Principal Types of Vegetative Shoot Apex Organization in Vascular Plants1
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Plant Terminology
PLANT TERMINOLOGY Plant terminology for the identification of plants is a necessary evil in order to be more exact, to cut down on lengthy descriptions, and of course to use the more professional texts. I have tried to keep the terminology in the database fairly simple but there is no choice in using many descriptive terms. The following slides deal with the most commonly used terms (more specialized terms are given in family descriptions where needed). Professional texts vary from fairly friendly to down-right difficult in their use of terminology. Do not be dismayed if a plant or plant part does not seem to fit any given term, or that some terms seem to be vague or have more than one definition – that’s life. In addition this subject has deep historical roots and plant terminology has evolved with the science although some authors have not. There are many texts that define and illustrate plant terminology – I use Plant Identification Terminology, An illustrated Glossary by Harris and Harris (see CREDITS) and others. Most plant books have at least some terms defined. To really begin to appreciate the diversity of plants, a good text on plant systematics or Classification is a necessity. PLANT TERMS - Typical Plant - Introduction [V. Max Brown] Plant Shoot System of Plant – stem, leaves and flowers. This is the photosynthetic part of the plant using CO2 (from the air) and light to produce food which is used by the plant and stored in the Root System. The shoot system is also the reproductive part of the plant forming flowers (highly modified leaves); however some plants also have forms of asexual reproduction The stem is composed of Nodes (points of origin for leaves and branches) and Internodes Root System of Plant – supports the plant, stores food and uptakes water and minerals used in the shoot System PLANT TERMS - Typical Perfect Flower [V. -
Untangling Phylogenetic Patterns and Taxonomic Confusion in Tribe Caryophylleae (Caryophyllaceae) with Special Focus on Generic
TAXON 67 (1) • February 2018: 83–112 Madhani & al. • Phylogeny and taxonomy of Caryophylleae (Caryophyllaceae) Untangling phylogenetic patterns and taxonomic confusion in tribe Caryophylleae (Caryophyllaceae) with special focus on generic boundaries Hossein Madhani,1 Richard Rabeler,2 Atefeh Pirani,3 Bengt Oxelman,4 Guenther Heubl5 & Shahin Zarre1 1 Department of Plant Science, Center of Excellence in Phylogeny of Living Organisms, School of Biology, College of Science, University of Tehran, P.O. Box 14155-6455, Tehran, Iran 2 University of Michigan Herbarium-EEB, 3600 Varsity Drive, Ann Arbor, Michigan 48108-2228, U.S.A. 3 Department of Biology, Faculty of Sciences, Ferdowsi University of Mashhad, P.O. Box 91775-1436, Mashhad, Iran 4 Department of Biological and Environmental Sciences, University of Gothenburg, Box 461, 40530 Göteborg, Sweden 5 Biodiversity Research – Systematic Botany, Department of Biology I, Ludwig-Maximilians-Universität München, Menzinger Str. 67, 80638 München, Germany; and GeoBio Center LMU Author for correspondence: Shahin Zarre, [email protected] DOI https://doi.org/10.12705/671.6 Abstract Assigning correct names to taxa is a challenging goal in the taxonomy of many groups within the Caryophyllaceae. This challenge is most serious in tribe Caryophylleae since the supposed genera seem to be highly artificial, and the available morphological evidence cannot effectively be used for delimitation and exact determination of taxa. The main goal of the present study was to re-assess the monophyly of the genera currently recognized in this tribe using molecular phylogenetic data. We used the sequences of nuclear ribosomal internal transcribed spacer (ITS) and the chloroplast gene rps16 for 135 and 94 accessions, respectively, representing all 16 genera currently recognized in the tribe Caryophylleae, with a rich sampling of Gypsophila as one of the most heterogeneous groups in the tribe. -
Studies on Seed Germination, Seedling Growth, and in Vitro Shoot
HORTSCIENCE 44(3):751–756. 2009. plantlets are detached from the mother plant that are dried and planted. However, seed propagation is more feasible and recommen- Studies on Seed Germination, Seedling ded for survival of rare species (Van Wyk and Smith, 1996). If this species has to be Growth, and In Vitro Shoot Induction propagated on a large scale by means of seed or tissue culture methods, then currently there of Aloe ferox Mill., a Commercially is no basic information available on these aspects. Aloes are succulent and warm-cli- mate plants, where both temperature and Important Species water play an important role in establishing Michael W. Bairu, Manoj G. Kulkarni, Rene´e A. Street, Rofhiwa them. This study was therefore conducted to B. Mulaudzi, and Johannes Van Staden1 examine 1) the effects of different temper- atures, growth-promoting substances, and Research Centre for Plant Growth and Development, School of Biological watering frequencies on seed germination and Conservation Sciences, University of KwaZulu-Natal Pietermaritzburg, and seedling growth of A. ferox; and 2) to Private Bag X01, Scottsville 3209, South Africa assess the applicability of an in vitro propa- gation protocol developed for other Aloe spp. Additional index words. cytokinins, growth regulators, multiplication rate, smoke solutions, temperature, tissue culture Materials and Methods Abstract. A study was done to investigate the effects of some physical and chemical factors on growth and development of Aloe ferox ex vitro and in vitro. The effects of light, Seed collection. Dried seeds of A. ferox temperature, and smoke–water on seed germination, ex vitro seedling growth require- were collected between the middle to the end ments, and effect of germination medium and cytokinins on shoot induction and of August from the Botanical Garden, Uni- multiplication in vitro were investigated. -
Chapter 5: the Shoot System I: the Stem
Chapter 5 The Shoot System I: The Stem THE FUNCTIONS AND ORGANIZATION OF THE SHOOT SYSTEM PRIMARY GROWTH AND STEM ANATOMY Primary Tissues of Dicot Stems Develop from the Primary Meristems The Distribution of the Primary Vascular Bundles Depends on the Position of Leaves Primary Growth Differs in Monocot and Dicot Stems SECONDARY GROWTH AND THE ANATOMY OF WOOD Secondary Xylem and Phloem Develop from Vascular Cambium Wood Is Composed of Secondary Xylem Gymnosperm Wood Differs from Angiosperm Wood Bark Is Composed of Secondary Phloem and Periderm Buds Are Compressed Branches Waiting to Elongate Some Monocot Stems Have Secondary Growth STEM MODIFICATIONS FOR SPECIAL FUNCTIONS THE ECONOMIC VALUE OF WOODY STEMS SUMMARY ECONOMIC BOTANY: How Do You Make A Barrel? 1 KEY CONCEPTS 1. The shoot system is composed of the stem and its lateral appendages: leaves, buds, and flowers. Leaves are arranged in different patterns (phyllotaxis): alternate, opposite, whorled, and spiral. 2. Stems provide support to the leaves, buds, and flowers. They conduct water and nutrients and produce new cells in meristems (shoot apical meristem, primary and secondary meristems). 3. Dicot stems and monocot stems are usually different. Dicot stems tend to have vascular bundles distributed in a ring, whereas in monocot stems they tend to be scattered. 4. Stems are composed of the following: epidermis, cortex and pith, xylem and phloem, and periderm. 5. Secondary xylem is formed by the division of cells in the vascular cambium and is called wood. The bark is composed of all of the tissues outside the vascular cambium, including the periderm (formed from cork cambium) and the secondary phloem. -
Tunica), the Cells of Which Show a Preferred Plane of Cell Division
APICAL MERISTEMS OF VEGETATIVE SHOOTS AND STROBILI IN CERTAIN GYMNOSPERMS BY ERNEST M. GIFFORD, JR.,* AND RALPH H. WETMORE BIOLOGICAL LABORATORIES, HARVARD UNIVERSITY Communicated April 26, 1957 There has been considerable interest of late in the transition of seemingly vege- tative shoots of angiosperms into "reproductive" axes with their appendages. The flowering shoot formed in this manner is assumed to be a reflection of a change or changes which must have taken place in the activity of the apical meristem. It is well known for certain angiosperms that a proper period of photoinduction will bring about flowering. While a considerable amount of information has accumu- lated on the responses of plants to photoperiodism, very few correlative anatomical studies have been made of the same plant material. Descriptive studies do exist for some angiosperms, but most of these are unrelated to controlled experimental studies on the effects of photoperiodism. Not all inflorescences or flowers are clearly transformations of shoots that were previously vegetative, for buds continue to be formed in some species after the plant has been induced to flower. Whether the apices of these later-formed buds exhibit initially a structure comparable to the parent vegetative shoot apex or whether they are, from their inception, fundamentally different in organization has not been elucidated. For many years the structure of shoot apical meristems in angiosperms has Leens described in terms of planes of cell divisions. In most angiosperms there is a dis- crete outer layer or layers (tunica), the cells of which show a preferred plane of cell division. New cell walls are perpendicular to the surface of the shoot apex. -
A Comparative Study of the Primary Vascular System Of
ArneI'. J. Bot. 5.5(4): 447-457. 1968. A COMPARATIVE STUDY OF THE PRIMARY VASCULAR SYSTElVI OF CONIFERS. 1. GENERA WITH HELICAL PHYLLOTAXISl KADAMBARI K. N AMBOODIRI2 AND CHARLES B. BECK Department of Botany, University of Michigan, Ann Arbor ABSTRACT The primary vascular system of 23 species belonging to 18 genera of conifers with helical phyllotaxis has been investigated with the intent of determining the architecture .f the system. Special attention has been given to nodal and subnodal relations of the vascular bundles. The vascular system seems to be composed solely of relatively discrete sympodia, that is, axial vascu lar bundles from which leaf traces branch unilaterally. Although the discreteness of the syrn podia is not immediately apparent because of their undulation and lateral contacts with neigh boring ones, close examination, including a statistical analysis of the tangential contacts, seems to reveal that each sympodium maintains its identity throughout. Although two traces may be apparent at nodal levels, the trace supply to a leaf originates, in all species, as a single bundle. An analysis is made of the relationship between the vasculature and the phyllotaxis. It is ob served that the direction of trace divergence can be accurately predicted when the direction of the ontogenetic spiral, the angle of divergence of leaf traces, and the number of syrnpodia are known. THE ORIGIN and evolution of gymnosperms that of the ferns by reduction (Jeffrey, 1902, are significant problems that deserve increased 1917). Consequently, he considered the leaf gap attention. There have been few modern compara of seed plants to be homologous with that of tive studies of extant gymnosperms, and most the ferns. -
Plant Pathology
Plant Pathology 330-1 Reading / Reference Materials CSU Extension Fact Sheets o Aspen and poplar leaf spots – #2.920 o Backyard orchard: apples and pears [pest management] – #2.800 o Backyard orchard: stone fruits [pest management] – #2.804 o Bacterial wetwood – #2.910 o Cytospora canker – #2.937 o Diseases of roses in Colorado – #2.946 o Dollar spot disease of turfgrass – #2.933 o Dutch elm disease – #5.506 o Dwarf mistletoe management – #2.925 o Fairy ring in turfgrass – #2.908 o Fire blight – #2.907 o Forest fire – Insects and diseases associated with forest fires – #6.309 o Friendly pesticides for home gardens – #2.945 o Greenhouse plant viruses (TSWV-INSV) – #2.947 o Honeylocust diseases – #2.939 o Juniper-hawthorn rust – #2.904 o Juniper-hawthorn rust – #2.904 o Leaf spot and melting out diseases – #2.909 o Necrotic ring spot in turfgrass – #2.900 o Non-chemical disease control – #2.903 o Pesticides – Friendly pesticides for home gardens – #2.945 o Pinyon pine insects and diseases – #2.948 o Powdery mildew – #2.902 o Roses – Diseases of roses in Colorado – #2.946 o Russian olive decline and gummosis – #2.942 o Strawberry diseases – #2.931 o Sycamore anthracnose – #2.930 CSU Extension Publications o Insects and diseases of woody plants of the central Rockies – 506A Curriculum developed by Mary Small, CSU Extension, Jefferson County • Colorado State University, U.S. Department of Agriculture and Colorado counties cooperating. • CSU Extension programs are available to all without discrimination. • No endorsement of products named is intended, nor is criticism implied of products not mentioned. -
2021 Plant List
2021 Plant List New items are listed with an asterisk (*) Conifers Pinus thungerbii Abies koreana 'Horstmann's Silberlocke' Pinus x 'Jane Kluis' * Chamaecyparis nootkatensis 'Pendula' Sciadopitys vert. 'Joe Dozey' Chamaecyparis noot. 'Glauca Pendula' Sciadopitys vert. 'Wintergreen' Chamaecyparis obtusa 'Chirimen' * Taxodium distichum 'Pendula' Chamaecyparis obtusa 'Gracilis' -Select Taxodium distichum 'Peve Mineret' Chamaecyparis obtusa 'Kosteri' Taxus cuspidaata 'Nana Aurescens' Chamaecyparis obtusa 'Nana' Tsuga con. 'Jervis' Chamaecyparis obtusa 'Nana Gracilis' Chamaecyparis obtusa 'Spiralis' Ferns Chamaecyparis obtusa 'Thoweil' Adiantum pedatum ….Maiden Hair Chamaecyparis obtusa 'Verdoni' Athyrum filix-femina 'Minutissima' Juniperus procumbens 'Nana' Athyrium 'Ghost' Larix decidua 'Pendula' Athyrum niponicum 'Godzilla' Larix decidua 'Pendula' -Prostrate Form Athyrum niponicum 'Pictum' Picea abies 'Hasin' * Athyrum niponicum pic. 'Pearly White' Picea abies 'Pusch' * Dennstaedtia punctilobula Picea omorika 'Nana' Dryopteris ery. 'Brilliance' Picea omorika 'Pendula' Dryopteris marginalis Picea orientalis 'Nana' Matteucciastruthiopteris var. pensylvanica Picea orientalis 'Shadow's Broom' * Osmunda cinnamomea Picea pungens 'Glauca Globosa' Polystichum acrostichoides Pinus mugo 'Mughus' - Rock Garden Strain Polystichum polyblepharum Pinus mugo 'Slowmound' Pinus nigra 'Hornibrookiana' Grasses Pinus parviflora 'Aoi' These are but a fraction of the grasses we'll be Pinus parviflora 'Glauca Nana' offering this year. Many more to come. They'll -
Tecophilaeaceae 429 Tecophilaeaceae M.G
Tecophilaeaceae 429 Tecophilaeaceae M.G. SIMPSONand P.J. RUDALL Tecophilaeaceae Leyb., Bonplandia JO: 370 (1862), nom . cons . Cyanastraceae Engler (1900). Erect, perennial, terrestrial herbs. Roots fibrous. Subterranean stem a globose to ellipsoid corm, 1- 4 cm in diameter, in some genera with a membra nous to fibrous tunic consisting of persistent sheathing leaves or fibrovascular bundles . Leaves basal to subbasal, or cauline in Walleria, spiral; base sheathing or non-sheathing, blades narrowly linear to lanceolate -ovate, or more or less petiolate in Cyanastrum and Kabuyea; entire, glabrous, flat, or marginally undulate; venation parallel with a major central vein. Flowers terminal and either Fig. 122A-F. Tecophilaeaceae. Cyanastrum cordifolium . A Flowering plant. B Tepals with sta mens. C Stamens. D Pistil. E solitary (or in small groups) and a panicle or (in Ovary, longitudinal section. F Capsule. (Takh tajan 1982) Walleria) solitary in the axils of cauline leaves. Bracts and bracteoles (prophylls) often present on pedicel. Flowers 1- 3 cm long, pedicellate, bisexual , trimero us. Perianth variable in color, zygomor fibrous scale leaves or leaf bases or the reticulate phic or actinomorphic, homochlamydeous, ba fibrovascular remains of these scale leaves (Fig. sally syntepalous; perianth lobes 6, imbricate in 2 123). The tunic often continues above the corm, in whorls, the outer median tepal positioned anteri some cases forming an apical tuft. Corms of orly; minute corona appendages present between Walleria, Cyanastrum, and Kabuyea lack a corm adjacent stamens in some taxa. Androecium aris tunic (Fig. 122). ing at mouth of perianth tube, opposite the tepals Leaves are bifacial and spirally arranged. -
Red Shoot Disease of Cranberry
Cranberry Pest Management Red Shoot Disease of Cranberry Red shoot, caused by the fungus Symptoms and spindly with red or yellow Exobasidium perenne, is a rela- leaves that are slightly more tively minor disease of the culti- Signs round than typical oblong cran- vated American cranberry Symptoms Þrst appear in the berry leaves (Figures 1Ð3). By (Vaccinium macrocarpon) and the spring on current yearÕs shoots mid summer the lower surfaces wild small cranberry (Vaccinium that occur singly or as a cluster of leaves become covered with oxycoccus). The red shoot fun- arising from a node on a buried white, powdery fungal spores. gus is related to the fungus that runner. Affected shoots do not Shoots wither after the fungus causes red leaf spot. Red shoot produce ßowers but rather are has shed its spores. Diseased has been reported in Wisconsin, shoots break off easily from the Massachusetts, the PaciÞc Northwest, and in the Atlantic maritime provinces of Canada. Red shoot is not an economi- cally important disease, and speciÞc control recommenda- tions have not been developed. However, the distorted, red or yellow shoots have sometimes been misidentiÞed as a weed, making growers think that an herbicide should be applied. The fact that red shoot is a dis- ease has been conÞrmed by tak- ing fungal spores from infected plants and inoculating them Figure1. Clusters of red shoot in a bed of ‘Searles’. The orangish-rust col- onto healthy plants; the inocu- ored shoots to the left are not affected by red shoot, but rather a different lated plants then developed red problem such as upright dieback. -
Vegetative Vs. Reproductive Morphology
Today’s lecture: plant morphology Vegetative vs. reproductive morphology Vegetative morphology Growth, development, photosynthesis, support Not involved in sexual reproduction Reproductive morphology Sexual reproduction Vegetative morphology: seeds Seed = a dormant young plant in which development is arrested. Cotyledon (seed leaf) = leaf developed at the first node of the embryonic stem; present in the seed prior to germination. Vegetative morphology: roots Water and mineral uptake radicle primary roots stem secondary roots taproot fibrous roots adventitious roots Vegetative morphology: roots Modified roots Symbiosis/parasitism Food storage stem secondary roots Increase nutrient Allow dormancy adventitious roots availability Facilitate vegetative spread Vegetative morphology: stems plumule primary shoot Support, vertical elongation apical bud node internode leaf lateral (axillary) bud lateral shoot stipule Vegetative morphology: stems Vascular tissue = specialized cells transporting water and nutrients Secondary growth = vascular cell division, resulting in increased girth Vegetative morphology: stems Secondary growth = vascular cell division, resulting in increased girth Vegetative morphology: stems Modified stems Asexual (vegetative) reproduction Stolon: above ground Rhizome: below ground Stems elongating laterally, producing adventitious roots and lateral shoots Vegetative morphology: stems Modified stems Food storage Bulb: leaves are storage organs Corm: stem is storage organ Stems not elongating, packed with carbohydrates Vegetative -
Pruning Fruit Trees: Rules of Shoot Growth
Pruning Fruit Trees: Rules of Shoot Growth Pruning fruit trees is usually perceived as a very difficult task for the inexperienced. This is due primarily to the uncertainty involved in deciding what to prune, where to cut, and how much wood to remove. Pruning decisions can be made much easier if we know the way a tree grows naturally. The purpose of this article is to present some basic principles and observations on shoot growth. While these rules were developed with tree fruit in mind, they are generally applicable to deciduous trees in the landscape. Rules of Shoot Growth 1. If two branches are identical in length, and diameter, and they emerge from the tree trunk at the same location, they will have the same growth potential. Such limbs have comparable hormone levels, leaf area, and they will receive comparable nutrients and growth promoting substances from the roots. 2. The steeper the angle of a branch, the more vigorously it will grow, provided all the other conditions are identical. This rule is especially important since it gives you, the pruner, a glimpse into the future about growth potential of limbs in a tree. Upright limbs, which form an angle with the trunk of 30 degrees or less, are apt to be too vigorous, grow inappropriately large, and by the nature of their vigor, be quite unproductive. Conversely, a limb that is flat and forms an angle of 90 degrees or more with the trunk will be weak and display little if any terminal growth. Further, vigorous upright shoots may grow from these flat limbs, and pose contin-ual problems in the future.