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Home , Acanthacris, Austracris, Catantopinae, Fenestra (grasshopper), Fer (insect), Gastrimargus

Evolutionary Systematics 2 2018, 21–30 | DOI 10.3897/evolsyst.2.22127

An annotated catalogue of the types of short-horned (, ) housed in the Zoological Museum Hamburg (ZMH)

Lara-Sophie Dey1, Martin Husemann2

1 General Zoology, Martin-Luther University Halle-Wittenberg, Hoher Weg 8, 06120 Halle (Saale), Germany 2 Centrum für Naturkunde, University of Hamburg, Martin-Luther-King Platz 3, 20146 Hamburg, Germany http://zoobank.org/9C76A3B3-5F8C-4B6A-8A32-AD2FC3A07E24

Corresponding author: Martin Husemann ([email protected])

Abstract Received 28 November 2017 Accepted 19 January 2018 Zoological collections remain the main archive of life on earth, especially in Published 19 March 2018 times of mass extinctions. The most important of these collections are the type specimens, which represent the original reference for a species. While a full digitization Academic editor: of most museum collections currently remains utopic, lists of types are an essential re- Danilo Harms source for researchers to perform taxonomic revisions. Here, we provide an updated type catalogue of the short-horned grasshoppers (Caelifera) housed in the Zoological Muse- um of Hamburg (ZMH), now part of the Center for Natural History Hamburg (CeNak). Key Words This collection comprises 209 type specimens belonging to 128 species: 65 holotypes, three lectotypes, one syntype and a large number of paratypes. We also list topotypes biodiversity of two species for reasons of completeness but note that they have no formal status. CeNak Overall, types of 13 species were added to the collection since the last catalogues in Godeffroy 1968 and 1977 and several corrections to the last catalogues were made. We provide a grasshoppers full bibliography for these species, aiming to provide a comprehensive taxonomic guide museum to this collection. type catalogue ZMH

Introduction the DORSA website (https://www.dorsa.de/) and have also been implemented in the Orthoptera Species File With 209 type specimens of the suborder Caelifera and (Cigliano et al. 2017). However, since the last update of more than 100 species of (Weidner 1966, new the type catalogue 40 years have passed, rendering a new type catalogue in preparation) the collection of the Zoo- list necessary. Hence, we present a new and updated list logical Museum Hamburg (ZMH), now part of the Center of type specimens of the suborder Caelifera housed in the of Natural History Hamburg (CeNak), is among the most ZMH with updated nomenclature, and a bibliography of important collections in Germany. The collection is cur- the original descriptions of all species. rently not digitally available, but the former curator Her- The ZMH Orthoptera collection is of special value as bert Weidner had presented a first list including types of many specialists have worked in the museum and depos- 116 species and a second publication with four additions ited material: the museum houses the type material of (Weidner and Wagner 1968, Weidner 1977; some of these many species collected by Hans Fruhstorfer, who main- were missing in the collection, when Weidner wrote his ly worked in Asia and South America (Baur and Coray catalogue; further, the status of some species was unclear 2004). Further, Yngve Sjöstedt, a Swedish biologist, who back then). As part of the DORSA project (Ingrisch et collected material during expeditions to Africa (Sjöstedt al. 2004), the types of the collection were examined and 1925), deposited many types in Hamburg. Carl Friedrich photographed in 2003; these pictures are available from Brunner von Wattenwyl, one of the first specialists of the

Copyright Lara-Sophie Dey, Martin Husemann. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 22 Lara-Sophie Dey & Martin Husemann: An annotated catalogue of the types of short-horned grasshoppers... *7 *10 *23 *28 *22 *22 *22 *22 *21 *21 *26 *28 *24 *32 *22 *16 *22 *22 *27 *27 *22 *36 *36 *36 *36 *16 *27 *27 Ref. Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y N Weidner Sanaa ? Stat. 109 Paraguay Location Turkey, Elazig Turkey, Indonesia, Flores Perak and Pahang Australia, Edel Land Arabia, Yemen, Sanaa Arabia, Yemen, Arabia, Yemen, Arabia, Yemen, Sanaa Arabia, Yemen, Sanaa Arabia, Yemen, Uruguay, Monte Video Uruguay, India, Hinterland of Assam India, Hinterland of Australia, Kimberley district Australia, Queensland, Tolga Australia, Queensland, Bowen Australia, Queens Land, Gayndah Sulawesi, Samanga, South Celebes Sulawesi, Samanga, South Celebes Sulawesi, Samanga, South Celebes Australia, Queensland, Peak-Downs Australia, Queensland, Peak-Downs Sulawesi, Patunuang, South Celebes Australia, Queensland, Rockhampton South-West Australia, north of Subiaco, South-West Angola, Bengo Province, Piri (Dembos) Angola, Bengo Province, Piri (Dembos) Indonesia, Camp Jor, watershed between Indonesia, Camp Jor, Australia, New South Wales, New Castle Wales, Australia, New South New Castle Wales, Australia, New South 8 07.09.1905 30.04.1921 30.04.1921 24.09.1952 24.09.1952 Stockholm, Mjöberg, Stockholm, Mjöberg, Boltze, 1902 Collector, year year Collector, Stal, No. 15844 C. Rathjens, 1931 Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Godeffroy J. Bohls, 10.01.1895 C. Rathjens, Aug. 1931 C. Rathjens, Aug. 1931 C. Rathjens, C. Rathjens, 25.05.1931 H. Fruhstorfer, Jan. 1896 H. Fruhstorfer, A. Demirsoy, 15.07.1971 A. Demirsoy, H. Fruhstorfer, Nov. 1895 Nov. H. Fruhstorfer, 1895 Nov. H. Fruhstorfer, 1895 Nov. H. Fruhstorfer, H. Fruhstorfer, 06.03.189 H. Fruhstorfer, Australien, alte Sammlung Hamb. Angola Expedition, Hamb. Angola Expedition, Alb. Grubauer, 30.12.1902 Alb. Grubauer, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Museum Godeffroy, No. 80011 Museum Godeffroy, Hamb. S. W. Austr. Expedition, Austr. W. Hamb. S. Museum Stockholm, 01.09.1922 Hamb. S. W. Austr. Expedition 1905 Austr. W. Hamb. S. Stal, 1861 Kevan, 1955 Kevan, 1955 Walker, 1870 Walker, 1870 Walker, Author, year Author, Uvarov, 1936 Uvarov, 1936 Uvarov, Uvarov, 1936 Uvarov, Uvarov, 1936 Uvarov, Sjöstedt, 1928 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1920 Sjöstedt, 1920 Sjöstedt, 1932 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Willemse, 1931 Willemse, Willemse, 1936 Willemse, 1936 Willemse, 1936 Willemse, 1936 Willemse, Demirsoy, 1979 Demirsoy, Giglio Tos, 1895 Giglio Tos, 2♀ 1♀ 1♂ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♂ 1♂ 1♀ 1♀ 1♀ 1♂ 1♂ 2♀ 1♀ 1♀ 1♂ 1♂ 4♂ 1♀ 1♂ 1♂ Sex 2♂ 1♂ 1♀ 1♂ 2♂ 5♀ 1 1 1 1 1 1 1 1 1 1 1 4 2 2 1 7 1 1 1 2 1 1 1 1 4 1 1 1 Count P P P P P P P P P P P P P H H H H H H H H H H H H H H H Type ------Acrida exaltata Urnisa rugosa Clepsydria historio Desertaria cinnam Apotropis tricarinata Apotropis tricarinata Apotropis Cedarinia vermiculata Cedarinia vermiculata Current Current Heteracris punctata Heteracris punctata SYN SYN Asphingoderus elazigi SYN SYN SYN SYN Acanthacris ruficornis africanus Gastrimargus SYN SYN SYN SYN bohlsii africanus orientalis scabra variegata fuliginosa cinnamomea cliens includens includens marginata amiculi scabra costata affinis affinis punctata punctata elazigi uvarovites guttulosa talawensis affinis ruficornis yemenita curta tuberculatus gracilis gracilis gracilis incompleta affinis Species The table shows all type specimens currently housed in the collection of the ZMH including the original and species name, and the currently valid name (SYN = the species is now name and the currently valid genus and species name, the original of the ZMH including housed in the collection currently type specimens shows all The table Fenestra Gastrimargus Cedarinia Cedarinia Cedarinia Clepsydria Desertaria Dichroplus Exarna Exarna Bibracte Caparra Caperrala Apotropis Apotropis Asmara Asmara Asphingoderus Austracris Apalacris Apalacris Apalacris Apalacris Apoboleus Apoboleus Acanthacris Acrida Adreppus Genus 22 21 14 15 16 17 18 19 20 10 11 12 13 7 8 9 4 5 6 No. 1 2 3 Table 1. Table = lectotype S = syntype, L in last column), reference as additional (author = paralectotype PL = paratype, P (H = holotype, status is presented The type in this line). stated synonym of the species The number of present type specimens is given in the column code). by the taxonomic not regulated for completeness, = topotype (recorded T reference in last column), (author as additional already had type the if displays ‘Weidner’, column The provided. are location and year collection collector, the as as well year, publication and Author (‘sex’). gender their by followed ‘count’, of each species; description (* Ref.) to the original we provide the reference Finally, = yes, N newly reported). Y 1977; Weidner 1968; Wagner & (Weidner been recorded in previous catalogues in case of lecto-, paralecto- or topotypes, author and year are presented (# Ref. number).

evolsyst.pensoft.net Evolutionary Systematics 2 2018, 21–30 23 #2 *2 *22 *34 *19 *21 *21 *22 *21 *22 *20 *10 *37 *21 *22 *23 *31 *23 *23 *23 *23 *23 *23 *23 *23 *23 *23 *27 *22 *22 *22 *22 *22 *37 Ref. Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y N Weidner Java 1200m Malke) Paraguay Location East India Massai land 1900-2100m bei Mundame Java, Kangean Togo, Misahöhe Togo, Indonesia, Flores Australia, Sidney Arabia, Yemen, Sanaa Arabia, Yemen, Salomon Islands, Buka Transvaal, Johannesburg Transvaal, ustralia, Kimberley District Australia, Queensland, Tolga Australia, Kimberley District A Australia, Kimberley District Australia, Victoria, Alexandra Australia, Victoria, Australia, Queensland, Bowen Australia, Queensland, Bowen Kamerum, Sikumba near Maputa Kamerum, Sikumba near Maputa Australia, Perth, north of Subiaco Australia, Queens Land, Gayndah Indonesia, Lombok, Sapit, 2000m Iran, Belutschistan, Iranshar, 800m Iran, Belutschistan, Iranshar, Jugoslavia (=Montenegro), (Sbrija- Australia, Queensland, Peak-Downs Australia, Queensland, Peak-Downs East Africa, Abessinien, Wonda East Africa, Abessinien, (near Wonda Australia, Queensland, Rockhampton Australia, Queensland, Rockhampton Sumatra, Westcoast, Tandjunggadang, Tandjunggadang, Sumatra, Westcoast, South Africa, Johannesburg, Transvaal Kosmet, Poc), Hajla-Cafe e Mustafes), Kamerum, Mukonjjefarm am Mungofluß 09 ? ? ? ? ? ? ? 1893 1928 30.04.1921 21.04.1954 30.04.1921 Boltze, 1902 09-16.05.1905 u Will, 04.10.1902 u Will, Collector, year year Collector, Le Moult, 1936 Le Moult, 1936 Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Godeffroy H. Rolle, 26.10.1903 Willemse, 04.08.1965 Willemse, Fra E. Jacobson, Feb. 1920 C. v. Erlangen, 7.12.1900 C. v. H. Fruhstorfer, 05.06.1899 H. Fruhstorfer, Dr. C. Rathjens, Aug. 1931 C. Rathjens, Dr. Richter and Schäuffele, 11- Richter and Schäuffele, Südsee-Exp. Wolf, 29.08.19 Südsee-Exp. Wolf, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Godeffroy, No. 15880 Museum Godeffroy, Hamb. S. W. Austr. Expedition, Austr. W. Hamb. S. Museum Stockholm, 30.04.1921 H. Fruhstorfer, May to June 1896 H. Fruhstorfer, Museum Stockholm, Mjöberg, Jan. Museum Stockholm, Mjöberg, Author, year Author, Uvarov, 1936 Uvarov, Ramme, 1931 Ramme, 1941 Sjöstedt, 1920 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1920 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1921 Sjöstedt, 1920 Sjöstedt, 1920 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1928 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Willemse, 1933 Willemse, Willemse, 1965 Willemse, Willemse, 1930 Willemse, Willemse, 1965 Willemse, Giglio-Tos, 1895 Giglio-Tos, Bey-Bienko, 1960 1♀ 2♂ 1♂ 1♂ 1♀ 1♂ 1♂ 1♀ 1♀ 1♀ 1♂ 1♂ 1♂ 1♂ 1♀ 2♀ 1♀ 1♀ 1♂ 1♀ 1♂ 1♀ 2♀ 1♂ 1♂ 1♀ 1♀ 1♂ 1♂ Sex 1♂ 1♀ 1♂ 1♀ 2♂ 1♀ 1♂ 1♀ 1 2 2 1 1 2 1 1 1 1 1 1 1 1 1 1 1 2 1 1 1 1 1 1 2 1 3 1 2 1 1 1 1 Count P P P P P P P P P P P P P P P P P P P T H H H H H H H H H H H H H Type ------ vocans Desertaria lepida Tauchira rufotibialis Tauchira Goniaeoidea modesta Goniaea opomaloides Gastrimargus Stenopola bohlsii Goniaea opomaloides Current taxonomy Current Gastrimargus crassicollis Gastrimargus Gastrimargus lombokensis Gastrimargus Gastrimargus crassicollis Gastrimargus SYN SYN Phlaeobella lombokensis SYN SYN SYN SYN SYN Gastrimargus verticalis Gastrimargus SYN Gastrimargus africanus Gastrimargus africanus Gastrimargus Gastrimargus africanus Gastrimargus africanus Gastrimargus africanus Gastrimargus africanus Gastrimargus Gastrimargus determinatus arabicus Gastrimargus SYN SYN SYN SYN SYN SYN SYN SYN SYN SYN SYN nanum lombokensis relicta brevialata picta coeruleipes africanus orientalis zebrata africanus v. pulchra rubripes salomona bohlsii elegans elegantula ampla affinis Species zebrata africanus v. zebrata africanus v. zebrata africanus v. zebrata africanus v. abessina v. brevipes clepsydrae floresensis musicus v. kimberleyensis transvaalensis volkensi arabicus acuta glaucipes macronotum vinaceipennis jucunda insularis respondens grandis turcmena Phlaeoba Podismopsis Perbellia Perbellia Loiteria Opiptacris Oxyblepta Oxytauchira Paratraulia Peakesia Peakesia Gastrimargus Gastrimargus Lobopteria Pardillana Genus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Gastrimargus Goniaea Goniaea Goniaea Goniaea Goniaeoidea Heteropternis Hyalorrhipis 49 50 47 48 46 38 39 40 41 42 44 45 No. 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 43

evolsyst.pensoft.net 24 Lara-Sophie Dey & Martin Husemann: An annotated catalogue of the types of short-horned grasshoppers... #1 #1 *9 *9 *22 *22 *24 *29 *22 *22 *24 *22 *21 *21 *21 *38 *38 *38 *22 *22 *27 *38 *27 *27 *22 *22 *22 *22 *16 *10 *18 *27 Ref. Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Weidner 109 Paraguay Location 84°14‘25W Java, Semarang Australia, Sidney Java, Penandjoeng Java, Radjamandala Java, Mt. Djampang Arabia, Yemen, Sanaa Arabia, Yemen, Arabia, Yemen, Sanaa Arabia, Yemen, Hadja Arabia, Yemen, Indonesia, Aru-Islands Caroline Islands, Ruck Australia, Tubiaco forest Australia, Tubiaco Costa Rica, Punta Arenas Costa Rica, Punta West Australia, Fremantle West Australia, Fremantle West Buba to Monti, 300-500m Lisboa, 30km to Bela Vista Lisboa, 30km to Bela Australia, New-South-Wales Australia, New-South-Wales Australia, New-South-Wales Australia, Kimberley District Australia, Kimberley District Australia, Kimberley District Australia, Queensland, Bowen Arabia, Yemen, Sanaa-Manakha Arabia, Yemen, Australia, Queens Land, Gayndah Central America, Corinto, 12°29’N Central Australia, Queensland, Rockhampton Africa, Tanganyika, Lake Tanganyika Africa, Tanganyika, Africa, Adamaua, near Garua and Rei Africa, Australia, Perth, north of Subiaco, Stat. Angola, Sanguengue, highlands of Nova ? 0.04.1921 3 30.04.1921 30.04.1921 12.05.1905 30.04.1953 12.-14.03.1928 Collector, year year Collector, Le Moult, 1936 Houy, Oct. 1912 Houy, W. Wölting, 1907 W. Wölting, 1907 W. Nelles, 24.05.1898 Nelles, 24.05.1898 Museum Godeffroy Museum Godeffroy Museum Godeffroy Le Moult, May 1936 Le Moult, May 1936 Le Moult, May 1936 H. Rolle, 14.06.1905 C. Rathjens, Apr. 1931 Apr. C. Rathjens, C. Rathjens, Sep. 1931 C. Rathjens, Aug. 1931 C. Rathjens, R. Paessler, 04-06.05.1912 R. Paessler, 04-06.05.1912 R. Paessler, Hamb. S. W. Austr. Exped. Austr. W. Hamb. S. Museum Godeffroy, No. 7989 Museum Godeffroy, Museum Godeffroy, No. 2719 Museum Godeffroy, Brunner von Wattenwyl, 1889 Wattenwyl, Brunner von Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Museum Godeffroy, No. 15893 Museum Godeffroy, Hamb. Angola Expedition, Nr 2, Hamb. Hamb. S. W. Austr. Expedition 1905 Austr. W. Hamb. S. Dirsh, 1974 Dirsh, 1974 Kevan, 1955 Author, year Author, Uvarov, 1936 Uvarov, Uvarov, 1936 Uvarov, 1936 Uvarov, 1936 Uvarov, Ramme, 1929 Sjöstedt, 1921 Sjöstedt, 1932 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1932 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1920 Sjöstedt, 1921 Sjöstedt, 1920 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Weidner, 1964 Weidner, Willemse, 1968 Willemse, 1968 Willemse, Willemse, 1968 Willemse, 1968 Willemse, Giglio Tos, 1895 Giglio Tos, ♀ ♀ ♀ ♀ ♂ 1♂ 1♀ 1♀ 1♀ 1♂ 1♀ 1♀ 1♀ 1♂ 1♀ 1 1 1♀ 1♀ 1 1♀ 1♀ 1♀ 1♀ 1♀ 1♂ 1♂ 1♀ Sex ♂2 ♂1 2 1 1♂ 1♀ 2♂ 1♀ 2♂ 1♀ 1♂ 1♀ 2♂ 2♀ 1 1 1 2 1 1 1 1 1 1 1 1 3 3 1 1 4 2 1 2 1 1 1 1 4 1 1 1 1 1 Count P P P P P P P P P P P P P P P P P P P P P L H H H H H H H PL Type ------Rapsilla fusca Stropis nigrovitellina Stropis Macrocara conglobata Macrocara Pseudogmothela pallida SYN Current taxonomy Current Rhammatocerus brunneri SYN Trachypetrella anderssonii Trachypetrella SYN SYN carolinensis Valanga nigricornis v. nigricornis v. aroensis gutulosa minor gutulosa minor irregularis carolinensis signata v. irregularis modesta kosswigiana picta pulchra nigrovitellina viridis cornelii centralis centralis flavovittatus rathjensi cornelii cornelii cornelii gracilis fusca fusca testacea modesta gracilis gracilis nana megalocephala brunneri diluta Species pis Valanga Urnisa Urnisa Valanga Valanga Valanga Terpillaria Terpillaria Trachypetrella Terpillaria Terpillaria Stro Stropis Stenocatantops Schistocerca Schistocerca Schizobothrus Sphodromerus Stenocatantops Stenocatantops Pycnodictya Rapsilla Rapsilla Rapsilla Rhitzala Pseudogmo Pseudostauronotus Pycnodictya Pycnodictya Pycnodictya Porraxia Genus 71 72 67 68 69 70 66 64 65 63 57 58 59 60 61 62 56 52 53 54 55 No. 51

evolsyst.pensoft.net Evolutionary Systematics 2 2018, 21–30 25 *5 *6 *6 #1 *6 #1 #1 #2 *1 #2 *1 #2 *1 #2 *1 *17 *22 *22 *22 *22 *22 *22 *33 *35 *25 *25 *24 *38 *21 *24 *25 Ref. Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y N N N N N Weidner Stat. 139 mberley District Java Peru Chile Mioko Location , Lion Mill, Java, Kangean Kraeng, 5000m near Alta Gracia (Bügelek Kayasi) Java, Penandjoeng Java, Radjamendal Java, Mt. Djampang Anatolia, Erciyas Dag Caroline Islands, Ruck Australia, Ki Australia Australia, Bunswick, Stat. 139 Australia, Queens Land, Gayndah Australia, Queens Land, Gayndah Australia, New-South-Wales, Sidney Australia, New-South-Wales, Australia, Queensland, Rockhampton Australia, Queensland, Rockhampton Australia, Queensland, Rockhampton Australia, Perth, Karrakatta, Stat. 139 Turkey, Erzincan-Kamaliye-Yuvaköyü Erzincan-Kamaliye-Yuvaköyü Turkey, Sulawesi, Samanga, South Celebes, Bua- Turkey, Antalya, Tahtali Dagi, ca. 1500m Tahtali Antalya, Turkey, Argentina, Province Cordoba, La Granja, Argentina, ? ? 1964 J. Hubrich 30.04.1921 Collector, year year Collector, Le Moult, 1936 Le Moult, 1936 Le Moult, 1936 oelckler, 06.02.1899 oelckler, Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Godeffroy Museum Goddefroy Museum Goddefroy Le Moult, Oct. 1936 Le Moult, Oct. 1936 O. V A. Demirsoy, 17.07.1949 A. Demirsoy, H. Fruhstorfer, Feb. 1896 H. Fruhstorfer, D. Hasan , 20.06.1968 Brunner von Wattenwyl, 1889 Wattenwyl, Brunner von Museum Stockholm, Mjöberg, Museum Stockholm, Mjöberg, Zoologisch Botanische Exkursion Hamb. S. W. Austr. Expedition 1905 Austr. W. Hamb. S. Expedition 1905 Austr. W. Hamb. S. Expedition 1905 Austr. W. Hamb. S. Liana, 1980 Brunner von Author, year Author, Serville, 1838 Serville, 1838 Serville, 1838 Serville, 1838 Sjöstedt, 1936 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1936 Sjöstedt, 1936 Sjöstedt, 1932 Sjöstedt, 1920 Sjöstedt, 1936 Sjöstedt, 1936 Sjöstedt, 1932 Willemse, 1931 Willemse, Willemse, 1935 Willemse, Willemse, 1968 Willemse, Demirsoy, 1973 Demirsoy, Demirsoy, 1973 Demirsoy, Demirsoy, 1973 Demirsoy, Wattenwyl, 1890 Wattenwyl, 1♀ 1♀ 1♂ 1♀ 2♀ 1♀ 1♀ 1♂ 1♀ 1♂ 1♀ 1♂ 1♀ 6♀ 1♂ 2♂ 2♀ 2♀ 1♀ 1♂ 1♂ 1♀ 1♀ 1♀ Sex 1 1 1 2 1 1 1 1 1 1 1 1 1 6 1 2 2 2 1 1 1 1 1 1 Count P P P P P P P P P S L L T T T T H H H H H H H PL Type ------Nocaracris Aucacris eumera Heide amiculi Amangu parva SYN Nocarodes bicoloripes Nocarodes Swanea laticornis Current taxonomy Current SYN Paranocarodes anatoliensis Paranocarodes Valanga nigricornis carolinensis Valanga SYN SYN peruvianus kosswigi mirabilis amiculi laticornis obscura parva hebardi citripes bicoloripes f. marani fieberi anatoliensis major major soror viridipes dirshi humilis humilis humilis humilis flavonigra signatus soror nobilis miokoana rubrispinarum Species Anchotatus Paranothrotes Altograciosa Callitala Moraba Moraba Moraba Moraba Aucacris Paranocaracris Paranocarodes Callitala Valanga Valanga Xenocatantops Xenocatantops Xenocatantops Xenocatantops Zebratula Eupatrides Valanga Valanga Valanga Genus 90 91 89 82 83 84 85 Ommexechidae 86 Pamphagidae 87 88 Chorotypidae 80 Morabidae 81 76 77 78 79 74 75 No. 73

evolsyst.pensoft.net 26 Lara-Sophie Dey & Martin Husemann: An annotated catalogue of the types of short-horned grasshoppers... *8 *3 *3 *3 *4 *3 *5 *5 *5 *5 *22 *22 *27 *27 *22 *22 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 *17 Ref. Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y N N N N N N N N N N N N N N N N Weidner Concepción Chile Parana Location Venezuela oterillos near Mendoza Theresopolis Bolivia, Oruro Peru, Callanga Chile, Coronel Bay, Cape Town Bay, Ecuador, Zaruma Ecuador, Ecuador, Zaruma Ecuador, Chile, Bolivia, Rio Sango Brasilia, Petropolis Brasilia, Petropolis Phillippines, Patria? Africa, Massai-Land W Austr., Serpentine Austr., W Arabia, Yemen, Sanaa Arabia, Yemen, Sanaa Arabia, Yemen, Costa Rica S Paulo, Tibirisá Costa Rica S Paulo, Africa, Tanganyika, Kihenga Africa, Tanganyika, Australia, Queensland, Bowen Peru, Alto Amazonas Province South Peru, Cayara, ca. 3200m South Africa, Natal, Marianhill South Brazilia, St. Catharina Province South-West-Australia, Serpentine South-West-Australia, Brasil, Rio de Janeiro, 900-1000m Cordillera, Poterillos near Mendoza Cordillera, P Australia, New-South-Wales, Sidney Australia, New-South-Wales, Australia, Queensland, Rockhampton Costa Rica S. Paulo, Tibirisá, near Rio Tibirisá, Costa Rica S. Paulo, Africa, Cape Province, Flats, False Brazilia, St. Catharina Province, Joinville ? ? n?, 07.11.1889 H. Rode F. Ohaus F. F. Ohaus F. 11.12.1889 Fischer, 1884 Fischer, 23-25.09.1905 23-25.09.1905 Collector, year year Collector, Alte Sammlung Museum Godeffroy Museum Godeffroy C. Bock, 23.07.1903 F. Ohaus, 03.12.1926 F. F. Ohaus, 20.10.1926 F. Ohaus, 20.10.1926 F. F. Ohaus, 20.09.1905 F. Ohaus, 20.09.1905 F. E. Heyne, 23.05.1900 Jacobsen?, 12.05.1899 Killman C. Rathjens, Aug. 1931 C. Rathjens, Aug. 1931 C. Rathjens, Stuhlmann, 12.09.1888 R. Paessler, 17.03.1914 R. Paessler, E. R. Helwig, 03.01.1939 H. Fruhstorfer, 12.02.1889 H. Fruhstorfer, R. Paessler, 02-08.10.1914 R. Paessler, Dr. O. Staudinger, 20.05.1899 O. Staudinger, Dr. Meteor Expedition, Dec. 1925 Meteor Expedition, Dec. 1925 Sammelreise 1936, 21.04.1936 Hamb. S. W. Austr. Expedition, Austr. W. Hamb. S. Hamb. S. W. Austr. Expedition, Austr. W. Hamb. S. Hamb. Geographische Gesellschaft, Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1980 Liana, 1973 Liana, 1973 Liana, 1973 Liana, 1980 Liana, 1980 Brunner von Brunner von Brunner von Brunner von Kevan, 1963 Author, year Author, Uvarov, 1936 Uvarov, 1936 Uvarov, Bolivar, 1904 Bolivar, Bolivar, 1904 Bolivar, Bolivar, 1904 Bolivar, 1904 Bolivar, Bolivar, 1905 Bolivar, Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Sjöstedt, 1921 Wattenwyl, 1890 Wattenwyl, 1890 Wattenwyl, Wattenwyl, 1890 Wattenwyl, Wattenwyl, 1890 Wattenwyl, 1♂ 1♂ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♂ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♀ 1♂ 1♀ 1♀ 1♂ 1♂ 1♂ 1♀ 1♀ 1♀ 1♂ 1♀ Sex 1♂1♀ 1♂1♀ 1♂1♀ 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 2 1 1 1 1 2 1 1 1 1 1 1 2 Count P P P P P P P P P P P P P H H H H H H H H H H H H H H H H H H H Type ------Anchotatus griseus Anchotatus griseus Protanita longiceps Protanita Current taxonomy Current Anchotatus weidneri Greyacris profundesulcata Greyacris SYN Apioscelis bulbosa Pseudoanchotatus jacsewskii SYN mastax yemenita yemenita minuta longiceps maculicornis latevittata ligata cordata grisea f. brunni pulchra spumans v. teretrirostris jaczewski longirostris propinquus saburrae antistes tibarisae tibarisae tibarisae plicatum plicatum brunneri grisea grisea weidneri subtilis subtilis gracilis carbonelli similis similis Species Monistria Paraspena Paraspena Pyrgomorpha Tanita Monistria Monistria Monistria Dictyophorus Monistria Dictyophorus Prosarthria Pseudoanchotatus Tetanorhynchus Tetanorhynchus Xenium Brunniella Corynorhynchus Corynorhynchus Corynorhynchus Astroma Astroma Callangania Cephaloscopia Cephaloscopia Cephaloscopia Corynorhynchus Corynorhynchus Anchotatus Apioscelis Astroma Anchotatus Genus 114 115 116 113 111 112 109 110 108 103 104 105 106 107 101 102 100 95 96 97 98 99 93 94 No. 92

evolsyst.pensoft.net Evolutionary Systematics 2 2018, 21–30 27 *4 *4 *4 *11 *30 *28 *14 *12 *22 *15 *13 *13 *13 *13 *13 *12 Ref. Y Y Y Y Y Y Y Y Y Y Y Y Y Y Y N Weidner Java Java Patria? D. Irwati Location Okahandja North Borneo North Borneo Peru, Callanga Java, Buitenzorg Perak and Pahang Ecuador, Santa Inez Ecuador, Canary Islands, Tenerife South West Afrika, Okundua near West South Sulawesi, Samanga, South Celebes Borneo, Kinabalu, 1500m, Watersadt Borneo, Kinabalu, 1500m, Watersadt Borneo, Kinabalu, 1500m, Australia, Queensland, Rockhampton Nieder-Burma, near Rangoon a. Unterl. Nieder-Burma, Indonesia, Camp Jor, watershed between Indonesia, Camp Jor, Collector, year year Collector, T. Becker, 1901 Becker, T. H. Schrader, 1913 H. Schrader, H. Rolle, 25.11.1904 H. Rolle, 25.11.1904 W. Krieg, 1933-1934 W. A. Speyer, 20.11.1899 A. Speyer, Patria, Alte Sammlung R. Haensch, 30.04.1903 A. Grubauer, 30.12.1902 A. Grubauer, H. Fruhstorfer, Nov. 1895 Nov. H. Fruhstorfer, O. Staudinger, 20.05.1899 O. Staudinger, H. Fruhstorfer, 05.06.1899 H. Fruhstorfer, H. Fruhstorfer, 10.02.1894 H. Fruhstorfer, Museum Marburg, 24.07.1901 Museum Marburg, K. Kraepelin, 24.02-12.03.1904 Museum Goddefroy, No. 15853 Museum Goddefroy, Walker, 1870 Walker, Author, year Author, Bolivar, 1905 Bolivar, 1905 Bolivar, Bolivar, 1905 Bolivar, Sjöstedt, 1921 Günther, 1937 Günther, 1994 Günther, Günther, 1939 Günther, 1939 Günther, 1939 Günther, 1939 Günther, 1938 Günther, 1938 Günther, Günther, 1939 Günther, Karaman, 1965 Willemse, 1930 Willemse, 1♂ 1♂ 1♀ 1♂ 1♀ 1♂ 2♂ 1♀ 1♀ 2♂ 2♀ 1♂ 1♀ 1♀ Sex 2♂1♀ 1♂ 2♀ 1 1 1 3 1 1 1 1 2 1 1 2 2 1 1 1 Count P P P P P P P P P H H H H PL PL PL Type ------Systella siccifolia meridionalis Current taxonomy Current Systolederus ophthalmicus SYN SYN dusmeti dusmeti philippensis meridianus borneensis dusmeti amplus meridionalis weidneri carli celebensis brunneri kraepelini otumboides difficilis compressa overbecki perakensis Species Systella Systella Systella Systella Afrotridactylus Systella Paratettix Paratettix Systolederus Bolivaritettix Chiriquia Hyboella Hyboella Loxilobus Loxilobus Allotettix Genus 129 130 126 Tridactylidae 127 Trigonopterygidae 128 124 125 118 119 120 121 122 123 No. 117 Literature: #1 Key (1958); #2 Willemse (1964); *1 Audinet-Serville (1839); *2 Bey-Bienko (1960); *3 Bolívar (1904); *4 Bolívar (1905); *5 Wattenwyl (1890); *6 Demirsoy (1973); *7 Demirsoy (1979); *8 Dirsh (1963); *9 Dirsh (1974); *10 (1963); *9 Dirsh Dirsh (1979); *8 (1973); *7 Demirsoy (1890); *6 Demirsoy (1964); *1 Audinet-Serville (1839); *2 Bey-Bienko (1905); *5 Wattenwyl (1904); *4 Bolívar (1960); *3 Bolívar #2 Willemse (1958); #1 Key Literature: (1980); *18 Ramme (1955); *17 Liana (1929); *19 Ramme (1931); *20 Ramme (1941); *21 Sjöstedt (1965); *16 Kevan (1994); *15 Karaman (1939); *14 Günther (1938); *13 Günther (1937); *12 Günther (1895); *11 Günther Giglio-Tos (1931)a; (1930)b; *32 Willemse (1930)a; *31 Willemse (1964); *30 Willemse (1870); *29 Weidner (1936); *28 Walker (1851); *27 Uvarov (1936); *26 Stål (1932); *25 Sjöstedt (1928); *24 Sjöstedt (1921); *23 Sjöstedt (1920); *22 Sjöstedt (1968) (1965); *38 Willemse (1936); *37 Willemse (1935); *36 Willemse (1933); *35 Willemse (1931)b; *34 Willemse *33 Willemse

evolsyst.pensoft.net 28 Lara-Sophie Dey & Martin Husemann: An annotated catalogue of the types of short-horned grasshoppers...

Figure 1. Shown are a) a cotype of Gastrimargus musicus v. kimberleyensis Sjöstedt, 1928; b) the holotype of Adreppus tuber- culatus Sjöstedt, 1921; c) the holotype of Callangania brunneri Liana, 1980; and d) the tentative type of a Tetrigid, identified as unknown, but not formally described by Kurt Günther and their labels. order Orthoptera and member of the Museum Godeffroy, Weidner and Wagner (1968) (Cephalocoema teretiuscu- worked with the collection of the ZMH (Weidner 1967). la, Brunner von Wattenwyl, 1890; Rhipipteryx striatipes, Hence, major parts of the Caelifera collection goes back Chopard 1954; Tridactylus emarginatus, Chopard 1954). to the Museum Godeffroy, a private collection belonging Further, the status of some species was not clear in Weid- to a leader of a big trade company in the 18th century. ner’s catalogues and some species were recorded twice in Shortly after the Godeffroy Company bankrupted, most the list; 115 of the species recorded by Weidner are also of the collections of the museum were sold and the zo- recorded here. Type material of 13 species has been de- ological material was largely transferred to the ZMH in posited in the collection since the last catalogue. 1885 (Weidner 1967). Additional material now housed in In addition to ZMH types reported here, other specimens the ZMH was collected during several expeditions by sci- originally designated as type, but not formally published, entists of the Museum Hamburg (e.g. to Africa (Fischer are present in the collection (Fig. 1d); this material is not 1884) and southwestern Australia (Michaelson 1908)) or listed here. An example comes from the genus Paratettix: was donated from researchers or private collectors. Many Günther (1979) mentioned material of the genus, which other important orthopterists worked with the collection likely belongs to an undescribed species. This material is and designated and deposited types; these include Boris marked as such in the collection, but a new species was Petrovitch Uvarov, D. Keith Mc. E. Kevan, Carl Stål, never formally described. Further, it is likely that addition- Grigory Yakovlevich Bey-Bienko, Willi Adolf Theodor al types, not marked as such, are present; especially the Ramme, V. M. Dirsh, Jean-Guillaume Audinet-Serville, material of the Collection Godeffroy is likely to contain ad- Ali Demirsoy, Anna Liana, Ignatio Bolivar, and Cornelis ditional type specimens. Experts of the respective groups and Willemse. are needed to investigate and work with the collection. This The collection currently houses 209 type specimens of paper is one of the first in a row of reports to be published Caelifera belonging to 128 species; 65 species are repre- in the next years to present updated type catalogues of the sented by holotypes (Table 1), three by lectotypes and one collection of the ZMH (now part of the CeNak). by a syntype. Besides a large number of paratypes and paralectotypes, the collection also houses topotypes of two species, which are not regulated by the International Acknowledgement Code of Zoological Nomenclature, but are listed here for completeness. Type material of 120 species was recorded We want to thank Holger Braun, Bruno Massa and an in previous catalogues presented by Weidner and Wag- anonymous reviewer for valuable comments on a previous ner (1968) and Weidner (1977). All previously recorded version of this manuscript. This project was funded by the types were recovered, except the material of three spe- Orthopterists’ Society and by the Gesellschaft der Freunde cies, which was already recorded as burned or missing by und Förderer des Zoologischen Museums Hamburg.

evolsyst.pensoft.net Evolutionary Systematics 2 2018, 21–30 29

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evolsyst.pensoft.net 30 Lara-Sophie Dey & Martin Husemann: An annotated catalogue of the types of short-horned grasshoppers...

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