'?urpeelsrur esrm:eq1oro en8e,req ol reedduleql suorlcellocplo ruo{ setlllecoleuos Surprp8arsrppue suort.ollo. luoca.Ituo4 elep eq1Sunurodrocur 'pepr,rord osp erc sduruuoqnqrJlsrp A\eN snl€ls uorlul.resuoc 'l€trqPq 'uorsr,\eJ pue eruq Suue,\\ou s€ qcns eql ur pare,\oclou seeJ€ol pu€ s1[ur1cueue8 pe:elp eq1 o1 3uqe1u elap,r,reuo1 ,{1uuu pelculser sr €xe1paqrrcsap ,4ysnor,rerd eq1 :oy eJaquelr8 uorlerutoJur eq1 reded srql ur pel€r1snllrpue peqrJcsepsr qcrq,r'sercedso aisp)tlaN L\eu euo rol ldecxe serceds eql IIB Jo suorleJlsnlppue suoqducseppelrcgep sepnord oqu uorsrl3l cnuouoxul s,Jrunhl (g16l) Jrunr{ ur Ir€lap ur petueseJdsr equl eqt ;o srequteruyp go ,(Jotsrqarll pu€lsueen| ol tseeerlur1snv 'eueptsdos,{q4 urelseTyyuro4 Surpuetxelspl eql pulr €rtur1snvuJelse,r-qlnos ol cruapuo o^\t tsJrJeqt eqrrl o1 Sur8uoleq €Jeue8 pet€lsJ flesolc ere DnaJSDo$aNpuB s{.q)otsouqJq7 'slsdos,tqd

'slsdosKl.ldot ollalso)MaN uJot! satcedsea;q1 Suure;sue4 fq setpuroue eseql sessupperaded srql Jaqlou€euo ol rJe'4lsarceds .or1a1socuap, snoleul-S eql Jo euo lseelte ol pet"lel ,{1eso1ceroru aq o1peJ€edd€ r{loq pue srsdos(r74ur pepnlcurueeq puq 'sta,rog snoreru-y,{puatsrsuoc 'serceds qll.4dqloq o,rl f,1ug o4atsocua6lJo (edfl oqt Surpnlcur)sJeqluelx leqto rxojl sjelcuruqJ 'sJe/$ou leJe^esur peJeJJrpta,{ snoreul-g,{lpnsn lraq}tdecxesJalc€.rDqc IIE ur ,slsdo,sqdqaleru ot punoJ plpJSD)$aNtrt 'aptsoJa\aN eJe/ pepnlcur,{lpuourpeJlsercads,reJ V pur s(l.lJolsouq)D1u pepr.Iaur se4tlua cgrcadsur;ureqt osp puu srsdostq4 snue8eql ,1ouoUetlullep eqt qtmr sureyqordpepe,rer eeecuqlueolq3,(pue; aql ur .{luoud uoq"^:esuocyo sguuyduerp4sny u:e1se.,1;o,{a,\lns luarat V

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lJeJlsqv

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(966r)a0l-6a:(l) lr xrsl{nN 80 NuytsiaVol.I l. No. I (1996)

Materials and methods

Apart from a few specimensborrowed from DNA and type specimensseen at MEL, all matedal examined in this study is housed at PERTH. A few recently collected specimensare cited to supplementthe specimenscited previously by Munir (1978); all of them are representedat PERTH and many of them have duplicatesat other herbaria.

'spike' For simplicity, the term is usedthroughout for the spike like inflorescencesfound in the three genera,but is not strictly correctas the cymesand flowers aresubsessile or shortly stalkedrather than sessile. The botanicalprovinces cited for WesternAustralia follow Beard (1980). Each symbol used on the geographicaldistributions indicates the presenceof the speciesin an area measuring 0.25 degreelatitude by 0.25 degreelongitude. Conservationcodes are thoseadopted by the Western Australian Department of Conservationand Land Managementfor its GazettedRare and Priority Flora List; they are defined at the end of this Nuytsia issue.

Taxonomic treatment

Table 1 comparesthe geographicaldistributions and morphological charactedsticsof the five generacurently recognizedin tribe Physopsideae.It should be noted that genedc and infrageneric boundariesin the Dicrastylis Mallophora group need further investigation,but that is beyond the scope of this paper.

Key to genera of tribe Physopsideae

1. Style with dendritic hairs on basal part, the hairs 0.3-0.7 mm long; upper part of style divided into 2 branchesor distinctly 2Jobed (in Mallophora rugosfolia Munir, the lobes are sometimesonly c. 0.2 mm long). Calyx tube much to slightly shorter than, or rarely equalling, the calyx lobes 2. Flowers alf or mosdy 5-merousor (in Dicrastltlis archeri Munir and D. capitelLataMunir) with the calyx mostly 5lobed but corolla lobes and stamensmostly 4. Style hairy on more than half of the undivided portion and sometimeson the branchesas well; branchesusually about as long as undivided portion but sometimesalmost reachingbase of style or as little as 1/4 lengthof sty1e...... Dicrastylis 2. Flowers all or mostly 4-merous. Style hairy only on the basal half of the undivided portion; branchesor lobes short, less than 1/3 the length of thestyle...... Mallophora 1. Style glabrous throughout or with simple or 2-branchedhairs on basal half, the hairs up to 0.3 mm longi apex notchedor 2 lobed. Calyx tube greatly to slightly exceeding,or rarely equalling,the calyx lobes 3. Upper leaf surfaceslightly to distinctly rugose,becoming glabrous. Corolla yellow or white; lobes 4 or 5, broadly obtuse,either with prominent reticulate yeinsor thickenedand with no veinsvisib1e...... Physopsis 'I eLqPlpue ,{J{ Jqt ur pJlEJrpursE 'aqr4 eq1ur ereua8reqlo uJo4 siqJrlsouqrp1 qsrn8urtsrpsratce:uqc luuorlpps lure,reg eeprsdos,{q4 aqr.r1ur enbrun eJD slalrull?qc o,rl eseql peapur :IJE^o oJuBu pelleJ-Z str puu seqol €lloroc 'elrl€leJ tuesquot peonpsJ,{lerrsr Lq'otTatspcuaN lsesolcs1r urorl peqsrn8uqsrps s{qJotsouq)oI

(1 a:n3rg) acur^ord lucruttog ueouursrg3qt otur FpqBIIsSurpualxa pue euozrelul uJelse,r-r0nospu€ ecur,\oJdlecrrclog lse,r-qlnos eql ul'eluelDl qlr,l^pelercosse ueuo 'slros 'BrlBJlsrlv ,4puesuo ,{yr.rrurrrdSuurnaJo uJalse,r-qlnosur cruJapueserceds xrs 3.osnua8 y

.x,{[uc 'pepees-Z ',(rp luelsrsredeq1 ur pesolcue ,{leJurJo -l luecsltlopur rlnrg peqclou xede :esBqauraJtxe 'JIeq eql13 srrtrl cruJpuapqlr.r osP serunelxos IesEquo srrcqpaqcuurq-7 xldeap ro eldurrslE1npuu13 -uou 'pegesxa 'flunlrru qtlr Jo snoJq€13 ,{peer8ai,Ost llec qcveur saln,roT qlr,,n lu pelyac-TSunuoceq 'peUesxe-8uol'aqu fuoto eseqpepuudxe uu r0r,r JepuelstueluElrJ :snoJqEIS eql Io uoq?nuquoc 'sqnl e uroJ ot Sulreeddepue (tuese:d ueq,$) seqol ulloroc eql qlr.,nSuuuuretp IEJoUJo trtuuns aql 1Bpauesul suawojs elqtst^,{Fea1tsura,r IEretBI ,\leJ e ,{1pnsnpue luaurruordure^pru eqt qlr^\ 'peJntxet-uqt 'snorqelS'ot€cun4 tsorul€ ot etnc€ 'reln8ueul ,{l,rolpqs ro e1ea.o-pesserdep'sueruetrs 'pnba 'pecnper puEeqnlelloJo3 u€q1:elroqs qcnur Sueserdueq,n ,{re^.ro lu€sq" s€qol:eprstno snoJqelS 'e1d:nd deep o'! e^neut ollolo1 eqnl u€qt re oqs soqol :eprsursnoJq€18 'eptslno Xrreq-cqrrpuap 'seqol 'snoJeur ,{lesuepr{lr, sueruutspup (tuesaldJr) saqol elloroc xftuc Jo slequrnulenbe qtr,r -(ZI)01-g(t) s.raaroJgs:rcd elrssncep-elrsoddour 'pere,,nog-g,{lpnsn seur,{c:(,{tuntEur t€ seru^3Jo srrcd luacefpe use^\teq sopouJaturpoqs suq ,{1ensn ytryt 'ottloqoua sKqcDlsouqJajur ldecxe) esuep SururprueJ'pnqur esuep se)rds :slelqauBrq,{;ua1 Suqeurrurel se{ds e1d41muro,{ru111os ur 'peanpeJ pe8uerreserufc elrsseslsorulE Jo aJuaJsatorful e€llnq aql ue€A\leqluelsrsJad lnq sl[rJruns eDfinqeql uro{ peqsseurqeuos sJrpq eq]'eEIInq eqt eprque+o qJrq,r'sIEq JIr.rpuJplElnpuElS uou 'elallnq qtl$ pere^oc ,{1deepo1 ,{I,,no1pqs acuJrns leddn :sJrpqcrtrJpuep JElnpu€I8-uou 3:o urnlueurnpur lualstsredasuep e qllt ec€JrnsJs,rol :oJrlue'lua.uncap seurrler.uos 'elrsses 'elessncep elrsoddo sa,ruaT 'sJIEqcltlJpuep relnpuu13-uou;oumluer.rJnpur esusp ,{lo^ u qlr^\ 'pelSuu-t puu pa8prr-7 3ur:vedde (luerncep sa^BelJr) Jo e)eretswa$ Saro; Sutpuerdsrc )xeJe'sqnlqs pezrstunrpelll lo IIETUS

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s1!do s ttt d puti o tpl sn MDN's {rpDt souI 2L1' etd 1 I 82 Nuyrsiavol. I I, No. I (1996)

Table 1 Comparison of the geographicaldistribution and morphologicalcharacteristics of the five genera of tribe Physopsideae.

Mallophora Dicrastylis Physopsis Newcastelia ltchnostachys

Distribution main area south-west south-west south-west arid zone south-west & arid zone Leaves upper surface bullate rugoseto smoothto bullate bullate bullate rugose glandular absent usually usually absent absent dendritichairs absent present

Flowers no. ofparts 4(5) 4,s(6) t\ (4)5-8 (4)s-10(12)

Calyx tube length < lobes S lobes > lobes > lobes > lobes

Corolla colour white white or yellow or puryle,blue purple coloured white or white lobes obvious obvious obvious obvious absentor inconspicuous lobesapex obtuse obtuseor broadly attenuate to acute to subacute obtuse caudate truncate lobesmidvein+ not rarely not prominent prominent prominent prominent

Style exsertion exserted exserted included included or exserted exserted indumentum denddtic dendritic absentor absent absent,simple simple or 2-branched apex distinctly branched distinctly notched or notched lobedor lobed slightly branched lobed

Mature ovary ovules/cell I

* Themidvein is termed 'prominent' when it ismore obvious orextending much further than the lateral veins and 'not prominent' whenthere is a Drominentreticulato Dattem ofboth tvDesofveins or whenthe veins are not visible. 0I6I eroJequro{ 6tcpq3rq^\}o Ip'spro.ar lnJ}qnop olucpul ol pesnerE sloqu^s uedo '7 '1 L'ltolrsDqra^ n^ DltoJi^Dqnr pue f owlnrtuod ru^DllotrDqla^ 7 - q:Dat!3tura{ J'. a o{LtoqouaT pue '7 pue j-V r'fi1rr]sor{rryJo uotnqutslp eln8ld a DftalrDrq 7-g: a rlsualp.rD8loor f suDilq, setx?ils FcqdrfoaD I

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NuytsiiVol. ll. No.1(1096)

Key to Lachnostqcftys species

1. Leaves narowly ovate to almost circular, not decurrent(the stemsterete) 2. Leaves usually acute;upper surfaceshallowly bullate. Inside of corolla tube glabrous. Ovary hairy but not glandular...... L, albicans 2. Leav€s usually obtuse;upper surfacedeeply bullate. Inside of corolla tube with dendritic hairs. Ovary both hairy and glandular 3. Bracts ovate or broadly ovate, with a purple or brown indumentum. Flowers (6)7-10 merous; corolla tube hairy inside almost to the sunmit ...... L. ferruginea 3. Bracts narrowly ovate or ovate, with a white or off-white indumentum. Flowers (4)5-7(8)-merous;corolla tube usually glabrous at summit inside (although glabrous portion may be overlaid with hairs that are inserted lower in the tube).. L. verbascifolia 1. Leaves linear and/or decu ent (the stems4 ridged or appearing4-angled) 4. Leaves narowly oblong to ovate: upper surfacedensely hairy. Flowers 5- or 6-merous,with dendritic hairs inside corolla tube. Style glabrous. Occurringin the Sandstone-PaynesFind-Coolgardie area ...... L. coolgardiensis 4. Leaves linear; upper surfacebecoming glabrous on the tuberculateor rugosesummits of the bullae. Flowers7-12-merous, with simpleor 2-branchedhairs inside corolla tube. Stylehairy nearbase. Occurring from Murchison River south eastto Lake King 5. Flower spikes severalto many in an opposite-decussatearrangement terminating each branchlet,olten becominginterrupted. Inside of corolla glabrousexcept near base...... L. eriobotrya 5. Flowers spikes solitary (each on a separateleafy branchlet),dense. Insideof corollahairy throughout...... L, bracteosa

Lachnostachysalbicans Hook. (Hooker l84l: t.414). Type: Swan River Colony lwestern Australial,1839, J. Drummonds.n. (holo: K n.v.).

Illustrations.Diels & Pritzel(1904: Figure 57,4,8); Munir (1978:Figure 48).

Description. Munir (1978: 646 650). Additionalcharacters: leaves with simpleglandular hairs mixed with longer non-glandulardendritic hairs; corolla lobes shallowlytriangular or depressed ovate.0.3-0.6 mm lone.

Selectedspecimens examined. WESTERN AUSTRALIA: 28 km W of Lake Grace, Tarin Rock Reserve,2 Oct. 1984,J.M. Brown 306; FrazerRd, Banjup,18 Sep. 1979,R. Cqrey Badgingarra National Park, 8 km W along CaddaRd off Brand Highway, 8 Oct. 1985,M. Carter 40'7;2 km W of Muja, Collie region,9 Oct. 1981,C."/. Keighery 4083;5 km E of Tincurrin,26 Oct. 1982,K.H. Rechinger59106.

Distribution. Extendsfiorn Badgingarrasouth to Collie and south-eastto Tarin Rock, in the South- westBotanical Province of WesternAustralia.

Habitat. Occvs in sandysoils, commonly over laterite,in Eucalyptusmarginata forest on the eastern sideof the DarlingRange and associatedhills, or in shrublandsin otherareas. (euozJ€luIu.retsa,{\ '(3cur^ord -qlnos aql ur)'raplnog ot ts€eqlnos puBIrBd IEuortENurqq€roog ot qlnos lDcluElog '?uotspues lse,r-qlnos aql eprsurlsnl) {rea13 ol lse,trqlnos (acul^ord l?Jluvtog useDuIeJAeql uI 'PIlu4snV Jo quou) uortuls ee8ptg uor; Sutpualxe ujelsel\ uJeqlnosuI sJnax) uollnqltJslo

'gp9 '0961 'purg lacotg 7, ^lnf 0I seu,4e4;oS tx{ 9 9 :0288bqua11 y y 'I86I 'peetsetuoH '8861 'uot1u15 des /t rpInEIJoAUqBZ'.\LELna{uDr) f A deg71 )lJouSurllEi[ '.lZ6I 'serzueti{lo uuo) Tg'9861 dos € S uq 9Z r'e1yroo81ey3o5iru1 ,Ol'$a og g'2661 d'as8Z 's)rou aurqlrpuD lo N [ur{ 9 0] lu 009 ;VI'lvUISnV NUAJSA,^N paulwora suawDadspaQaPS

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'(uees '[BIlP4snV uolt€rtsnlll'^ ?r869901 A\SN) Jrls42l,1J 7'0061 'lJrJtsrp uratsalAl orpre8tooJ :adKJ (O0l:€061 erooyO aloot{ S s.rsua.rpJeSloors,{qeulsouqcul

'lues?Jd lE Isrr tu aq ol pereprsuocloN si/lrJs uollDMasuo)

'rsqtueceq reqrueldag pouad 8uua$oll

'spuElqruqsur pepJoceJ'la,ler8 cquelel qlr,r Jo glrJelvluo uauo'slros ^puts ur sJncao iDJ?gDl1

'ErlEISnV urelssid Jo 3cur^ord lDcIu€]oglse,l\-qlnos eql '8ur) ur e{p'I ot lsBe-qtnospu€ uruorJ a)€-I Jo q ou ol tsEeuep,{H leeu uIo.IJspue$g uollnqu$lo

'Drqn! 'v'l 'w '99L2 x Duanwv uoqg t o '986I '.96L8 des,'p€o1 ep8ep,reNo1 3ury e1e1eq1qtl\\ pd ooJEJJo uorlresralul Kaq aN UX 'I86I 'uruorJ des €I e{u.I Jo N url 0I :VI'MJS6V NdAJSa,46 pawruuo suaulrads papaps

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'satcadss(t1tolsoutlrr? Jeqlo ur uuq] snor,\qoerou ,{leraueB ere qcrq,{ 'seqol Elloroc 'pa?nper,{ra,r q8noq}lu 'truqsrp sDqs,{u,tp sercedssrql satoN

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!t\.1o!\qdpv nptcDJuN iqru.au4 'Dl e\\-tE 86 NuytsiaVol.11,No. 1 (1996)

Hqbitat. Ocg1asin sandysoils ofred, yellow or othercolours, on sandplains,dunes or rarely on lateritic rises, in spinifex grasslandsor in shrublands,sometimes in open woodlands.

Flowering period. June-October.

Consenation status. Not consideredto be at dsk

Nofes. Previouslytwoforms of I'achnostachyscoolgardiensls were recognised, f. brevispicqtahaving broaderleaves and larger spikesthan f. coolgardiensis(Munir 1919:645). As in many other species, there is a tendency for leaf size and inflorescencesize to be corelated, but both charactersshow continuousvariation with no clearcutdivision into forms. Such charactersmight also be expectedto vary greatly within populationsand individuals partly as a result of minfall fluctuationsfrom year to year, which can be extreme in arid environments

Lachnostachys eriobotrya (F. Muell.) Druce (Druce 1917: 630). - Walcottia eriobotrya F. Muell (Mueller 1859: 241-242).- lachnostachJswalcottiiF.Muell. (Mueller 1861: 140). rype: Murchison River, Western Australia, P. Wqlcott (lecto: MEL 41166,Jide Munir (1978: 673))

Lachnost.tchysrodwayana W. Fitzg. (Fitzgerald l9O4: 29). Type: Atino, Western Australia, September1903, W.V. Fitzgerald (PER'IH 01607367)

PycnolachneledlfuliaTtrcz. (Turczaninow 1863: 214-215) Iyper SwanRiver [Westem Australia], J. Drummondcoll.'7, n.220 (holo: KW 4.v.,photograph PERTH 04134559).

Illustrations. Diels & Pdtzel (1904: Figure 57C-E); Munir (i978: Figure 53)

Description. Munir (1978: 672-679). Additional characte$: leavesoften decurent; corolla lobes absentor depressedovate and up to 0.6 mm long; stamens7-12, most commonly 8-10.

Selectedspecimens examined. WESTERN AUSTRALIA: 10km S ofEneabbaalongBrand Highway' 24 lan. 19'79,B. Barnsley 893; Man Rd, Watheroo National Park, I I Oct. 1985' M. Cattet 456; 13.2 km W of Coomberdale,on CoomberdaleWest Rd, 17 Oct. 1'993,B. Conn 3851 & M.E. Tozer', Port Gregory road, 14.6 km NW of Northampton, 27 Sep. 1985, N. HoyLe473; Eurady Station, 26 Sep. 1988,P. Roberts9O7.

Distfibution. occurs mainly in the south-west Botanical Province of westem Australia, extending from north ofEurady Station(near Murchison River) south-eastto nearMoora, with an isolatedrecord from Kellerbenin. There are also a few recordsfrom inland areas,including Mount Magnet and Sturt Meadows Station, in the EremaeanBotanical Province

Habitat. Occursin sandysoils, often over latedte,in low to tall shrublandsor woodlands,sometimes dominatedby Banksia or Eucalyptr{Jspecies.

Flowering period. August-November.

Conservationstatus. Not consideredto be at risk. 'J 'eul€u ur Suoyeq seop'Dllolsntqo oJplnJluDd rB^ sD IeleID^ srql repun ,{lpur8r:o peqlJcsep 'Je,\e,roH 'otloftJsDqiaa uox"l e oau8nttal 7 1o eBueru,,nou1 eqt puo^eq lle,t srnJJo )r pue 7 seqqeurslcadser:eqlo llE uI lnq unlusrunput x.{1ecgoqs qt urDauISnuaI 7 seq)tew otolnttuod tc11 'e.req pesropueuoucaJJoa e'uptr)soqn^ Z sp (€068092nquv'A 3 a) uoxel stqt1o suaurtrads '^llu€ceJ lure,\?s peurulletepeJs€q Jruntri eJol,^l (8/6 1) Jlunt{ ,{q sercedsslql o1 perleJsuurlse^\ 'otyo!:twqtat 'DtDl T sercedspe1e1er ,{1eso1c eql tepunpequcsep,(lluu t8uo se,lrqJt q^\ n)luDd Jut\zluluu 'oaut8nttal lElerJEAeql sKt[co\souq)t/llo srx,{uou,{ssE e^oqB patsrl sulloJ eql ol uoDIppEuI .taloN

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NuytsiaVol. 1 l, No.I (1996)

L. ferruginea, rtrs from an areawhere therehave been many collectionsof L ?rruginea but norleof L. verbascifolia.

Of the forms previouslyrecognized in llu chnostachysferruginea, f. acutfolia andf. reticul.tta arc separatedfrom f. /erruginea by leaf characters,v./hile f . obtusifolid is separatedby its inflorescence consistingof more spikes.Two of the taxa,f. obt6ifolia and,f. reticulata, werc namedoriginally from single specimens.since then,other specimensof both taxahave been collected from the type localities or nearby,as well as intermediatespecimens from theseor other sites. As lnchnostachysferruginea showscontinuous variation in eachof the charactersused to distinguishthe forms, with considerable vadation apparentlyoccuning within populations,there appears to be nojustification for maintaining theseforms as distinct entities.

Lschnostachys verbascifulia is very closely related to L. ferruginea and, owing to the great variability of thesetaxa, particularly the former, key charactersthat are reliable throughoutthe range of both speciesare hard to find. Apart from the differencesgiven in the key, l. verbascfolia can generally be distinguishedfrom L. ferruginea by its longer leaveswith a greaterlength/width ratio. Although both species have inflorescencesvarying from one to many spikes terminating each branchlet,l,. verbascfolia almostinvariably has more than one spike per branchletwhereas solitary spikes are fairly common in L. feruginea. Of the two varieties of L. verbasclfulia, only var. verbascifulia o'terlapsin geographicalrar'ge with L.ferruginea, andthis varietyis readily distinguished by its longer calyx indumentum with fine antrorsebranches on the dendritic hairs.

yerbascifolia Lachnostachys F. Muell. (Mueller 1868: 158-159). Zlpe. lwestern Australia], J. Drummondcoll.5, n.237 (lecto:MEL 41202,fide Munir (1978;663)).

Description.Munir (1978: 663 669). Additional characters:flowem (4)5-7(8)-merous;calyx indumentum white or rarely very pale ferruginous.

Dist bution. Occurs mainly in the South-westBotanical Province of Westem Australia, extending from Pinjar south to Collie and Cranbrook and from there east to near Mt Coombaninya(south of Balladonia) and Mt Ragged. Also extendsinland to Noongal Station (nearyalgoo) in the Eremaean Botanical Province, with a few other doubtlul inland records. llablrat Occursin sandysoils, rarely associatedwith laterite,in shrublandsor woodlands,sometimes dominated by Eacall2lls species.

Floweringperiod. June-November.

Nores. Specimensfrom the north (in the Pinjar-Yalgoo-WonganHills area) always have a long indumentum on the stems and leayes, whereasthose from the south-east(from near Grass patch eastwards)have a relatively short indumentum,and there is a mixture of short- and long-haired specimensin intermediateareas. The south-easternvariant differs from other variantsof the species in its shorter calyx indumentum with more spreadingbranches on the dendritic hairs; it has been recognized both at the species level, as lnchnostachys dempsteri, and at the varietal level, as L. verbascifulia yar. pqniculata. Varietal rank is curently accepted.

Mostflowers are5-7-merous,but occasional8-merous flowers have been observed on aoredominantlv 7 merousspecimen (A.5. George 6851) of yar.verbascifolia and 4-merousilowe,, ,,r. ", .orn-on u, (l g ernSIC:816I) runl/,{ -uotiDrtsnnl

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slsdos{UJ puc olt atsnftaN s{lpDtloulrr\'et}I 1 A 90 NuytsiaVol. 11,No. I (1996)

Selectedspecimens examined. WESTERN AUSTRALIA: 42 km NE of Coujinup Hill, 27 krn N of Rollands Rd on Carmody Rd, 25 June 1983, M.A. Burgman 1505 & S. McNee; Noongal Station, 9 Aug. 1993, S. Dawson 929;75 km NE of Ravensthorpe,4 km W of Dunn Swamp, 21 Sep. 1979, J. Taylor 167, M.D. Crisp & R. Jackson',E Perenlori, 25 July 1995,J. Trox.

Distribution. Extendsfrom Pinjar and Noongal Station(near Yalgoo) southto Collie and Cranbrook and south-eastto near the upper Young River.

Conservationst4trs. Widespreadand not consideredto be at risk.

Noles. Three O.H. Sargent specimensfrom SandSprings (south-eastof York) were examined,one labelled 909a (PERTH 03718247) and the other two unnumbered(PERTH 03717739, 03717992). Although thesewere collectedfrom the type locality of Lachnostachyscordifulia, they appearnot to belong to the type collection but to have been collectedshortly after the taxon was named,probably in 1924.

Newcastelia F. Muell.

Mueller (1857: 22). Type: Newcasteliacladotricha F. Muell.

Small or medium-sized shrubs, erect or spreading, with an indumentum of non-glandular dendritic hairs on the stems,leavesand calyces. Youngstem.r terete, densely hairy. Leavesopposite- decussateor rarely (occasionallyin N interrupta) in whorls of 3, sessileor shortly petiolate,entire; lower surfacewith a densepersistent indumentum of dendritic hairs; upper surfacebullate, covered with dendritic hairs,often also with sessileglands or simpleglandular hairs but thesegenerally hidden by the longer dendrrtlchairs. Inflorescenceof redtced, almost sessilecymes ananged in solitary or multiple spikes or dense heads terminating leafy branchlets; spikes dense in bud, sometimes developingshort or long internodesbetween the adjacentpairs ofcymes at maturity: cymes3-flowered or (1nN.cladotricha) reducedto a solitaryflower, in opposite-decussatepain. Flowers (4)5-8-merous, with equal numbersof calyx lobes,corolla lobes and stamens.Calyx denselydendritic-hairy outside, glabrousinside; lobes sho er than or rarely as long ascalyx tube. Corolla usually pale to deeppurple or blue, rarely white, glabrousoutside; tube longerthan or aboutas long asthe lobes,with a denselong antrorse indumentum inside but often padially glabrous; lobes equal, spreading,very nanowly to broadly ovate, glabrous,either nanrowthroughout or attenuateto a slenderterminal protrusion,thin- textured,with the midvein prominentand commonlya few lateralveins clearly visible, the main lateral veins becomingmore or lessparallel to the midvein towardsthe apex;terminal protrusionof eachlobe recurved or spreading,the apex obtuseand with a recurved margin. Stdmelr.ralternating with the corolla lobes,included to greatly exserted,glabrous; filament insertedin upper part of corolla tube or at summit of tube, expandedat the baseor slenderthroughout. Ovary 4-celled, with 1 ovule in each cell. Sty/e included to greatly exserted,glabrous; apex notched or slightly 2Jobed. Fruit dry, indehiscent, 1(2) seeded,enclosed in the persistentcalyx.

A genusof nine species,occuning primarily in arid regions,extending from Eighty Mile Beach in Western Austalia eastacross central Australia (Northern Territory and South Australia) to south- easternQueensland. In WesternAustralia, it occursmainly in the EremaeanBotanical Province,but also occursin the southernpart ofthe Northern BotanicalProvince and in the South-westernInterzone (Figures 2-4). Two species,Ne$,c6steliq interrupta Munir and N. velutina Munir, are endemic to Queensland;the remaining speciesall occur in WesternAustralia, where three of them are endemic. 'DrlrlJtotpods'onatrDr aN g'Drorldopt4?olptsorMrN - V srlltsnv urets.iA ur uollnqutsrc Z rnBLC

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This genusis closely relatedboth to l,achnostachJsard Physopris,but can be distinguishedftom the fomer by the number of ovary cells, ftom the latter by leaf charactersand from both generaby corolla characters.Newcasteliaspeeies differ ftom the othergenera in thattheir corollalobesare either narower tfuoughout or have a narrow apical protrusion

Key to Newcdsrelia species

1. Stamensinserted below summit of corolla tube and included or barely exserted;filament scarcelyenlarged towards the base. Style included to sho ly exse ed 2. Inflorescencewith long white or pink to purple hairs giving the flowers a shaggyappearance, each bract subtending I flower. Ovary glabrous, with sessileglands on summit. Occuring in Western Australia and No hem Territory N' cladotricha 2. Inflorescencewith short white hai$, each bract usually subtending 3 flowers. Ovary hairy on summit, sometimesalso with sessileglands. Occurring in Queensland. 3. Corolla tube hairy insid€ except at the base. Ovary lacking glands . ... . N' interrupta 3. Corolla tube hairy inside in basal half but not distal half. Ovary with sessileglands N' velutina 1. Stamensinserted at or just below summit of corolla tube, greatly exserted; filament gradually expandingtowards the base. Style greatly exserted' 4. Flowers usually 5 merous. Ovary denselyglandular throughout' glabrous' 5. Flowers usually in globular to obloid heads,rarely in densespikes Calyx indumentumc. 2 mm long, with two types of hairs of very differentsize, the largehairs with long anhorsebranches ...... N' cephalantha 5. Flowers in short to long, denseor interupted spikes Calyx indumentum 0.5-1 mm long; hairs variable in size but not of two distinct types, with short spreading branches 6. Leaf undersurfacewith a continuousdense indumentum. Inllorescenca of usually more or less continuousspikes, with a white or off-white indumentum N' sPodiotricha 6. Leaf undersurfacewith hairs concentratedalong the veins, the interyeinal spacesglabrous. Inflorescenc9of usually distinctly interruptedspikes, with a pink to red indumentumon the calyces N' roseoazurea 4. Flowers usually 6-8-merous. Ovary either glandular only on the summit or densely hairy at first in upper half (but glabrescentin fruit) 7. Leaves very narrowly ovate. Flower headsor spikessubglobular to shortly cylindric, with a bright yellow indumentumand white or lilac corollas. Bracts sessile,very narrowly ovate...... N. insignis 7. Leaves nanowly ovate to narrowly elliptic. Flower spikescylindric, with a white or off-white indumentumand purple corollas. Bracts very shortly petiolate,ovate or broadly ovate 8. Corolla lobes rather suddenlynarrowed to apical point Ovary g)abrous, with sessileglands on summit...... N. bracteosa 8. Corolla lobes very gradually attenuateto apical point. Ovary denselyhairy at first in upperhalf but glabrescentin fruit, alsowith sessileglands ...... N. hexarrhena .(900tt tgl{ :opq) satlD !,,lErlEr]snvqlnosl 'eBIO ty{ pue re^rd eueqlv uee,rleg :adtJ (V :9 L81Jellent{) Ilent{ d Bqguepqdar ellalseoarap

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st \dor{Id pue otpt sor^raN !&IJD ts.'uL! J rr l' r \\ 1 g 96 Nuytsiavol. 11.No. 1 (1996)

Newcasteli{!cephalantha vnt. oblongaMunir (Munir 1978: 634-635) ?ype; Hann Range,70 miles 1119kml north of Alice Springs,Northern Territory, 1 June 1"962,D.J. Nelson 372 (holo: AD n.v.; lso: PERTH 01607375).

Newcastelia cephalantha ,tar. queenslandica Domin (Domin 1928: ll01) Iype: Windorah, Queensland,1910, W. Rose (holo: PR 530702 n.v.).

Newcqstelia ceph.llantha var. tephropeplaMwir (Munir 1978: 635-636). I)pe About 46 km west of Musgrave Park Homestead,along tack to Mt Davies,western portion of MusgraveRanges, South Australia, 6 September1.963, Hj. Eichler 1'731O(holo: AD n.v.).

Illustrations.Diels & Pritzel(1904: Figure 56H,J); Munir (1978:Figure 45); Munir (1981:Figure 399); Munir (1986:Figure 5508).

Description. Munir (1978: 629'636).

Selectedspecimens ewtmined. WESTERN AUSTRALIA: Rudall River region,Iune 7987,R.P. Hart 648; 48 km N of Wiluna on road to Neds Creek,12 Sep. 1987,J.W. Green5355; ParooStation' 26 July 1983,A.A. Mitchell 1140;18 krn E of WoolnoughRange, 30 June 1984,G.J. Morse 224; 13 km N of Kumarina Roadhouse,l0 Aug. 1980, C I. Stqcey670.

Distribution. In Western Austmlia, distributedin the EremaeanBotanical Provinca' extendingfiom Telfer Mining Centre south to Albion Downs station (south of wiluna) and from Kumarina Station east to the border with Northern Territory. Also occurs in Northern Territory, South Australia and Queens)and.

Habitat. ()cctts in red sandy soils with spinifex grasslands,sometimes dominated by shrubs, cornmonly Acacia species,or by -Eucalyptusspecies with a mallee habit.

FIowe ring period. June-October.

Consenqtion status. Widely distributed and not consideredto be at risk

Nole.r. The varietiesof Newcasteliacephalantha arc r'olonger recognizedin WesternAustralia. The indumentumcolours and leaf measurementsused to distingtish var. tephropeplaJtomthe other two varieties are conflicting on some of the PERTH specimensannotated by Munir; for example A.S. George 9070 is a largeJeavedspecimen included under var. cephaLanthabut has white hairs rather than ferruginous ones. Var. oblonga was distinguishedfrom var. cephaLan/ia by its larger leaves and flower heads but there is considerableoverlap in these characterson the annotated specimens of the two varieties, a situation similar to that discussed above for the forms of lac hnos tac hy s co o Iga rdi en s i s.

Newcastelia cladotricha F. Muell. (Mueller 1851:22). Zlpe; Sturt's lsturt] Creek, towards Mt Mueller, [Western Australia], March 1856, F. MueIIer (holo: MEL 41017).

Illustration. Munir (1978:Figure 4l).

Description. Munir (1978: 615-619t. '(028r09I0 HJuad '(929 :oltalo.sr:uees uoqpJtsnllr :8t6I) JIUryNapq'080IZZ6L qy :oJral)6VBpztlrd ?'I06I Jaqolco 'erplsnv 'selzuehtr uretse,l!\ :adq (909:t06I Iszlrld T' sler() Ieaud g qu31su1ellalsBJ^{aN

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slsdost1t.Ipu€ Dlt a$or^a7i'stqcolsouqtn'a(>j 1 g 98 NuytsiaVol.11, No. I (1996)

Illustrations. Diels & Pritzel (1904: Figure 56E-G); Munir (1978: Figure 44).

Description. Munir (1978: 626-629). Additional characten: corolla Fobably varying frorn pale puryle to white, the lobes nanowly obtuseor acute.

Selectedspecimens examined. WESTERN AUSTRALIA: Doney Lagoon,Adelong Station, 22 Sep.1988, R.J. Cranfield 7590; 2.5 km NEof CometVale, 16Oct. 1987, G.J Keighery9680; 6.5 km SE of ArsusCorner, 26 Nov. 1986,D J. Pearson7094.

Distribution. Endemic to Westem Australia, reaorded fiom the Comet Vale area and from the northern boundary of QueenVictoria Spring Nature Reserve,the two areasabout 250 km apart.

Habitat. Ocaursin red, or occasionallyyellow, sandysoils. Recordedin spinifex grasslandsand/or in shrublands,sometimes dominated by Acacia ot Eucalyptus specLes.

Flowering period. September-NoYemb€r.

Conservattonstatus. CALM ConservationCodes for WesternAustralian Flora: Pliority 3. Recorded from about four localities.

"yellow Noles. Although the corolla colour is given as or whitish-yellow" in Munir ( 1978:628), there "lilac are only two specimensat PERTH with a tabel indicating corolla colour, and this is recordedas "white" or white" (CA. Gardner 11104) and (G.J. Keighery 9680). The bright yellow hairs on the calyx give the flowers and fruits on the dried specimensa strong yellow appearanceoverall, but the dried corolla colour is pale to medium brown and there does appearto be a slight purplish tinge on some specrmgns.

Newcastelia roseoazurea Rye, sp. /lov.

Newcasteliaespodiotrochae alt\nis sedindumento folii paginaeinferioris minusdenso, indumento alabrstrorumfu\co rosecvel rubro.

Typus:South of RudallRiver camp, Western Australia, 12 August 1971' Wilson 104'15(.holo: PERTH 03771148; iso: CANB, MEL, NSW).

Shrubs 0.3-1 m high, often rounded,with an indumentumof patent dendritic hairs on the stems, leavesand parts ofthe flowers, the many-branchedhairs often broaderthar long Stemindumentum 0.5-0.8 mrn long, white or ferruginouson the vegetativestems and pinkish on the inflorescenceaxes tedves opposite-decussate,sessile or very shortly petiolate,ovate to very broadly ovate or oblong- elliptic,8-38 x 4-19 mm, acuteor obtuse,with recurvedmargins, green below the indumentumbut often tinted by the white or ferruginouscolour ofthe indumentum,the upper leaf surfaceusually grey- green throughout or grey-greentowards the baseand ferruginoustowards the apex; lower surface reticulate-veined,with hairs concentratedalong the veins and with scatteredsessile glands, the veins almost white or yellow to pale ferruginousiupper surfacedeeply bullate, with severalhairs towards the summit of each bulla; hairs 0.2 0.5 mm long on upper surface,slightly longer on lower surface Inflorescencerp to 200 mm long, usuallycomprising a terminal spike and two or more arillary spikes in an opposite-decussatealrangement, rarely (in depauperatespecimens) reduced to a single terminal spike typically 25-35 mm long; pedunclesup to 50 mm long but usually5-20 mm; spikesup to 'DaJIZDOASOt N 'peldnre]ur ueql soqolplloroJ ro,^\orlPurltr^ sro,^\oU.JeBuol sEq ,{lpnsn ,Lplrlolpods N pu€ ,{ltcurlsrp 'snonurguoc ,{1pnsnsr acuecsaroL+ureq)oalnzooasol N ur s€aleg,r ssalro aroru,{1pnsn sr ecuecseJol+uI eqt Dq)uplpods N uI spnq J€,ro[+ €l{t uo untueunpur elrq,r JJo .ro e]rq,r pup ecpJrnsrepun yuey,{rruq ,{lasueperou slr ur sJoJJrpq"rq/ 'oqcltlorpods t2IaJsDJ$aNol pel€leJ ,{lesolJ ,taloN

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-t 'ttJo.{4d pw:ntl)ttD$aN :.(4'DtlouPvt ,Ad E 100 NuytsicVol. I l. No.I (1996)

Figwe 5 - Netrca$elid roseoazurea. A - floweilngstem (x I ), B - portion of undersurface ofleaf (x9), C portion of upper surface ofleaf(x9), D -enlargementofbulla€ (sideview) from uppersurface of leaf (x9), E - topandside views offlower(x4), F stamen and adjacentcorollalobes(x6). Drawn from R.P d.lrt 977.

Newcasteliaspodiotricha F. Muell.(Mueller 1862: 21 22). Type:Between Victoria River and Gulf of Caryentaria,lNorthern Territory], J. McDouallStuqrt (holo: MEL 41041)

Illustrations.Munir (1978: Figure 38); Munir (1981: Figure 401); Munir (1986: Figure 550C).

Description.Munir (1978: 603-609).

Selectedspecimens examined. WESTERN AUSTRALIA: Trackto RedBluff, S of MoffettahWell' 9Dec.1991.,H.N. Foole 300; Gardner Range, 190 km SEof HallsCreek, 6 July 1995,K. Coate3188; J^€q tTtdor'(qd (srleq r€lnpue[3eldurs s" II3,^\s3) s.rleqcl]upuep JEInputlSe,\Bq ol pe^J?sqou3aq lo tuorJ pu" "(o{ cuaue8 eql ul sarceds1y sialce:eqcpuoDtppB l€Jeles pu? es"ql Kq skqDlsouqrrry rq uPJstsdosttl6 pesn sJetcpJ€qcelloJoc puu JBaleql f,q '\1weu m1alsrtcualguror; peqstn8uttsrp

'rg JJnBrlrJJut^old lcJtueloB ucJEuJl3 aqt.1o sceLc luorefpn olur ,4pq8qsspuetxe tnq euozJeluluJelse,\\-qlnoS puu ecul^oJdlecluclog lsa'{\-qlnoSeql uI ^lrll?lulJo 'selJad:^ sJncco s?sda,t&/d'EllE.IlsnV uJalsel\ Jo ls3,{\-qlnoseql ol clur€puo J^U Jo snu33 V

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'puEtsueenopue erlellsnVqlnos ^Jolr.uJl uJJquoNur sJnccooslv (eSuua uognqml6Jo lsa.&-qlnos)s3loq1con uu33n1^tr reau ol qlnos scuL\o.IdlE3luelog ueeurueJaeqr;o ged uJelsre-quoueq1 q8norqt (ecul^ord lecluulog ulaquoN Jql Jo lrcd uraqlnos 'EIIPllsnV uolnqlJlsl(f aqt ur sa8uuAre8pg Jo quou) $lcod euroJCleeu u]o4 spuelxe uJe]selduI

'601 '9861 'seBuEU 's{3ou 3ulll!to|Z lco 9 IPBpg 'uot1e15 aruorgsp:e.uol SutssorJ leer3,{lF8?eCJo ldur{ t S :62lI .raddog q'5'9961 dag 97 sumo6 'uorSeX rerdurequo arog qSnHrhtrI oNJo IASSu;q 89'2'.168rnH'd U'066I rc1r167 rarq llepng

g IOI s1sdost4Jpwr olp|sDrt aN'siu2o1souq2try'ef'x1 102 NuytsiaVol. 11, No. 1 (1996) on very young leaves,usually on the upper suface or margin of the lanina, although thesemay not be present on all specimens. Such hairs are rare in the remainder of the family but do occur in DicrasfJlis incanaMunir (tdbe Physopsideae)and Pityrodia scabraA.S. George(tribe Chloantheae). Sessileglands and simple glandularhairs are commonly found on very young leavesin other genera but are generally not obvious becauseof the much larger non-glandulardendritic hairs.

Other characterspresent in Pftysopsisbut uncommonin the remainderof tribe Physopsideae,are the relatively smooth and glabrous upper surface of mature leaves, the very short stamens,the filaments being more reducedrn Physopsischrysophylla and P. spicatq than in any other members of the tribe, and the short styl€ with a distinctly lobed apex.

K€y to Pft)sopsis species

1. Leaves with a very short petiole 0.5-3 mm iong (petiole may only be clearly visible when the leaf is removed from the stem); upper surfaceof lamina usually fairly flat, the margins often narrowly recurved. Flowers 1 per bmct. Stamenssessile or subsessile.Style basewith sessileglands or glandular hairs 2. Leayes with a white indumentumon undersurface. Inflorescencewith a white indumentum and yellow corollas. Calyx lobes much shorter than tube. Corolla tube 4-7 mm long; lobes 4, with small simple glandular hairs inside towardsthe base...... P. spicata 2. Leaves with a yellow indumentumon undersurface. Inflorescencewith both the indumentum and corollas yellow. Calyx lobes slightly shorterthan tub€. Corolla tube c. 2 mm long; lobes (4)5, with long simple non-glandularhairs insidetowards base ...... P. chrysophylla 1. Leaves sessile,with strongly recurvedto revolute margins. Flowers mostly 3 per bract. Stamenswith a short filament 0.3-0.5 mm long. Style glabrous, not glandular 3. Leaves mostly opposite,narrowly ovate to ovate or elliptic. Corolla tube hairy insidetowards the base;lobes 4 ...... P. lachnostachya 3. Leaves mostly in whorls of 3, more or less linear. Corolla tube hairy inside throughout; lobes (4)5 4. Stems,leaves and flowers with a bright yellow indumentum. Calyx indumentum0.4-0.5 mm long ...... P. chrysotricha 4. Stems,leaves and flowers with a white or off-white indumentum. Calyx indumentum0.7-1.2 mm long...... P. viscida

Physopsis chrysophylla (C.A. Gardner)Rye, comb. nov.

Newcasteliq chrysophylla C.A. Gardner (Gardner 1964: 62). Type: About 45 km N of Murchison River, Western Australia, 3 January 1959, C.A. Gardner 12054 (lecto: PEP''|H 01607804, here designated;isolecto: CANB, MEL, PERTH 01607812).

Illustration. Munir (1978: Figure 40).

Description. Munir (1978: 612 615). Additional characte$: calyx indumentum 0.5-l mm long; corolla apparentlyyellow, the lobes very prominently reticulate-veined. 'd eq ',{a1 eqt uI pesn sJelcEreqJ3ql ruo{ l.l€dv oq)tJ|osfuq) d ol ueql o|or|ds ol ^lu€llulls 'sercads rel€eJ8,roqs ol sIl]adde op(4tlosfuqc 2'q8noqlp o,t1 eq1uee,tleq dtqsuollElaloqt suJ.IIJuoc fpnls srql peJeJJIpBxBt o,lr1 eql ^\oq ^luo Suqou :elleq slq] loJ suosu3lsnl uo 3lBroq€lelou plp lnq 'DqJuwsfuqJ (slsdostrryJ,tou) D4a$p)4 aN ol peleleJsP,^ selceds srql lEql pelDts(t961) leuppC

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80r sls do sKq d puaDu a$nr Aa N'stq ro tsoq r al' ef,ij'l g 104 Nuytsiavol. 11,No. 1(1996) similarities betweenP. chrysophylLaand P. spicata rncludetlteir long corolla lobes, which are hairy on the thickened base, and their ovary, which is either hahy thoughout or with hairs extending somewhatbelow the densely glandular summit. The other three Physopsisspecies have glabrous corolla lobes and the ovary with a mixture of hairs and sessileglands only on the surnmit.

Physopsis chrysotricha (F. Muell.) Rye, comb. nov.

Newcastelio chrysoticha F. Muell. (Mueller 1876: 15-16). Z)pe. Near Victoria Springs lQueen Victoria Spring, Westem Australial, "/. Young(lecto: MEL 41013,fide Mtnir (1978: 636); isolecto: MEL 41012).

Illustration. Munir (1978: Figure 46).

Description. Munir (1978: 636-639). Additional characters:calyx indumentum0.4-0.5 mm long; corolla colour not recorded but almost certain to be either yellow or white, the lobes prominently reticulate-veined.

Distribution. Recorded from the south-eastof Great Victoria Desert in the EremaeanBotanical Province and ftom nearQueen Victoria Spring in the South-westemInterzone of WesternAustlalia.

Habitat. Nohab\tat detailsrecorded, but the speciesoccurs in a more arid enyironmentthan any other membersof the genus.

Flowering period. September.

Conseruationstqtus. CALM ConservationCodes for WesternAustralian Flora: Pdority l Pi)sopsis chrysotricha has not been collected since 1891, suggestingthat it is very rare or extinct. However, the absenceof recentcollections may be due partly to the remotenessof the areawhere the taxon has been recorded; a field survey specifically for this speciesis clearly needed.

Noler. Very closely relatedto Phlsopsls vlsclda, which possibly should be regardedas a subspecies of P. chrysotricha, but readily distinguishedby its bright yellow indumentum. P. chry;otricha also appearsto have more leafy and more interruptedspikes with a shorter indumentum. However, it is known ftom only two collections,neither of which is in fruit. More material is neededof this very ooorlv known taxon to determinewhether thesecharacter differencas are reliable.

Physopsislachnostachya C.A. Gardner(Gardner 1939; t.3384). Type:Gravelly hills, Kukerin, westem Australia,9 November1935, C.A. Gardner s.n. (holo; PERTH01607863; probable iso: PERTH01607855).

Illustrations. Gardner(1939: Tablet 3384); Munir (1978:Figure 35).

Description. Munir (1978:585-688). Additionalcharacte$: calyx indumentum1.2-1.8 mrn long; corolla lobes prominently reticulate-veined.

Selectedspecimens examined. WESTERN AUSTRALIA: Tarin Rock Reserve,28 km W of Lake Grace, 2 Oct. 1984, l.M. Brown 305;22 km S of Hyden on road to Newdegate, 18 Sep. 1984, D.B.Foreman1T5;23 km S ofHyden,23Nov. i985, D.B.Foreman 1177,Kananda Farm, Kulin East, 24 Ocl 1994. Kulin Kondinin Herbarium 14. 'luaserd 1u4srr 1eeq ol pereprsuocloN snioJsuouDMasuoo

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stsdos&ldpueotpts t dN s(qJotsou| 't/t'z()l 1A 106 NuytsiaVol.11, No. 1(1996)

Notes. This speciesdiffers from other membersof the genusin its very long calyx and corolla tube, its corolla lobeslacking obviousveins but havingglandularhairs atthe base,and alsoin the occurrence of hairs and sessileglands on the base of the style. Its nearestrelative appean to be Physopsis chrysophylla.

Physopsis viscida (E. Pdtzel) Rye, comb. nov.

Newcastelia viscidaE. Pritzel (Diels & Pdtzel 1904: 505). Type: Coolgardie goldfields, Westem Australia, October 1901, E. Pritzel 858 (lecto: AD, fide Mun[ (1978: 640), n.v., illustration seen; tso lect o : PEP.TH 01607847).

Illustrations.Diels & Pritzel(1904: Figure 56A-C); Munir (1978:Figure 47); Munir (1981:Figure 4O2).

Description. Munir (1978: 639-643). Additional characters:calyx indumentum0.7-1.2 mm long; corolla white, the lobes prominently reticulate-veined.

Selectedspecimens examined. WESTERN AUSTRALIA: c. 25 km W of Mulline on Menzies to Diemals road, 17 Sep. 1989, B.l. Conn 316l & J.A. Scolt; 7 km N of Emu Well, Adelong Station, 22 Sep. 1988, R."/. Cranfield 7578 c. 6 km NE of Bungalbin Hill, 28 Sep. 1995, B.J. Lepschi 2074; Melita Station, 4 Nov. 1988, H. PringLe2210; N of Leonora, Sep. 1994, T. Read'l .

Distribution. Recordedmainly trom the South-westInterzone, but extendingslightly north into the EremaeanBotanical Province of WesternAustralia. Extendsfrom north of Lake Barlee eastto near Elora Soak and south eastto Hampton Hills Station (south of Kalgoorlie).

Habitat. Occuxsmainly in red sandor sandyclay, also sometimesin yellow soils, recordedon rocky hillsides, sanddunes and on plains, often with lateriteor lateritic gravel, rarely associatedwith other types of rocks. The vegetation is usually recorded as shrublandsor woodlands dominated by Eucallptus or a variety of other genera.

Flowering period. Mainly September-November.

Conservationstatus. Not consideredto be at risk.

Notes. ln the protologue, Diels & Pritzel (1904: 505), not€d that this specieshas a similar lloral structure to Newcastelia (now Physopsls) chrysotricha, Newcastelia cladotricha ar'd Physopsis spicata. The first of theseis undoubt€dlyits closestrelative (see notes under P. chrysotrtcha),and all three speciesare similar to P. vlscida in having short stamensand style. Most Newcastelia species ^nd all lachnostqclys specieshave prominently exsertedstamens and style.

Discussion

The transfer of three prirnarily 5-merousspecies from Nqecastelie to Physopsis,has resultedin a clearer separationof the two genera. Previously,the sole chzuacterused to distinguishPrlsopJiJ and Newcasteli0, was the number of floral parts. This was, in any case, a somewhat incomplete difference,separating species with consistently4-merous flowers from thoseproducing mainly 5- but LZLt6l :lt)gt nocsot\satsrtunrrN durl .os llng lDldu.sepulenbsncnq eDllnuuou alerHodotrl\lla aeoccueqre^ (t981) N'^\oulutz.rnl '8t t ,ZlUZnor.or\srrrlcrnrrNdrrrlJoS llng LUnrullnuuou ulruenadssnqruoqducsep srlrsfpl runrotdgcsepuou anbsnrnqurnruucld unleu.S lrlYassi.ac (6'81) N'^toulutr7.rnl (:prctJpvrJruud r\o9) 0o1.081 ,id tuDd l.vrlclsnv qtnosJo Erotd,. (sPe) UH'urllso.LT 31'dossal:t4 aeer.qlueolqJtrlurJ (9861)VV'rlunll (treup^S:$toogpaaU)3ggt67dd..tqcasny1wu.JloBroH,,(pa)df'dossef:r./aco.eqlucolqJI6(1861)VV'rlunt"'l 269-aoi:l rruounlg oreplsdos,{tldqu .ee.xqluDolqJJo uolsl^er.ItLIouo xcl (8161)V'V rrunN ('eurnoqtew:reruud l^oD) 0l-l IoA ,. e[llrrlsnv erlqdrl3ol{qd.trlucruilrrC,,(9/8l 6981)C r0ll.nN iz-il:6 csrl,r{prDD ^\e) tog fs,ralooH xlretepe$rxadnu sn.doryrtul eEIlEDsnVs€vld uI seJorrDrlonbrrc scl.rrds]3xraue8 e^oN (LSsI) C'lellenl^l .6nZ 912.69 log t slu€lduDIFrsnV tser''\ r'ncN (lz6t)'r\01 S'croot{ 'l0t 86:I'rog I It ?lplsnv ulorJslurld.\\eN (t06I) N.l S'croolr',{ 09t-10' 1l:(6)sId uorls.relooH (lt8l) f A|r.looH -tg-ig :at rrry4snvuolsey'A ros.{oU rordt t IIIX slFruaplrroaxlFrlsnv aflou sauorl nqul uoJ (t961) VJlrupJDD 00t€-91,t€11:(t)it Id uorls,la)iooH rq (6[6]) VJ reuprrD 9€-t:i sos lsrHleN IeIsnV/A I rrouu€lpJlsnvlsa.'\\arllolsuoDlppv(t0ot) A/A Plcrr8zlu 09-99:'Z lsnvrnltNurlrol.t^ IIIA oN rllDqsnvjoelou oqlor uoqnquluoc (206I) f v'llu.^\g (,ttolLueluroquoN eqlJo uorssttutuoJ uoqD^r:suo)) .. rllxrsnV',&oltucl ureqlroN aqtJo stu? td rElfl.se^ eqt lo lslllraqJ.,(9661) g ClqrarqtvT d l'eI.(oJ'f t'rlllleg')d'zln'I"fDrq.xel'U:)'dolunc 't99 109 dd Z Iddnssetqlug cosqnlJ qtyA rog uDIIersnVpur urrl4v f$erq.:seloulDlnlrl3ueuloN (al6l) J D'..nrc '82II-€t0t:t(9)68=l IZ log qlollqla suerle.lsnveqdwSoa8uazu?Ud puoYrolCrnz.3€!Icg (8261) )'uluoC 'lazlud 'l 'slelc '829-99:9€lsls qrqrt log sllelucplssoarrlerlsnv ceIqdDJSo qd!]ueluilll (t061) g T 8S-Z€:tsetoN seu qleH IeIsnV r\d ?rt€lsnv ulcrse,\{Jodxtu.IqduSoeSoltrtld ^\3s V (0861)S't'plteg

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