, so that nobody Herbarium of Martin Luther´s University in Halle – Witten- could find them. Therefore the exact localities had not been berg (HAL) – e. g. original material of Agrimonia gry- always cited, neither in his work, nor on the herbarium posepala, Crepis polymorpha, Erythraea angustifolia, Hal- sheets. On the labels often only abbreviations were men- imus pedunculatus, Rosa marginata, Vicia multicaulis, tioned, e. g. Thg., Thr., Thrg.,Thurg., Thuring. [Thuringia] Rumex sylvestris and Orobanche rubens, and also in or Fl. hal., agr. Hal. [Region of Halle]. Not even his name Haussknecht Herbarium in Jena (JE), e. g. original material was always indicated. But he had usually never forgotten to of Agrimonia procera, Artemisia seriphium and Armeria add m. (most labels), mihi, mei, W., Wallr. or exceptionally halleri. The labels had the same characteristic features, Wallroth behind each name of his new taxon. The original often without locality or date, as labels in Wallroth´s collec- labels are also missing when there was too much material of tion in Prague. some individual species in the original cover. Then the To the purchase of the herbarium material, Wallroth´s material has been divided or rearranged on several sheets manuscripts were added. This literary estate was placed with only one original label. The labels were exceptionally separately in the archives of the Botany Department of the absent.

6 Text-fig. 1. Wallroth´s original handwriten page dealing with Valeriana collina WALLROTH.

7 Text-fig. 2. Title page of Wallroth´s unpublished work on roses.

8 According to the conventions of the 19th century, Wall- Orobanche reticulata, Sparganium minimum and some roth was an economical man, he used for the labels and species of the genus Potamogeton. envelopes, the overleafs of official requests and letters, con- nected with his post as regional physician. These pieces of Explanatory note letters documented that sometimes the labels were written The following list of Wallroth´s type specimens pre- much later than the names of new taxa were published. This is e. g. the case with Pinguicula gypsophila, published in served in the herbarium of the National Museum in Prague 1822, but provided with a label made from a letter from is arranged alphabetically. 1830. The secrecy of localities was probably the reason why Every taxon is treated according to the following pat- Wallroth wrote some of the labels later. tern: In the past only a few original specimens were selected 1/ Name of Wallroth´s taxon, the original publication with and placed in the special type collection of PR, the majori- quotation – part, page and year of publication (if possi- ty of Wallroth´s types remained in the general herbarium. ble the number of table with illustration) and the num- Not until a few years ago, more than one hundred type spec- ber of type collection in PR herbarium. If there existed imens of various taxonomical levels were selected and more sheets of the same species, they were designated A, incorporated into a separate collection of types in PR. B, C etc. Wallroth´s material from Thuringia and Harz Region 2/ Original diagnosis. From Wallroth´s literary work only contained rather frequently his new, but never published the most important diacritical features are mentioned, names, which appeared only on the labels, not to mention because the original text was very detailed and often it many, not validly published names of varieties. did not help to distinguish individual species. In Wallroth´s collection there were specimens from all 3/ Locality cited from Wallroth´s work. In fact, they were over Europe and also a small number from overseas. For not always stated. some of them Wallroth suggested new names, which were 4/ Brief notes on the herbarium specimen. A very short mentioned directly on the labels, but they were never pub- description of plants on the sheets, including the pres- lished. Thus he named e. g. a few new species of genus ence or not of Wallroth´s original envelope with remains Plantago – some of them originating from Russia (P. tauri- of flowers or fruits. ca), Greenland (P. gronidda), Italy (P. marginata), Cape The quantity of plants on the sheets often present some Town Region (P. hians), North America (P. longifolia), problems. The specimens in the herbarium cover have Brazil (P. curvirostris), Australia (P. polyphylla) or from usually only one of Wallroth´s original labels. The Juan Fernandes (P. berteroi). plants were sometimes so numerous, that it was neces- Only the species Plantago uniglumis WALLR. ex WALP., sary to divide them, either previously or during this collected by W. S. J. G. Besser, is mentioned in R. K. F. Pil- study. The duplicate sheets were designated with letters ger´s monograph on Plantago: „Hochgebirge von Chile: (A, B, C etc.). In this case the lectotype was usually Walpers l. c. gibt an: in uniglumis Wallr. Monogr. Plant. selected on the basis of the presence of an original label ined.! Besser, Pl. Chil. 105!... (Engler Das Pflanzenreich, on the sheet. A further important factor for designation 1937, IV. 269/102 Heft: 113)“. The specimen of Plantago of the lectotype was the condition of plant. uniglumis, mounted on the sheet PR 3952 contained one 5/ Label. The text on the labels has been transcribed small leafy plant (1,5 cm) with two flowers. The label was between quotation marks („ „), complying as far as pos- very brief: „Plantago uniglumis m. / Chili. Besser“. sible, with the original writing. If more than one label A large number of Wallroth´s type specimens, including had been attached to the same sheet, each of them is many varieties, are not presented in our article below, but numbered. The labels contained the transcription of the they had been already selected and placed in the separate name of the taxon (not printed in italics according to botan- collection of types. Digitalised images of further Wallroth´s ical convention), the locality and the date of collection taxa as e. g. Agrimonia gryposepala, Agrimonia platycarpa (when present) including full stop, exclamation and ques- (two fruits only), Agrimonia serrifolia, Anthemis hetero- tion marks. phylla, Anthemis×sulphurea, Armeria halleri, Avena×villis From the text on the labels we suggest the place of occ- (only seeds exist, the material was sent before World War II. as a loan to German botanist Kurt Wein and it was not urence with contemporary orthography or more detailed returned), Cerastium gracile, Conopsidium platantherum, geographical location, added in brackets [ ]. Crepis capillaris, Crepis polymorpha, Dictamnus sessilis, Further symbols were used for a new line ( / ) and for Dictamnus suffultus, Erysimum divaricatum, Hieracium untranscribed text because of illegibility ( ... ). pedunculatum, Hieracium versicolor, Lapsana sylvatica, Morphological features and literary data written on the Lathyrus viciaeformis, Nasturtium tentaculatum, Ono- labels have not been transcribed in most cases, because brychis decumbens, Polygonum nervosum, Reseda tenuifo- they were often identical with the text in the original lia, Rumex cristatus, Rumex sylvestris, Sonchus fallax, Xan- publication. thium pungens will be presented in the near future on Inter- The text was mostly written in Wallroth´s own hand, net. when not, it is mentioned. In spite of the fact that a great number of Wallroth´s 6/ Selection of type specimens has been also indicated by types originating from Thuringia or the Harz Region have some specialists, who had seen Wallroth´s material. been deposited in the herbarium PR, there remain many 7/ Identification of material is based on recent taxonomic type specimens which have had not been found there, e. g. research.

9 Wallroth´s selected type specimens preserved lens, but these names of infraspecific taxa are not present in in herbarium PR the herbarium PR. For plants placed on sheets A and B, Wallroth used different, unpublished names of varieties – α. Aconitum bernhardianum WALLROTH phoenicea and γ. leucantha. Sched. Crit.: 250-252, Tab. II, 1822. – PR 11761. Locality for varieties α, β: Inter seget- es hybernas ad Rossleben, Querfuth, Lodersleben [SW of Original diagnosis: Cuculli calcare brevi Querfurt]. spirali, labio ovato-lanceolato bifido recto, galea conica longe unguiculata antrorsum in rostrum protracta a petalis Brief notes on the specimens: On the lateralibus subspathulatis obliquis retracta, petalis inferi- first sheet A three complete plants are mounted – the first oribus lanceolatis obtusis deflexis .... (52 cm long), with ripened achenes, the second (63 cm) with achenes and flowers and the third one (22 cm) in full flower. Locality: In nemore Loderslebensi [Lodersleben, SW of Querfurt] passim .... L a b e l : „Adonis maculata / α. phoenicea / mihi“. Two labels with some morphological features were added. B r i e f n o t e s o n t h e s p e c i m e n : Upper Locality is not given. part of plant (ca 32 cm) with flowers and leaves. The her- Two small simple plants (13 and 19 cm long) are on the barium plant fully corresponds with the plant on the picture second sheet B, one of them with a flower, the second dam- published in Schedulae criticae. aged plant had a brokenoff flower. L a b e l : „Aconitum Bernhardianum mihi / four syn- L a b e l : „Adonis maculata / γ. leucantha / fl. hal.“. onyms followed / Fl. Har.“ [Harz]. The precise locality is not indicated. The sheet PR 11761 is selected here as a lectotype. It is not appropriate to select the lectotype from Wall- This plant corresponds to Aconitum variegatum L. roth´s material as it was published as a new name (nomen N o t e : In Wallroth´s material there is one specimen novum). obviously originating from Sprengel´s collection. The text on All plants correspond to Adonis aestivalis L. the label: „Aconitum neomontanum Lodersleb.“ is written probably by Sprengel, who used this name for plants collect- Alchemilla glaucescens WALLROTH ed near Lodersleben (see Wallroth Sched. Crit.: 251, 1822). Linnaea 14: 134, 1840. – PR 4400. Adonis anomala WALLROTH Text-fig. 3. Sched. Crit.: 273-276, 1822. – PR 11782 A, B. Original diagnosis: Hätten Sie, wie ich in Original diagnosis: Floribus tripetalis, petal- Willdenow´s Sammlung die Reihenfolge der verschiedenar- is oblongis planis obtusiusculis patentibus ungue concol- tigsten Exemplare von Alchemilla montana gesehen: so oribus, calycibus ovatis acutiusculis extus pilosis... würden Sie die am Harz vorkommende und von mir zuerst in meinem Annus bot. als A. montana beschriebene Art Locality: Auf dem Schachtberge inter Bottendorf nimmermehr als A. montana Willd. n. 225 aufgestellt ha- et Rossleben, Freyburg [NW of Naumburg], Querfurth et ben. Ueber dieselbe werde ich mich aber anderwärts erk- similibus locis minus frequens. Extra fines inter Tilledam lären und bemerke nur noch, dass ich dieselbe längst als A. [Tilleda] et Borksleben [Borxleben, NW of Artern]: in arvis glaucescens aufbewahrt und ausgegeben habe. juxta collem Mittelberg dictum [a hill NE of Nordhausen], Wallroth´s above mentioned diagnosis as A. montana prope Auleben [Kyffhäuser]. from Annus botanicus (1815: 26) is the following: Foliis Brief notes on the specimens: On the reniformibus, novemlobis subtus caule petiolisque pilosis first sheet A, there are five complete plants (10–22 cm), sericeis, floribus fastigiatis glomeratis sessilibus. three of them flowering and two plants with more or less Locality was not given. ripened fruits. Brief notes on the specimen: Two ro- L a b e l : „Adonis anomala mihi“. settes of leaves with 2 and 6 flowering stems. Three flowering plants are on sheet B. The original label L a b e l : „Alchemilla glaucescens m.“ / some differ- is absent. ences followed / „Inter Leimbach et Steigerthal [E of Nord- The specimen PR 11782 A is selected here as a lecto- hausen] in pascuo, / den Plan dicto .... / Juni 46“. type. In 1937, W. Rothmaler designated this specimen on the The plants correspond to the species Adonis flammea revision label as a type of Alchemilla glaucescens WALLROTH, JACQ. identical with the species Alchemilla hybrida MILL. Now the species A. glaucescens is considered as an independent one. Adonis maculata WALLROTH The specimen PR 4400 is selected here as a neotype. Sched. Crit.: 270–273, 1822. – PR 11781 A, B. Original diagnosis: Petalis suboctonis obo- Alchemilla glaucescens var. β. alpestris WALLROTH vato-oblongis planis basi maculatis calycibusque glabris obtusis, carpellis quinquefariis conoideo-prismaticis stylo Linnaea 14: 549, 1840. – PR 4398 and PR 4399. recto concolori rostratis supra commissuram marginato-cir- Original diagnosis: Caulibus adscenden- cumscissis... tibus petiolisque villosis, foliis glaucescentibus utrinque Wallroth divided this species into three varieties α. mini- subsericeis .... calycibus e basi obtusa rotundatis estriatis ata (floribus miniato-phoeniceis), β. ochroleuca and γ. pal- lacinias ovatas aequantibus pedicellisque lanatis.

10 Text.-fig. 3. Lectotype specimen of Alchemilla glaucescens WALLROTH.

11 Locality: In Laubwäldern des südöstlichen Harzes. Label: „Arabis brassicaeformis m. / Erysimum Brief notes on the specimens: One com- alpinum Baumg. DC. / ex loco Thalii / Sondershausen plete plant with two flowering stems are on the sheet PR 4398. Thurg.“ T w o l a b e l s : 1. „Alchemilla glaucescens m. / var. The specimen on sheet PR 2330 is selected here as a lec- alpestris / A. montana Willd. pr. p. / Breitenstein [NE of totype. Nordhausen] 1812“ and 2. „Quid vero! Harz ..... / in coryl- According to J. Štěpánek (in Kubát 2002) the correct etis ad Breitenstein Hercyn / legi“. name of this species is Arabis pauciflora (GRIMM) GARCKE. Two complete plants with flowers and one flowering stem separately are on the sheet PR 4399. Artemisia mertensiana WALLROTH L a b e l : 1. „Alchemilla montana / In nemoribus pr. Sched. Crit.: 466, Tab. V, 1822. – PR 11785 A, B. Lauterberg [Bad Lauterberg im Harz] / Majo med.“ and Original diagnosis: Foliis demum glabrius- 2. „Alchemilla glaucescens m. / var. alpestris / Lauterberg culis eglandulosis, inferioribus bipinnatifidis .... caulinis 1826“. subsimplicibus linearibus, floribus globosis racemosis bre- The sheet PR 4398 is designated here as a lectotype. vissime pedunculatis nutantibus, receptaculo nudo, caule According to Rothmaler´s revision label (1937), speci- simplicissimo. mens on both sheets are Alchemilla hybrida MILL. Locality: Prope Borksleben [Borxleben], pagum ab oppido Artern [Kyffhäuser]. Alectorolophus parviflorus WALLROTH B r i e f n o t e s o n t h e s p e c i m e n s : Wall- Sched. Crit.: 318-320, 1822. – PR 11790 A, B. roth´s abundant material was mounted on two sheets, the Original diagnosis: Caule immaculato, cor- locality was not mentioned on the labels. ollae tubo recto laciniis ovatis breviore, labio superiore On sheet A there are five plants with 1–2 flowering obsolete bidentato, dentibus, pistillo incluso bracteisque stems (7–15 cm) and one rosette of leaves. The original concoloribus. envelope with remains of inflorescences is attached. Locality: In pratis editioribus ad Lodersleben, L a b e l : „Artemisia mertensiana m. schd. cr. / A. la- Ziegelrode [SW of Querfurt], ad Nietleben, Doelau [Niet- ciniata Koch et auct. fl. germ. nec Willd. / A. laciniata γ. leben and Dölau on W outskirts of Halle-Neustadt]. Mertensiana DC. prodr.V. 111 (recte VI. 111) / Bertero in Brief notes on the specimens: On lit.“ Further some distinguishing features from A. laciniata sheet A there are four complete plants (25–44 cm) with are added. fruits and flowers, further one fructiferous plant without On sheet B there are eight flowering plants with rosettes lower parts (45 cm).The original envelope, made from a let- of leaves and with flowering stems (6–16 cm long) in vari- ter, containing fruits was added. ous states of damage. L a b e l : „Alect. parviflorus / var. speciosa / hybrida L a b e l : „A. Mertensiana / mihi“. ex utos. consortio / Koenigshütte bei Lauterb. [S periphery The sheet PR 11785 A is selected here as a lectotype. of Bad Lauterberg] / i. Flussbette“. The plants correspond to Artemisia laciniata WILLD. On sheet B five flowering leafy plants (23–33 cm) and According to information from J. Štěpánek, a further one lower part of a plant (50 cm) without inflorescence large quantity of Wallroth´s original material of Artemisia were mounted. The original envelope containing fruits was mertensiana is preserved in the herbarium in St. Petersburg added. (LE). L a b e l : „Al. parviflorus“. The specimen PR 11790 A is selected here as a lecto- Atriplex ruderalis WALLROTH type. Sched. Crit.: 115-116, 1822. – PR 427. All specimens correspond to Rhinanthus minor EHRH. Original diagnosis: Foliis hastato-deltoideis dentatis oppositis omnibus subconformibus, summis sen- Arabis brassicaeformis WALLROTH sim minoribus, valvulis ovatis integerrimis semen aequan- Sched. Crit.: 359-361, 1822. – PR 2330. tibus in paniculam subramosam congestis. Original diagnosis: …. Caulinis cordato- Locality: In ruderibus prope Lodersleben [SW of sagittatis amplexicaulibus integerrimis subtus cauleque Querfurt]. Extra fines ad fossas circa Goersbach [Görsbach, stricto glaucescentibus, siliquis erectis obsolete quadran- SE of Nordhausen]. gulis stigmate brevissimo coronatis. B r i e f n o t e s o n t h e s p e c i m e n : One lea- Locality: In aggeribus sylvaticis montosis ad Sch- fy plant broken into three pieces (30, 20 and 13 cm) with mon [SW of Querfurt] et ad Biebram [Bad Bibra, NW of fruits. Naumburg] .... ad Sondershusam [Sondershausen] …. am T w o l a b e l s a r e p r e s e n t : 1. „Atr. ruderali Straussberge [Straussberg W of Sondershausen], supra mihi“ and 2. „Atriplex ruderalis m. / A. microsperma W. et Haynrodam [Hainrode NW of Sondershausen]. K. / potius Atr. patulae var. / microtheca / Goersbach“. B r i e f n o t e s o n t h e s p e c i m e n s : Four The specimen on the sheet PR 427 is selected here as a leafy plants with fruits (23, 29, 38, and 40 cm), one lower lectotype. part of a stem without any flower or fruit and one broken According to personal communication with B. Mandák, stem with one leaf. a specialist on the genus Atriplex, the plants correspond to

12 Atriplex prostrata DC. subsp. latifolia (WAHLENB.) RAU- inferioribus supra basin petioliformen sublyratis lanceolatis SCHERT. .... achaeniis ovato-oblongis striatis subcalvescentibus .... Locality: Auf unfruchtbaren Grasplätzen, in ver- Camelina sylvestris WALLROTH fallenen Steinbrüchen des südwestlichen Harzes stellen- Sched. Crit.: 347, 1822. – PR 11788 A, B. weise im Spätherbst. Text-fig. 4. Brief notes on the specimens: Two Original diagnosis: Siliculis ovato-rotun- branched leafy plants (42 and 50 cm) with cauline leaves datis umbonatis in basin attenuatis margine compressius- linear or lanceolate, lower leaves pinnately lobed, involucre culis, caule foliisque sagittatis subintegris hirsutis. ovoid-globose. Locality: In tectorum, murorum grumis, collibus L a b e l : 1. „Centaurea phaeocephala m. / achaeniis apricis vicinisque agris sterilibus ad Bennstaedt [Bennstedt pappo brevis coronatis / striatis. / Ad vias publ. der Sch- near Halle], Schmon and Lodersleben [SW of Querfurt], erns“ [Schern, SW of Nordhausen] and 2. „Achaenia pappo Rossleben et alii circa urbem locis. coronata / Centaureae m. phaeocephalae“. The text of the B r i e f n o t e s o n s p e c i m e n s : The plants second label is written on a small label, where two achenes originaly placed in one common cover were later divided are also stuck. On the original envelope with achenes and onto two sheets with Wallroth´s label placed on sheet A. the remains of florets is the following legend: „Centaurea / On sheet A is one well-developed plant (60 cm long) Achaenia striata, pappo brevi .... / Schern bey Northus.“ without basal part, with ripened fruits and two whole small The specimen PR 11794 is selected here as a lectotype. plants (25 and 30 cm) with leaves, fruits and roots. In 1970, Czech specialist on the genus Centaurea, J. Five upper parts of fruiting stems (19–22 cm long) with Dostál identified these plants as Jacea pannonica (HEUFL.) empty capsules were on sheet B. SOJÁK ssp. pannonica. L a b e l : „Camelina sylvestris Wallr.“ A few morpho- logical features and six synonyms followed according to the Chenopodium microspermum WALLROTH text in Schedulae criticae. The locality was not mentioned. Sched. Crit.: 112 and 507, 1822. – PR 11784 A, B. In 1968 M. Smejkal on the revision label designated the O r i g i n a l d e s c r i p t i o n : Foliis triangularibus sheet 11788 A as „verisimiliter holotypus of C. sylvetris in basin subrhomboideam attractis parce dentatis, racemis WALLR.“ and he classified this species as Camelina micro- ramosi erectis subaphyllis folio brevioribus axillaribus ter- carpa ANDRZ. subsp. sylvestris (WALLR.) HIIT. minalibusque, seminibus minutissimis fuscis. The designation of the specimen PR 11788 A as the holotype is not correct. We consider it as a lectotype. Locality: In aggeribus campisque salsis circa Halam; ad Seeburg [W of Halle]; extra fines ad Artern [Kyffhäuser], auctumno. On p. 507 the following remark Campanula thaliana WALLROTH was added: folia carnosa: locum natalem: in pratorum sali- Sched. Crit.: 86, 1822. – PR 11763 A, B. norum aggeribus vor Langenbogen [W of Halle]. Original diagnosis: Caule simplici folioso Brief notes on the specimens: One up- basi foliisque subtus incano-pubescentibus inaequaliter per part of leafy plant (36 cm) in full flower is on sheet A. dentatis .... pedunculis in racemum longissimum digestis, L a b e l : 1. „Chenopodium microspermum / for. dentibus calycinis linearibus brevibus denticulatis. carnosi / Cf. Schd. cr. 507 / Agr. hal.“ and 2. „Ch. micros- Locality: In collibus herbidis ad Lettin [ NW of permum m. / for. carnosa“. Halle], Rossleben, an der Steinklippe [ near Wendelstein, A whole leafy plant (70 cm) with inflorescences and one SE of Rossleben], et aliis agri hal., thuring. et hercyn. locis small broken branch (10,5 cm) were mounted separately on rarius. sheet B. Brief notes on the specimens: On sheet L a b e l : „Chenopodium microspermum / mei.“ A one well-developed leafy plant (78 cm long) with inflo- The specimen PR 11784 A is selected here as a lecto- rescence is present. One leaf is mounted separately. type. The plant corresponds to the species Chenopodium La b e l : „Campanula Thaliana m.“ urbicum L. One fertile leafy plant (74 cm long) with ripened cap- sules is mounted on sheet B. Dianthus schizopetalus WALLROTH L a b e l : „Eadem fructifera / Ex eodem loco, in collis Linnaea 14: 570, 1840. – PR 11753. / aprici vicini vepretis lata.“ The plant PR 11763 A is selected here as a lectotype. Original diagnosis: Caulibus caespitosis ela- The plants correspond to the species Campanula bonon- tis .... dentibus calycis linearibus cuspidatis, petalis ex iensis L. ungue lineari in aream lanceolatam sensim abeuntibus in lacinulas angustissimas illa sexies longiores partitis .... Centaurea phaeocephala WALLROTH Locality: In trockenen Vorwaldungen des südli- Linnaea 14: 640, 1840. – PR 11794. chen Harzes stellenweise sehr häufig. Original diagnosis: Caule erecto anguloso Brief notes on the specimens: Two rigido foliisque arachnoideo-lanoso, radicalibus caulinisque complete plants, about 60 cm long, in full flower.

13 Text-fig. 4. Lectotype specimen of Camelina sylvestris WALLROTH.

14 L a b e l : „Dianthus schizopetalus m.“ In addition, Brief notes on the specimens: The many synonyms follow according to the text in Linnaea. lower part of a branched plant with root (9 cm), two fertile Down at the corner of the label „Thrg.“ is written. branches (ca 30 cm) with flowers and one broken branch The specimen PR 11753 is selected here as a lectotype. (8 cm) with one leaf. The plants are identical with Dianthus superbus L. L a b e l : Four labels were attached. One of them was probably written by Schaerer: „Cheiranthes pallens Hall. Erigeron corymbosus WALLROTH fil./ Hall. hist. n. 450 / Simplon“. Three further, very simi- Linnaea 14: 642-643, 1840. – PR 11557 A – D. lar labels were writen by Wallroth: 1. „Er. Schaererianum mihi. / E. helvet. DC. / Simplon Helvet.“; 2. „Erysimum Original diagnosis: Caule erecto corymboso Schaererianum / W. Schd. cr. 365 / Ex Helvetia“ and 3. folioso, foliis inferioribus e basi tenuata oblongis .... „Erysimum Schaererianum m. / W. Schd. cr. 365 / Ex Helv. achaeniis oblongis adpresso-pilosis politis utrinque carti- m. Simplon / am. Schaerer“. lagineo-costatis pappo exalbido triplo brevioribus. In 1974, A. Polatschek, a specialist on the genus Erysi- Locality: An Vorbergen des südlichen Harzen .... mum, on the revision label, designated this specimen as B r i e f n o t e s o n t h e s p e c i m e n s : On sheet A a lectotype and identified it as Erysimum rhaeticum (SCHLE- there are three fertile plants (62, 70 and 75 cm long) with ICH. ex HORNEM.) DC. rich inflorescences. L a b e l : „Erigeron corymbosum m. / caule simplici ela- Fumana pinifolia WALLROTH to / Koenigskopf [near Stempeda, NE of Nordhausen] Thr.“ Linnaea 14: 583, 1840. – PR 11769 A, B. Two fertile plants (33 cm and 42 cm) are on sheet B. Wallroth divided the species Helianthemum fumana L a b e l : „Erigeron corymbosum m.“ MILL. into two new species Fumana pinifolia and F. erici- On the further two sheets C and D, there are fertile plants folia. In herbarium PR only specimens of Fumana pinifolia (45–75 cm) with rich inflorescences, without any label. are preserved. Based on information on the labels, sheet PR 11557 A Original diagnosis: Suffruticosa, exstipula- was selected by specialist O. Šída as a lectotype in 2002. ta .... pedunculis axillaribus solitariis, fructiferis reflexis The sheets PR 11557 B, C and D he designated as isolecto- folia superantibus, calycibus clausis ovato-rostratis convo- types. lutis, foliolis calycinis scabriusculis, externis linearibus According to Šída the plants correspond to the species acuminatis, internis ovatis acutis. Erigeron muralis LAPEYR. (= E. serotinus WEIHE). Locality: Wurde von mir vor 30 Jahren an sonni- gen Gypsvorbergen, eine Stunde weit von der südlichen Erysimum cheiriflorum WALLROTH Harzgränze entdeckt. It was probably the locality of He- Sched. Crit.: 367, 1822. – PR 3931. lianthemum fumana, which Wallroth already mentioned in his work Annus botanicus (1815: 72): „in montibus cal- O r i g i n a l d e s c r i p t i o n : Foliis inferioribus careis inter arcem vetustam Rothenburg et Auleben [Kyff- oblongis obtusis remote-sinuatoque dentatis utrinque pube häuser] Thuringiae“. tripartita hirsutis, petalis subrotundis (speciosis) e basi obo- vata in ungues calycibus longiores abrupte attenuatis .... Brief notes on the specimens: Very rich collection – eight fertile dwarf shrubs, richly branched Locality: In collibus gypsaceis unweit Neustadt with leaves and capsules with 7–8 seeds. The plants origi- infra arcem Hohnstein [the village and the castle N of Nord- nally lying together in one common cover are now divided hausen]. onto two sheets A and B, with two of Wallroth´s labels Brief notes on the specimens: Two mounted on sheet A. upper parts of leafy plants in full flower (28 and 37 cm). Five fertile shrubs are placed on sheet A. Two original T w o l a b e l s a r e a t t a c h e d : 1. „Erysimum small envelopes made from old newspapers with remains of cheiriflorum m. / in sylvis pr. Neustadt“ and 2. „Ex collibus capsules were added. apricis gypsaceis / des Hohnstein citra / Nordhusen et Neu- L a b e l : 1. „H. fumana Auleben Thersdorf „ and stadt“. 2. „Fumana pinifolia m.“ Further detailed description with In 1980, specialist A. Polatschek on the revision label diagnostic morphological features in Latin follows. designated this specimen as an isotype of Erysimum cheiri- Three fertile shrubs and one small leafy shrub are florum WALLR. and identified the plant as Erysimum odora- mounted on sheet B. tum EHRH. Sheet PR 11769 A is selected here as a lectotype. The plants are identical with Fumana procumbens (DU- Erysimum schaererianum WALLROTH NAL) GREN.et GODR. [ = Helianthemum fumana MILL.]. Sched. Crit.: 365, 1822. – PR 11158. Galium agreste WALLROTH Original diagnosis: Foliis radicalibus lance- olato-linearibus subintergerrimis cauleque subramoso pube Sched. Crit.: 59, 1822. – PR 11766 A - E. bipartita hirsutiusculis, petalis ovato-subrotundis abrupte Original diagnosis: Foliis senis octonisve angusteque unguiculatis .... lanceolatis mucronatis basi attenuatis supra scabris margine Locality: .... olim ab amicissimo Schaerero, Ber- carinaque retrorsum hispidis .... germine minoribus, nate, in monte Simplon Helvet. fructibus reniformibus laevibus hispidisve.

15 Locality is not indicated, sensim angustioribus, capsulis obverse cuneiformibus Wallroth divided this species into two infraspecific taxa: bicornibus complanatis longe pedicellatis. α. echinospermon (fructibus aculeolis aduncis parce obsitis) Locality: Ad salinas Arterenses [Artern, Kyffhäu- and β. leiospermon (fructibus laevissimis); later in Linnaea ser] et Numburgenses [Naumburg] extra fines abundat. (Linnaea 14:548, 1840) he mentioned these varieties in Brief notes on the specimen: One reverse order (an orthographic variant β. echinospermum complete leafy plant in full flower with roots (23 cm). and α. leiospermum). Only the specimen α. echinospermum is present in the PR herbarium. L a b e l : „Halimus pedunculatus W. / Ad salinas Num- B r i e f n o t e s o n t h e s p e c i m e n s : A bunch burgenses“. of plants with leaves and echinaceous fruits are on sheet A. I have seen (B.S.) Wallroth´s original specimen of the The original envelope containing seeds was added. same species in the herbarium of Martin Luther University in Halle (HAL) in 2004. There is here one well preserved leafy L a b e l : „Galium agreste / var. echinosperma W.“ plant with a few branches on the sheet with the following text A further four duplicate sheets (B, C, D and E) are lying on the label: „Halimus pedunculatus / mihi / Wallroth.“ in one herbarium cover without any label. According to the The lectotype was not selected by us, because in this stage of development they all originated from the same col- case it is only a combination; Wallroth quoted the name lection. Atriplex pedunculata L. as a synonym. The sheet PR 11766 A is selected here as a lectotype of The plant corresponds to Atriplex pedunculata L. Galium agreste WALLROTH. The plants correspond to Galium spurium L. Heracleum ursinum WALLROTH Galium gracile WALLROTH Linnaea 14: 560, 1840. – PR 11759. Sched. Crit.: 57 and 503, 1822. – PR 11765 A – E. Original diagnosis: Foliis scabro-hirsutis pinnatis, pinnis lobatis, umbellis radiantibus, ovariis tenu- Original diagnosis: Foliis reflexis antrorsum iter pubescentibus, fructibus ovato-subrotundis apice emar- hispidis, inferioribus ovatis subquaternis, superioribus senis ginatis dorso umbonato hirsutiusculo tristriato vittas e basi lanceolatis duplo majoribus .... partialibus inaequalibus tenuata subclavatas ultra mediam fructus partem propexas bifidis aphyllis fructu ruguloso duplo longioribus. monstrantibus .... Further there is a note: „Steht zwischen Locality: In agris bey der Windmühle inter Teut- H. sphondylium L ..... und H. asperum M. Bieb.“ with very schen-Thal et Opphausen [Teutschenthal and Obhausen, detailed description of the fruit. SW of Halle], in arvis sterilibus inter Gleina et Lauchstaedt [ Laucha, NW of Naumburg] passim. In Addenda (p. 503) a Locality: Auf Wiesen des südlichen Harzes stel- supplement is added: In arvis arenosis hinter dem Zorgs lenweise. abundat [Zorge, S Harz]. Brief notes on the specimens: The mi- Brief notes on the specimens: Five ddle part of a hirsute stem (65 cm long) with only one in- sheets were originally placed together in one herbarium complete leaf, one small part of a leafless stem (17 cm) and cover, but only two of them were provided with a label. All one small complete leaf are mounted separately. The origi- plants were collected in the same state of development. nal envelope full of fruits with Wallroth´s inscription: „Her- On sheet A there is a bunch of stems with tiny leaves and acleum ursinum W. / Heringen“ [Heringen, N of Sonder- fruits. shausen] and further remarks concerning the synonyms L a b e l : „Galium gracile Wallr. / In arvis sterilibus / were added. inter Kleina [Gleina] et Lauchstaedt / 1817“. L a b e l : „Heracleum ursinum Wallr.“ On a further sheet B there is also a mixture of fertile plants. The plant on the sheet PR 11759 is selected here as L a b e l : „Galium gracile / Teutschen-Thal / 1812“. a lectotype. It is interesting that both the above mentioned labels The plant corresponds to Heracleum sphondylium L. were cut-out from one letter dated 6. Oct. 1819, which proved the fact that labels were occasionally written much Hieracium lactucella WALLROTH later than the plants were collected. Sched. Crit.: 408–410, 1822. – PR 5892 A, B. On sheet D there is the mixture of fruiting plants and a Text-fig. 5. label with locality: „Hinter d. Zorgs.“ Two remaining sheets (C and E) with a similar bunch of Original diagnosis: Stoloniferum, foliis rad- plants are without any label. icalibus lanceolatis acutis integris glabris glauco-viridibus, The specimen on the sheet PR 11765 A is selected here stolonum subspathulatis adscendentibus basi margineque as a lectotype. parce pilosis pedicellis calyces subaequantibus, caule scapi- All these plants are identical with the species Galium formi inferne foliolo lineari aucto 1–4 floro. parisiense L. Locality: …. ad Lodersleben [SW of Querfurt] haud longe ab ipso pago, ad Ziegelrode [near Rossleben]. Halimus pedunculatus WALLROTH B r i e f n o t e s o n t h e s p e c i m e n s : Seven Sched. Crit.: 117, 1822. – PR 11789. flowering plants with rosette of leaves, flowering stems Original diagnosis: Foliis caulinis inferiori- (10–25 cm) and stolons, inflorescence with 1 – 3 capitula, bus ovatis obtusis in petiolum attractis, summis oblongis are on sheet A.

16 Text-fig. 5. Lectotype specimen of Hieracium lactucella WALLROTH.

17 L a b e l : „Auricula L. Sp. pl. / Hieracium Lactucella L a b e l : „Lamium rubrum / Wallr. / Fl. Megap.“ m. / Thuring“. [Flora Megapolitana = Flora of Mecklenburg]. On sheet B seventeen more or less rather small plants On sheet B there are seven leafy stems with flowers with rosette-leaves, flowering stems (3–12 cm) and 1–3 (24–30 cm) and the lower part of one bunch with two flow- capitula in inflorescences are mounted. ering stems (18 and 27 cm). This is probably a mixture of L a b e l : „H. auricula L. / Hieracium Lactucella m. / plants with violet and white flowers. / var. angustifolia. / In ericetis udior inter Gr. Wechsung L a b e l : „Lamium rubrum m. / cum var. albiflora / m.“ [Großwechsungen] et Kemstedt“ [SW of Nordhausen]. On the third sheet C, five flowering plants (26– 36 cm) J. Chrtek jun., specialist on the genus Hieracium, in 2007, are mounted. Wallroth’s original label is absent. selected the plants on the sheet PR 5892 A as a lectotype. The plant on the sheet PR 11760 A is selected here as a The plants correspond to the present concept of this lectotype. species. All plants correspond undoubtedly to the species Lami- um maculatum L. N o t e : A further two sheets with Wallroth´s authentic specimens of H. lactucella are preserved in the herbarium Malva adulterina WALLROTH PR, and bear the following text on both labels: 1. „Hieraci- um lactucella / lusus Surcatus“ and 2. „Ad... H. lactucellam Linnaea 14: 611, 1840. – PR 11791. mei / / praestat / Ex Alp. Salisb.“ Text-fig. 7. Original diagnosis: Caule prostrato, foliis cordato-subrotundis 5–7 lobis, pedunculis aggregatis Hieracium rothianum WALLROTH demum porrecto-declinatis petiolisque adpresso-pilosis, Sched. Crit.: 417-419, 1822. – PR 5894. petalis calyce triplo longioribus, valvulis acute-marginatis Text-fig. 6. reticulato-rugosis. Original diagnosis: Foliis lancolatis in basin Locality: Am südlichen Harze hier und da. attenuatis utrinque cauleque ramoso paucifloro pilis confer- Brief notes on the specimens: One tis rufescentibus hispidis, pedicellis unifloris repetito- leafy stem (27 cm) with flowers, fruits and four remnants of dichotomis patentibus calyces pilosos .... branches at the bottom, two upper parts of leafy stems (5 Locality: In pineto halensi vicinisque locis, in and 10 cm) with flowers and fruits. rupibus ad Kröllwitz [NW suburb of Halle] passim. L a b e l : „Malva hybrida“ / and above this name is Brief notes on the specimens: Two written: „adulterina m. / „pedunculis adpresso pilosis / / bottom parts of plants with hairy leaves and stems (ca 23 cm) Thuring“. Some distinguishing features with M. rotundifo- without flowers. Two broken stems with remains of capitu- lia, M. neglecta and M. sylvestris were added. la (only sepals preserved) are mounted separately. The specimen PR 11791 is selected here as a lectotype. L a b e l : 1. „Hieracium echioides Lumn. / var. ramosa / The plants correspond to the hybrid Malva×adulterina H. rothianum Wallr. Schd. / Ex agro halensi.“ and 2. In WALLROTH (M. neglecta×M. pusilla). a rather long text concerning names, probably written by A. W. Roth, the locality „Haide bay Halle“ [NW suburb of Malva equina BRUNFELS EX WALLROTH Halle] is mentioned. Wallroth added (by pencil) on the Linnaea 14: 611-612, 1840. – PR 11793 A, B. label: „H. rothianum mihi / cymosum Spr.“ Original diagnosis: Foliis omnibus basi Botanists L. Čelakovský, G. H. E. W. Hieronymus and intacta cordatis stipatis conformibus, inferioribus leviter-, P. Ascherson had written supplementary notes on further superioribus profunde quinquelobis, lobis ovato-ellipticis enclosed labels. obtusiusculis inaequaliter dentatis .... petalis cuneatis pro- In 2007, J. Chrtek jun. selected the plants on the sheet funde emarginatis pedicello duplo brevioribus .... PR 5894 as a lectotype. The plants correspond to the pres- Locality: … am südlichen Harze stellenweise häufig. ent concept of this species. Brief notes on the specimens: One le- afy plant (ca 30 cm long) with roots, two branches with Lamium rubrum WALLROTH flowers and fruits are on sheet A. Sched. Crit: 300, 1822. – PR 11760 A, B, C. L a b e l : „peduncular pubes duplex! flore purpur. / Original diagnosis: Calycibus patentibus in Malva equina Brunf. / In locis lutosis pagi Nordhausen“. verticillos 10–12 floros laxe digestos, dentibus calycinis On sheet B the branch of a leafy plant with fruits, bro- inaequalibus, superioribus majoribus, foliis cordatis irregu- ken into four parts is mounted. lariter inciso-serratis, floralibus breviter petiolatis, imis The label is not present. remotis. The original specimen PR 11793 A is selected here as a lectotype. Locality: Ad sepes pomariorum, inter dumetorum The plant corresponds to the species Malva sylvestris L. vepreta ad Lodersleben [SW of Querfurt], Ziegelrode [Ziegelroda near Rossleben], Rossleben et alibi. Malva neglecta WALLROTH Brief notes on the specimens: On sheet A there is one well-developed plant with 5 flowering leafy Syll. Pl. Nov. 1: 140-141, 1824. – PR 11792 A, B, C. stems (22–28 cm), two leafy stems and the remains of three Text-fig. 8. lower parts of plants. Original diagnosis: M. seminum arillis

18 Text-fig. 6. Lectotype specimen of Hieracium rothianum WALLROTH.

19 Text-fig. 7. Lectotype specimen of Malva×adulterina WALLROTH.

20 Text-fig. 8. Lectotype specimen of Malva neglecta WALLROTH.

21 pubescentibus immarginatis dorso latusculisque laevigatis, foliolis obovatis irregulariter dentatis, caule basi ramosissi- calyculi foliolis linearibus, laciniis calycinis ovatis corolla mo adscendente. duplo brevioribus receptaculum lenticulare attingentibus, Locality is not mentioned. pedunculis fructiferis declinatis .... Brief notes on the specimens: On Locality is not given. sheet A one branched plant (48 cm long) with yellow flow- Brief notes on the specimens: One le- ers and young fruits is mounted. afy, branched stem (ca 80 cm long) with fruits is on sheet A. L a b e l : „Melilotus arvensis Wallr.“ L a b e l : „Malva neglecta W. / caulibus prostratis elon- On sheet B there are two upper parts of fruiting stems gatis“. (32 and 33 cm). Wallroth´s original envelope made from a One leafy stem with flowers and fruits (24 cm) is on sheet B. letter, containing seeds, and with the inscription: „Mel. L a b e l : „Malva neglecta W. / var. leucantha W.“ arvensis“ was added. On sheet C there is one branched plant (22 cm) with L a b e l : „Melilotus arvensis Wallr. / var. legum. apice flowers and fruits, the petioles 7–11 cm long. productiori / ancipiti / Thuring“. L a b e l : „Malva neglecta W. / var. multiflora“. One flowering stem broken into three parts is on sheet C. The specimen PR 11792 A is selected here as a lectotype. L a b e l : „Melilotus arvensis m. / var. latifolia m. / The plants correspond to the present concept of this stipules .... inciso-dentatis! / Kohnstein“ [a hill NW of species. Nordhausen]. The mentioned infraspecific name var. latifo- lia has not been published. Medicago stipularis WALLROTH The specimen PR 11752 A is selected here as a lecto- Linnaea 14: 630, 1840. – PR 11768. type. The plants are identical with Melilotus officinalis (L.) Original diagnosis: Foliolis late-obovatis PALLAS. subrhomboideisve a media inde parce crenatis apice ro- tundato-retusis .... leguminibus inermibus compressis reni- Melilotus tenella WALLROTH formibus parce-venosis pilosis, stylo illius diametro duplo breviore coronatis. Linnaea 14: 617, 1840. – PR 11751. Locality: Unter dem Sommergetreide auf steini- Original diagnosis: Caulibus erectis simpli- gen Aeckern des östlichen Harzes stellenweise häufig, cibus, stipulis heteromorphis .... racemis laxis .... seminibus anderwärts gar nicht. (helvolis), hilo punctiformi exilissimo. Wallroth added the distinguishing features in German: Brief notes on the specimens: One le- „Unterscheidet sich jedoch durch einen einfachen, schlan- afy plant (ca 65 cm long) with flowers and very young ken, nur oben in Blüthenäste zertheilten Stengel; breitere fruits, one broken part of stem (15 cm) with leaves and flo- Blättchen; doppelt kleinere Blüthen, deren Fahne von ganz wers. The fruit are practically glabrous. gleicher Länge mit den übrigen Blüthentheilen ist; zugerun- L a b e l : „Medicago stipularis Wallr. / Inter segetes pr. dete, sitzende Kapseln und mit einem sehr kleinen Nabel Biebram“ [Bad Bibra NW of Naumburg]. versehene Saamen.“ The plant on the sheet PR 11768 is selected here as Locality: Auf fruchtbaren Gypsvorbergen des a lectotype. südlichen Harzes stellenweise. The plant is identical with Medicago lupulina L. subsp. lupulina. Brief notes on the specimens: Six N o t e : Another of Wallroth´s species – Medicago plants, with simple stems (30–38 cm long), with 3-4 inflo- mniocarpa Wallr. ex Ser. is preserved in the herbarium PR rescences in different stages of flower are on the sheet. The and placed on two sheets. On the first sheet there are two flowers are small (ca 4 mm) and white. fertile plants with the label: „Medicago mniocarpa Wallr. / L a b e l : „Melilotus tenella m. / In promont. gypsaceis M. lupulina γ. unguiculata DC. / Ex Belgio pr. Athum [Ath, d. Kohnsteins“ [a hill NW of Nordhausen]. SW of Belgium] 1814“. On the second sheet four plants are The specimen PR 11751 is selected here as a lectotype. present and two labels with following text: „Medicago At present these plants are evaluated as small flowered mniocarpa W. / Ex Thuring. pr. Hrgen [ Heringen/Helme] / plants within the variability of the species Melilotus albus Medicago lupulina L. / monstrum unguiculatum“. De Can- MED. dolle (Prodr. 1825, 2: 172) mentioned Wallroth´s name: „Trigonella mniocarpa Wallr! in litt. Medicago mniocarpa Monotropa hypophegea WALLROTH Wallr! in litt. (v. v. et s.).“ within his taxon M. lupulina γ. Sched. Crit.: 191, 1822. – PR 11750 A, B. unguiculata (Ser. mss.). In this case it was probably con- nected with the monstrosity of the fruit. Text-fig. 9. Original diagnosis: Racemo paucifloro, nubili conferto nutante, fructifero surrecto laxo, floribus Melilotus arvensis WALLROTH ovatis eroso-laceris, staminibus styloque obconico tereti Sched. Crit.: 391-392, 1822. – PR 11752 A, B, C. glabris, stigmate subinfundibuliformi sub4-gono capsu- Original diagnosis: Leguminibus racemosis laque subrotunda glaberrimis. subdispermis ovatis acutis transverse-rugosis nudis demum Locality: .... in humo laxa faginetorum umbrosorum helvolia, alis vexilloque aequalibus carinam excedentibus, saltus Ziegelrodensis [Ziegelrode near Eisleben] et alibi.

22 Text-fig. 9. Lectotype specimen of Monotropa hypophegea WALLROTH.

23 Brief notes on the specimens: On sheet Brief notes on the specimens: The A there are ten glabrous plants in different stages of devel- plants mounted on sheets A and B were originally placed opment – from flowering to fertile plants (13–25 cm long). together probably as parts of one specimen with one com- L a b e l : „Monotropa hypophegea W.??“ After a short mon label. On sheet A there is one leafy plant (56 cm long) diagnosis, the isolated and conspiciously underlined word: with many fruits. On sheet B there are three parts of sterile „verte“ is written. Locality is not given, only biotope is leafy ascending stems. mentioned: „Ad fagor. radices“. Furthermore, a few of L a b e l : „Nasturtium aquaticum m. / Ad lacum citra Wallroth´s morphological notes were added on the front moleadivum ....“ [perhaps Mühlhausen /T.]. side of the label. Some doubts concerning identification One fertile plant (ca 130 cm) with roots, broken into two were written probably by L. Čelakovský on the back of this parts is on sheet C, the leaves are mounted separately. label. L a b e l : „Nasturtium aquaticum / α. integrifolium W. Seven flowering plants (12–23 cm) and one rhizome in schd. cr.“ The locality is not indicated. separate bags are mounted on sheet B. On sheet D there is one lower leafy part of a plant L a b e l : Text on label B was written by L. Čelakovský: (45 cm) and an upper branched part (27 cm) with fruits. „Monotropa hypopitys Lin. / var. glabra / M. hypophegea L a b e l : „N. aquaticum mihi / α. integrifolium“. Wallr. / Ex herb. Wallrothii“. Wallroth´s original label is absent. The specimen PR 11771 A is selected here as a lecto- The specimen PR 11750 A is selected here as a lectotype. type. At present such glabrous plants are evaluated as Mo- This species is identical with Rorippa amphibia (L.) notropa hypophegea WALLROTH or Monotropa hypopitys BESSER. L. var. glabra ROTH. Nasturtium brachystylum WALLROTH Montia arvensis WALLROTH Linnaea 14: 606-607, 1840. – PR 11772. Linnaea 14: 547, 1840. – PR 11764. Original diagnosis: Caulibus decumbentibus Text-fig.10. ramossisimis .... siliquis linearibus subcompressis pedicello Original diagnosis: Seminibus subrotundis a declinato duplo fere brevioribus, stigmate sessili capitato margine obtusato inde convexis in basin abrupte brevi- diametrum siliquae aequante coronatis. terque tenuatam sensim declinatis, tuberculis verruci- Locality: An Flussufern des südwestlichen Harzes. formibus hemisphaericis conformibus circa basin mole sen- Brief notes on the specimens: Two sim decrescentibus iisque punctiformibus nigrorem opacum branched, well developed upper parts of plants (ca 25 and mentientibus scabris, umbilico fusco. 30 cm long) with fruits and some flowers. Locality: Auf feuchten Aeckern des südlichen L a b e l : Two of Wallroth´s labels are present: 1. „Nas- Harzes stellenweise. turtium brachystylum m.“ / some notes on synonymy and Brief notes on the specimens: The morphology follow / „Ex agro northus.“ [Nordhausen] and original sheet contained two different groups of plants. 2. „Nasturtium brachystylum m. / hybrida forsan progedies These plants were revised and divided by specialist O. Nils- ex N. palustri / et N. sylvestre, saltem ex habitu / Ex agro son into two parts: 1. Three fertile bunches (2–4 cm long) heringensi“ [Heringen]. were designated in 1967 by Nilsson as Montia verna NECK- Due to the existence of two different localities on the ER and 2. One bunch of flowering and fertile plants (8–9 cm two different labels it is not appropriate to select the lecto- long) were designated as Montia fontana L. Two new type here. envelopes with a few seeds, made probably during the revi- This taxon is now considered as a hybrid Rorippa×bra- sion, were added. They were provided with O. Nilsson´s chystyla (WALLR.) LAWALRÉE (R. palustris×R. sylvetris). inscriptions: M. minor and M. fontana. L a b e l : „Montia arvensis m. / minor Gm.“ Locality Nasturtium riparium WALLROTH was not given. Sched. Crit.: 373-374, 1822. – PR 11774 A, B. The first mentioned specimen (the bunch of plants) des- Original diagnosis: Foliis variis, inferi- ignated as Montia fontana by Nilsson, is selected here as oribus pinnatis integrisve, caulinis superioribus ovato- a lectotype. oblongis inaequaliter serratis in basin cordato-amplexicau- lem attenuatis, siliquis ovato-oblongis utrinque attenuatis Nasturtium aquaticum WALLROTH compressis stylo duplo brevioribus. Sched. Crit.: 371-372, 1822. – PR 11771 A – D. Locality: Wallroth mentioned that the localities are Original diagnosis: Foliis variis, inferiori- the same as for the above mentioned species Nasturtium bus submersis multifidis, emersis serratis lyratisve, superi- aquaticum. oribus sessilibus conformibus integrisve, siliquis siliculae- Brief notes on the specimens: One formibus subglobosis styllo illis breviore coronatis pedicel- leafy plant (32 cm long) and two small branches (14 and lo multoties longiore suffultis, caulibus inflatus stoloniferus. 18 cm) with flowers and fruits are mounted on sheet A. Locality: In sulcis pratorum inque aquis limpidi- L a b e l : „Nasturtium riparium m. / N. anceps oribus stagnatibus circa Halam [Halle], Rossleben et alibi Wahlenb. et Reichb.“ The undated revision label is added: passim. „N. amphibium × silvestre / N. riparium Wallr.“ According

24 Text-fig. 10. Lectotype specimen Montia arvensis WALLROTH.

25 to the written note by E. Bayer, curator of the herbarium exists in the herbarium PR – it is a longitudinal section of during the period 1903 –1927, the revision label was writ- the flowering plant and also a fruiting one. The label: „Oro- ten by the German botanist Kurt Wein. banche coerulea / comosa“. On the second sheet B there are three leafy plants (23, 25 and 33 cm) with flowers and fruits. L a b e l : „Nasturtium riparium / W. / anceps Wahlb.“ Orobanche nudiflora WALLROTH On the right top of the label is written and crossed out in Sched. Crit.: 310, 1822. – PR 11787. unknown handwriting: „armoracioides Tausch?“ Original diagnosis: Scapo simplicissimo The specimen PR 11774 A is selected here as a lecto- inferne squamato, squamis bracteisque ovatis extus hirsuto, type. floribus ventricoso-subcampanulatis glaberrimis …. calyce The plant corresponds to Rorippa amhibia (L.) BESSER. profunde 4-fido, dentibus inaequalibus, superioribus maj- oribus e basi ovata lineari-setaceis. Nuphar systylum WALLROTH Locality: Ad radices Pruni spinosae auf dem Linnaea 14: 582, 1840. – PR 11754. Schachtberge prope Rossleben. Original diagnosis: Rhizomate cicatrisato Brief notes on the specimens: Four valido foliifero .... antheris linearibus elongatis, styli disco well-developed flowering plants (17–27 cm long) and the orig- orbiculato integerrimo profunde umbilicato stigmatibus inal envelope made from a letter with the remains of flowers. pulvinatis confluentibus adpressis breviter radiantibus L a b e l : 1. „Orob. nudiflora / W.“ (the label is mount- praeter ambitum integerrimum obliterato. ed directly on the left plant); 2. „Orobanche minor SUTT. / Locality: In einen langsam fliessenden Wiesen- Ex agro Lausannensi / Gag!“ and 3. „Orobanche nudiflora / graben, unmittelbar an der südöstlichen Gränze des prope Lausanne / Gag. / (S. O. minori titulo)“. The further Harzes. illegible revision label by an unknown author, written in German was added. B r i e f n o t e s o n t h e s p e c i m e n s : One young The very left plant with label (no. 1) is selected here as leaf, the blade with a short piece of petiole (13 cm) and one a lectotype. strongly damaged flower with peduncle 15 cm long. According to the revison label of Pusch from January L a b e l : 1. „Nuphar systylum Wallr! / Ex agro hal. / 2000 these plants correspond to the species Orobanche legit 1846!“ (the date was written in pencil probably later) minor SM. and 2. „Nuphar systylum W. / Thuring“. Some morpholog- ical features, also written in pencil, were added. The specimen PR 11754 is designated here as a neotype Orobanche rubens WALLROTH owing to the date on the label, the year 1846. Sched. Crit.: 307, 1822. – PR 11756. We consider this Wallroth’s species to be identical with Text-fig. 11. the species Nuphar luteum L. Original diagnosis: Scapo simplicissimo bracteisque colorato, floribus ventricosis in spicam Orobanche comosa WALLROTH oblongam congestis, staminibus intus hirsutiusculis, stylo parce glanduloso .... dentibus carinatis, superioribus obtu- Sched. Crit.: 314, 1822. – PR 11757. siusculis obsoletis tubi plicam attingentibus. Original diagnosis: Scapo simplicissimo, Locality: In collibus editioribus ad Biebram [Bad bracteis squamisque furfuraceo-hirsutis, floribus tubulosis Bibra NW of Naumburg] – Extra fines ad Auleben [Kyff- medio constrictis in spicam oblongam condensatis tribra- häuser] et Sondershusam [Sondershausen]. cteatis, stigmate bilobo (flavescente) exserto, calyce conna- to 5-fido .... Brief notes on the specimens: One well-developed plant (45 cm) with inflorescence and one Locality: Ad Genistae tinctoriae radices in colle upper part of rather small plant (12 cm). Wallroth´s original herbido der Steinklippe [near Wendelstein, SE of Ross- envelope with the remains of flowers and seeds is attached. leben]. L a b e l : „O. rubens m. / Ex agro Sonderhu / sano B r i e f n o t e s o n t h e s p e c i m e n : One upper [Sondershausen] 1820“. part of a fertile plant (23 cm) without basal part. The well-developed plant on the sheet PR 11756 is to be T w o l a b e l s : 1. „Orobanche comosa m. / Thg“. considered as a lectotype for this selection. The label is mounted directly on the plant and 2. According to J. Zázvorka, specialist on the genus „Orobanche comosa schd. cr. / Diask. 59. / Steinklippe pr. Orobanche, Wallroth´s species named as O. rubens corre- Wendels. [Wendelstein, SE of Rossleben] / 1808“. Seeds sponds to Orobanche lutea BAUMG. and flowers are attached in the original small envelope. Wallroth´ s original specimen of Orobanche rubens in The specimen PR 11757 is selected here as a lectotype. the herbarium of Martin Luther University in Halle-Witten- According to the January 2000 revision by German berg (No. HAL 90591) was already designated as type by botanist J. Pusch this specimen corresponds to Orobanche H. Uhlich. The text on Wallroth´s original label: „Oro- arenaria BORKH. banche rubens mihi Wallroth“ and on the revision label: „O. N o t e : A further specimen O. comosa from Wallroth´s lutea Baumg. / Det. Rev. Confirm.: Holger Uhlich / Typus! collection, originating probably from the Mediterranean, 17. 9. 2000“

26 Text-fig. 11. Lectotype specimen Orobanche rubens WALLROTH.

27 N o t e : Wallroth´s rich material of this species, origi- Locality: Prope Osterhausen [S of Eisleben] inter nally placed in the same herbarium cover together with the Brassicam Napum ad Heringen. Aug. chosen lectotype (PR 11756) and probably originating from B r i e f n o t e s o n t h e s p e c i m e n s : Three the same locality Sondershausen, is mounted on five further leafy plants (20, 30 and 32 cm) with a few capsules, two sheets. The original labels are absent and only small pieces broken upper parts of plants are on the sheet PR 2436. of paper with embossed stamps provided with the inscrip- L a b e l : Three labels are attached: 1. On the large tion: MUS. BOH. COLL. WALLROTH were added to original label the name „Papaver trilobum m.“ is written every sheet. and some remarks concerning distinguishing features fol- low. The locality is not mentioned; 2. Wallroth wrote by Orobanche sparsiflora WALLROTH pencil on the small label: „Papaver trilobum / Ann. bot.“ Sched. Crit.: 309, 1822. – PR 11785 A, B. and 3. „Vide: Dr. Wallroth´s Sendschreiben / an den Original diagnosis: Scapo simplici, squa- Apotheker H. Ernst Hampe / zu Blankenburg in Linnaea / mis bracteisque oblongo-lanceolatis concoloribus, floribus Bd. 14 (1840), pag. 81“ – this text was not written by Wall- ventricosis hirsutis remotis in spicam elongatam laxe dis- roth. positis. In Wallroth´s text only one variety was mentioned: Three leafy plants in flower (ca 25 cm) are on the sheet „β. pauciflora, scapo humili paucifloro“ (without any other PR 2437. notes). L a b e l : The original Wallroth´s label is absent, only the remark written probably by L. Čelakovský: „In der That Locality: In collibus et promontoriis graminosis ausgezeichnet. / Wird in DeCand. Prodromus I, p. 119 / als sylvaticis inter Bottendorf [near Rossleben] et Schweinsro- Pap. trilobum Spreng! fl. hal. / suppl. / angeführt.“ The da [Schönewerda, NW of Rossleben]. same text: „P. trilobum (Spreng! fl. hal. suppl.)“ was given Brief notes on the specimens: On by De Candolle (Prodr. 1824,1:119). sheet A there is one flowering plant, 25 cm long. The specimen on the sheet PR 2436 is selected here as L a b e l : The label is mounted directly on the plant: a lectotype. „Orob. caryophyllacea / v. sparsiflora / Thg.“ Wallroth All specimens represent Papaver rhoeas L. var. tri- wrote by pencil on the top right of the label: lobum (WALLR.). „O. sparsiflora“. The original envelope, made from a letter, containing one young fruit was added. Pinguicula gypsophila WALLROTH On sheet B there are two flowering plants (28 and Linnaea 14: 533-534, 1840. – PR 11762 A – E. 17 cm). Original diagnosis: Calycis (exigui) seg- T w o l a b e l s : 1. „Orobanche caryophyllacea / β. mentis patentissimis, superioribus trilobis, lobis ovatis bre- sparsiflora Diask. 38. / O. sparsiflora schd. cr. / Thuring.“ vissimis patentibus porrectis obtusiusculis ab excisione and 2. „sparsiflora“ (the name is written in pencil on the corollae superioris retractis .... stigmate emarginato (albo), label, mounted on the left plant). capsula (exigua) ovata mucronata ventricosa calyce duplo The specimen PR 11758 A is selected here as a lecto- breviore, seminibus reticulato-cellulosis .... type. Locality: Auf quelligen Moospläten zwischen According to Zázvorka this specimen corresponds to Gypselfelsenklüften des südlichen und südwestlichen Orobanche caryophyllacea SM. Harzes hier und da. N o t e : In Wallroth´s collection there are more than Brief notes on the specimens: On 40 sheets of the genus Orobanche, named as O. alba, sheet A there are seven flowering plants (5–9 cm long) with O. arenaria, O. caryophyllacea, O. flava, O. gracilis, O. 1–2 flowers. purpurea, O. ramosa, O. rubens. Most of them originated from Thuringia. The specimens from other parts of Ger- L a b e l : „Pinguicula gypsophila m. / Prope Stempe- many were not collected by Wallroth himself (e. g. by An- dam. Juli“ [Stempeda, NE of Nordhausen]. gerstein from Hiddensee, by Gottl. (?) near Bonn, by Braun Sheet B contains four complete fertile plants (8–14 cm), in Munich). In spite of the very rich material of Orobanche, three of them with four and one with three fruiting stems. further type specimens of species, described by Wallroth in L a b e l : „fructif. Pinguicula gypsophila“. Some mor- Schedulae criticae (O. apiculata, O. micrantha) and in phological features were added. Orobanches generis diaskene (O. cumana, O. reticulata, O. The inscription on Wallroth´s original envelope contain- variegata, O. violacea) have not yet been found in the ing seeds: „Pinguicula gypsophila m.“ herbarium PR. Two fertile plants (6–8cm), each with two fruiting stems are mounted on sheet C. Broken remains of two plants are placed separately in a bag. All material was originally pre- Papaver trilobum WALLROTH served in an enclosed envelope, made from a letter dated Annus Botanicus p. 149, 1815; Sched. Crit. Tab. I, 1822. 20 February 1830. On this envelope Wallroth added: „Pin- – PR 2436 and PR 2437. guicula gypsophila / Kohnstein“ [a hill NW of Nordhausen]. Original diagnosis: Capsulis obovatis cau- Eight small one-flowered plants (4 – 7 cm) are on sheet leque ramoso glabris, calycibus petiolisque ciliatis, foliis D without any label. trilobis obtusis integris glaucis basi cuneata in petiolum One fertile plant (ca 16 cm) with three fruiting stems is attenuatis. mounted on sheet E, also without any label. Two small orig-

28 inal envelopes with the remains of fruiting stems are pres- The specimen PR 10515 A is selected here as a lecto- ent. type. The plant on the sheet PR 11762 A is selected here as All plants correspond to Rhinanthus major L. a lectotype. To the sheet PR 11762 A and B, the revision label was Senecio fluviatilis WALLROTH attached: „Pinguicula vulgaris L. / rev. J. Schindler 1907“. All plants correspond to this species. Linnaea 14: 646, 1840. – PR 11803 A – E. Original diagnosis: Foliis glaberrimis mar- Potentilla hybrida WALLROTH gine tenuiter cartilagineo dentibus inaequalibus squarrosis Sched. Crit.: 247, 1822. – PR 11783 A, B. brevissime mucronatis serratis, infimis e basi longe protrac- Text-fig. 12. ta integerrima, involucro ovato foliolis filiformibus 4 illo duplo brevioribus succincto, ligulis 7–8 patentibus ungue Original diagnosis: Foliis radicalibus 3–4 pilosis, achaeniis glabris. natisve, foliolis obovato-oblongis apice infraque illum con- niventi-serratis supra glabriusculis subtus glaucescentibus Locality: An den Ufern langsam fliessender Flüsse sericeis, laciniis calycinis ovatis acuminatis petala obcorda- der südlichen Harzgränze. ta aequantibus. Brief notes on the specimens: The Locality: Inter virgulta rechts von der Steiger- plants, which were in all probability originally lying in one thaelschen Mittelmühle [Steigerthal, NE of Nordhausen], common herbarium cover, have been divided onto five extra agr. hal. fines. sheets. One upper part of a leafy plant with an inflorescence Brief notes on the specimens: On the first sheet A there are four plants (ca 20–25 cm) with rather big (57 cm long), broken into two parts is now on sheet A. On leaves, one broken leaf and one flower are mounted separately. a further four sheets there are also leafy plants with inflo- rescences which are more or less withered – one plant L a b e l : „Potentilla hybrida m. / Ex agro erford. [Er- (60 cm long) broken into two parts and one upper part furt?] – in hortis Bilzii mei culta et exhibita“. (24 cm) are on sheet B, one plant (56 cm) is on sheet C and On the second sheet B there are four small plants one plant (55 cm) broken into two parts is on sheet D. Two (5–15 cm), each of them with an individual label: 1. „Poten- upper parts of leafy stems (32 cm) with early buds and two tilla hybrida m. / attulit ex. fl. erfordana -?“; 2.“P. hybrida sterile shoots (33 and 16 cm) are on sheet E. mea“; 3. „Potentilla hybrida Schd. cr. / 247. / P. splendens Ram.“ and 4. „specimen carpelophorum!“ Wallroth´s origi- L a b e l : Wallroth´s only original label is mounted on nal envelope made from a letter and containing one seed is sheet A: „Senecio fluviatilis m. / Senecio sarracenicus Jacq. attached. Reichb. et / auct. nec. L. / Ad ripas Viprae [Wipper] inter The specimens on the sheet PR 11783 B are selected Wollers / leben et „Nohram“ [SW of Nordhausen]. here as a lectotype. At the turn of the year 1984–1985, J. Herborg had elect- J. Soják, specialist on the genus Potentilla, considered ed the specimen on the sheet PR 11803 A as a lectotype, the all these plants as the hybrid Potentilla alba×P. fragarias- further four sheets (B, C, D and E) he designated as isolec- trum with correct binomial name Potentilla×hybrida WALLR. totypes of Senecio fluviatilis WALLR. (Herborg 1987:184). According to Herborg´s monograph on Senecio Rhinanthus buccalis WALLROTH nemorensis-complex (1987: 187) these specimens corre- spond to the species Senecio sarracenicus L. Flora 25/II, Nro. 32: 504, 1842. – PR 10515 A, B, C. N o t e : A further two sheets with rather damaged Original diagnosis: Caule simplici superne plants of Senecio fluviatilis are preserved in Wallroth´s col- in ramos floriferos patentes diviso, foliis lanceolato-lin- lection. For both of them, only one label exists: „Senecio earibus, calyce e basi rotundata ad apicem usque aequaliter fluviatilis m. S. sarracenius auct. nec L. – Merse Julio ad inflato molliter pubente .... ripas Helmae?? lecta arenosa“. Locality was not mentioned. Brief notes on the specimens: The plants were originally lying together in one herbarium cover; they Senecio fontanus WALLROTH were later divided into three sheets and Wallroth´s original Linnaea 14: 647-648, 1840. – PR 11802 A, B, C. label was placed on sheet A. Five fertile plants are mounted on sheet A, three fertile plants (29–60 cm) and one damaged Original diagnosis: Foliis inferioribus inter- sterile plant (30 cm) on sheet B. mediisque hinc leviter decurrentibus, superioribus ses- silibus, corymbo polycephalo effuso, bracteis linearibus hir- L a b e l : „Rhinanthus buccalis m. / Inter segetes hye- sutiusculis, involucri foliola dorso hirsuta tricostata in males pr. Northusen [ Nordhausen] / juxta viae ad Neosbod- apicem parce sphacelatum sensim excurrentia subaequan- ium“ [probably Neustadt]. tibus, radio 5–8 ligulata .... On the third sheet C there are mounted two lower parts Locality has not been indicated. of plants (4–20 cm), six upper parts of leaves, fruits and seeds with inflorescences and a small envelope containing Brief notes on the specimens: On sheet the remains of plants. All these remains of plants were orig- A there is one leafy stem (55 cm) with inflorescence. inaly placed in one large envelope (enclosed at the back of T w o l a b e l s : l. „Senecio fontanus W.“ (the species the cover) with the inscription: „Rhinanthus buccalis m.“ name had been originally written as „amplexicaulis“, than it

29 Text-fig. 12. Lectotype specimen of Potentilla×hybrida WALLROTH.

30 was crossed out) / „α. pentaglossus / Ex nemore Lodersleb. Senecio salicifolius WALLROTH [Lodersleben, SW of Querfurt] / ad fontes. 1819“ and 2. Sched. Crit.: 478-481, 1822. – PR 11801. „Senecio amplexicaulis W. / α. pentaglossus / Ex nemore Original diagnosis: Foliis petiolatis oblon- Lodersleb.“ go-lanceolatis acutis cartilagineo-dentatis, omnibus con- One leafy stem (52 cm) broken in two parts with inflo- formibus, summis sensim angustioribus, floribus radian- rescence is on sheet B. tibus corymboso-umbellatis, pedunculorum bracteis lineari- T w o l a b e l s : 1. „Senecio fontanus W. / β. octo- filiformibus glabris. glossus / Ex palude Lodersleb. fl. / halensis. 1812“ and Locality: Ad Wiehe [S of Rossleben], proper ripas 2. „Senecio octoglossus DC / pr. VI. 354 / S. germanici var. der Unstruth [ Unstrut, river in Thuringia]. In agro thuring. δ. W. / Schd. cr.“ et hercyn. infer. abundat. The upper part of a leafy stem with inflorescence Brief notes on the specimens: Two le- (27 cm) is mounted on sheet C. afy stems with inflorescences (44 and 60 cm). T w o l a b e l s : 1. „Senecio fontanus W. / α. penta- T w o l a b e l s : 1. „Senecio salicifolius m. var. poly glossus m. / Ex Sudetis Silesiae.“; and 2. „Senecio Jacqui- (6–8)-glossa. / Kohnstein [a hill NW of Nordhausen] e reg- nianus REICH. / ..... Sudetae. Wagner.“ iore des / Kupferhamers. Aug.“ and 2. „var. angustif. / lusus Herborg (1987:104) considered the name Senecio 6–8 glossus / Kohnstein“. fontanus WALLROTH as an illegitimate one (…nom. illeg., The specimen PR 11801 is selected here as a lectotype. gibt u. a. S. jacquinianus RCHB. (1828) als synonym an). According to Herborg´s revision label (1985) these According to Herborg´s revision labels from 1985–1986 specimens correspond to the species Senecio ovatus the specimens placed on these three sheets 11802 A, B and (P. GAERTN., MEY. ET SCHERB.) WILLD. C correspond to the species Senecio germanicus WALLR. N o t e : A further large sample of Wallroth´s material of Senecio sarracenicus α. latifolius and β. longifolius (Lin- Senecio germanicus WALLROTH naea 14: 645) existed in the herbarium PR, which J. Her- Sched. Crit.: 476-478, 1822.– PR 11800 A, B, C. borg also identified as S. ovatus. Text-fig. 13. Original diagnosis: Caule foliisque subtus Solidago chrysantha WALLROTH margineque tenuiter pubescentibus .... pedunculis elongatis Linnaea 14: 643, 1840. – PR 10635 A – D. umbellato-corymbosis, bracteis linearibus, laciniis calycinis Original diagnosis: Caule striato racemoso- alternis latioribus costatis. paniculato .... capitulis ovatis multifloris demum patulis, Locality: In dumetis collinis inque nemoribus .... involucri foliolis linearibus omnibus conformibus, ligulis ad Schmon [ SW of Querfurt], Lodersleben [SW of Quer- (10) ovato-ellipticis utrinque obtusatis obsolete striatis .... furt], Biebra [Bad Bibra NW of Naumburg]: in nemoribus Locality: An den sonnigen Gypsvorbergen des des Alten-Stolberg´s [a hill E of Nordhausen] extra fines südwestlichen Harzes stellenweise häufig .... abundat. Brief notes on the specimens: The plants B r i e f n o t e s o n t h e s p e c i m e n s : Three were originaly placed in one herbarium cover with only one sheets of species Senecio germanicus are present. On sheet label; later they were divided into four sheets and the orig- inal label was mounted on sheet A. A there is an upper leafy stem with inflorescence (45 cm On sheet A there is one plant (64 cm) broken into two long). parts with rich, shortly branched inflorescences and the T h r e e o r i g i n a l l a b e l s : „1. Senecio germa- above mentioned label. nicus mihi / Senecio nemorensis Linn. ex Syn.“ / some dis- L a b e l : „Solidago chrysantha m. / Kohnstein am tinguishing features followed / „Ex dumetis apricis collis Kalkbruche“ [a hill NW of Nordhausen]. herbidi Bibraensis / M. Thr. 18“. The further two labels One lower part of a leafy stem (50 cm) with a leafy were stuck on the previous one: 2. „Senecio germanicus W. rosette and a flowering branch and one leafy branch (58 cm) / β. rigida / Biebra“ and 3. „Senecio nemorensis L. ? / caulis with dense inflorenscences are mounted on sheet B. flexuosis .... / S. ovatus.“ One plant (135 cm long) without the base, broken into three parts is on sheet C. On sheet B there are two upper parts of leafy stems One leafy rosette and one plant (56 cm) with leaves and (32 and 58 cm) with achenes. many inflorescences is placed on sheet D. L a b e l : „Senecio germanicus Wallr.“ The specimen on the sheet PR 10635 A is selected here Two leafy stems with inflorescences (52 and 55 cm) and as a lectotype. one lower part of a leafy stem (45 cm) are on sheet C. We evaluated these plants with dense inflorescences as a variation within the varibility of the species Solidago vir- L a b e l : „Senecio germanicus m. / α. pentaglossus“. gaurea L. The name of this variety has never been published neither in Schedulae Criticae, nor in Linnaea. J. Herborg on the revision label in 1985 selected the Thalictrum bauhinianum WALLROTH sheet 11800 A as a lectotype of Senecio germanicus WALL- Sched. Crit.: 264, 1822. – PR 11755. ROTH, it was later published (Herborg 1987:103). Original diagnosis: Caule stricto simplici All specimens correspond to this species. angulato-striato, foliorum inferiorum petiolatorum foliolis

31 Text-fig. 13. Lectotype specimen of Senecio germanicus WALLROTH.

32 lanceolato-linearibus, superiorum sessilium adpressorum B r i e f n o t e s o n t h e s p e c i m e n s : Four foliolis lineari-filiformibus convolutis .... leafy plants (30–58 cm long), mostly with basal parts of Wallroth divided this species into three infraspecific stems and fruitless inflorescences. Two original envelopes taxa: α. seselioides, β. cnidioides and γ. galioides. Only one with fruits are present, each with a different inscription: 1. specimen γ. galioides is present in the PR herbarium. „Bennstaedt 21“ and 2. „Carpella Thalictri montani asperi“. Brief notes on the specimens: Two T w o l a b e l s : 1.“Thalictrum minus L. / Ex vineis / unbranched leafy plants (28 and 20 cm) with inflorescences. Benstadiensis [Bennstedt] Aug. 20. / 21“ and 2. „Th. mon- tanum / α. virens mihi“. L a b e l : Two of Wallroth´s labels with nearly the same The specimen PR 2172 A is selected here as a lectotype text are attached: 1. „Th. bauhinianum / var. γ. / Th. galioides of Thalictrum montanum WALLROTH (nom. illeg.) var. virens Nstl.“ and 2. „Th. Bauh. γ. galioides / Th. galioides Nestl“. WALLROTH. The locality is not given – neither in his work Schedu- The plants correspond to the species Thalictrum minus L. lae criticae, nor on the labels. The plant on the sheet PR 11755 is selected here as γ ALLROTH a lectotype of Thalictrum bauhinianum WALLROTH. Thalictrum montanum . glandulosum W R. Hand (2001) considered this name as a synonym of (PR 2172 B) Thalictrum simplex L. subsp. galioides (DC. in LAM. et Original diagnosis: Foliolis subrotundis DC.) KORSH. leviter trifidis utrinque (magis minusve) glaucis, illis, vagi- nis, petiolis carpellisque glandulosis. Thalictrum collinum WALLROTH Locality: In collibus gypsaceis, calcareis ad Fran- Sched. Crit.: 259, 1822. – PR 2174. kenhausen [Bad Frankenhausen], Sondershausen, Botten- Original diagnosis: Caule basi folioso dorf [NW of Rossleben] inque jugo meridionali arenoso der internodiis approximatis intercepto, foliis utrinque viridibus Rothenburg, infra arcem, ubi maxima hujus Th. luxuries et glaberrimis, foliolis e basi ovata subcordatave subrotundis mutabilitas; in similibus locis der Steinklippe [SE of Rossleben]. obtuse-trifidis, floribus paniculatis cernuis, carpellis ovatis aequaliter costatis concoloribus stigmate recto coronatis. Brief notes on the specimens: Two parts of well-developed plants (32 and 34 cm) with many Locality: In collibus editioribus circa Rossleben, leaves and only a few fruits. Auleben [Kyffhäuser] et alibi. L a b e l : „Th. mont. γ. glandulos. / Ex collibus gypsa- B r i e f n o t e s o n t h e s p e c i m e n s : Three ceis herbidis / Frankohusani [Bad Frankenhausen] Juni 21“. well developed plants (43, 45 and 49 cm) with leaves espe- A further two, more or less scraps of paper with some mor- cially on the lower part of stem. Two plants have mostly phological remarks are attached. fruitless inflorescenses, the third one is with flowers. The specimen PR 2172 B is selected here as a lectotype L a b e l : Three labels with Wallroth´s original hand- of Thalictrum montanum WALLROTH (nom. illeg.) var. glan- writing are present: 1. „Th. collinum mihi“; on a further two dulosum WALLROTH. labels, some morphological features are mentioned. The locality is not given. Hand (2001:170) cosidered it as Thalictrum minus Specimen PR 2174 is selected here as a lectotype. L. subsp. saxatile DC. ex CESS. in CATTANEO. We consider this Wallroth´s specimen to be Thalictrum minus L. Thalictrum neglectum WALLROTH Linnaea 14: 590, 1840. – PR 2175. Thalictrum montanum WALLROTH Original diagnosis: Caule e basi vaginata Sched. Crit.: 255–259, 1822. – PR 2172 A, B. aphylla erecto striato, nodis foliiferis remotis intercepto Original diagnosis: Caule flexuoso supra foliisque epruinosis utrinque viridibus .... foliolis remotis basin vaginatam aphyllam folioso .... floribus umbellato- ovato-subrotundis, obtuse tri- s. quinque fidis, carpelis paniculatis cernuis, carpellis oblongis aequaliter costatis ovato-subrotundis, costa externa prominula in stigma costae utrinque attractis, illis nervis foliolorum vaginisque interiori gibbosae inclinatum excurrente. coeruleo-pruinosis. Walroth divided this species into three infraspecific Locality: Auf unfruchtbaren Aeckern und in den taxa: α. virens, β. roridum and γ. glandulosum, which dif- benachbarten Buschölzern des südöstlichen Harzes .... fered especially in colour and indumentum of the leaves. In B r i e f n o t e s o n t h e s p e c i m e n : One plant Wallroth´s collection two names are present: 1. α. virens (73 cm high) broken into two parts with leaves and fertile and 2. γ. glandulosum. inflorescence, only a few fruits are present. One original Hand (2001:148) considered Wallroth´s name Thalic- envelope containing fruit was added. trum montanum as an illegitimate one, hence we are not L a b e l : Two labels are attached. 1. „Th. neglectum m. selecting the lectotype from Wallroth´s material. / Th. elatum sched. cr. / nec Jacq. / Ex agro hal.“ and 2. „Th. elatum Murr. / α. mihi“. Thalictrum montanum α. virens WALLROTH The specimen PR 2175 is designated here as a lectotype. (PR 2172 A) We suppose that this plant corresponds to Thalictrum Locality: In vineis saxosis sterilissimis ad Benn- minus L. subsp. minus; Hand (2001:155) considered it as staedt [Bennstedt near Halle]. Thalictrum minus L. subsp. elatum (JACQ.) STOJ. ET STEF.

33 Tormentilla parviflora WALLROTH tum oblongum e basi rotundata tenuatum laevigatum pedi- Linnaea 14: 580, 1840. – PR 11786 A, B. cello longiorem constituentibus .... Locality: Auf salzhaltigen Triften der südlichen Original diagnosis: Caulibus adscenden- Salinen .... tibus ramosis, foliis radicalibus longe-, caulinis brevius petiolatis s. subsessilibus alternis oppositisve ternatis …. B r i e f n o t e s o n t h e s p e c i m e n s : Eight pedunculis filiformibus elongatis, phyllis calycinis subcon- fertile stems (7–18 cm) and one broken bottom part of formibus ovatis acutiusculis, petalis obovatis calycem a stem, three sterile leaves and a small rosette of leaves. aequantibus .... L a b e l : Two labels are present: 1. „Triglochin salina Locality: In Birkenwäldern des südwestlichen Wallroth 1816 / Triglochin intermedia / Wallr. in lit. 1813 / Harzes stellenweise. Ad salinas Aerenser Thrg. [probably Artern, Kyffhäuser] Wallroth divided this species into two varieties: α. ses- 1812“ and 2. „Triglochin salina intermedia Wallr.“ (the for- silifolia and β. petiolata. Both of them are deposited in the mer designation intermedia was crossed out) / „salina 1816 herbarium PR. / a Triglochine maritima, affinis ubique differt“. A further seven different features follow. The sheet PR 11767 is selected here as a lectotype. Tormentilla parviflora var. α. sessilifolia WALLROTH The plants correspond to the species Triglochin mariti- (PR 11786 A) mum L. Original diagnosis: Caulibus decumbentibus tenuibus, foliis sessilibus, omnibus alternis, stipulis ovatis. Valeriana collina WALLROTH Brief notes on the specimens: Eight Linnaea 14: 537-538, 1840. – PR 10118 A – F, PR 10122. tufts with leaves and fruiting creeping stems (up to 14 cm long) Original diagnosis: …. foliis multi(10–12)- with various degrees of damage, bearing only a few fruits. jugis, omnibus subconformibus, foliolis approximatis lin- L a b e l : „Tormentilla micrantha m.“ – above this eari-lanceolatis subintegerrimis obtusiusculis basi juxta name is added: „parviflora α. sessilifolia“. Further some rhachin foliaceo-canaliculatam decurrentibus .... synonymical remarks follow. Down at the right bottom: „Ex Locality has not been mentioned. agro heringensi“ [Heringen]. Wallroth divided this species into three infraspecific Specimen PR 11786 A is selected here as a lectotype of taxa α. simplex, β. stolonifera, and γ. monstr. vesicarium. Tormentilla parviflora var. α. sessilifolia WALROTH. He further divided the variety V. collina β. stolonifera into The plants correspond with the species Potentilla erecta a. leiocarpa and b. eriocarpa, both of them are preserved in (L.) RÄUSCHEL (Soják 1995). the herbarium PR. Wallroth´s original remarks published in Linnaea did not correspond accurately with his labels. Tormentilla parviflora var. β. petiolata WALLROTH Brief notes on the specimens: Two (PR 11786 B) well-developed flowering plants (62 and 57 cm), with Original diagnosis: caulibus erectis 1–2 6 cauline leaves and 22–24 linear entire leaflets are on the pedalibus, foliis breviter petiolatis, intermediis oppositis, sheet PR 10118 A. foliolis stipulisque oblongis duplo majoribus. L a b e l : „Valeriana sylvestris Dodon.“ The designa- Brief notes on the specimens: Two tion „sylvestris“ was crossed out and obviously later Wall- creeping leafy stems (above 45 cm long) with long petioles roth rewrote it as: „Valeriana collina / var. stolonifera m“. (3–4 cm), bearing a head of achenes. Below this inscription is written: „Valeriana collina m. / γ. eriocarpa“ and four synonyms identical with published text L a b e l : Above the name „Tormentilla micrantha m. / were added. The locality follows: „Possen“ [probably var. oppositifolia“ there is written: „parviflora β. petiolata Possenheim near Naumburg]. W. / stipularia integritate varians / In nemusculo hinter / u. Five further sheets (PR 10118 B – F) without any labels, über der Hirschteichen / pr. Salsam“ [stream Salza on evidently originated from the same locality. All plants are W outskirts of Nordhausen]. well collected and preserved. The small label, written on The specimen PR 11786 B is selected here as a lectotype a typewriter with the text: „Herbarium Wallroth / sine sche- of Tormentilla parviflora WALLROTH. da / verisimiliter isotypus/ Valerianae collinae/ eriocarpae“ Specialist on this genus, J. Soják, considered the speci- was stuck on all these sheets. The label was probably added mens on sheet B to be Potentilla erecta (L.) RÄUSCHEL var. when the original material was divided and mounted in the dacica (BORBÁS EX ZIMMETER). second half of the 20th century. One plant (ca 90 cm) with an unbranched stem, a few Triglochin salina WALLROTH leaves and inflorescence and four upper parts of stems Linnaea 14: 567, 1840. – PR 11767. (24–30 cm high) with inflorescences are on the sheet PR Original diagnosis: Rhizomate transversali 10122. The middle cauline leaves with ca 15 entire linear tunicis fibrosis obducto, scapo basi adscendente foliorum leaflets are sporadically dentate. vaginis ovato-rotundatis involuto bulboso superne laxe bre- L a b e l : „Valeriana collina m. / α. leiocarpa / Kohn- viterque spicato, rhachi teretiuscula, capsulis 6 oblongo-lin- stein“ [a hill NW of Nordhausen]. earibus triquetris dorso costato planis acute marginatis in The specimen PR 10118 A is selected here as a lectotype apicem stigmate reclinato coronatum sensim tenuatis, fruc- of species Valeriana collina WALLROTH.

34 According to botanists J. Holub et J. Kirschner (1997) According to Holub et Kirschner (1997) the name Vale- the name V. collina is illegitimate and they considered this riana multiceps is a synonym for Valeriana officinalis L. taxon to be Valeriana stolonifera CZERN. subsp. angustifo- lia SOÓ. Valeriana procurrens WALLROTH Linnaea 14: 540-541, 1840. – PR 10123 A, B. Valeriana multiceps WALLROTH Original diagnosis: Radice praemorsa fibras Linnaea 14: 539-540, 1840. – PR 10119 A, B, C and PR 10121. prolixas stolonesque plurimos longe procurrentes ex artic- Original diagnosis: Radice praemorsa fib- ulis radicantes in comam foliosam desinentes undique spar- rosa multicipite stolonibus foliosis destituta, caule stricto gente,caule elato, foliis pauci-(4–6)jugis, foliolis radicalium simplici elato supra basin profunde sulcato, foliis pa- innovationumque ovato-oblongis .... uci–(3–7)jugis .... impari obovato-oblongo apice profunde Locality: In schattigen Hainen und auf feuchten tridentato .... Abhängen des südwestlichen H. [obviously Harzes] stellen- Wallroth divided this species into two infraspecific taxa: weise. α. dentata (fol. a basi inde dentatis) and β. incisa (fol. pro- Brief notes on the specimens: One funde acuteque incisi-dentatis), both of them are represent- ed in Wallroth´s material. plant (110 cm long) was divided and mounted onto two sheets A and B with one original label placed on sheet A. The upper part of the stem with inflorescence and a few Valeriana multiceps α. dentata WALLROTH cauline leaves and middle part with two pairs of leaves are (PR 10121) on sheet A. On sheet B there is the lower part of the same Locality: An fruchtbaren Bergabhängen des plant with one well developed pair of leaves. südlichen Harzes .... L a b e l : Valeriana procurrens m. / Elfelder Thal“ Brief notes on the specimen: One [probably near Ilfeld, N of Nordhausen]. well-developed plant (105 cm high), divided into three The specimen PR 10123 A is selected here as a lecto- parts. Basal part of stem with three pairs of leaves, middle type. part of stem with two pairs of leaves and an upper part with Holub and Kirschner (1997) evaluated this taxon as an inflorescence in full flower. Leaves with 5–7 pairs of Valeriana excelsa POIRET subsp. procurrens (WALLR.) leaflets. HOLUB. L a b e l : „Valeriana multiceps m. / var. dentata / Kohn- stein [ a hill NW of Nordhausen] Aug.“ Vicia onusta WALLROTH The specimen PR 10121 is selected here as a lectotype of the species Valeriana multiceps WALLROTH. Linnaea 14: 623, 1840. – PR 11770. Original diagnosis or description was not given. Wall- Valeriana multiceps β. incisa WALLROTH roth mentioned only a comparison with related species: .... (PR 10119 A, B, C) als eine schmallblätrige Form der Vicia cracca …. Locality: Am südlichen Harzen nur stellenweise Locality is not given. häufig .... B r i e f n o t e s o n t h e s p e c i m e n s : Three B r i e f n o t e s o n t h e s p e c i m e n : One leafy original sheets of this variety are present. One plant plant (57 cm), with inflorescences. (125 cm high) is divided onto two herbarium papers A and B, and the original label is mounted on sheet A L a b e l : „onusta Wallr.“ (this is written on the very top Two parts of a stem with three pairs of cauline leaves, above the name tenuifolia) / „Vicia tenuifolia Roth“ (the 6–7 pairs of deeply dentate leaflets and the upper part of a name Roth is crossed out and changed for DC. nec). The stem with an inflorescence in full flower are on sheet A. On species name „onusta Wallr.“ had obviously been written the second sheet B there is the basal part of stem with one above later. The locality is not mentioned. incomplete leaf, three leaves are mounted separately. The specimen PR 11770 is selected here as a lectotype. L a b e l : „Valeriana multiceps m.“ (the former desig- This species is identical with Vicia tenuifolia ROTH with narrow leaflets. nations „sylvestris DOD.“ and „paludosa“ were crossed out) / „var. β. incisa m. / Ad nemoris .... infra Petersdorf“ [ near Nordhausen]. All these changes were written by Wallroth. On the third sheet C there is one complete plant divided Acknowledgements into three parts – the remains of large basal leaves with roots and bottom part of stem with one leaf, middle part of This work was funded by the Grant Agency of the cauline stem with two pairs of leaves and upper part of stem Czech Republic (grant No. 206/07/1555). with an inflorescence. We would like to express very special thanks to Dr. L a b e l : „Valeriana multiceps m. / var. incisa / Peters- Z. Pouzar for his valuable comments and help with prepa- dorf“. ration of this paper. Without his assistance the work could The specimen PR 10119 A is selected here as a lectotype have never been finished. We also thank RNDr. J. Štěpánek of the variety Valeriana multiceps WALLROTH var. incisa for very thorough revision and Mgr. O. Šída for making the WALLROTH. photos.

35 References

Anonymus (1858): Wallroth´s Sammlungen. – Bot. Zeitung Soják J. (1995): Potentilla. – In: Slavík B. et al. Květena 16/30: 224. České republiky 4. – Academia, Praha, p. 312. Anonymus (1859): Sammlungen. – Oesterr. Bot. Z. 9/4:141. Sprengel K. (1806): Florae Halensis tentamen novum. – Barthel K.J. et Pusch J. (2005): Die Botaniker des Halae saxonum. Kyffhäusergebietes. – Jena. Stafleu F. A. et Cowan R. S. (1988): Taxonomic literature, Hampe E. (1836): Prodromus Florae Hercynicae. – Halle. 7: 43–45, Utrecht. Hand R. (2001): Revision der in Europe Vorkommenden Štěpánek J. (1992): Arabis. – In: Slavík B. et al: Květena Arten von Thalictrum subsectio Thalictrum (Ranuncu- České republiky 3. – Academia, Praha, p. 134. laceae). – Bot. Naturschutz Hessen, 9: 1–358, Frankfurt Rauschert S. (1965): Zur Flora Thüringens und der nor- a. M. dostlich angrenzenden Gebiete. 2. – Wiss. Z. Univ. Herborg J. (1987): Die Variabilität und Sippenabgrenzung Halle math. nat. R.,14: 495–496. in der Senecio nemorensis-Gruppe (Compositae) im Wallroth F. G. (1815): Annus botanicus, sive supplemen- europäischen Teilareal. – Dissert. Bot., 107, Berlin- tum tertium ad Curtii Sprengelii Floram Halensem. – Stuttgart. Halae. Holub J. et Kirschner J. (1997): Valeriana. – In: Slavík B. et Wallroth F. G. (1822): Schedulae criticae de plantis florae al.: Květena České republiky 5. – Academia, Praha, Halensis selectis. – Halae. p. 516–527. Wallroth F. G.(1824): Ad Hoppeum, virum in re herbaria Kellner K. (1979): Die floristische Erforschung der Südhar- peritissimum, de Malvis germanicis epistola Frederici zlandsachaft um Nordhausen. – Beitr. Heimatk. Stadt Wallrothii. – Syll. Pl. Nov. 1: 140–142, Ratisbonae. Kreis Nordhausen, Heft 4: 50–61. Wallroth F. G. (1825): Orobanches Generis Diaskene. – Kison H. U. et Sacher P. (1995): Ernst Hampe Flora Her- Francof. cynica. – Quedlinburg Wallroth F. G. (1828): Rosae plantarum generis historia Kubát K. et al. (2002): Klíč ke květeně České republiky. succincta. – Nordhusae. – Academia, Praha. Wallroth F. G. (1840): Scholion zu Hampe´s Prodromus Kützing F. T. (1857): Dr. Carl Friedrich Wilhelm Wallroth. Flora Hercyniae. – Linnaea 14: 1–158, 529–704. – Bonplandia 5/9: 147–148. Wallroth F. W. (1842): Zur Naturgeschichte des Rhinanthus Osswald L. (1896): Aus dem Leben Wallroth´s. – Mitt. Alectorolophus der deutschen Floristen. – Flora 25/II, Thüring. Bot. Ver. N. F. 9: 14–27, Weimar. Nro. 32: 497–506. Schroeter W. (1991): Karl Fridrich Wilhelm Wallroth. Zur Wallroth F. W. (1842–1844): Beiträge zur Botanik. Eine Wiederkehr seines 200. Geburtstages. – Beitr. zur Sammlung monographischer Abhandlungen über Heimatkunde aus Stadt und Kreis Nordhausen, Heft 16: besonders schwierige Gewächs-Gattungen der Flora 89 –112. Teuschlands. – Leipzig.

36 The list of Wallroth´s original material presented in this paper

name page name page

Aconitum bernhardianum 10 Montia arvensis 24 Adonis anomala 10 Nasturtium aquaticum 24 Adonis maculata 10 Nasturtium brachystylum 24 Alchemilla glaucescens 10 Nasturtium riparium 24 Alchemilla glaucescens var. β. alpestris 10 Nuphar systylum 26 Alectorolophus parviflorus 12 Orobanche comosa 26 Arabis brassicaeformis 12 Orobanche nudiflora 26 Artemisia mertensiana 12 Orobanche rubens 26 Atriplex ruderalis 12 Orobanche sparsiflora 28 Camelina sylvestris 13 Papaver trilobum 28 Campanula thaliana 13 Pinguicula gypsophila 28 Centaurea phaeocephala 13 Potentilla hybrida 29 Chenopodium microspermum 13 Rhinanthus buccalis 29 Dianthus schizopetalus 13 Senecio fluviatilis 29 Erigeron corymbosus 15 Senecio fontanus 29 Erysimum cheiriflorum 15 Senecio germanicus 31 Erysimum schaererianum 15 Senecio salicifolius 31 Fumana pinifolia 15 Solidago chrysantha 31 Galium agreste 15 Thalictrum bauhinianum 31 Galium gracile 16 Thalictrum collinum 33 Halimus pedunculatus 16 Thalictrum montanum 33 Heracleum ursinum 16 Thalictrum montanum γ. glandulosum 33 Hieracium lactucella 16 Thalictrum neglectum 33 Hieracium rothianum 18 Tormentilla parviflora 34 Lamium rubrum 18 Tormentilla parviflora var. α. sessilifolia 34 Malva adulterina 18 Triglochin salina 34 Malva equina 18 Valeriana collina 34 Malva neglecta 18 Valeriana multiceps 35 Medicago stipularis 22 Valeriana multiceps β. incisa 35 Melilotus arvensis 22 Valeriana procurrens 35 Melilotus tenella 22 Vicia onusta 35 Monotropa hypophegea 22

37