Index

Page numbers in italics refer to Figures and page numbers in bold refer to Tables. acanthodians 74, 75, 77, 78 ostracode case study 106,107, 108-111 accuracy and database management 172, 173 Astraspis 77 acrotetides 26 athyridides 27 Actinodonta 41 Atlantic, Caribbean and East Pacific (ACEP) realm Actinostereon gregareum 133 153, 160, 163, 164 age of , reliability assessment 173,174 atrypides 27 Allonychia 43 Australia 63,183 Aloconconcha 38 Australia/New Guinea block collision 160-162 alpha diversity 2, 99 Autolamellibranchia 39-45 Amadeus Basin 72 Avalonia 19 Ambonychia 43 biodiversity 86, 92, 95 Ambonychiopsis 43 brachiopod diversity profiles 31 Amorphognathus Biofacies 90 conodonts 88, 89, 91 amphibian diversity 183-187,188, 188,189, 191,192, bivalves 36 193 Ordovician palaeogeography 28 analytical time averaging 174 palaeolatitudes 95 Ananterodonta 41 anaspids 74, 75, 78 Anatolepis 70, 72, 76, 79 Babinka 4142 Andean Basin bivalve populations Baltica diversity 128, 134-136 biodiversity 92, 95 effect of Hispanic Corridor 131-134 conodont provinces 90 extinction rates 129, 130 Ordovician 18-19, 19-20, 28 immigration rates 131 rynchonelliformean brachiopods 15, 18 Angarella 19 Baltoscandian conodont provinces 86 angiosperms 1, 5, 5-6 Bambachian megaguilds 25 Cenozoic biodiversity 158 Bavarilla hofensis 56, 58 radiation 141,143-144 Belodella 90 Antarctic Peninsula 145, 146-147 Belodina 90 Anomalocoelia 43 Bennettitales 144,145 44-45 beta (between habitat) diversity 2, 99, 157 Anomalodonta 43 Billingsella Association 15, 16,17, 20 anoxic events and planktonic foraminifera 122 biodiversity in Cenozoic 154, 158-164 Antarctic Peninsula biofacies Cretaceous flora 144-147 architecture 88-91, 93-95 plate setting 141-142 biogeographic cladogram, trilobites 61, 62 role in floristic radiation 147-149 biogeography, Palaeozoic 53-54, 76-78 Antarctica, geographic isolation 154-156 biotic provincialism 1 Anti-Atlas (Morocco) bivalves 35, 36 birds radiation 1 Apatobolus micula 101,102 bivalve molluscs 35-36 Apedolepis 72 Andean Basin - NW Europe study Aphelognathus-Oulodus Biofacies 90 database and analytical methods 128-130 Appalachignathus 90 results 128, 130, 131-134 Arabia, species diversity 183 heteroconch evolution 5 aragonitic foraminifera 121 -Cenozoic diversity 154 Arandospis 69 Ordovician 4,105 Arca spp. 37, 40 Autolamellibranchia 39-45 Arcomytilus pectinatus 133 Protobranchia 36-39 Arctic Ocean isolation 156 Black Mountain eustatic event 20 Arctochedra spp. 16, 17 Bohemia, rynchonelliformean brachiopods 15 Arenigomya 44 Bornholm 90-91 Armorica and rynchonelliformean brachiopods 15 Brachilyrodesma 40 Asaphopsis province 53 brachiopods Asaphopsis- Taihungshania-Neseuretus Fauna 55 Ordovician ash deposition effects 4, 101-104 diversity 105 202 INDEX

mid Wales range chart 110 Coninae 5 radiation 19-20 conodonts 4, 70, 72, 73, 74, 75, 76 rhynchonelliformean 4, 13, 15, 16, 26 see also euconodonts 14-16 Conoglobigerina 118, 121 distribution 14, 15 continental breakup and radiation 1, 6 migration potential 19 Copidens 41 Ordovician 16-18, 25, 26-27, 29-31 Corallidomus 44 radiation and palaeogeography 19-20 corals, Cenozoic diversity 160-163 Breviorthodonta 45 Cosmogoniophorina tenuicostata 44 Bryantodina 89 Coxiconcha 42 bryophytes 145 craniformeans 26 bryozoans 5, 19, 26,101,105 Cretaceous Burgess Shale Lagerst/itte 72 floristic radiation 143-144,145, 146-149 Burj Formation 16 Gondwana breakup 141-143 ocean circulation 159 planktonic foraminifera 115,116, 117, 120, 121, Cadomia 39 122-123 Calymenacean-Dalmanitacean Province 53 Cretaceous-Recent biodiversity 1, 5-6, 7 Calymenella 56, 58, 62, 63 Creux de l'ours 118 Calymenesun spp. 56, 57, 58, 63 crocodilians 176, 179 Cambrian Crozonaspis 54 bivalves 35, 45 crustacean evolution 5 diversity decline 26 Cryptolichenaria 19 mid-late period sea level 19 Ctenodonta 38 palaeogeography 70 Ctenostreon rugosum 133 rynchonelliformean brachiopods 14-16 Cuneamya 44, 45 skeletonized fauna 13 Curtognathus 90 vertebrates 69, 72, 74, 76, 78-79 Cycloconcha 41, 42 Cambrian Evolutionary Fauna 13, 14, 26 Cymatonota 45 cameralloideans 18. 19 Cymbithyris 16 Camnantia 42 Cyrtodonta 43 Camptonectes laminatus 133 Cyrtodontula 43 carbonate deposition in Ordovician 20 cyrtomatodonts 26-27 Cardiolaria 39 Cardoilarioidea 39-40 Caribbean Cenozoic coral diversity 160, 163 Dacryornya lacryma 133 Carminodonta 41 Dactiloceras 118 Carotidens 43 Dalmanellidina 26 Catamarcaia 42 databases 6, 180-182 Celtic faunas 27, 28, 29 application 175-177 Celtoconcha 42 use in diversity analysis 182-190 Cenozoic biodiversity and climate 154-164, 160-163 use in palaeoclimate analysis 190-197 Central American Isthmus (CAI) uplift 156, 163-164 scale 172-175 Central Asia, rynchonelliformean brachiopods 15, 18 structure 169-172 Cephalochordata 74, 75 taxonomy 175 Ceratomya concentrica 133 decapod evolution 5 cheilostome bryozoan, evolution 5 Deceptrix 37 Chengyiang Lagerst~tte 72 delta diversity 2 Chile, Jurassic bivalves 132 deltidoidonts 26-27 China fauna 63.75, 77, 86 Denticelox 43 chondrichthyans 74, 75, 78 Derenjal Formation 16, 17 Cimmeria 61 fauna 2, 25, 78, 79 cladograms, Reedocalymeninae 60, 61, 62, 64 Diaphelasma 18 Clarkella Fauna 15, 17, 18, 20 differentiation diversity 2 Classopolis 148 Diraphora Association 15, 16 Claueonychia 43 discinoideans 26 Clavatipollenites hughesii 148 dispersal potental 19 Cleionychia 43 Distomodus 92, 93 climate and biodiversity 4, 154-157, 162-165, 179,184 diversification and Ordovician volcanism 99-100, clitambonitidines 18, 19 103 Colpantyx 44 case study 106, 107, 108-111 Colpocoryphe 55 ecology 101-103 Compactogerina stellapolaris 122 gene pool 103-104 Coniferales 145 Dob's Lin 86 INDEX 203

Drake Passage opening 156 glaciation 4, 86 Dulcineia 41 glacioeustasy and diversity 163 global warming and floristic radiation 149 Globigerinina 122 East Uralian microcontinent 19 Globuligerina spp. 122 echinoderms and Ordovician diversity 18, 26,105 Glyptarca 39, 40, 42 Ectillaenus 55 Glyptonichia 43 Ekaterodonta 38 Glyptoria spp. 16,17 energy flux and diversity 180, 195 Glyptotrophia 18 Entolium corneolum 133 Gondwana Eocene area cladogram data 61, 63 Messel Shale 190-191,195 brachiopods 15, 16, 18, 20, 31 ocean circulation 155,159 breakup 141-143 Eocene-Oligocene ocean cooling 154, 156 Cambro-Ordovician faunal replacement 20 Eohomalonotus 54 floristic provincialism 146, 148 Eopecten spondyloides 133 Ordovician bivalves 36 epsilon diversity 2 Ordovician palaeogeography 28 Eridodychia 43 trilobites 53-54 Erismodus 90 vertebrates 74, 77 Eritropis 39-40 Goniophorina tenuicosta 44 escalation, Cenozoic 158 grain 172, 173-175 euconodonts 72, 73, 76 graptolite blooms 101,102 presence-absence matrices analysis 86-95 graptoloids, Ordovician diversity 105 Euramerican vertebrate phylogeny 75 Great Basin of Laurentia, brachiopod radiation 25, 26 Europe 183 Gresslya peregrina 133 Jurassic bivalve population gymnosperms 145 diversity 128, 136-138 effect of Hispanic Corridor 131-134 extinction rates 129,130 habitat islands 179-180 immigration rates 131 Haeuslerina helvetojurassica 122 extent defined 172 Haikouella 72 extinction events Haikouichthys 72 late Ordovician 78 Hamarodus-Dapsilodus-Sabbardella (HDS) Pliensbachian-Toarcian 127 Biofacies 90 methods of analysis 128-130 Hanchungolithus 55 results 130-131 Harding Sandstone 69, 70, 72 testing hypothesis 131-134 Harpoceras 118 extinction rates in bivalves 128-129,130, 131-133 Hemiprionodonta 40, 42 Hepatophytes 145 Heteroconchia 40-42 Falcatodonta 43 40 Favusella 120, 122 heterostracans 77, 78 Fidera 36 Heterostraci 74, 75 fish, teleost 5 Hirnantian glaciation 4 flat file format 169,170 Hispanic Corridor 5, 128 floristic development in Cretaceous 1, 5, 5~6, effect on immigration of bivalves 129, 131-133, 134 143-144,145, 146-147 Homilodonta 37 role of Antarctic Peninsula 147-149 Huenella spp. 17, 18 role of global warming 149 Huenella-Palaeostrophia Association 15 foraminifera (planktonic) 5 evolution 121-123 Jurassic-Cretaceous distribution 118,120, 121 Iapetus Ocean 27-28, 28-29, 71, 86 Mesozoic development 115,116,117 Iberocoryphe 54 origins 118-121 Icriodella 92, 93 Fordilla 35 immigration of bivalves 129, 131 role of Hispanic Corridor 131-133 Inaequidens 40 galeaspids 74, 75, 77, 78 Indo-Pacific gateway 160,161 gamma diversity 2, 99 Indo-West Pacific (IWP) realm 153, 160, 164, 165 genetic diversity and ash falls 103-104 glacioeustasy and diversity 163 geographical range, role of 3 coral diversity 160-162 geograpic information systems (GIS) 169-170 Inoperna sowerbyana 133 Gervillaria daeformis 133 inventory diversity 2 ghost ranges, vertebrates 76, 78 island biogeography theory 179 204 INDEX

Jamesella 16 Modiolopsis 44 Johnmartinia 39 Modiolus imbricatus 133 Jurassic Modiomorpha 44 bivalve study in Andean Basin-NW Europe molluscs 163 database and methods of analysis 128-130 see also bivalve molluscs results 128, 130, 131-134 mongolepids 72, 76, 77 planktonic foraminifera 115,116,117 Montagne Noire bivalves 35, 36 distribution 118,120 Moridunia 41 evolution 121-122 Moyeronia 19 origins 118-121 Myllokunmingia 72 Pliensbachian-Toarcian extinctions 127, 128 Myodakryotus 43 Myoplusia 39 Myxinoidea 74, 75, 76 Kerfornella 54 Kerguelen Plateau 143 Koagash Formation 17 Na Mo Formation 63 Kockelella 92 Natasia 38 Kutorginata 14-16 negative difference vegetation index 185, 186, 187, Kutorginide Fauna 15, 16 188, 190 Kutorina 16 neogastropod evolution 5 Kyjandy Formation 16 climate change 162-165 Neseuretinus turcicus 58 Neseuretus spp. 54, 56, 57, 58, 62, 63 Lamellodonta 35 Neseuretus Association 54-56 land bridges 5 Nicaniella voltzii 133 latitude and biodiversity 153, 154 Nileus 63 Laurentia Nimra Formation 16 biodiversity 86, 91, 95 Noradonta 40 conodonts 86, 87, 89, 91 North America species diversity 183 Iapetus margin 71 North Atlantic Deep Water (NADW) 156 Ordovician North Atlantic Province (NAP) conodont biofacies brachiopod radiation 15, 18, 19-20 88-89 faunal replacement 18-19 Nuculana ovum 133 palaeogeography 28 Nuculites 39 vertebrates 74, 76, 77 Nuculoida 36-39 Leconychia 43 Lingulide Fauna 15 linguliformeans 26 Obelellata 14-16 linguloideans 26 Oberhauserella quadrilobata 118, 119 Liomegalaspides 54 Obolid Fauna 15 lithofacies preservation potential 78-79 ocean circulation patterns 154-156,159 Llangynog (Wales), bivalves 36 ocean gateways 5 Lophoconcha 39 Ogyginus 54 lycophytes 145 Old Red Sandstone continental assembly 70, 71 Lyrodesma 40 Olenty Formation 18 Opisthoptera 43 Opsiconidion nudum 101 mammals 1, 5,183, 187-188, 186, 189, 190, 191-192 orbital forcing and range dynamics 163 Marnes de Villers 122 Ordovician Maryonychia 43 biogeography and biodiversity 3-5 Merlinia 54 bivalves 35-45, 46, 47 Mesozoic-Cenozoic boundary and biodiversity 1, brachiopods 16-18 5-6, 7 development in Iapetus 29 Miaopu Formation 63 diversification 26-27 Messel Shale 190-191,195 diversity profiles 30, 31 Microcachyridites antacticus 148 radiation 19-20 Midcontinent Province, conodonts 86 carbonate deposition 20 migration potenta119 diversification 85, 99-100, 105 Milankovitch cyclicity 6, 162-163 effect of ash on 101-104,106,107, 108-111 Miocene euconodonts presence-absence matrices analysis coral diversity 160-162 86-95 ocean circulation 155,159 faunal replacement 18-19 Miocene-, global cooling 154 glaciation 86 Modioliodon 44 palaeoplate faunal exchange 13 INDEX 205

Ordovician (continued) plate tectonics and biodiversity 4 plate reconstructions 37, 55, 70 plectambonitiodeans 18, 26 radiation 1, 7, 25-26 Plectodina 90 sea-level change 19, 20, 30 Plegagnathus 91 tectonic activity 19 Plethocardia 43 trilobites 99, 53-54 Pliensbachian-Toarcian extinctions 127, 128 vertebrates 69-70, 72-73, 74, 76-77, 78-79 Pliocene ocean circulation 157 organophosphatic forms 26 Pojetaia 35 Orthida 26 Polyplacognathus 90 Orthide-Polytoechoidean Association 15 polytoechiides 18 Orthide-Syntrophiodean Association 15 polytoechinoideans 16 orthides, endopunctate 18, 19 Porophoraspis 69, 72 Orthidina 26 Posidonia 118 orthotetides 16 Posidonotis semiplicata 132 Ortonella 43 potental evapotranspiration (PET) 180,185, 191,196 Osteichthyes 74, 75 Pradoella spp. 58, 62 osteostracans 74, 75, 77, 78 Praeanomalodonta 43 ostracoderm 72, 76-79 Praegubkinella spp. 118, 119, 120 ostracodes and Ordovician volcanism 106,107, Praenucula 36 108-111 precipitation and diversity 184, 186, 187, 188, 190 Oulodus 91, 92 precision defined 172 oxygenation and ash falls 102, 103 predator pressure 132 Ozarkodina 92 presence-absence matrices analysis 86-95 Prolobella 43 Propeamussium pumilum 132 palaeoclimate and diversity 190-197 Protambonites 17, 18 Palaeoconcha 36 Prothyris 45 palaeogeography Protobranchia 36-39 early Palaeozoic 37, 55, 70-71, 70 Protocalymene mcallisteri 56, 58 role in radiation 1, 6 Protopanderodus 90 Palaeoheterodonta 40 Protopanderodus-Periodon Biofacies 86 palaeolatitude and planktonic foraminifera 122-123 Protorthide Fauna 15, 16 Palaeoneilo 39 Protorthis 16 palaeontological uncertainty principle 174 provinciality changes 2 Palaeonucula hammeri 133 Pseudarca 40 Palaeopteria 43 Pseudolimea duplicata 133 Palaeostrophia 18 Pseudolonchodina 92 Palaeozoic Evolutionary Fauna 13, 16, 18, 19, 26 Pseudopecten barbatus 133 Parallelodon hirsonensis 133 Psiloconcha 45 Parallelodus 44 Psilonychia 43 Paramytilarca 43 Psiloria 16 parental imprinting 104 Pteridophytes 145 Paulinea 36 Pteriornorphia 42-44 pelromyzontids 76 Pteronychia 43 Pensarnia 36 Pterospathodus 92 Pentamerides 26, 27 pyroclastics see ash Perunica, brachiopod diversity profiles 31 Petromyxinoidea 74, 75 Pharcidoconcha 43 radiation and diversity 184, 186, 187, 188, 190 Pharostomina oepiki 56, 58 Ramsey Island (Wales), bivalves 36 Pholadomya fiducula 133 Rapoport's rule 180 Phragmodus 90 Redonia 41 Phragmodus-Icriodella-Plectodina Biofacies 90 Reedocalymeninae 4, 56-61 phylogenetic analysis, trilobites 56-61 relational databases 169,170 phylogeny, vertebrates 73-76 reptile diversity 183,186, 187, 187, 191,192,193 Pipiscius 72 resolution see grain Pirchanchaspis rinconensis 72-73 Rhipidognathus 89 Pituriaspida 74, 75 rhynconelliformean brachiopods see under placoderms 74, 75, 77, 78 brachiopods Plaesiacomia 54 Rhytimya 44 Plagiostorna rodburgensis 133 planktonic blooms 102, 103 planktonic foraminifera see foraminifera Sacabambaspis 69 plate reconstructions see palaeogeography Salta Province (Argentina) bivalves 35, 36 206 INDEX

Salterocoryphe 55 Toquima-Table Head faunas 27, 28, 29 Sarrabesia spp. 58, 62, 63 Tornquist Sea closure 71 scale problems in databases 172-175 Trecanolia 42 sea level tree diversity 189 change and biodiversity 4-5, 163,165 Tremadoc 19, 20-21 Palaeozoic 19, 20, 30 Trematobolus 16 relation to tectonism 104 Trematosia 16 Selenopeltis Province 53 Trigonioida 40 Semicorallidomus 44 Trigonoconcha 38 Shirgesht Formation 17, 18 trilobites Siberia asaphide 18 Cambrian fauna 16 Cambrian Evolutionary Fauna 26 conodont provinces 86 Neseuretus Association 54-56 early Palaeozoic 71, 75, 77 Ordovician 4 Ordovician fauna 18-19, 19-20, 28 diversity 99,105 Siberoctenia 38 provinciality 53-54 siliciclastic input and diversity 4 phylogenetic analysis 56-61 triplesiides 19 diversity 92-93 Tritoechia 18 endemism and cosmopolitanism 79 Tritoechia-Protambonites Association 15 palaeogeography 70 Tromelinodonta 40 provincialism 2 Tuarangia 35 vertebrates 73, 74, 77-78 Tunicata 74, 75 Similodonta 38 Turkey, trilobites 61, 62, 63 siphonotretides 26 Turukhansk-Irkutsk facies 16 skeletonized fauna 13 Tuva, vertebrates 75, 77 Skiichthys halsteadi 69 Tuvaella 77 snakes radiation 5 Solemyoida 39 Solenomorpha 45 uncertainty principle 174 Sort Tepe Formation 63 Uralian margin uplift 19, 20, 21 South Africa species diversity 183 Uskardita 42 South America 63, 148,183 spiriferides 27 Stairway Sandstone 69 Vanuxemia 43 Sthenodonta 39 Veneridae evolution 5 stratigraphic reliability codes 174 vertebrates stromatoporoid reefs 25 biogeography 76-78 Strophomenata 16 early Palaeozoic 69-70, 72-73 Strophomenida 26 phylogeny 73-76 strophomenides 16, 19 Vietnam, trilobites 63 sympatric speciation 104 Vietnamia douvillei 57, 58, 63 Synchomalonotus 62 Villicumia 38 Synhomalonotus 55, 56 volcanism and diversity 4, 19, 101-104 synonymization 175 ostracode case study 106,107, 108-111 syntrophiidines 18 syntrophioideans 18, 19 Warburgia 43 Welsh Basin 19, 99, 101 Taconic Orogeny 4 ostracode development 108-111 Taihungshania 54 Whiterock Fauna 4, 26 Tancrediopsis 38 within-habitat diversity see alpha diversity taphonomic time averaging 174 taxonomy, role in database management 175 tectonism, role in radiation 1, 6 Xestoconcha 44 teleost fish evolution 5 Xidazoon 72 temperature and diversity 184, 186, 187, 188, 190,192, 197 Tethyan Ocean closure 156, 160 Yangtze Platform, trilobites 63 Tetralobula 18 Yunnanozoon 72 tetrapod diversity 182-183,185, 186,197 thelodonts 72, 74, 75, 76, 77, 78 Thorslundia 43 Zadiemrodia fastigata 41 Thysanotos-Leptembolon Fauna 15 Zeehania 39 Tironucula 38 Zel&zkella 55