ACTA ZOOLOGICA BULGARICA Acta zool. bulg., 59 (1), 2007: 33-39

A Contribution to the Pselaphinae Fauna (Coleoptera: Staphylinidae) of the Vitosha Mountain

Rostislav Bekchiev1*, Maria Shishiniova1

18 Dragan Tsankov Bvld., Department of Zoology and Anthropology, Faculty of Biology, Sofia University, 1164 Sofia, Bulgaria, e-mail: [email protected]

Abstract: Twenty-three species from subfamily Pselaphinae (Coleoptera, Staphylinidae) of the Vitosha Mountain have been reported. From them 17 species are new to the mountain fauna and one species (Bibloporus mayeti Guillebeau, 1888) is new to the Bulgarian fauna. The total number for the Vitosha Mountain fauna is 24, that is near 23% from all known species of the Bulgarian Pselaphin fauna.

Key words: Pselaphinae, faunistics, new locality, Vitosha Mountain, Bulgaria

Introduction Material and Methods

In Bulgaria subfamily Pselaphinae is poorly known The research was made in different localities in the and there is a lack of regular investigations on it. The Vitosha Mountain (Table 1) from September 2003 to reported total number of species for Bulgaria until November 2003 and from March to November 2004. this moment is about 93 species (Löbl, Besuchet The specimens were collected by sifting leaf, soil 2004). and wood litter, samples were taken every month. The area of the Vitosha Mountain is not studied The collected material was placed in a Tullgren or yet and the information for Pselaphinae species is Winkler – Mokzarski extractor. A hand collecting scanty and sporadic. First data about this subfamily under stones, bark and ant nests was made at the were given by Rambousek (1909), who reported same time. Fifty soil traps were also used. Totally 4 species (Trimium caucasicum Kolenati, 1846, 623 individuals were collected. Bryaxis ullrichii Motschulsky, 1851, Bryaxis simo­ All individuals were pinned and the aedeagi ni (Reitter, 1880), Tychus rufus Motschulsky, of the male specimens were mounted following 1851) and by Drenovski (1942), who added Bythinus the procedure explained by Besuchet (1957). The sp., too. Later data for pselaphid fauna were given entire collection is preserved in the Departement of by Besuchet (1999), who described the species Zoology and Anthroplogy, Faculty of Biology, Sofia Dicentrius rousi Besuchet, 1999. The latest data University. were given by Lapeva-Gjonova (2004), who reported The collection material of Musèum d’Histoire another 2 species (Euplectus signatus (Reichenbach, Naturelle de Genève, (Suisse) – MHNG and the 1816), Trimium puncticeps Reitter, 1880). private collection Peter Hlaváč (Slovakia) – PHC,

*Corresponding author

33 Bekchiev R, M. Shishiniova

Table 1. Localities and leaf litter samples in the Vitosha Mts.

Altitude Locality Leaf litter GPS (m) above Knyazhevo Village 660 Fagus sylvatica N 42°39’062” E 23°14’347” place „Byala voda” 900 Fagus sylvatica N 42°38’515” E 23°13’850” place „Byala voda” - place „Kopitoto” 1100 Fagus sylvatica N 42°38’372” E 23°13’659” place „Kopitoto” 1300 Fagus sylvatica N 42°38’053” E 23°14’389” „Momina skala” Hut 1432 Picea abies N 42°37’708” E 23°14’851” „Tintyava” Hut 1600 Fagus sylvatica N 42°37’067” E 23°15’444” place „Gyoreva cheshma” 1700 Fagus sylvatica N 42°36’923” E 23°15’704” „Bor” Hut 1600 Fagus sylvatica N 42°36’999” E 23°15’262” place „Zlatnite mostove”-Vladaya Village 1250 Fagus sylvatica N 42°36’875” E 23°13’419” above Vladaya Village 970 Fagus sylvatica N 42°37’135” E 23°12’615” above Dragalevtsi Village 910 Fagus sylvatica N 42°37’253” E 23°18’207” above Dragalevtsi Village - place „Bai Krastyo” 1210 Fagus sylvatica N 42°36’562” E 23°17’868” place „Bai Krastyo” 1450 Pinus peuce, Fagus sylvatica N 42°36’217” E 23°17’929” place „Bai Krastyo” – „Aleko” Hut 1500 Picea abies N 42°36’028” E 23°17’932” place Bai Krastyo - Aleko Hut 1600 Fagus sylvatica, Salix sp. N 42°35’824” E 23°17’893” „Aleko” Hut 1800 Picea abies N 42°34’704” E 23°17’985” reserve „Bistrishko branishte” 1700 Picea abies N 42°34’452” E 23°18’555” reserve „Bistrishko branishte” 1600 Picea abies N 42°34’349” E 23°18’687” reserve „Bistrishko branishte” 1400 Fagus sylvatica N 42°34’032” E 23°19’116” reserve „Bistrishko branishte” 1300 Fagus sylvatica N 42°34’205” E 23°19’645” Fagus sylvatica, Pinus Zhelezhnitsa Village - place „Brezite” 1100 N 42°31’639” E 23°21’900” silvestris place „Brezite” - place „Yarema” 1200 Fagus sylvatica, Picea abies N 42°30’870” E 23°20’579” place „Yarema” 1300 Fagus sylvatica N 42°30’587” E 23°19’706” near place „Lashkova mogila” 1500 Fagus sylvatica N 42°30’714” E 23°19’059” „Lashkova mogila” - Kechek summit 1600 Fagus sylvatica N 42°30’946” E 23°18’767” Kechek summit 1650 Fagus sylvatica N 42°31’946” E 23°18’767” Kechek Summit - Barite Summit 1634 Fagus sylvatica N 42°31’341’ E 23°19’015” above place „Bukite” 1451 Fagus sylvatica N 42°31’734” E 23°20’331” Zhelezhnitsa Village near the river 1200 Fagus sylvatica N 42°31’975” E 23°21’ 683” near Bosnek Village 900 Fagus sylvatica - Bosnek Village 1000 Fagus sylvatica, Quercus sp. - near „Duhlata” Cave 1000 Fagus sylvatica N 42° 29’777” E 23°11’855” Bosnek Village - Chuipetlovo Village 1000 Fagus sylvatica - Chuipetlovo Village 1300 Fagus sylvatica, Salix sp. - Marchaevo Village 1000 Fagus sylvatica, Quercus sp. N 42° 35’46” E 23°10’02” above Rudartsi Village 976 Fagus sylvatica N 42° 35’23” E 23°10’40” Kladnitsa Village 1100 Fagus sylvatica N 42°34’068” E 23°11’587” Kladnitsa Village – „Selimitsa” Hut 1150 Fagus sylvatica N 42°35’21” E 23°10’56” „Selimitsa” Hut 1240 Fagus sylvatica N 42°34’306” E 23°12’146” „Selimitsa” Hut –„Ostritsa” Hut 1520 Salix sp. N 42°35’172” E 23°12’283” „Rudnichar” Hut 1620 Picea abies N 42°35’357” E 23°13’071” „Rudnichar” Hut- place „Yavorov” 1670 Picea abies N 42°35’615” E 23°13’750” „Radioto” Hut 1550 Fagus sylvatica N 42°36’201” E 23°14’064” place „Zlatnite mostove” 1300 Fagus sylvatica N 42°36’539” E 23°14’320”

34 A Contribution to the Pselaphinae... were checked and the data from them were also Bibloporus minutus Raffray,1914 included in this work. New records: Bistritsa Village, in leaf litter of Due to the impossibility to determine the female Picea abies L., 07.3.2003, 1 ♀; between place “Byala exemplars of some species from the genera Bryaxis voda” and place “Kopitoto”, 19.5.2004, under bark Kugelann, 1974 (11 ex.) and Bythinus Leach, 1817 of F. sylvatica, 1 ♂, 1 ♀; above place “Byala voda”, (83 ex.), information about them is not mentioned in in rotten wood of old F. sylvatica tree, 31.8.2004, 1 the present article. ♀; under bark, 22.10.2004, 1 ♀. The general distribution of species is given in Distribution: Europe, Asia. alphabetical order by Löbl, Besuchet (2004). Trychonix sulcicollis (Reichenbach, 1816) Varna, Rila Monastery (Karaman 1969). Results New record: place “Byala voda”, in pitfall trap, 900 m, 23.06-21.07.2004, 1 ♀. List of Species Distribution: Europe Claviger testaceus Preyssler, 1790 Trimium carpathicum Saulcy, 1875 New records: Chuipetlovo Village, under stone Jalna, Osogovo, Sofia (Karaman 1967). in Lasius flavus (Fabricius,1782) nest, 1300 m, New records: place “Byala voda”, 20.3.2004, 28.4.2004, 1 ♂, 2 ♀♀; 29.5.2004, 1 ♂. 2 ♀♀; place “Kopitoto”, 19.5.2004, 1 ♂; 31.8.2004, Distribution: Europe, Asia (Turkey). 2 ♂♂; near hut „Momina skala”, 22.4.2004, 1 ♀; Euplectus kirbii Denny, 1825 place “Zlatnite mostove”, under trunk, 1300 m, Varna, (Karaman 1969). 22.4.2004, 5 ♀♀; „Bistrishko branishte” Reserve, New records: Kladnitsa Village, in rotten wood 23.7.2004, 1 ♂; near „Duhlata” Cave, 28.4.2004, 2 of Populus sp. tree, 1000 m, 02.9.2004, 1 ♂, 2 ♀♀; ♂♂, 2 ♀♀; 06.6.2004, 4 ♀♀; in moss, 09.10.2004, 1 05.10.2004, 1 ♀. ♀; near Bosnek Village, in leaf litter of F. sylvatica, Distribution: Europe 900 m, 29.5.2004, 1 ♂; in leaf litter of Quercus sp., Euplectus karstenii Reichenbach, 1816 03.8.2004, 1 ♂, 2 ♀♀; 23.8.2004, 1 ♂; Chuipetlovo Sliven (Rambousek 1909). Village, 1300 m, 03.8.2004, 3 ♀♀; between Kladnitsa New records: place „Bai Krastyo”, 30.6.2004, Village and „Selimitsa” Hut, 05.10.2004, 1 ♂; above 1 ♂; Kladnitsa Village, in rotten wood of Populus sp. Rudartsi Village, 31.5.2004, 1 ♂; 25.6.2004, 2 ♀♀; tree, 1000 m, 02.9.2004, 1 ♂, 2 ♀♀. Marchaevo Village, 25.6.2004, 1 ♀; 30.7.2004, 2 Distribution: Europe ♀♀; Zheleznitsa Village, place „Brezite”, 03.7.2004,

Plectophloeus fischeri( Aubé, 1833) 3 ♀♀; 22.9.2004, 1 ♀; in moss, 01.8.2004, 1 ♀; Rila - Borovets, Varna (Rambousek 1909, 22.9.2004, 1 ♀. Karaman 1962, 1969). Distribution: Central and Eastern Europe New records: Vitosha, 1928, 2 ♂♂, leg. Pl. Trimium puncticeps Reitter, 1880 Fodor (MHNG); place „Byala voda”, in moss, 900 Osogovo, Bermakov, Samokov, Trevna m, 01.11.2003, 1 ♀. (Karaman 1967), Vitosha – Bistritsa (Lapeva- Distribution: Europe. Gjonova 2004). Bibloporus mayeti Guillebeau, 1888 New records: Bistritsa Village, 12.5.2002, 1 New records: Kladnitsa Village, in rotten wood ♀; 02.1.2003, 1 ♀; Boyana Village, 09.3.2003, 1 ♀; of Populus sp. tree,1000 m, 05.10.2004, 1 ♂. Vitosha 05.05.2003, 1 ♀, leg. D. Gradinarov; side Distribution: Europe. of the Boyansko ezero Lake, 08.2.2004, 1 ♀; place

35 Bekchiev R, M. Shishiniova

„Byala voda”, 22.10.2003, 1 ♀; in moss, 01.11.2003, Distribution: Central Europe, Ukraine, Mace­ 1 ♂; 20.3.2004, 1 ♀; 23.6.2004, 1 ♀; 21.7.2004, 1 donia, Romania, ex Yugoslavia (present Serbia and ♀; 20.10.2004, 1 ♀; Marchaevo Village, 31.5.2004, Montenegro). 1 ♂, 5 ♀♀; 25.6.2004, 4 ♂♂, 3 ♀♀; 30.7.2004, 1 Bryaxis carinula (Rey, 1888) ♂, 1 ♀; in pitfall trap, 25.06 – 30.07.2004, 1 ♂, 3 Varna, Shumen (Karaman 1969, 1972). ♀♀; Zheleznitsa Village, place „Brezite”, 16.3.2004, New records: Chuipetlovo Village, 28.4.2004, 1 ♂; 30.3.2004, 6 ♀♀; 27.5.2004, 1 ♂, 1 ♀; in moss, 1 ♂, 2 ♀♀; near Bosnek Village, 900 m, 03.8.2004, 01.8.2004, 2 ♂, 3 ♀♀; 27.8.2004, 1 ♀; 22.9.2004, 1♂; Marchaevo Village, 25.6.2004, 1 ♂; 02.9.2004, 2 ♀♀; 24.10.2004, 1 ♂; above Rudartsi Village, 2 ♀♀; place „Byala voda”, 30.9.2004, 1 ♂. 31.5.2004, 4 ♂♂; 25.6.2004, 2 ♂♂, 7 ♀♀; 30.7.2004, Distribution: Central and Eastern Europe. 5 ♂♂, 9 ♀♀; 02.9.2004 , 2 ♂♂, 5 ♀♀; near Bosnek Village, 21.3.2004, 1 ♀; 29.5.2004, 1 ♀; 03.8.2004, Bryaxis roumaniae Raffray, 1904 4 ♂♂, 2 ♀♀; 23.8.2004, 1 ♂,1 ♀; 09.10.2004, 1 ♂; Vratsa, Maslen nos (Karaman 1972), Iskera­ near „Duhlata” Cave, 06.6.2004, 1 ♂, 1 ♀; 28.4.2004, novo, Kuleftse (Besuchet 1999 a), Belasitsa (Lapeva- 1 ♀; 23.8.2004, 1 ♀; 09.10.2004, 1 ♀; Chuipetlovo Gjonova 2004). Village, 28.4.2004, 1 ♂, 2 ♀♀; 23.8.2004, 1 ♀; Yarlovo New records: Chuipetlovo Village, 21.3.2004, Village, in leaf litter of F. sylvatica, 30.3.2004, 5 ♀♀; 1 ♂, 1 ♀; place „Byala voda”, 22.10.2003, 1 ♂; above Vladaya Village, 22.4.2004, 2 ♀♀; between 22.4.2004, 1 ♂; 18.9.2003, 1 ♂; between place place „Byala voda” and place “Kopitoto”, 22.4.2004, „Zlatnite mostove”and Vladaya Village, 19.5.2004, 2 ♂♂, 10 ♀♀; place “Kopitoto”, 19.5.2004, 1 ♂, 4 1 ♂, 2 ♀♀; between „Selimitsa” Hut and „Ostritsa” ♀♀; between place „Brezite” and place „Yarema”, Hut, in leaf litter of Salix sp., 05.10.2004, 1 ♀. 27.5.2004, 1 ♀; 03.7.2004, 1 ♀; Zheleznitsa Village, Distribution: Bulgaria, Greece, Romania, ex riverside of Selska reka River, 01.8.2004, 1 ♀; Yugoslavia (present Serbia and Montenrgro). 27.8.2004, 1 ♂; Kladnitsa Village, 05.10.2004, 1 ♂, Bryaxis curtisii orientalis (Karaman, 1952) 3 ♀♀; „Bor” Hut, 22.4.2004, 1 ♀. Rila – Samokov, Rila Monastery, Pleven, Distribution: Albania, Bosnia Herzegovina, Rhodopes – Lepenitsa, Zabardo (Karaman 1953, Bulgaria, Croatia, Hungary, Macedonia, Moldavia, 1957, 1969, 1972). Romania. New records: above Dragalevtsi Village, 24.9.2004, 1 ♂. Brachygluta fossulata (Reichenbach, 1816) Distribution: Central and Eastern Europe Sofia, Maglije, Varna, (Karaman 1961, 1969, 1972), German (Rambousek 1909), Tarnovo Bryaxis simoni (Reitter, 1880) (Netolitzky 1912). Vitosha, Rila – Borovets (Rambousek 1909), New records: near Zheleznitsa Village, in hay Stara Planina (Karaman 1957, 1969). litter, 1200 m, 27.4.2004, 1 ♂. New records: Vitosha, 28.05.1983, 1450 m, 1 Distribution: Europe, Asia. ♂, leg. S. Andreev (MHNG). The species is very common in almost all of the Bryaxis nodicornis Aubé, 1833 studied locations, from 660 m up to 1880 m. In view German Monastery (Rambousek 1909). New records: place „Byala voda”, 22.10.2003, of that we summarized the information and present 1 ♂; 20.3.2004, 2 ♂♂; 22.4.2004, 1♂; 19.5.2004, the data by months: March – 1 ♀; April – 1 ♂, 4 ♀♀; 1 ♂; 21.7.2004, 1 ♂; 30.9.2004, 1 ♂; 20.10.2004, May – 6 ♂♂, 8 ♀♀; June – 3 ♂♂, 14 ♀♀; July – 10 1 ♂; between place “Zlatnite mostove” and ♂♂, 18 ♀♀; August – 13 ♂♂ , 49 ♀♀; September – Vladaya Village, in leaf litter, 22.4.2004, 1 ♂; place 14 ♂♂, 34 ♀♀ ; October – 8 ♂♂, 19 ♀♀; November “Kopitoto”, 22.4.2004, 1 ♂. – 1 ♂, 2 ♀♀.

36 A Contribution to the Pselaphinae...

Distribution: Bosnia Herzegovina, Bulgaria, Bythinus leonhardinus Reitter, 1913 Greece, Macedonia, ex Yugoslavia (present Serbia Rila – Samokov, Varna, Longoza (Karaman and Montenegro). 1954, 1957, 1969, Besuchet 1999 a). New records: аbove Rudartsi Village, 31.5.2004, Bryaxis ullrichii Motschulsky, 1851 1 ♂; between Kladnitsa Village and „Selimitsa” Hut, Vitosha - Knyazhevo (Rambousek 1909). 31.5.2004, 1 ♂. New records: between Dragalevtsi Village and Distribution: Bulgaria, Greece, Macedonia. place „Bai Krastyo”, 24.4.2004, 1 ♂; in pitfall trap, 25.08 – 24.09.2004, 1♂. Tychus rufus Motschulsky, 1851 Distribution: Central and Eastern Europe Vitoscha-Knyazhevo (Rambousek 1909), Varna – Rusko selo (Karaman 1969). Bythinus lunicornis Reitter, 1884 New records: Bosnek Village, riverside of the Rila - Borovets, Samokov (Karaman 1954, Struma River, in leaf litter, 1000 m, 21.3.2004,1 ♂; 1957, 1969). place “Byala voda”, 30.9.2004, 1 ♂; 20.10.2004, 1 New records: place “Byala voda”, 21.7.2004, 1 ♂. ♂; „Momina skala” Hut, 19.5.2004, 2 ♂♂; between Distribution: Albania, Bosnia Herzegovina, place “Zlatnite mostove” and Vladaya Village, Bulgaria, Croatia, Greece, Hungary, Italia, Romania, 23.6.2004, 1 ♂; place „Yarema”, 27.5.2004, 1 ♂; Slovenia, ex Yugoslavia (present Serbia and 22.9.2004, 4 ♂♂; 24.10.2004, 3 ♂; near place Montenegro) „Lashkova mogila”, 27.8.2004, 1 ♂; 24.10.2004, 1 ♂; Chuipetlovo Village, 03.8.2004, 2 ♂♂; 29.5.2004, Tychus niger (Paykull, 1800) 1 ♂; 03.8.2004, 1 ♂; 09.10.2004, 2 ♂♂; between German Monastery (Rambousek 1909), Razgrad Kladnitsa Village and „Selimitsa” Hut, 31.5.2004, (Markovitch 1909), Rila – Samokov (Karaman 1 ♂; 30.7.2004, 1 ♂; between „Selimitsa” Hut 1955). and „Ostritsa” Hut, 25.6.2004, 1♂; place „Zlatnite New records: Bosnek Village, riverside of the mostove”, 02.9.2004, 1♂. Struma River, in moss, 1000 m, 09.10.2004, 1 ♂. Distribution: Bosnia Herzegovina, Bulgaria, Distribution: Europa, Asia (Turkey).

Crete, Greece, Hungary, Macedonia, Romania, ex Dicentrius rousi Besuchet, 1999 Yugoslavia (present Serbia and Montenegro). Vitoscha (Besuchet 1999 b).

Bythinus seidli Karaman, 1952 New records: place “Kopitoto”, 20.3.2004, 1 New records: place “Byala voda”, 19.5.2004, 2 ♂; „Tintyava” Hut, 23.6.2004, 1 ♂; place „Gyoreva ♂♂; 20.10.2004, 1 ♂; near place „Lashkova mogila”, cheshma”, 23.6.2004, 1 ♀; in pitfall trap, 21.7.2004, 27.8.2004, 1 ♂. 1 ♀; place „Zlatnite mostove”, 31.8.2004, 1 ♂; Distribution: Bulgaria, Rumania, ex Yugoslavia between place „Zlatnite mostove”and Vladaya Village, (present Serbia and Montenegro). 19.5.2004, 1 ♂; pitfall trap, 23.06 – 21.07.2004, 1 ♂; above Dragalevtsi Village, 24.4.2005, 1 ♀; 25.08.2004, Bythinus acutangulus lunifer Karaman, 1948 1 ♂; between Dragalevtsi Village and place „Bai Rila – Samokov, Rhodopes – Bachkovo Krastyo”, 25.5.2004, 1 ♂; 30.6.2004, 1 ♀; 23.7.2004, (Karaman 1969). 2 ♂♂, 1 ♀; 24.9.2004, 1 ♀; place „Bai Krastyo”, New records: Zheleznitsa Village, place „Brezi­ 25.5.2004, 4 ♀♀; 30.6.2004, 3 ♂♂, 2 ♀♀; 02.4.2004, te”, 03.7.2004, 3 ♂♂; 22.9.2004, 1 ♂; аbove Rudartsi 1 ♀; 15.11.2003, 1 ♀; „Aleko” Hut, 25.5.2004, 1 ♂; Village, 30.7.2004, 1 ♂. „Bistrishko branishte” Reserve, 1300 m, 24.4.2004, 1 Distribution: Bulgaria ♀; 25.5.2004,1 ♂; in pitfall trap in P. abies forest, 23.07

37 Bekchiev R, M. Shishiniova

– 25.08.2004, 1 ♀; pitfall trap in F. sylvatica forest, References 23.07 – 25.08.2004, 1 ♀; “Kechek” Summit, 27.5.2004, Besuchet C. 1957. Une technigue nouvelle pour la preparation de 1 ♂; between “Kechek” Summit and “Barite” Summit, l’edeague des Microcoleopteres. – Mitteilungen der schwei­ 27.5.2004, 1 ♀; Vitosha, near Bistritsa Village, 1200 zerishen entomologichen Gesellschaft, 30: 341-342. Besuchet C. 1999a. Psélaphides paléarctiques. Notes taxonomiqu- m, 2.7.1998, 1 ex., leg. Bayer (PHC). es et faunistiques (Coleoptera Staphylinidae Pselaphidae). Distribution: Bulgaria – Revue suisse de Zoologie, 106(1): 45-67. Besuchet C. 1999b. Le genre Dicentrius (Coleoptera Staphy- linidae Pselaphidae). – Mitteilungen der schweizerishen Conclusions entomologichen Gesellschaft, 72: 221-233. Drenovski К. 1942. Vierter Beitrag zum insektenfauna Bulgariens As a result of this study, twenty-three species from und Macedoniens (Apterygogena und Käfer). – Mitteilun­ gen der Bulgarischen Emtomologischen Gesellschaft in subfamily Pselaphinae of the Vitosha Mountain were Sofia, XII: 1-14. (In Bulgarian). determined. They belong to 12 genera and from them Karaman Z. 1953. Über neue Coleopteren aus Jugoslavien, 5 genera were new to Vitosha Mountain fauna. insbesonders aus Mazedonien. – Acta Musei Macedonici Scientarum Naturalium, 1(5): 91-116. A new species to the Bulgarian Fauna is Karaman Z. 1954. Über einige neue Coleopteren der Balkanfauna. Bibloporus mayeti. – Fragmenta balcanica, 1(6): 45-55. Karaman Z. 1955. Revision des Tribus Tychini (Col. Psel.) mit New to the Vitosha Mountain fauna are the besonderer Berücksichtigung der balkanischen Arten. following 17 species: Claviger testaceus, Euplectus – Acta Musei Macedonici Scientarum Naturalium, 3(4): 105-144. kirbii, Euplectus karstenii, Bibloporus minutus, Karaman Z. 1957. Die balkanischen Bythininen (Col. Pselaphi- Plectophloeus fischeri, Trychonix sulcicollis, dae). Ihre Systematik, Zoogeographie und Phylogenie. Trimium carpathicum, Brachygluta fossulatta, – Bioloski Glasnik (Periodicum biologorum), 10(3-4): 161-208. Bryaxis curtisii orientalis, Bryaxis carinula, Bryaxis Karaman Z. 1961. Neue Gruppierung der Arten des Genus Bra- nodicornis, Bryaxis roumanie, Bythinus lunicornis, chygluta Thomson (Coleoptera Pselaphidae). – Acta Musei Macedonici Scientarum Naturalium, 7(7): 127-174. Bythinus seidli, Bythinus acutangulus lunifer, Karaman Z. 1962. Über das Genus Plectophloeus Reitter (Eu- Bythinus leonhardinus, Tychus niger. plectini Psel. Col.). – Jahrbuch der Land und Forstwirst- chaftlichen Fakultät der Universität in Skopje, 15 (1961- Euplectus signatus and Trimmium caucasicum 1962): 145-153. (In Serbo-Croatian, German abstract). were not found, they had been reported for the Karaman Z. 1967. Revision der Gattung Trimium Aubé (Coll. Vitosha Mountain before. Pselph.). – Acta Musei Macedonici Scientarum Naturalium, 10(6): 131-173. Тhe presence of Trimium caucasicum in the Karaman Z. 1969. Über einige neue und interessante Pselaphiden Vitosha Mountain is uncertain. Possibly the question vertreter der Balkanhalbinsel. – Fragmenta balcanica, 7(3): 9-20. is about Trimium puncticeps, Rambousek (1909) Karaman Z. 1972. Vier balkanische und eine italienische neue did not use the aedeagi for identification, and since Pselaphiden Arten. – Fragmenta balcanica, 9(4): 37-51. the two species were morphologically similar, they Lapeva-Gjonova A. 2004. Pselaphinae (Col. Staphylinidae) from ant nest (Hym. Formicidae) in southwestern Bulgaria. could have been confused. – Acta zoologica bulgarica, 56(1): 67-73 The total number of established species from Löbl I., C. Besuchet 2004. Pselaphinae. – In: Löbl I., A. Smetana (eds.): Catalogue of the Palearctic Coleoptera. Vol. 2. subfamily Pselaphinae for the Vitosha Mountain Stenstrup. Apollo Books, 272-329 p. fauna is 24, near 23% from the all known species of Markovich А. 1909. Contribution for insect fauna in the neigh- borhood of Razgrad Town. – In: Sbornik za umotvorenia, the Bulgarian Pselaphin fauna. nauka i knizhnina. Sofia,XXV : 1-2. (In Bulgarian). Netolitzky. F. 1912. Eine sammelreise nach Bulgarien. – Coleo­ pterologische Rundschau, Wien, Bd. 1: 156-163. Acknowledgements: The authors thank to Mr. Claude Besuchet, Rambousek J. 1909. Příspěvek k poznaní bulharských Pselaphidů Mr. Peter Hlaváč, Mr. Stanislav Vit, Mr. Plamen Mitov and Mrs. a Scydmaenidů. – Časopis československé společnosti Albena Lapeva - Gjonova for their help and friendship. This entomologické, VI: 16-24. study was supported by fund “Scientific Research” of the Sofia University “St. Kliment Ochridski”, contract № 29/2004. Received: 24.09.2006 Accepted: 17.01.2007

38 A Contribution to the Pselaphinae... Принос към фауната на подсем. Pselaphinae (Coleoptera: Staphylinidae) на Витоша

Р. Бекчиев, М. Шишиньова

(Резюме)

Двадесет и три вида от Pselaphinae (Coleoptera, Staphylinidae) се съобщават за Витоша. От тях 17 са нови за фауната на планината, един вид (Bibloporus mayeti Guillebeau, 1888) е нов за фауната на България. Общият брой видове за Витоша е 24, което е около 23% от пселафината фауна на България.

39