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Doctoral and Master’s Programs @ Penn W a te r c olor by

Canopy walk, Tiputini Biodiversity Station, Ecuador Shena z K hi m j i 0 Our program, ranked 6th nationally, 4 offers a four-field approach to the study of . At Penn, Biological Anthropology students interested in and human behavior also benefit from interactions with faculty in Monogamy, Paternal other departments, such as Drs. Dorothy L. Cheney (Biology), Robert Kurzban Care and Pairbonds (Psychology), and Robert M. Seyfarth (Psychology). in Neotropical

Ph.D students are offered 5 years of full Primates support through the Benjamin Franklin Fellowships.

For questions regarding the Graduate Department of Anthropology Program contact the Graduate University of Pennsylvania 3260 South Street Coordinator Zoe Beckerman at (215) Philadelphia, PA 19104-6398 Primatology and 746-0409. Phone: 215-898-7461 www.sas.upenn.edu/anthro/ Anthropology @ Penn Field sites Some current collaborations: Genetics of monogamy and paternal care ( with Dr. T. Schurr Anthropology, Penn)

Owl Monkeys of the Argentinean Gran Chaco Principal A multidisciplinary project on the behavior, Investigator: Hormonal correlates of monogamy ecology, demography and conservation of owl and paternal care, (with Dr. C. Dr. Eduardo Fernandez-Duque monkeys in the Argentinean Chaco. Valeggia, Anthropology, Penn) Some of the current studies are: ƒ Male and female contributions to territoriality, parental care, and pair bond maintenance Chronobiology studies ƒ National program for the conservation of owl (with Drs. H, Eckert, Univ. Tubingen and monkeys and gallery forests in the Argentinean H. De la Iglesia, University of Gran Chaco Washington) ƒ Behavioral and hormonal correlates of biparental care I am a biological ƒ Paternity analyses and population genetics anthropologist interested in understanding the evolution of social Biodemography of systems. My research examines male-female red howlers relationships, pair bonding and paternal care in (with Dr. Rudy Rudran non-human primates. Smithsonian Although all the species that I study are socially Institution) monogamous, there are some fascinating differences in their degree of physical sexual dimorphism, the extent of affiliation between partners, the amount of paternal care provided and their activity patterns (i.e., nocturnal or diurnal). These differences make them an Monogamous Primates of the Ecuadorian Amazon excellent model to evaluate alternative hypotheses for the evolution and maintenance A comparative project on the behavioral and of monogamy and paternal care in primates and ecological strategies of four different species of early humans. pair-living primates. In collaboration with Dr. Anthony Di Fiore ( University), we are comparing the owl monkeys of the Argentinean Contact information: Gran Chaco with the owl monkeys, titis and sakis in , Phone: 001-215- 898-1072 the Amazon forests of Yasuní National Park in Social behavior and parenting Webpage: http://www.sas.upenn.edu/~eduardof/ Ecuador. (with Dr. S. Evans, Dumond Conservancy)