<<

Rivista Italiana di Prleontoloeia e Stratigrafia volume 1l0 no. l , pr IlI ;' ,t'.;r; I .a,prirzoo+

THE RUSSIAN PLATFORM AS A KEY REGION FOR VOLGIAN/ CORRELATION: A REVIE\tr OF THE MEDITERRANEAN FAUNAL ELEMENTS AND AMMONITE BIOSTRATIGRAPHY OF THE VOLGIAN STAGE

MIKHAIL A. ROGOV'

Receit-ted September 19, 2002; accepted October 3, 2003

Key uords. , Volgian, Tithonian, Boreal-Teth1'an correla- Introduction tion, amnronites, biostratigraphy, Russian Platform. Over the last few decades several papers devot- Abstract. A review of the ammonite faunas and biostratieraphy ed to the analysis of mixed Boreal-Tethyan associations of the uppern.rost Kimmeridgian - lowermosr Middle Volgian of Central of , and Tithonian-Volgian correlations (Zeiss Russia is presented. The major role of the Sub-n-rediterranean ammonites in the Volgian assemblages is described, providins additional detail to 2001Kutek & Zeiss 1997;Hoedemaeker 1991; Sachs et the existine scheme of Tithonian-Volgian correlations. The base of the a\.1979: Mesezhnikov 1989; Sey tr Kalacheva1993,1.997; Tithonian and the base of the Volgian are coincident, based on the asso- Schweigert 1.993, 1.994, 2000; Scherzinger Er Schweigert ciation of Neoc/retocerds steraspis-Lingulaticeras soLenoides.The Klimovi 1999) have been published. Unfortunately, material from Zone corresponds to nrost of the Hybonotum Zone. However, the top the Russian Platform was either not studied during those of the Hybonotum Zone, based on the distribution of Paralingulati- ceras, Íalls within the Sokolovi Zone.The allocation of r Tenuicostata investigations, or only ammonites never described and il- Subzone (with two horizons: neoburgense and puschi) in the Pseudo- lustrated before were mentioned. scythica Zone is proven. The neoburgense horizon may be correlated with the cíliata md penicillatum horizons of Neuburg and, probably, with the Semiforme Zone. The findings oÍ Lingulatìceras blaschkei, Pseu- Material and Methods dolissoceras, rnd Sutneria in the Panderi Zone of the Middle Tìthonian suggest that the base of the Middle Volgian may not be younger rhan This paper is based on the author's field investigations and col- the Middle Tithonian Fallauxi Zone. lections, and on data collected by PA. Gerasimoq M.S. Mesezhnikov and VV Mitta. The ammonites illustrated are kept in the Geological Institute Riassunto. Viene fornita una revisione delle faune ad ammoniti of Russian Academy of Sciences (RAS) (author's collection, no. GIN e della biostratigrafia dalla sommità del Kimmeridgiano alla parte più MIV MK), Paleontological Institute of RAS (PA. Gerasimov's collection, bassa del Volgiano medio della Russia centrale. Viene descritto il ruolo no. PIN 4861) and in the State Vernadsky Museum (VV Mitta's collec- principale degli ammoniti submediterranei nelle associazioni Volgiane. tion, authors collection, no. SGM BX 17) in Moscow. Unfortunately, the Ciò permette di specificare I'esistente schema di correlazione Titonia- maiority of the collection of M.S. Mesezhnikov (S. Petersburg, VNIGRI) no-Volgiano. Fondandosi sull'associazion e N eochetoceras sterasp is - Lin - remained inaccessible. Also, most of the Kimmeridgian-Middle Volgian gulaticeras solenoides, la base del Tironiano e la base del Volgiano coin- ammonites with Tethyan affinities, quoted by M.S. Mesezhnikov (Me- cidono. La Zona a Klimovi corrisponde alla maggior parte della Zona sezhnìkov et a|.1977; Blom et al. 1984; Mesezhnikov 1989; Yakovleva ad Hybonotum. Tuttavia la parte sommitale della Zona ad Hybonotum, 1993) have not yet been recovered from his collection. basata sulla distribuzione di Paralingulaticeras, ricad,e nella Zona a So- Hitherto, only one outcrop of the Russian Platform (Gorodish- kolovi. Nella Zona a Pseudoscythica viene dimostrata la ripartizione di chi, Fig. 1,3) yielding an Upper Kimmeridgian-Middle Volgian ammonite una Sottozona a Tenuicostata (con due orizzontí: neoburgense e puscbi). succession composed of Sub-mediterranean ammonites has been discov- lorizzonte a neoburgense può essere correlato con gli orizzonti a cj- ered. Last year all the levels containing Tethyan ammonites, identified liata e penicillatzz di Neuburg e, probabilmente, con la Zona a Semi- in the Gorodishchi section, were traced through a number of localities forme. I ritrovamenti nellaZona a Panderi dei medio-tìtonìani Lingu- in the middle Volga area (Fig. t), since it is important to ascertain that laticeras blaschkei, Pseudolissoceras e Sutneria suggeriscono che la base the sequences of Sub-mediterranean ammonites in different secrions del Volgiano medio può non essere più giovane della Zona a Fallauxi are coincident. As a result, the wide distribution of these ammonites del Titoniano medio. throughout the middle Russirn Sea became apparent.

Geological Institute of Russian Academy of Sciences, Pyzhevskii Lane,7,119017, Moscon, Russia. E-mail: [email protected] 322 l!. A. Rogn'

Llr[]q IIlz TFf ro x .lllll :ffirr 1 ITTTilTI + ffi rz LYSKO\'O llilllllìlltìllT s [-,] t: B iostratigraphy,: 6 PN - Parrderi Zone lH F_E r+ PU - pust'hilrorizotr [illlfllT]lrllll,lllll s z ll t\t 15 N - neoburgetT.rc horizon [l]llllllllllllllrillllllillll t : P - PseLrdoscythica SLrbzone ffi8 lllllllTllllTllTllTfl :s S - Sokolovi Zone Strr.:r llffl]l rss L - efinroti horizon K \\,\S1l N - N.srera.rprs -L.solenoides Beds lfffffffffl] n Fl, F2 - Fallax Subzone lflffllfltr 20 E Tflfl 168 ifEffiT8r ffiilr0r aIT------ffi -r Tttrl I ffi "

PI Pì N 't. >\." | \l \ì\tì L | , S krn 7 L N

Iol crnrost Miclc{lc Vrlgirn Rr'.rpino,6 Murzicr'. Z- lsrdr:

r.,i.i). lì - Istaritcs. l3 - Littgrlaticcr-.r.r, l-{ - ParalingtÌaticertslfotttttntcsicllr, l5 - outcrops Ioc;rlin'.

Ammonites, zonal nomenclature and correlation cr:rnrplc scc Kutek & Zciss 1997. p1.20. tir. l). M.rcroconchs h;rrc;r lol'er point of rib irrrc;rtion thiìt nlrkcs thcnr ve'rv sinril,rr to IlotLaiskyt klinot'i Since there ;ire different rìpprorches in conrprring (llov.riskr' IM]. As thc evolutionrrrv tcndcncies of rlicro-,rncl nrrrcroconchs the brse of thc Tithonian rnd Volgi'rn stirscs (Schwcisert in thc.t.rrrrutirTrltìnttes Ilot;'aisÀ1,,r ìine.rge rrre virirrble (sc'c';rlso dat;r on thc Klinrovj Zonc). gcrrus 1/oi;.ri,,,(-l'.r c:rn bc'subclivicìccl furthcr into 199-{t H:rntzpclsue ct .il. l99S). ir is nece sr.rrr to beqin rhe nricro- rncl rttlcroconchilte suLrgcner,t, rìlso rrcccptecl ior cocr',rl pcrisphinc- this re',,ierv with r description of thc ch:rrircteristics of ticl rr:rnrorite s ol Southcrn Ccrrrrrrrr' (Schu'cigc'rt 1996). F-rccpt tor S.lr- the uppenlos t Ki n-rr.nerid si;rn F:rl I ;rx Subzone (Auti ssio- natisphínacs sp. nor:. this Ievel inclucles corìlnron Lingulatic'cras solenoidcs dorcnsis Zone). (Quc'nstcdt) (lìoeov 2002b. pl.. iir. l) nttl Naot'l:ctocarts cx !t'. subnutt"t- lrnr (lrrrrt.) (lìogov2002e.pl. t,iir. 1,2):Ocbetoccr,T.ic\gr.zro (Oppel) Upper Kimnrcridgian (Pl. l. ii*. l51.,4spidoceras sp. intl., Gnrle.v,r sp. ind. tn(1 AulrcostelhLúils Arrtissiodorcnsis Zonc Zicgler, l9 62 ci. jasottoidcs (l)rrvlov), thor-rgh thcil presc'ncr'. up to the brsr. oi tlrc Lol cr Fallax Sr,rbzone Iloveiskv, 1941. This subzorrc ìs n idch'clisrribrrtcd \trlgirn. is cxtrcnrelv r.rrc. It is inrport:rnt to notc th,rt Ocy'rcloccr;r-,, rs we]l in thc nritl.llc Volgr lrc.r. lJ,rscrl ttn thc tlistributron of,rnr:ronites, rìris sutr- x Attl"ttostcplutta-t do rrot cxtcntl into thc Tìthonien ;rnri c,rrr Lre consiclered zonc nl;ì\' bc sLrbdividetl inro tl'o ;rrrts. s hìch c,rn be iurthcr consicle,rc.tl .rs ,rs gcncra charlctcristic oi the'Kinrnreridsi:rn (Schs'eìecrt, pc'r'. conrnr. (for' î,run,rì horizons. IJoth loels ilre lin()\\'n rrlso ixnr l)ollntl (Kuteli & Zciss Otlntotcras)\. Anroug thc oppcliids it is possibìc to notc thc plc\cncc ()f 1997). but the .rrtnronite rrssocirtions ol tìrc lìussirrn plrrttornr.ìrc nlorc ?T,n'ancllicct',ts tii.Jt',utt'iscttunt (l"ont;rnrrcs) (Corodishchi, lsrtlv sections; divcrsc. Thc ìorr cl prrt ol thc I:rrllrr SuLrzonc c()ntiì;ns rrunrcrous .S.lrlllIr lìogor l00lr. pl. l, iir. l; Pl. l. iig. 7) tttlNeotlrtoc'?rds ci.rcboilleti"tlutnt spltinctcsJìtlla.t (llovriskr') l\,l.nrl,,1r1,rcosta/,lttrtusstrbtrndorircll).rrlor),--1. (Bcrckhcnrcr & Hiilder) ([sach scction, Pì. l. lig.S; r'erv closc ro N. rcbou- jasottoitlcs (l).rvlorr).J. cr gr..llll-s-,ior1or.cl-.1-s (Cortcru), nrorc r,rrclv Nco- !cri.rtttiltt irr\elru.i1.rrJJ00.1'l.l.ri.l.i'rrc.rrrlre r{,1)(,irhr'lrllrrsrrh- r'lrctoterts c[. sttbnttdttrttn 1l:ont.7. LingtÌttittrts solenoitlas (Qucnstctlt) rrncl 't.onc. Ncotlx,tot'er.ts ci. rt'bouletitnntn (not illustrete cl) s,rs rrlso cluotccì unîreque nt,'i.'7i116131il.i sp. Thc uppcr part oi thc subzonc contit;ns nl:rinlv ironr the Uppe r Kinrnrerirlgi.rn oi the North Subc.rspiln rcgion (Bogcllnov S,tnnatispltittttcs sp. nor: IM, nr]. trlicrocorrchs ol this sf.ccics cli[icr ironr 193-{) pcrrritting thc corrclation ol thc uppcnrost lìalì;rr Subzone lith .S./.r1lr.r (lloveiskr') lnrl in its lol riLr coeificicnt (usurlll rbour 2.51 iol rhc raboulatinttlr horizon oi thc Southern (lcnrrrnr' (Rogov 2002r). Upper Jurrtssic of tbe \/olgt area 323

Fig.2 - Schenre of the Tithonian- Submeditemaoean Province (after Geyssmt, 1997; Sch- Subboreal Province (qmmorite succ€ssion Volgian correl;.rtion based on weiger! 1999; Schweigert et al., 199Q. Middle Titho ofthe Russian Plate) ammonite distribution. Ab- HORIZON brevi.rtions: U.Kirnrn. Up- ORBEDSWTIII - uer Kinrnreridgien, Autiss. - î fz Au t is s io doren s is. =F-: -l f J Z v>tr l-: z G -) tv iT F = F 1> ! J .:, I o =L -) F 4 +3 <= fz z >Z o O =i J J zf '; - Y2 al E q F )< Trì F >z Mocnrs- - frl J hcimcnsis l F - -J z Ruepcì lianunr t.l o : T [ì icclcnsc , nsrftrsrri ll Y

q v U lmcnsc O - u (pail) l q c0 a

Lower Volgian In]] ch:ìrirctcriz-cd bv,r vcry high point of rib furcatìon. The first appear- Klimovi Zone Michailov, 1964 mce ol Subdicboto,nocerLls (Sphinctoceras) r'Llso occurs in this horizon. In thc nriddle Vrlga art':r thc Klinrovi zonc'is subdivicled into Russirrrr hoirlizguLtticeras tnd Fontitnnesielln dilfer fronr the European the Neocltetotu'Lts stcl'Ltlfris Lingul"tticarus solutoides Bcris (rt the brrse gcner,r in thc;rbsc'ncc of ventrolatcnl tubercles and in the small shell of thc zone) tncl iltc lithograplicunt eJ'intot'i horizon. size. Ns'ertheless, t1'pe of ribbing rnd spectrum of variability with- . N. .tlcril-r?l.i L. solenoides Becls Roqori 2001. N. -\Ic'ir.'?l.i in Russi,rn and tvpical Mc'dìtc'rranc'an lmnronites rre the sxne. Thus, (Oppel) (Pl.l, fir. l0-11) occurrcnccs irre nunrcroLrs, u'hilst 1/or',ri.sÀ'tz Russirrr specìnrens nl;.rv be considered ls endernic species of the genera klintoti (If or,;riskr.), Lingulctticer,ts solenoitles (Qr"re nsteclt), Sunerid if. P,tralingul,rticer"ts a,nd FoiltLtnilesie ll"t.The efintoxi horìz,on corresponds bracheri Bc'rckhcnrer ;ìl'r' nlol'e rale (Pl. l, fig. l).These berls nr,rv cor- appro\inrrtelv to the cf. e)'stettense-noentsheinensis horizons of South-

respond to tha rueppelliLlttutlt tnLl rietllittgensis horizorrs lncl ln urr- e rn Gerrr.rrrr.rl, (Irir.2). The level rvith nunlerous Paralingulrtticeras is ab- nrnrecl horizon (Schs eigc'rt Ei Zeiss 1998) bets'een rhc ruappcllianutt scnt in Srvirbir (Schrvcigcrt, pers. cournr.) end Polrrnd. Probably these tnd eigeltingense horizons of Southcrn Gr'nlliìn\' (Ssrbie). A nunrbcr rnrnrorrites penetrilted into the nriddle Volg;r rrer froÌl the Northern of horizons frorl Soutlrcrn Gernrrrnr, :rlso cont:rin Nc,oty';cfot,rir-r ex gr. CrLrcrsus, n,lrere dimorphs o'i Paralingulaticer"ts - Fontannesiella pro-

steraspis (Oppel) tnd Litryul,tticetLts solatoidcs (Quenstc'clt) (Schwcigert, I ithogrdp hi cu nt (Fonurn nes) occur. pers. conrnr.), l,lrile 2l',r/ilgrrlaticerts;rre iLrsent ol'\'crv l'irre (rs ;n rhc riedlingensi,s horiz-on). The sepiìrrìtc lo'c'l with N. pracurrsor Zciss Sokolovi Zone Ilovaisky, 1941 (eigeltingense horizorr) h,rs not Lrccn found vct in thc Russirn Plrtiornr, Evcntu;rlhi it will be possible to subdivide this zone rnto two but rn rnr:ronite verv closc'to N. notlu/osunr Berclihenrer & H

Pseudoscythica Subzone Ilovaisky, 1941. In the nrajority of ; i"{ ? sections, <>nly I. pseudoscl,t/:ica (llovaisky) [M, m I is known from this level. I. schascbkozae (llor,aisk1,) ancl L (Ilovaisky) (Kutek & a 7r.jú;J ianscbini aì z (, .= c Al\1r\r()\lll I2 -= Zetss 1997) probablv occur in the sanre subzone. t> E==ll I "rsena o ' pl., fiia.10) iìre rArer. Vith the exception of the anrrronites, other r.nol- i r= ;-tE I ll lusc,r frr:rnr the neoburgense horizon h;rve a Sub-rlediterrlne.rn origìn. Belenrnites ,rre represented only bv Hil:olithes, lnd :r ver), divcrse bivelve ^c EE !i;itl asscnrbl;rsc irrcludes con.ìmon M),ophorelLt, and l:rrge Liosn"ea. For rhe I level. îirst tirre since the Procerìtltium is found at this This I =-H t =iiìi Neoburgense event is not defined in Poland and, probably, is connected to rn increasing ìnfluence of the North-Caucrsian Basin.The neobur- hr a ú,. { ;tr;Èi ll gelse horizon is s,ìdely distributcd jn Centr,rl Russìa. In edclition to the Vof gr r.Lre;r, A. neoburgense (Oppcl) is knorvn rrlso ironr the Ural River Li luEllri'l (llovrìskv & Florensky 1941, pI.23, fis.42,42a) and frorr the Moscow ;; ;rrea (S:rsonov 1953; not illustrated). The prcsence neoburgense z iÈ;t _,e of the,4. (Oppcl) lnd S. dsenLl (Oppel) neoburgense horizon is correlated with o l :E I ìÉ.ii ,ii* oJ I lr: -=:Éiì^:5i- the penicillatun md ciliata horizons of Neuburg (Fig.2). This horizon, probably corresponds to the Semiforme Zone of the other Tethyen areas, in r.hich the species,4. neoburgense (Oppel) a,nd S. .tserna (Oppel) also xre very nun'rerous (Kutek & \Wierzbowski 1986; Fòzy et al. 1994). ú, ., a o . puschi Horiz-on Kutek and Zerss, 1974. The index species of the Puschi Zonc oî Polend (Kutck & Zeiss 197.1) was later replaced by J , the Tenuicost:rtunr (Zeiss 1977),6ut the first nanre is more preferable l**È:sììllPiF' in Rnssir, x P tenuicostatun (Michailov) trlso occurs ìnrhe neoburgense Z horizon. It is probable that Mesezhnikov's (1982) opinton rh:;. P. tenu S O icosttttun (Michailov) occurs throughout the Pseudoscythica Zone was rvrong. In the Gorodishchi section P tenuicostatunt (Michailov) and P puschi Kutek & Zeiss (Rouoi' 2002, pl., fir.l3) occur approxirnately in q ìdentic;rf anrounts, but in the Polevye-Bikshiki secrion P. tenuicost"ttunt jur: :l (Michailov) stronuly prevrils. Danubisphinctes sp. (Pl.l, fig.1Z) also I'lillì1il occurs sporldic,rlly in the Polewe-Bikshikì sections and rare Pseudor:ìt'ga- tites a[f. seorsus (Oppel) are knorvn fronr the Gorodishchi section (Irie.1, 3). Based on the presence o'i Danubispbinctes spurius (Schneid) in the A I coeval deposits of Poland, the upper part of the Tenuicostata Subzone can be compared rvith the uppernrost Prlmatus Subzone of Neuburg. 2 5 I Middle Volgian Panderi Zone Rosanov, 1906 6 Some researchers (e.g. Callomon tr Birkelund 1982; Hantzpereue et al. 1998; Scherzinger & Schweigert 1999) suggest the presence of a gap between the Lorver and Middle Volgian. However, Fig. - Arnrlonite I rlnse charts in the Gorodishchi section (from due to the lÌradual chanses in the Pseudovigratitinae-Virgatitinae line- the uppermost Kimmeridgi;rn ro lorverr.r.rost Middle Voleian). rge, thìs is considered ìmprobable. In the lorvest 0.5 nr of the Panderi Abbre vi;rtions: A. - Aulacostephatrus; A.n. - neoburgense ho- Zone in the Gorodishchi section, Dorsopllnitinae occur together with rizon; An - Anaspidocerds; F - Fontannesielld; F2 - Fallax amnronites, which are closer to Pseutlooirgatites, than to Zar"tiskites. Subzone, uppcr prrt; I Ilotaisk)'a:L- Linguhticer,tslT - Therefore, we cannot be sure th:rt the base of the Panderi Zone tn Titrrtrn e I I i cera s : N - N e o c lt et o cerzr ; N.s.- L. s. N e oc b et o cerrts Russia ,rnd the base of the Scythicus Zone of Poland are coincident. steraspis- Lingulaticeras so/enoides Beds; P - Parz/ inguLtticeras; The allocirtion of the detached level rvith Zaraiskites contadictionis P.e. - efimovi horiz-on; P.p. - puschi horizon; Ps. - Pseudovir- (llovaisky) andZ. tliprosopa (Ilovaisky), as proposed by Zeiss (2OOl), gatites; S - Stntt,ttisphinctcs: St. - Sutilerid. ìs questionable. These anrmonites rre found together wirh Zaraisbites Litholosy: 1 - drrk-grri' cley; I qr,rl crlc.rreous clay; 3 - - s cyt b i c u s (Vischniakoff ), D ors o p I an it e s p an deri (d' O rbigny), Padovia silt: a - er;r1,, b - d:rrk-r.:r.ty;'{ - nr.rrl; 5 - bituminous shal1, paz:loti (Michrlski) (Mikhailov 1964) and correspond closely to the clay, 6 phosphorite nodules. r';rriabilitv of Z. scytbicus (Vischniakoff). Only the trmnronites named by Ilovaisky (19a1, p.128, pl. 26, fig. 50) as Z. contradictionis var. B differ appreciably front Z. settbicus (Vischniakoff). These .rnrntonites, which resenrble Ilouaiskya, are found in the phosphate nodules ho- rizon in VetÌianka. It is possible that they rvere redeposited from the Pseudoscythica Zone Ilovaisky, 1941 Lower Voleian. The Pseudoscythica Zone is subdivided ìnto trvo subzones The Pancleri Zone oÍ the Central Russia still cennot be subdi- Pseudoscvthicrr and Tenuicosrrrunr. vided into f;runal horizons. Hoq,ever, the presence of Zaraishites regu- Upper J urassic of the Volga area 325

laris Kutek in the bituminous shales of the Gorodishchi section and Z. deriZone cannot be younger than the Middle Tithonian Fallauxi Zone quenstedti (Rouillier), Z. scythicus (Vischniakoff) in the lower part of (Rogov 2002b). On the other hand, findings oí Zaraiskites regularis Panderi Zone suggests that in the future it will be possible to trace also Kutek in the Upper Tithonian of the Carpatho-Balkan zone allows in Central Russia the horizons identified in Poland by Kutek (1994). Z. us to compxre the uppermost part of the Panderi Zone with the part regularìs Kutek occurs also in the northern areas of the Russian plat- of Upper Tithonian Calpionella.Zone A (Kutek 1994; Kurek & Zeiss fornr (Sachs 1976, pl. 22, fig. 2). 1997). Ar present x more precise correlation of the Panderi-Virgatus In addition to Zaraishites, tlso Dorsoplanites panderi boundary with the Tethyan zonal succession is not possible, (d'Orbigny), D . tlors op la n is (Vischniakoff ), Pa.o loo ia p av looi (Michal- skl),Acuticostites acuticostatus (Michalski),2{. bitrifurcatus Mitta (Mitta I 993) are found in the Panderi Zone. Anrmonites with a Tethyan affin- Achnowledgemenrs. I wish to thank Dr. G. Schweigert (Stutt- ity are very rare. According ro Mesezhnikov (Mesezhnikov et al. 1977; gaft) for providing so much information concerning the ammonite suc- Blom et al. I 984; Yakovleva I 993), in the Gorodishchi section the Pan- cession of South Germany, including unpublished data. Dr. V V Mit- deri Zone contains unfrequent Pseudolissoceras, Sutneria, . ta and Dr. O. A. Erlanger from Moscow kindly loaned specimens of Unfortunately, these ammonites have yet to be found in his collec- Lingulaticeras blaschkei from the Panderi Zone. Biostratigraphic stud- tion stored in VNIGNI (S. Petersburg). R:r'e Lingulaticeras blaschkei ies of the Kimmeridgian-Volgian deposits and the presentation of the Cecca Er Enay (Pl.l, fig. l3,la) xre known from the Panderi Zone of results was nrade possible with the help of Dr. S. A. Kurenkov (GIN Chuvashiya and the Tver area (Rogov & Egorov 2002, fig. 1.2 a, b), as RAS, Moscorv). Valuable suggestions were made by Dipl.-Ing. A. Scher- well as in the Fallauxi Zone (Richteri Subzone) of Ardèche (Cecca Ec zinger (Hattingen) and by reviewers, Drs. G. Schweigert (Stuttgart) and Enay 1991). The presence of this species and anrmonites of the genera J.C.V Cope (Cardiff). This publication has been supported by RFBR Pseudolissoceras r'nd Sutneria, indicates that the lower oart of the Pan- grant no.03-05-64297.

PLATE I

All illustrated in natural size and collected by the author, except for marked specinrens.

l. Sutneria {f . bracheri Berckh. Klinrovi Zone, N. steraspis - L. solenoides Beds. Gorodishchi. (x2) GIN MIV 667/l;2. Sutneria cf . asema (Opp.). Pseudoscythica Zone, Tenuicostatum Subzone,neoburgense horizon. Gorodishchi (x2). GIN MK 576; 3. Sutneria asema (Opp.). Pseudoscythica Zone, Tenuicostatum Subzone, neoburgense horizon. Gorodishchi (xa). GIN MK 609; 4, 5. Lingulaticeras blaschhei (Cecca, & Enay). Panderi Zone. 4 - Per-vonrayskoe. SGM BX 17/1. Collected by VV Mitta; 5 - Kimry (Tver area). PIN 4861/25. Collected by PA. Gerasim ov;6. Haploceras sp. ,uv. Klimovi Zone, eJìmooi horizon. Gorodishchi. SGM 572-7 (x2); 7. ?Tarantelliceras franciscanum (Font.). Autissiodorensis Zone, Fallax Subzone. Isedy. GIN MIV 285; L Neochetoceras cf. rebouletianun (Berckh. t{. Hólder). AutissiodorensisZone, Fallax Subzone. Isady. GIN MIV 284; 9. Ne- ochetoceras nodulosum Berckh. Er Hólder. Klimovi Zone, not in situ (?N. ster.tspis-L.solenoid.es Beds). Murzicy. GIN MIV 630; 10. Neochetoceras cf. steraspis (Opp.). Klimovi Zone, N. stertlspis - L .solenoides Beds. Polevye-Bikshiki. GIN MIV 655; 17. Neochetoceras cf. steraspis (Opp.). Kli- movi Zone, N. steraspis - L. solenoides Beds. Gorodishchi. SGM 572-22: 12, 13. Paralingulaticer"ts efimooi Rogov Klimovi Zone, efimovi horizon. Gorodishchi. l2 - SGM 572-21; 13 - Holotype SGM 572-l l: 14. Fontannesiella sp., Klimovi Zone, efimoai horizon. Gorodishchi. SGM 522-6; 15. Ocbetocer,ts tf zio (Opp.). Autissiodorensis Zone, Fallax Subzone. Gorodishchi. GIN MIV 692/l;16. Anaspidoceras neobu,Eense (Opp.) [m], Pseudoscythica Zone, Tenuicostatum Subzone,neoburgense horizon. Polevye-Bikshiki. GIN MK 607;17. Danubìspbinctes sp. juv. Pseudoscythica Zone, Tenuicostatum Subzone, puscbi horizon Polevve-Bikshiki. GIN MK 589; 18. Pseudooirgatites tenuicostittzrr (Michailov), Pseudoscythica Zone, Tenuicostrtum Subzone, puschi horizon Polerrre-Bikshiki. GIN MK 582. 326 M. A. Rogot,

',t"{ ,l |II ;€l7?: I *$ I l5 18

PI,ATI.- I Uppar J rtr,t'sit' oJ'the \ltlg arcd 327

REFEIIENCES

Blonr G.1., Kuznetz.rx';r K.l. & Mcscz.hnikov M.S. (1934) - Mcsczhnikor' lvt.S. (1939) - Tithonirrn, Volsirrrr rìnd Portlxndi;rn Jur:issic-Cret:rccous boundrrrl bccls in the Miclcllc Vilg.ì st;lses (geologicrl trrrcl biologicrl cvcnts, corrclltion). In ;rncl Rjrs;rn ;rrciìs. txcLrrsion 0(r0. 27 Intern;rtionrrl Gcol. Soliolov B.S. (Ed.): Scclinrentarv cover of thc L,arth ir-r Congr., Moscoui 198-1. Guicle booli of thc excursiorrs 059. spi.rcc i.rnd timc. Strrtisrrrphl' rncl p;1le ontolos\': 100- 107. 060, 0(r6: 33--19, Moscori'. Moscos'. Iin Russien] Boqci:rnov A.A. (1934) - Hvdrochloric clo:rcs oI the l-ou'cr Za- Mcscz-hniliov N,l.S., D.rin l-.G., Kuznctzor';r K.I. "\ Y;rkovlcv:-r volz.hie . Ilrtll. Soc'. Natur. Alosc'o u. Scct. Góo1., l2: 315- S.lì (1917) - Jur:rssic/Crctiìceolrs bouncl,rrv bcds in thc 367, Moscos'. Iin Iìussiirn] Middlc Volg;r Arc;r (A Prospc'ctus to Gcological Excur- Crllorlon J.H. & Birkclund T. (1982) - Thc rrr.r.rnronirc z.rr.rcs sion): -1-l pp., [-cningrrd. of tlre Borcrrl Vrl!Ìi,ìn (Uppe r JLrrassic) in E:rst Grce n- Mikh.rilov N.P (196+) - Borc;rl l-:rtc Jur:rssic (l-orvcr Volgi:rn) land. In lrrrbrv A.li. E: Balkri,ill H.R. (Eds.): Arctic Gc- iìrìrnronites (Virg;rtosphinctin:rc). T1'dttsttct. Geologic,rI ologv ,rrrcl Gcopìrvsics. A'lcnt. Cdrtad. Soc. Pctntl. Gaol., Irtst., 107: Z-33. Moscou,. Iin Russien] 8: 3-19-369, C.rl garr,'. Mitt.r V.V. (1993) - Anrr.r.ror.ritcs ;rncl zon;rl str':rtiqrrphy of thc Cccc;r F. c\ Enav R. (199 l) - Les;tr.t.tr.nonitcs clcs z-r'rr.tc i Scnri- Middlc Volgien clcposits oi tl.re Ccntrrrl lìussir. 133 pp. fornrc ct ì Frrllauxi clu Tithonicltrc clc I'Arcle'chc (Sucl- Kiev. Iin Russien] [:st dc la lìr:rncc): strrtiqr:rphic, peleror-rtolouic, prlcro- Roeov M.A. (2002r) - Ar-rtissioclorensis Zonc (Upper Kim- biogéogrephie . Palaunttogr,rpltittt. A., 2 I 9. I -3: I -tì 7, nrcriclgirn) of thc Volgtr ;ì.rcr'r: iullnl()r1;tc rrssembhges, Stuttg:rrt. biostr:rti gr:ìph)i corrcl;rtion. In. : Boqd:rnc>r' N.A., Vierz-- Ijijzv I. (1988) - Upprcr Juressic f;rcics :rncl :Ìululonrtc sLrcccs- bitsl

I 7: 97- 102, Prtris. t lt i c'cras : its distribution, r.ni gr;rtior-rs ;rnd s i gnificance for Hrrrtz-pcrsuc P, B:ruclin Ir., Mitt;r V., Olfcricv A. & Z:rkh:rrol thc Borcrrl-Tcthvrn corrclltion. In: Bogchnov N.A., Vi- V. (1998) - The Upper Jur;rssic of thc VolrÌa Lr:rsirr: rrnr- crzbitskv VL,., Vishnevsk.tv:r VS. et al. (L,ds.): Modcrn r.r.ronitc biostr,rtigrtrphv iÌnd occurre ncc of orqillic-c.ì|- (ìucstions of geologr': 325-329, Moscorv. [in Russian]. bon rich facics. Correl:rtions Lrctl,ccrr IJore:rl-Subborcal Sachs VN. (Ed.) (I976) -Str:rtigr:rph1' of thcJur,rssic Systcnr of ;rnd Submcclitcrr;ìncrrr.ì Provitrccs. In Cr:-Lsquin-Solc:ru thc north of thc USSR. 136 pp. Moscorv. [irr Russian] S., Barrier E. (Eds.): Pcri-Tcthls Mcnroir -l: cpicr.rtonic Sachs V.N., Mcsczhniliov M.S. & Shulginr N.l. (1979) - Stretis- brsins oi Pcri-Tethr':rn pl;rtforurs. tllén. .1[us. nath. Hist. raphv of the Jur;rssic-Crct:rccous boundlrv beds in thc 1t,tt., | 79: 9-.1.1, P,1f i5. Borc:rl Bclt. In Sechs VN. (Ed.): UppcrJurrssic and their Hocdcnr:rcker Ph.J. (199 1) - Tethvan-bore;rl correl:rtions;tncl boundrrv n.ith Ce rt:rccous S1'sterl: 93- 102, Nor,osibirsk. thc Jr"rrr-Lssic-Crct;rccoLrs bou ncl;rrr'. Ncii',s/. Srllrigr;, 35. Iin Russì;rn] l: 37-6A, Bcrlirt & Stuttq:rrt. S;ìson()\'N.T. (1953) - Stretirlrephv of theJurrssic ilnd Ilov;riskv D.I. & Irlorcnskv K.P (19+l) - Lcs ;rrt'rn.ror.ritcs clu cìcposits of the Russi;rn pl:rtfomr. Bull. Soc. Ndtur. trloscou. Juressiquc sr,rpóricur clcs brrssins clcs riviòrcs Ourll e t Ilck. Scct. Géol., 28. 5: Zl-100, Moscos: [in Russi;u] Contribtftiorr d li cotnttlissttncc dc la géologic dc I'USSR.. So'I.l. & K;rhcho'a E.D. (1993) - Biostr;rtigr;rphicrl criteria of Nour'. Ser., l(5):7-lL)5, Moscou'. Iin Russirnl thc Jurrrssic-Crct:rce olrs bor-rndrrv for thc Russi:r. 60 pp. Kutck J. (1994) - Thc So'thicus Zonc (Midclle Volgien) in Po- S;rint-Petc'rsbure. Iin Russi:-rn] llnd: its ;ullnr()nitcs ;rncl biostrltigr;rphic snbclivisions. Scv Ll. 3. K;rl;rcho'i-t E.D. (1999) - Lol'cr Bcrriasian of Southe rr-r Ac'ta gcol. Poltntiu., ++/l-2. l-13. \Versz:ur';r. I)rinron'e (Frrr Lrrst Rtrssia) ,rncl tìrc problcrl of Borc;rl-

Kutck J. & Vicrzbos ski A. (1986) - A ne l' ;ìccount on thc Up- Tet h r';rn co rrcLrt io r.r. Pa l a a o gc ogr., Pt l tt a o c' l i n r., Pa l tt e o e ui., pcr Jurassic stretigrirph\' ,Lncl ,rnrrlorritcs of the Ctl>rsz- I 50: 19-63, An'rstcrrl:rrl. tvn succe ssion, Picne rrv Klippcn Be lt, Polrnd.,4 cta Caol. Scìrcrzinqcr A. & Schsciecrt G. (1999) - Die Anrnronrtcn- Polottic',t., 36/1: 29 | 3 15, W:rrszrtri';r. Iìr.rr rr en horiz-on tc clcr Ncr-rburg-Ii)nll;ìtiol'ì (Oberjurri, KutcliJ. & Ze iss A. (197+) - Tithoni:rn-Vrlsi:tn anrnronitcs frorl Sticllichc lir;rnkcnalb) :rncl ihrc Bcziclrungcn z-unr Vol- Brzost

Ac' tt C c o L P t t I t n t i c',t., 2 1/ 3 : 5 O 5- 5 12, \\'rrrs zrrs'rr. ll, Miirrcl'rcn. Kuteli J. & Zeiss A. (1992) - Thc hiehcst Kinrnrericlgi.rn encl Schu e igcrt G. (I99-1) - Subbore ,rlc Frlllncnclcnrcr.rtc' (Anrrno- l-orvcr Volgien in Ccntr:rl Pol:rncl; thcir ;rnunonites ,rncl noiclc.r) irl obcrcrr WeilÌjur:r (Oberliinrrlcrideiunr) dcr bicrstrrrtiurlphv. Act,t Geol. Priottitit.. 17/3-1: l0l-198. Schu;ibiscl.rcn Alb. Prrl'i1,5: l -l l- 155, Stutts:u't. W;lrszil\\'i.t. Sclrs'eiscrt G. (199+) - Ubcr ciniec bcnrcrlicns\\/crtc Ar.t.r.no- Mcse zhnikoi' M.S. (l9tì2) - Titl.ronian (Vrluian) Stese . Zorre s of nitcn inr Obcrliinrrleridgiur'ìl clcr Schl':ibischen Alb

the Jur:rssic Svstcrn in thc USSIì. Tr',ttts,tct. Itttcnlcpart. (Siìclrvestcletr tsch l.rnd ). S t t t ttgtrtt'r Il e i t r. N n t trktle., IJ. Sn,ttígr. Cont., lA:' 120-l+6. [-cningrecl. Iir-r Russien] l0.l: l- 15, Stuttgelt. 328 M. A. Rogoo

Schweigert G. (1996) - Die Hangende Bankkalk-Formation iirn A new approach to the correlation of the Upper Kim- schwàbischen Oberjura.Jber Mitt. obenbein. geol. Ver., meridgian Beckeri Zone across the Tethyan Sea. N. tb. N.F., 78: 281-308, Stuttgan. G e o l. Pakont., Abb., 202/ 3 t 3 4 5 -37 3, Stutt gart. Schweigert G. (1998) - Die Ammonitenfauna des Nusplinger Yakovleva S.P (Ed.) (1993) - Unified stratigraphical scheme of Plattenkalks (Ober-Kimmeridgium, Beckeri-Zone, Ul- the Jurassic deposits of the Russian Platform (1993).28 mense-Subzone, Baden-\liirttemberg). Stuxgarter B ein sheets, 71 pp., Saint-Petersburg. Naturkde., B. 267: 1-61, Stuttgart. Zeiss A. (1977) - Some ammonites of the Kletnice Beds (Up- Schweigert G. (2000) - New biostratigraphic data from the per Tithonian) and remarks on correlation of the Up- Kimmeridgian/Tithonian Boundary Beds of Srùl Ger- permost Jurassic. Aca Geol. Polonica.,27/3: 369-386, many In Hall R.L. 6c PL. Smith (Eds.): Advances in 'Warszawa. Jurassic Research 2000. GeoResearch Forurn,, 6: 195- Zeiss A. (2001) - Die Ammonitenfauna der Tithonklippen von 202,Zuerich-Ueticon. Ernstbrunn, Niederósterreich. Denhschr. Naturhist. Mus. rùlien. Schweigert G., Krishna J., Pandey B. 8c Pathak D.B. (199ó) - Wien., 6: I 15 pp.,