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Ediacaran) of Earth – Nature’S Experiments
The Early Animals (Ediacaran) of Earth – Nature’s Experiments Donald Baumgartner Medical Entomologist, Biologist, and Fossil Enthusiast Presentation before Chicago Rocks and Mineral Society May 10, 2014 Illinois Famous for Pennsylvanian Fossils 3 In the Beginning: The Big Bang . Earth formed 4.6 billion years ago Fossil Record Order 95% of higher taxa: Random plant divisions domains & kingdoms Cambrian Atdabanian Fauna Vendian Tommotian Fauna Ediacaran Fauna protists Proterozoic algae McConnell (Baptist)College Pre C - Fossil Order Archaean bacteria Source: Truett Kurt Wise The First Cells . 3.8 billion years ago, oxygen levels in atmosphere and seas were low • Early prokaryotic cells probably were anaerobic • Stromatolites . Divergence separated bacteria from ancestors of archaeans and eukaryotes Stromatolites Dominated the Earth Stromatolites of cyanobacteria ruled the Earth from 3.8 b.y. to 600 m. [2.5 b.y.]. Believed that Earth glaciations are correlated with great demise of stromatolites world-wide. 8 The Oxygen Atmosphere . Cyanobacteria evolved an oxygen-releasing, noncyclic pathway of photosynthesis • Changed Earth’s atmosphere . Increased oxygen favored aerobic respiration Early Multi-Cellular Life Was Born Eosphaera & Kakabekia at 2 b.y in Canada Gunflint Chert 11 Earliest Multi-Cellular Metazoan Life (1) Alga Eukaryote Grypania of MI at 1.85 b.y. MI fossil outcrop 12 Earliest Multi-Cellular Metazoan Life (2) Beads Horodyskia of MT and Aust. at 1.5 b.y. thought to be algae 13 Source: Fedonkin et al. 2007 Rise of Animals Tappania Fungus at 1.5 b.y Described now from China, Russia, Canada, India, & Australia 14 Earliest Multi-Cellular Metazoan Animals (3) Worm-like Parmia of N.E. -
Lineages, Splits and Divergence Challenge Whether the Terms Anagenesis and Cladogenesis Are Necessary
Biological Journal of the Linnean Society, 2015, , – . With 2 figures. Lineages, splits and divergence challenge whether the terms anagenesis and cladogenesis are necessary FELIX VAUX*, STEVEN A. TREWICK and MARY MORGAN-RICHARDS Ecology Group, Institute of Agriculture and Environment, Massey University, Palmerston North, New Zealand Received 3 June 2015; revised 22 July 2015; accepted for publication 22 July 2015 Using the framework of evolutionary lineages to separate the process of evolution and classification of species, we observe that ‘anagenesis’ and ‘cladogenesis’ are unnecessary terms. The terms have changed significantly in meaning over time, and current usage is inconsistent and vague across many different disciplines. The most popular definition of cladogenesis is the splitting of evolutionary lineages (cessation of gene flow), whereas anagenesis is evolutionary change between splits. Cladogenesis (and lineage-splitting) is also regularly made synonymous with speciation. This definition is misleading as lineage-splitting is prolific during evolution and because palaeontological studies provide no direct estimate of gene flow. The terms also fail to incorporate speciation without being arbitrary or relative, and the focus upon lineage-splitting ignores the importance of divergence, hybridization, extinction and informative value (i.e. what is helpful to describe as a taxon) for species classification. We conclude and demonstrate that evolution and species diversity can be considered with greater clarity using simpler, more transparent terms than anagenesis and cladogenesis. Describing evolution and taxonomic classification can be straightforward, and there is no need to ‘make words mean so many different things’. © 2015 The Linnean Society of London, Biological Journal of the Linnean Society, 2015, 00, 000–000. -
A Framework for Post-Phylogenetic Systematics
A FRAMEWORK FOR POST-PHYLOGENETIC SYSTEMATICS Richard H. Zander Zetetic Publications, St. Louis Richard H. Zander Missouri Botanical Garden P.O. Box 299 St. Louis, MO 63166 [email protected] Zetetic Publications in St. Louis produces but does not sell this book. Any book dealer can obtain a copy for you through the usual channels. Resellers please contact CreateSpace Independent Publishing Platform of Amazon. ISBN-13: 978-1492220404 ISBN-10: 149222040X © Copyright 2013, all rights reserved. The image on the cover and title page is a stylized dendrogram of paraphyly (see Plate 1.1). This is, in macroevolutionary terms, an ancestral taxon of two (or more) species or of molecular strains of one taxon giving rise to a descendant taxon (unconnected comma) from one ancestral branch. The image on the back cover is a stylized dendrogram of two, genus-level speciational bursts or dis- silience. Here, the dissilient genus is the basic evolutionary unit (see Plate 13.1). This evolutionary model is evident in analysis of the moss Didymodon (Chapter 8) through superoptimization. A super- generative core species with a set of radiative, specialized descendant species in the stylized tree com- promises one genus. In this exemplary image; another genus of similar complexity is generated by the core supergenerative species of the first. TABLE OF CONTENTS Preface..................................................................................................................................................... 1 Acknowledgments.................................................................................................................................. -
British Columbia Geological Survey Geological Fieldwork 1990
GEOLOGY OF THE MOUNT BRUSSILOF MAGNESITE DEPOSIT, SOUTHEASTERN BRITISH COLUMBIA (825/12, 13) By George J. Simandl and Kirk D. Hancock KEYWOKLIS: Industrial minerals, rconomic geology. mag- nesitr. Cathedral Formation. Middle Cambrian. dolumitiz- tion, porosity. base metal association, deposit model. INTRODUCTION Magnesite (M&O,) is an induswial mineral that can bc converted into either caustic, fused or dead-burned magne- sia. Dead-burned magnesia is used mainly in the manufac- ture of refractory products; caustic magnesia is used in treatment of water, in animal feedstuffs, fertilizers. magne- sia cements, insulating boards and wood-pulp processing, in chemicals and pharmaceuticals and us a curing agent in rubber (Coope. 19X7). Magnesium metal is produced either from magnesite or from caustic mapnesia. In the short-term future. production of dead-burned mag- nesia is cxpectcd to remain constant. however, demand for caustic magnesia is increasing (Duncan, IYYO). With the increasing trend toward the use of high-perfomxmce “mag- carbon” refractories. future demand for fused magnesia looks promising. A number of magnesite deposits are known in British Columbia (Grzant, 19X7). the most important of these is the Mount Brussilof orebody. It is hosted by dolomites of the TECTONIC SETTING Middle Cambrian Cathedral Formation. The Mount Brussilof deposit is located in the Forekm3 rectonostratiEnlphic belt and within the “Kicking Horse HISTORY Rim”, as defined by Aitken (1971, 1989). It is situated east of a Cambrian bathymetric feature commonly referred to as The Mount Brussilof deposit was discovered during the Cathedral escarpment (Fritz. 1990: Aitken xnd regional mapping by the Geological Survey of Canada Mcllreath, 19X4, 1990). -
Lifestyle of the Octoradiate Eoandromeda in the Ediacaran
Lifestyle of the Octoradiate Eoandromeda in the Ediacaran Authors: Wang, Ye, Wang, Yue, Tang, Feng, Zhao, Mingsheng, and Liu, Pei Source: Paleontological Research, 24(1) : 1-13 Published By: The Palaeontological Society of Japan URL: https://doi.org/10.2517/2019PR001 BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Complete website, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www.bioone.org/terms-of-use. Usage of BioOne Complete content is strictly limited to personal, educational, and non - commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. Downloaded From: https://bioone.org/journals/Paleontological-Research on 15 Feb 2020 Terms of Use: https://bioone.org/terms-of-use Access provided by Bing Search Engine Paleontological Research, vol. 24, no. 1, pp. 1–13, JanuaryEoandromeda 1, 2020 ’s Lifestyle 1 © by the Palaeontological Society of Japan doi:10.2517/2019PR001 Lifestyle of the Octoradiate Eoandromeda in the Ediacaran YE WANG1, YUE WANG1, FENG TANG2, MINGSHENG ZHAO3 and PEI LIU1 1Resource and Environmental Engineering College, Guizhou University, Huanx 550025, Guiyang, Guizhou, China (e-mail: [email protected]) 2Institute of Geology, Chinese Academy of Geological Sciences, 26 Baiwanzhuang Street 100037, Beijing, China 3College of Paleontology, Shenyang Normal Univeristy, 253 Huanhe N Street 110034, Shengyang, Liaoning, China Received May 14, 2018; Revised manuscript accepted January 26, 2019 Abstract. -
A Model of Mass Extinction
A model of mass extinction M. E. J. Newman Cornell Theory Center, Rhodes Hall, Ithaca, NY 14853. and Santa Fe Institute, 1399 Hyde Park Road, Santa Fe, NM 87501. Abstract A number of authors have in recent years proposed that the processes of macroevo- lution may give rise to self-organized critical phenomena which could have a sig- nificant effect on the dynamics of ecosystems. In particular it has been suggested that mass extinction may arise through a purely biotic mechanism as the result of so-called coevolutionary avalanches. In this paper we first explore the empirical evidence which has been put forward in favor of this conclusion. The data center principally around the existence of power-law functional forms in the distribution of the sizes of extinction events and other quantities. We then propose a new math- ematical model of mass extinction which does not rely on coevolutionary effects and in which extinction is caused entirely by the action of environmental stresses on species. In combination with a simple model of species adaptation we show that this process can account for all the observed data without the need to invoke coevo- lution and critical processes. The model also makes some independent predictions, such as the existence of “aftershock” extinctions in the aftermath of large mass extinction events, which should in theory be testable against the fossil record. arXiv:adap-org/9702003v1 12 Feb 1997 1 Introduction Building on ideas first put forward by Kauffman (1992), there has in re- cent years been increasing interest in the possibility that evolution may be a self-organized critical phenomenon (Sol´eand Bascompte 1996, Bak and Paczuski 1996). -
The Cambrian Substrate Revolution and the Early Evolution of Attachment in MARK Suspension-Feeding Echinoderms
Earth-Science Reviews 171 (2017) 478–491 Contents lists available at ScienceDirect Earth-Science Reviews journal homepage: www.elsevier.com/locate/earscirev The Cambrian Substrate Revolution and the early evolution of attachment in MARK suspension-feeding echinoderms ⁎ Samuel Zamoraa,b, , Bradley Delinec, J. Javier Álvarod, Imran A. Rahmane a Instituto Geológico y Minero de España, C/Manuel Lasala, 44, 9B, Zaragoza 50006, Spain b Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington DC 20013-7012, USA c University of West Georgia, Carrollton, GA, USA d Instituto de Geociencias (CSIC-UCM), c/José Antonio Novais 12, 28040 Madrid, Spain e Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW, UK ARTICLE INFO ABSTRACT Keywords: The Cambrian, characterized by the global appearance of diverse biomineralized metazoans in the fossil record Palaeoecology for the first time, represents a pivotal point in the history of life. This period also documents a major change in Evolution the nature of the sea floor: Neoproterozoic-type substrates stabilized by microbial mats were replaced by un- fl Sea oor consolidated soft substrates with a well-developed mixed layer. The effect of this transition on the ecology and Attachment evolution of benthic metazoans is termed the Cambrian Substrate Revolution (CSR), and this is thought to have impacted greatly on early suspension-feeding echinoderms in particular. According to this paradigm, most echinoderms rested directly on non-bioturbated soft substrates as sediment attachers and stickers during the Cambrian Epoch 2. As the substrates became increasingly disturbed by burrowing, forming a progressively thickening mixed layer, echinoderms developed new strategies for attaching to firm and hard substrates. -
Plated Cambrian Bilaterians Reveal the Earliest Stages of Echinoderm Evolution
Zamora, S., Rahman, I. A., & Smith, A. B. (2012). Plated Cambrian bilaterians reveal the earliest stages of echinoderm evolution. PLoS ONE, 7(6), [e38296]. https://doi.org/10.1371/journal.pone.0038296 Publisher's PDF, also known as Version of record Link to published version (if available): 10.1371/journal.pone.0038296 Link to publication record in Explore Bristol Research PDF-document University of Bristol - Explore Bristol Research General rights This document is made available in accordance with publisher policies. Please cite only the published version using the reference above. Full terms of use are available: http://www.bristol.ac.uk/red/research-policy/pure/user-guides/ebr-terms/ Plated Cambrian Bilaterians Reveal the Earliest Stages of Echinoderm Evolution Samuel Zamora1, Imran A. Rahman2, Andrew B. Smith1* 1 Department of Palaeontology, The Natural History Museum, London, United Kingdom, 2 School of Geography, Earth & Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom Abstract Echinoderms are unique in being pentaradiate, having diverged from the ancestral bilaterian body plan more radically than any other animal phylum. This transformation arises during ontogeny, as echinoderm larvae are initially bilateral, then pass through an asymmetric phase, before giving rise to the pentaradiate adult. Many fossil echinoderms are radial and a few are asymmetric, but until now none have been described that show the original bilaterian stage in echinoderm evolution. Here we report new fossils from the early middle Cambrian of southern Europe that are the first echinoderms with a fully bilaterian body plan as adults. Morphologically they are intermediate between two of the most basal classes, the Ctenocystoidea and Cincta. -
A Solution to Darwin's Dilemma: Differential Taphonomy of Ediacaran and Palaeozoic Non-Mineralised Discoidal Fossils
Provided by the author(s) and NUI Galway in accordance with publisher policies. Please cite the published version when available. Title A Solution to Darwin's Dilemma: Differential Taphonomy of Ediacaran and Palaeozoic Non-Mineralised Discoidal Fossils Author(s) MacGabhann, Breandán Anraoi Publication Date 2012-08-29 Item record http://hdl.handle.net/10379/3406 Downloaded 2021-09-26T20:57:04Z Some rights reserved. For more information, please see the item record link above. A Solution to Darwin’s Dilemma: Differential taphonomy of Palaeozoic and Ediacaran non- mineralised discoidal fossils Volume 1 of 2 Breandán Anraoi MacGabhann Supervisor: Dr. John Murray Earth and Ocean Sciences, School of Natural Sciences, NUI Galway August 2012 Differential taphonomy of Palaeozoic and Ediacaran non-mineralised fossils Table of Contents List of Figures ........................................................................................................... ix List of Tables ........................................................................................................... xxi Taxonomic Statement ........................................................................................... xxiii Acknowledgements ................................................................................................ xxv Abstract ................................................................................................................. xxix 1. Darwin’s Dilemma ............................................................................................... -
September 2009
Coaster Brook Trout (Salvelinus fontinalis) 12-Month Finding Decision Document SEPTEMBER 2009 0 Coaster Brook Trout 12-Month Finding Decsion Document TABLE OF CONTENTS PURPOSE ............................................................................................................................................................... 1 BACKGROUND ........................................................................................................................................................... 1 DECISION STRUCTURE ........................................................................................................................................... 4 DECISION PROBLEM .................................................................................................................................................... 5 OBJECTIVES ................................................................................................................................................................. 7 ALTERNATIVES ............................................................................................................................................................. 7 CONSEQUENCES ........................................................................................................................................................ 10 TRADE-OFFS .............................................................................................................................................................. 13 DISTINCT POPULATION ANALYSIS ................................................................................................................. -
A Stem Group Echinoderm from the Basal Cambrian of China and the Origins of Ambulacraria
ARTICLE https://doi.org/10.1038/s41467-019-09059-3 OPEN A stem group echinoderm from the basal Cambrian of China and the origins of Ambulacraria Timothy P. Topper 1,2,3, Junfeng Guo4, Sébastien Clausen 5, Christian B. Skovsted2 & Zhifei Zhang1 Deuterostomes are a morphologically disparate clade, encompassing the chordates (including vertebrates), the hemichordates (the vermiform enteropneusts and the colonial tube-dwelling pterobranchs) and the echinoderms (including starfish). Although deuter- 1234567890():,; ostomes are considered monophyletic, the inter-relationships between the three clades remain highly contentious. Here we report, Yanjiahella biscarpa, a bilaterally symmetrical, solitary metazoan from the early Cambrian (Fortunian) of China with a characteristic echinoderm-like plated theca, a muscular stalk reminiscent of the hemichordates and a pair of feeding appendages. Our phylogenetic analysis indicates that Y. biscarpa is a stem- echinoderm and not only is this species the oldest and most basal echinoderm, but it also predates all known hemichordates, and is among the earliest deuterostomes. This taxon confirms that echinoderms acquired plating before pentaradial symmetry and that their history is rooted in bilateral forms. Yanjiahella biscarpa shares morphological similarities with both enteropneusts and echinoderms, indicating that the enteropneust body plan is ancestral within hemichordates. 1 Shaanxi Key Laboratory of Early Life and Environments, State Key Laboratory of Continental Dynamics and Department of Geology, Northwest University, 710069 Xi’an, China. 2 Department of Palaeobiology, Swedish Museum of Natural History, Box 50007104 05, Stockholm, Sweden. 3 Department of Earth Sciences, Durham University, Durham DH1 3LE, UK. 4 School of Earth Science and Resources, Key Laboratory for the study of Focused Magmatism and Giant Ore Deposits, MLR, Chang’an University, 710054 Xi’an, China. -
Smithsonian Miscellaneous Collections
SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 116, NUMBER 5 Cfjarle* £. anb Jfflarp "^Xaux flKHalcott 3Resiearcf) Jf tmb MIDDLE CAMBRIAN STRATIGRAPHY AND FAUNAS OF THE CANADIAN ROCKY MOUNTAINS (With 34 Plates) BY FRANCO RASETTI The Johns Hopkins University Baltimore, Maryland SEP Iff 1951 (Publication 4046) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION SEPTEMBER 18, 1951 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 116, NUMBER 5 Cfjarie* B. anb Jfflarp "^Taux OTalcott &egearcf) Jf unb MIDDLE CAMBRIAN STRATIGRAPHY AND FAUNAS OF THE CANADIAN ROCKY MOUNTAINS (With 34 Plates) BY FRANCO RASETTI The Johns Hopkins University Baltimore, Maryland (Publication 4046) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION SEPTEMBER 18, 1951 BALTIMORE, MD., U. 8. A. CONTENTS PART I. STRATIGRAPHY Page Introduction i The problem I Acknowledgments 2 Summary of previous work 3 Method of work 7 Description of localities and sections 9 Terminology 9 Bow Lake 11 Hector Creek 13 Slate Mountains 14 Mount Niblock 15 Mount Whyte—Plain of Six Glaciers 17 Ross Lake 20 Mount Bosworth 21 Mount Victoria 22 Cathedral Mountain 23 Popes Peak 24 Eiffel Peak 25 Mount Temple 26 Pinnacle Mountain 28 Mount Schaffer 29 Mount Odaray 31 Park Mountain 33 Mount Field : Kicking Horse Aline 35 Mount Field : Burgess Quarry 37 Mount Stephen 39 General description 39 Monarch Creek IS Monarch Mine 46 North Gully and Fossil Gully 47 Cambrian formations : Lower Cambrian S3 St. Piran sandstone 53 Copper boundary of formation ?3 Peyto limestone member 55 Cambrian formations : Middle Cambrian 56 Mount Whyte formation 56 Type section 56 Lithology and thickness 5& Mount Whyte-Cathedral contact 62 Lake Agnes shale lentil 62 Yoho shale lentil "3 iii iv SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL.