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VOL. 30, N° 2, 2011 Revue de Paléobiologie , Genève (décembre 2011) 30 (2): 619-684 ISSN 0253-6730 The Oppeliidae of the Acanthicum Zone (Upper Kimmeridgian) from Mount Crussol (Ardèche, France): ontogeny, variability and dimorphism of the genera Taramelliceras and Streblites (Ammonoidea) Cyril B AUDOUIN 1* , Patrick B OSELLI 2* & Didier B ERT 3* Abstract The Kimmeridgian outcrops of the Mount Crussol (in Ardèche), already long known for their abundant ammonites, contain many Oppeliidae. The study of the Acanthicum Zone (Upper Kimmeridgian) enables us to conclude that the microconchs of Taramelliceras compsum ( OPPEL ) and Streblites weinlandi ( OPPEL ) correspond to forms previously described by the authors as Glochiceras (Lingulaticeras ) crenosum QUENSTEDT and Creniceras dentatum ( REINECKE 7KHVWXG\RIDVLJQLÀFDQWVDPSOHIURPDSUHFLVHKRUL]RQ leads to treat both T. compsum ( OPPEL ) and 7SVHXGRÁH[XRVXP ( FAVRE ) as two morphological elements from the same paleobiological species. This is the same for S. weinlandi ( OPPEL ) and S. levipictus ( FONTANNES ), which are morphologically very close. Finally, studying the genera’s variability highlights mechanisms underlying it (heterochrony of the development and “laws” of covariation). Keywords Ammonoidea; Oppeliidae; Taramelliceras ; Streblites ; Ochetoceras ; Upper Kimmeridgian (Upper Jurassic); Crussol, Ardèche (France). Résumé Les Oppeliidae de la zone à Acanthicum (Kimméridgien supérieur) de la montagne de Crussol (Ardèche, France): ontogenèse, variabilité et dimorphisme des genres Taramelliceras et Streblites (Ammonoidea).- /HV DIÁHXUHPHQWV GX .LPPpULGJLHQ GH OD montagne de Crussol (Ardèche), bien connus pour leur richesse en ammonites depuis longtemps, contiennent en particulier une DERQGDQWHIDXQHG·2SSHOLLGDH/HXUpWXGHSRXUOD]RQHj$FDQWKLFXP .LPPpULGJLHQVXSpULHXU SHUPHWG·pWDEOLUTXHOHVPLFURFRQTXHV de Taramelliceras compsum ( OPPEL ) et Streblites weinlandi ( OPPEL FRUUHVSRQGHQWjGHVIRUPHVGpFULWHVSUpFpGHPPHQWSDUOHVDXWHXUV comme des représentants respectifs de Glochiceras ( Lingulaticeras ) crenosum QUENSTEDT et Creniceras dentatum ( REINECKE ). L’étude G·XQpFKDQWLOORQLPSRUWDQWSURYHQDQWG·XQKRUL]RQUHVWUHLQWFRQGXLWpJDOHPHQWjFRQVLGpUHU T. compsum ( OPPEL ) et 7SVHXGRÁH[XRVXP (FAVRE FRPPHGHX[H[SUHVVLRQVPRUSKRORJLTXHVGHODPrPHHVSqFHSDOpRELRORJLTXH,OHQHVWGHPrPHSRXU S. weinlandi ( OPPEL ) et S. levipictus ( FONTANNES TXLVRQWGHPRUSKRORJLHVWUqVSURFKHV(QÀQO·pWXGHGHODYDULDELOLWpGHFHVJHQUHVSHUPHWGHPHWWUHHQ pYLGHQFHOHVPpFDQLVPHVTXLODVRXVWHQGHQW KpWpURFKURQLHVGXGpYHORSSHPHQWHW©ORLVªGHFRYDULDWLRQ Mots-clés Ammonoidea ; Oppeliidae ; Taramelliceras ; Streblites ; Ochetoceras .LPPpULGJLHQ VXSpULHXU -XUDVVLTXH VXSpULHXU &UXVVRO Ardèche (France). I. INTRODUCTION monographs, but in a very typological way and without a very accurate stratigraphic calibration because of The historical site of the Mount Crussol (Fig. 1 and 2) their ancient collection (no bed-by-bed tracking). Since has provided many Kimmeridgian ammonites to the then, several works have highlighted the importance of SUHYLRXV DXWKRUV ZLWK D VLJQLÀFDQW SDUW FRQVWLWXWHG WKH LQWUDVSHFLÀF YDULDELOLW\ ZLWKLQ WKH 2SSHOLLGDH IURP by the representatives of the family Oppeliidae various ages ( HÖLDER , 1955; PALFRAMAN , 1966), and DOUVILLÉ , 1890. This family was particularly studied by the existence of a probably sexual dimorphism for that DUMORTIER & F ONTANNES (1876) and then FONTANNES family ( PALFRAMAN , 1966; M AKOWSKI , 1962; Z IEGLER , (1879). Many species of the genera Taramelliceras DEL 1974; Q UEREILHAC , 2009; K EUPP & R IEDEL , 2009). In CAMPANA , 1904, Streblites HYATT , 1900, Creniceras the main case of Taramelliceras , HÖLDER (1955) brought MUNIER -C HALMAS , 1892, Glochiceras HYATT , 1900 together different species in large morphological and and Ochetoceras HAUG , 1855 were described in their stratigraphic groups illustrating the polymorphism of this 1 79, rue Pierre Julien, F-26200 Montélimar, France; [email protected] 2 57 bis, avenue des Patriotes, F-26300 Bourg-de-Péage, France; [email protected] 3 Université de Bourgogne, Laboratoire Biogéosciences, UMR CNRS 5561, 6 bd Gabriel, F-21000 Dijon, France; [email protected]. * Groupe d’étude en Paléobiologie et biostratigraphie des Ammonites (G.P.A), F-04170 La Mure-Argens, France. 620 C. BAUDOUIN , P. BOSELLI & D. B ERT )LJ *HRJUDSKLFVHWWLQJDQGVLPSOLÀHGJHRORJLFDOPDSRIWKH&UXVVRODUHD genus. This applies to the broad group of Taramelliceras Submediterranean realm developed by the French compsum ( OPPEL , 1863) at the Upper Kimmeridgian. Group for the Study of the Jurassic ( HANTZPERGUES In genus Streblites , ZIEGLER UHFRJQL]HG VRPH et al ., 1997), completed and updated with the works of microconchs corresponding to forms assigned to the SCHWEIGERT (1999) and BAIER & S CHWEIGERT (2001). species “ Creniceras dentatum ” ( REINECKE , 1818); he noted that the stratigraphic range of genus Streblites coincides exactly with that of “ C. dentatum ” ( REINECKE ). II. GEOLOGICAL AND BIOSTRATIGRAPHICAL SETTING The recent harvest (C.B. & P.B.) of many Oppeliidae, done with a good stratigraphic tracking in the lower part of The historic area of Mount Crussol is located in Ardèche, the Upper Kimmeridgian in Mount Crussol, now allows RQ WKH ZHVWHUQ ÁDQN RI WKH 5K{QH 9DOOH\ GRZQ VRXWK XVWRFRQVLGHUWKHLUSDOHRQWRORJLFDOUHYLHZIRUDVSHFLÀF RIWKH5K{QH,VqUHFRQÁXHQFHMXVWDFURVV9DOHQFHFLW\ level of the Acanthicum Zone (Upper Kimmeridgian) 'U{PH)UDQFH 7KLVPRXQWULVLQJXSWRPKLJKW WDNLQJ LQWR DFFRXQW WKH LQWUDVSHFLÀF YDULDELOLW\ DQG is a small calcareous massif surrounded by Quaternary modern concept of the species in paleontology. The DOOXYLXP RI WKH JUHDW SODLQ RI 9DOHQFH )LJ 7KLV biostratigraphic framework used in this work (Fig. 3) site has long been known for its fossils abundance, and UHVXPHV WKH .LPPHULGJLDQ VWDQGDUG ]RQDWLRQ RI WKH numerous Jurassic outcrops dating from Toarcian to The Oppeliidae of the Acanthicum Zone (Upper Kimmeridgian) from Mount Crussol 621 Fig. 2: Localisation of the outcrop sections of Mount Crussol (from ATROPS S 7KH0DOOHWTXDUU\LV) 622 C. BAUDOUIN , P. BOSELLI & D. B ERT Fig. 3: Zonation of the Kimmeridgian (from HANTZPERGUES et al ., 1997). Tithonian can be observed. However the massif bulk is However, the sections he published do not exceed the made of limestones and marls from the Oxfordian and top of the Divisum Zone. In our work, the most recent WKH.LPPHULGJLDQVWDJHV7KDQNVWRPDQ\ROGTXDUULHVDV levels of those sections were completed using the ZHOODVQDWXUDORXWFURSVWKHVHTXHQFHRIWKHVHOHYHOVFDQ same numbering (Fig. 4 and 5). According to ATROPS , be observed (Fig. 2). The Jurassic of Mount Crussol has the Acanthicum Zone (and thus the base of Upper EHHQWKHVXEMHFWRIQXPHURXVZRUNVGRQHDORQJWLPHDJR Kimmeridgian) starts from bed No. 175, but beds No. (SAUTIER , 1854; L ORY , 1860; O PPEL , 1865; H UGUENIN , 175 to 178 form a very thick and currently inaccessible 1874; D UMORTIER & F ONTANNES , 1876; F ONTANNES , set that cannot be studied in our work. The Acanthicum 1879; R ICHE & R OMAN , 1921; R OMAN , 1950 - see Zone continues at least to the bed No. 197, but its upper HÖLDER & Z IEGLER , 1959 for a complete biography) OLPLWVWLOOKDVWREHFODULÀHG and more recently by ATROPS (1982) who focused his research on the Lower Kimmeridgian. The Oppeliidae studied in this work are mainly from the The log sections given by ATROPS (1982, p. 275-291, orderly working of beds No. 193 and 195 which form table 46-47) are a valuable and indispensable basis in a thin and easily accessible bundle containing abundant the study of the Kimmeridgian of the Mount Crussol. fauna. These levels are particularly visible around spot The Oppeliidae of the Acanthicum Zone (Upper Kimmeridgian) from Mount Crussol 623 Fig. 5: Outcrop section of the Acanthicum Zone of the Mallet TXDUU\ 1RDQGDWWKHWRSRIWKHJUHDW0DOOHWTXDUU\ )LJ 2, 4-5). Oppeliidae and Aspidoceratidae are predominant in the ammonite fauna of those beds (Fig. 6 and 7): Oppeliidae such as Taramelliceras compsum ( OPPEL , 1863) ([M] & [m]), Streblites weinlandi ( OPPEL , 1863) ([M] & [m]), Ochetoceras canaliferum ( OPPEL , 1863), and Aspidoceratidae: Aspidoceras acanthicum ( OPPEL , 3O ;, ÀJ Orthaspidoceras lallierianum (D’O RBIGNY 3O;,,ÀJ Sutneria cyclodorsata (MOESCH 3O;,,ÀJ 7KHUHDUHDOVR Nebrodites hospes ( NEUMAYR 3O ;, ÀJ Nebrodites gr. agrigentinus ( GEMMELLARO 3O ;,, ÀJ 1), representatives of the genus Discosphinctoides OLORIZ 3O;,ÀJ3O;,,ÀJ DQGUDUHO\ Aulacostephanus phorcus ( FONTANNES , 1876) (Pl. XII, ÀJ Phylloceras praeposterium ( FONTANNES , 1875) 3O;,ÀJ P. aff. VD[RQLFXP NEUMAYR , 1871 (Pl. XI, ÀJ Holcophylloceras polyolcum ( BENECKE , 1866) (Pl. ;,,ÀJ Ptychophylloceras ptychoicum ( QUENSTEDT , 3O;,ÀJ Lytoceras polycyclum NEUMAYR , 3O;,,ÀJ DQGWKHUHDUHDOVRVRPHQDXWLOXV with sinuous septa [ Pseudaganides cf. pseudaganiticus (SCHLOTHEIM @ 3O;,ÀJ $IHZEHOHPQLWHV Fig. 4: Section of the Acanthicum Zone of Mount Crussol. gastropods and bivalves complete the fauna. 624 C. BAUDOUIN , P. BOSELLI & D. B ERT Fig. 6: The ammonites’ genera in bed No. 193 (N=212). III. GOALS AND METHODS name. This approach is particularly convenient if the intermediaries between various possible morphotypes are The choice of a “paleobiological” approach UHFRJQL]HG LQWUDVSHFLÀFSRO\PRUSKLVP 7KXVHYHQLIDW Species concept in paleontology has undergone ÀUVWJODQFHWKHSDOHRELRORJLFDODSSURDFKVHHPVWREHDQ VLJQLÀFDQW FKDQJHV VLQFH WKH ÀUVW SDUW RI WKH WZHQWLHWK apparent loss of signal in the morphological information, century when the “typological species”
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  • Occurrences, Age and Paleobiogeography of Rare Genera Phlycticeras and Pachyerymnoceras from South Tethys

    Occurrences, Age and Paleobiogeography of Rare Genera Phlycticeras and Pachyerymnoceras from South Tethys

    N. Jb. Geol. Paläont. Abh. 283/2 (2017), 119–149 Article E Stuttgart, February 2017 Occurrences, age and paleobiogeography of rare genera Phlycticeras and Pachyerymnoceras from South Tethys Sreepat Jain With 24 figures Abstract: New data on two rare genera (Phlycticeras and Pachyerymnoceras) from the Callovian (Middle Jurassic) sediments of Kachchh, western India are presented with an update on their South Tethyan occurrences. This paper documents the earliest occurrence of the genus Phlycticeras from the entire south of Tethys (P. polygonium var. polygonium [M]) from latest Early Callovian sediments (= Proximum Subzone, Gracilis Zone). Further, in light of the new taxonomic data, the previously recorded early Middle Callovian P. gr. pustulatum [M] is reevaluated as also all other Phlycticeras occurrences from the Indian subcontinent. Data suggests that in Kachchh, Phlycticeras has a long range from the latest Early to Late Callovian interval. Additionally, two new macroconch species of Pachyerymnoceras are also described and illustrated from Late Callovian sediments. A critical review of previous records suggests that in Kachchh, Pachyerymnoceras is restricted to the Submediterranean interval of the Collotiformis-Poculum subzones of the Athleta Zone. A note on the paleobiogeography and probable migratory routes of these two genera to India and elsewhere is also suggested. Key words: Kachchh, Middle Jurassic, Late Callovian, Pachyerymnoceras, Phlycticeras. 1. Introduction nent, stratigraphically precise data is scarce both for Pachyceratidae BUckMAN (WAAGEN 1873-1875; BUck- Kachchh (Fig. 1) is a prolific Jurassic ammonite area in MAN 1909-1930; SPATH 1927-1933; KRISHNA & THIErrY the Indo-Madagascan faunal Province (South Tethys) 1987; SHOME & BARDHAN 2005) and Phlycticeratinae which has been extensively studied for its taxonomic, SPATH (WAAGEN 1873-1875; SPATH 1927-1933; JAIN 1997; biochronostratigraphic and paleobiogeographic signifi- BARDHAN et al.
  • The Transversarium-Bifurcatus Zone Boundary at Rocha (Peral Area, East-Centralalgarve, Portugal)

    The Transversarium-Bifurcatus Zone Boundary at Rocha (Peral Area, East-Centralalgarve, Portugal)

    The Transversarium-Bifurcatus Zone boundary at Rocha (Peral area, East-CentralAlgarve, Portugal) B.MARQUES * F. OLORIZ ** F. J. RODRIGUEZ-TOVAR ** P. S. CAETANO * * - Centro Estratigrafia e Paleobiologia da UNL, Faculdade de Ciencias e Tecnologia, Quinta da Torre, P-2825 Monte de Caparica, Portugal ** - Dpto. Estr. Paleont., Fac. Ciencias, Univ. Granada and Inst. And. Geol. Mediterranea (LA.G.M.) 18002 Granada, Spain pp. 109-125 Ciencias da Terra (UNL) Lisboa NQll 1992 3 figs., 2 pI. zones Bifurcatus/Bimammatum a ete reconnue une autre discontinuite, correspondanta celledu type II quisepareles RESUMO cycles 4.3-4.4 de HAQ et al. (1987); cette discontinuite avait deja ete individualisee sur la marge sud de I'Iberie. Palavras-chave: Amonites - Macroinvertebrados Dans I' intervalle entre la partie superieure de la zone bentonicas - Biostratigrafia - Ecostratigrafia ­ a Transversarium et la partie inferieure de la zone a Descontinuidades-s--Jurdssico superior-Oxfordiano-:-­ Bimammatum,l'analysedel'evolutionecostratigraphique, Algarve- Sui de Portugal. faite a partir des spectres de la faune, a montre que dans I'ensemble des cephalopodes, les ammonites sont les Este trabalho representa 0 primeiro estudo organismesles plus tolerantsau stressecologique,provoque, pormenorizado do limite das Zonas de Transversarium­ soitparla diminution del'ecospace, soitpardes arrivees de Bifurcatus no Algarve. Este limite esta associado, na area materiel terrigene; dans ces conditions, la faune benthique do Peral, a uma descontinuidade estratigrafica cujo hiato diminue considerablernent. Dans l'ensemble de la faune afecta, parcialmente, as Zonas de Transversarium e de recoltce on signale la presence, relativement abondante et Bifurcatus. Uma outra descontinuidade foi reconhecida no diversifice, desDichotomoceras,auparavantpascommuns. limite das Zonas de Bifurcatus-Bimarnrnatum que, nesta area, coincide com a descontinuidade doTipo II que separa osciclos4.3-4.4 de HAQet al.
  • Pseudosonninia, a New Genus of Oppeliid Ammonite (Haploceratoidea) from the Callovian (Middle Jurassic) of the Neuquén Basin, Argentina

    Pseudosonninia, a New Genus of Oppeliid Ammonite (Haploceratoidea) from the Callovian (Middle Jurassic) of the Neuquén Basin, Argentina

    VOLUMINA JURASSICA, 2019, XVII: 39–48 DOI: 10.7306/VJ.17.3 Pseudosonninia, a new genus of oppeliid ammonite (Haploceratoidea) from the Callovian (Middle Jurassic) of the Neuquén Basin, Argentina Horacio PARENT1, Alberto C. GARRIDO2, Günter SCHWEIGERT3, Luciano BRAMBILLA4 Key words: ammonites, Callovian, Los Molles Fm., Chacay Melehué, Argentina. Abstract. From the uppermost part of the Los Molles Formation (upper Lower and lower Middle Callovian) in Chacay Melehué (Neu- quén Province, Argentina), a group of hecticoceratine ammonites with a conspicuous morphology not assimilable to any known genus of this subfamily has been recently discovered. These forms are inflated oxycones with a prominent, sharp keel developed from the juvenile phragmocone up to the adult peristome. The new genus Pseudosonninia is established on the basis of these ammonites, with Pseudoson- ninia chacaymelehuensis n. gen. n. sp. as its type species. The new genus seems to be endemic to the Neuquén Basin and is recorded here from the upper Lower and lower Middle Callovian of Chacay Melehué and Río de Los Patos. INTRODUCTION zons of the Lower Callovian Bodenbenderi and Proximum zones, as well as from the highest levels of the Los Molles The Lower-Middle Callovian oppeliids from the upper Fm., a few meters below the Tábanos Fm., which are situated part of the Los Molles Fm. are poorly known throughout the above the Proximum Zone. Among these ammonites there Neuquén Basin (Fig. 1A). Besides several papers mention- are hecticoceratine morphotypes mostly belonging to Hecti- ing different species (e.g. Keidel, 1910; Groeber, 1918, coceras Bonarelli, 1893, but others correspond to an unde- Groeber et al., 1953; Stipanicic, 1965; Westermann, 1967; scribed genus.
  • Author's Personal Copy

    Author's Personal Copy

    Author's personal copy Geobios 47 (2014) 45–55 Available online at ScienceDirect www.sciencedirect.com Original article § Ammonite aptychi: Functions and role in propulsion a, b c,d Horacio Parent *, Gerd E.G. Westermann , John A. Chamberlain Jr. a Laboratorio de Paleontologı´a, IFG-FCEIA, Universidad Nacional de Rosario, Pellegrini 250, 2000 Rosario, Argentina b 144, Secord Lane, Burlington, ON L7L 2H7, Canada c Department of Earth and Environmental Sciences, Brooklyn College of CUNY, Brooklyn, NY 11210, USA d Doctoral Programs in Biology and Earth and Environmental Sciences, CUNY Graduate Center, New York, NY 10016, USA A R T I C L E I N F O A B S T R A C T Article history: Seven previous proposals of aptychus (sensu stricto) function are reviewed: lower mandible, protection of Received 7 June 2013 gonads of females, protective operculum, ballasting, flushing benthic prey, filtering microfauna and pump Accepted 9 December 2013 for jet propulsion. An eighth is introduced: aptychi functioned to actively stabilize the rocking produced by Available online 23 January 2014 the pulsating jet during forward foraging and backward swimming. Experiments with in-air models suggest that planispiral ammonites could lower their aperture by the forward shift of a mobile cephalic Keywords: complex. In the experiments, the ventral part of the peristome is lowered from the lateral resting (neutral) Ammonoidea position by the added ‘‘ballast’’ of a relatively thin Laevaptychus to an angle < 258 from horizontal with Functional morphology adequate stability to withstand the counter-force produced by the jet of the recurved hyponome. However, Aptychus Protection of the shell forms tested, only brevidomes with thick aptychi, e.g., the Upper Jurassic Aspidoceratidae with Feeding Laevaptychus and average whorl expansion rates, were stable enough to swim forward by jet propulsion at Locomotion about Nautilus speed ( 25 cm/s).
  • Abbreviation Kiel S. 2005, New and Little Known Gastropods from the Albian of the Mahajanga Basin, Northwestern Madagaskar

    Abbreviation Kiel S. 2005, New and Little Known Gastropods from the Albian of the Mahajanga Basin, Northwestern Madagaskar

    1 Reference (Explanations see mollusca-database.eu) Abbreviation Kiel S. 2005, New and little known gastropods from the Albian of the Mahajanga Basin, Northwestern Madagaskar. AF01 http://www.geowiss.uni-hamburg.de/i-geolo/Palaeontologie/ForschungImadagaskar.htm (11.03.2007, abstract) Bandel K. 2003, Cretaceous volutid Neogastropoda from the Western Desert of Egypt and their place within the noegastropoda AF02 (Mollusca). Mitt. Geol.-Paläont. Inst. Univ. Hamburg, Heft 87, p 73-98, 49 figs., Hamburg (abstract). www.geowiss.uni-hamburg.de/i-geolo/Palaeontologie/Forschung/publications.htm (29.10.2007) Kiel S. & Bandel K. 2003, New taxonomic data for the gastropod fauna of the Uzamba Formation (Santonian-Campanian, South AF03 Africa) based on newly collected material. Cretaceous research 24, p. 449-475, 10 figs., Elsevier (abstract). www.geowiss.uni-hamburg.de/i-geolo/Palaeontologie/Forschung/publications.htm (29.10.2007) Emberton K.C. 2002, Owengriffithsius , a new genus of cyclophorid land snails endemic to northern Madagascar. The Veliger 45 (3) : AF04 203-217. http://www.theveliger.org/index.html Emberton K.C. 2002, Ankoravaratra , a new genus of landsnails endemic to northern Madagascar (Cyclophoroidea: Maizaniidae?). AF05 The Veliger 45 (4) : 278-289. http://www.theveliger.org/volume45(4).html Blaison & Bourquin 1966, Révision des "Collotia sensu lato": un nouveau sous-genre "Tintanticeras". Ann. sci. univ. Besancon, 3ème AF06 série, geologie. fasc.2 :69-77 (Abstract). www.fossile.org/pages-web/bibliographie_consacree_au_ammon.htp (20.7.2005) Bensalah M., Adaci M., Mahboubi M. & Kazi-Tani O., 2005, Les sediments continentaux d'age tertiaire dans les Hautes Plaines AF07 Oranaises et le Tell Tlemcenien (Algerie occidentale).
  • La Sous-Famille Des Taramelliceratinae

    La Sous-Famille Des Taramelliceratinae

    Carnets de Géologie / Notebooks on Geology – Mémoire 2009/02 (CG2009_M02) La Sous-Famille des Taramelliceratinae (Ammonitina, Haploceratoidea, Oppeliidae) de l'Oxfordien moyen et supérieur (Zone à Plicatilis, Sous-Zone à Vertebrale - Zone à Bimammatum, Sous-Zone à Berrense) du Nord de la Vienne, France (Province subméditerranéenne) 1 Philippe QUEREILHAC Résumé : Dans la région étudiée du Nord du Poitou, la Sous-Famille des Taramelliceratinae est présente dans tous les niveaux stratigraphiques, hormis la Sous-Zone à Schilli (très peu représentée) et la Sous-Zone à Stenocycloides dont la faune est constituée uniquement de Perisphinctidae et de Trimarginites sp. Bien que certaines d'espèces soient connues et citées en littérature, elles sont souvent mal interprétées : Taramelliceras (Taramelliceras) dentostriatum (QUENSTEDT), T. (T.) callice- rum (OPPEL), T. (Proscaphites) anar (OPPEL). La localisation stratigraphique des différentes espèces a été faite par l'association d'autres faunes de même récoltes, et par la présence (ou l'absence) de marqueurs stratigraphiques précis reconnus: par exemple, Neomorphoceras chapuisi (OPPEL) = Zone à Transversarium, Sous-Zone à Luciaeformis ; Taramelliceras (? Taramelliceras) colleti (LEE) = Zone à Transversarium, Sous-Zone à Rotoides ; "Epipeltoceras semimammatum" (QUENSTEDT) = Zone à Bimammatum, Sous-Zone à "Berrense", horizon à Semimammatum ; la disparition de Neoprionoceras lautlingensis (ROLLIER) présent dans la Sous-Zone à Parandieri, mais absent dans celle à Luciaeformis. D'autres qui ont été incorrectement positionnées stratigraphiquement: T. (T.) dento- striatum (QUENSTEDT), T. (T.) callicerum (OPPEL), T. (T.) externnodosum DORN, ont été restaurées à leur emplacement exact, déterminé à partir des collectes de l'auteur et de celles d'autres collectionneurs qui avaient pris note de leurs relation stratigraphiques avec d'autres espèces.