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Maquetación 1 HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez ADVERTIMENT. L'accés als continguts d'aquesta tesi doctoral i la seva utilització ha de respectar els drets de la persona autora. Pot ser utilitzada per a consulta o estudi personal, així com en activitats o materials d'investigació i docència en els termes establerts a l'art. 32 del Text Refós de la Llei de Propietat Intel·lectual (RDL 1/1996). Per altres utilitzacions es requereix l'autorització prèvia i expressa de la persona autora. En qualsevol cas, en la utilització dels seus continguts caldrà indicar de forma clara el nom i cognoms de la persona autora i el títol de la tesi doctoral. No s'autoritza la seva reproducció o altres formes d'explotació efectuades amb finalitats de lucre ni la seva comunicació pública des d'un lloc aliè al servei TDX. Tampoc s'autoritza la presentació del seu contingut en una finestra o marc aliè a TDX (framing). Aquesta reserva de drets afecta tant als continguts de la tesi com als seus resums i índexs. ADVERTENCIA. El acceso a los contenidos de esta tesis doctoral y su utilización debe respetar los derechos de la persona autora. Puede ser utilizada para consulta o estudio personal, así como en actividades o materiales de investigación y docencia en los términos establecidos en el art. 32 del Texto Refundido de la Ley de Propiedad Intelectual (RDL 1/1996). Para otros usos se requiere la autorización previa y expresa de la persona autora. En cualquier caso, en la utilización de sus contenidos se deberá indicar de forma clara el nombre y apellidos de la persona autora y el título de la tesis doctoral. No se autoriza su reproducción u otras formas de explotación efectuadas con fines lucrativos ni su comunicación pública desde un sitio ajeno al servicio TDR. Tampoco se autoriza la presentación de su contenido en una ventana o marco ajeno a TDR (framing). Esta reserva de derechos afecta tanto al contenido de la tesis como a sus resúmenes e índices. WARNING. Access to the contents of this doctoral thesis and its use must respect the rights of the author. It can be used for reference or private study, as well as research and learning activities or materials in the terms established by the 32nd article of the Spanish Consolidated Copyright Act (RDL 1/1996). Express and previous authorization of the author is required for any other uses. In any case, when using its content, full name of the author and title of the thesis must be clearly indicated. Reproduction or other forms of for profit use or public communication from outside TDX service is not allowed. Presentation of its content in a window or frame external to TDX (framing) is not authorized either. These rights affect both the content of the thesis and its abstracts and indexes. UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez Hominin-carnivore interaction during the Pleistocene: Development of new methods for the study of human behavior through a transdisciplinary and evolutionary approach (Doctoral Thesis, Universitat Rovira i Virgili, Tarragona) Edgard Camarós, 2016 All rights Reserved Cover illustration, Yolanda González, 2015 Printed by: Tratamiento Gráfico del Documento, S.L. Avda. de los Castros s/n (Edif. Interfacultativo) 39005 Santander [email protected] Printed in Santander, Spain First Edition: November 2016. ISBN 978-84-608-3696-4 UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez I STATE that the present study, entitled “Hominin-carnivore interaction duringDepartment the Pleis- d’Històriatocene: Development i Història de of l’Art new methods for the recovery of humanInstitut behavior Català de through Paleoecologia a trans- Humanadisciplinary i Evolució and evolutionary Social approach”, presented by Edgard Camarós for the award of the degree of Doctor, has been carried out under my supervision at the of this university and at the (IPHES). Doctoral Thesis Advisors, Florent Rivals, PhD Jordi Rosell, PhD Principal Advisor Advisor th Tarragona, 10 November 2015 UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez A en Teo. El meu petit homínid. La millor co-evolució. La unión en el rebaño obliga al león a acostarse con hambre (Proverbio africano) UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez Acknowledgements 9 UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez Agraïments Doctoral Thesis A l’hora d’agraïr l’ajut o el suport rebut al final d’una Tesi Doctoral, un ha de pensar molt bé quin ha estat el camí que l’ha portat fins al dia de la lectura. Un ha de saber identificar qui- nes han estat les variables que han fet possible el camí. Tot i així, segur que n’hi ha que se’ns obliden però que han contribuït de manera especial. Primer de tot m’agradaria agraïr als directors de la Tesi, els Professors Florent Rivals i Jordi Rosell de l’Institut Català de Paleoecologia Humana i Evolució Social, el seu suport i mes- tratge al llarg de tot el procés. El seu ajut ha estat total i ha fet possible la recerca que ha cul- minat en aquesta Tesi. També agraeixo al director de l’IPHES, el Prof. Eudald Carbonell, el seu suport i sincer interès en la meva investigació, i l’haver creat un context ideal per a fer cièn- cia. La Tesi és un compendi d’articles i no puc més que agraïr tot l’ajut i col·laboració dels co-au- tors per a desenvolupar la nostra recerca. Especialment a la Marián Cueto i a en Luis Teira, amb qui més estretament he col·laborat. Aquesta Tesi s’ha dut a terme gràcies a una beca predoctoral FI (DGR2013) de la Generali- tat de Catalunya i AGAUR a l’IPHES, en el marc del Doctorat de la Universitat Rovira i Virgili. També gràcies a una beca per a estades a l’extranger BE-AGAUR a la (Alemanya) (2012 BE 00971). He rebut el suport econòmic dels projectes finançats pel MINECO HAR2010-19957/HIST i HAR2013-48784-C3-1-P, i dels projectes de l’AGAUR 2014-SGR-900 i 2014/100573. Agraeixo els IP del projectes la sevaEberhard confiança, Karls-Uni- ja que senseversität l’ajut Tübingen econòmic d’aquest projectes no hagués pogut dur a terme la recerca. La Tesi s’ha desenvolupat a l’IPHES, i agraeixo la col·laboració de tot els col·legues i compa- nys del centre. També he realitzat dues estades fora de sis mesos cadascuna. Una a la amb el Prof. Nicholas J. Conard, i l’altra a la amb el Prof. Pablo Arias. A tots dos i a tothom amb qui he col·laborat d’aquestes universitats els hi agraeixo tot el que he après. Uni- versität Tübingen Universidad de Cantabria Vull agraïr també a diverses institucions que m’hagin permès analitzar diferents contextos i materials arqueològics. Gràcies al Museu de València, , Família PlassardMuseo de la Grotte de Huesca, de Rouffignac Museo de de Altamira, França, MuseuInstituto Etnogràfic Internacional de la de Investigaciones Prehistóricas de Cantabriai (IIIPC), Museo de Ar- queología y Prehistoria de Cantabria (MUPAC),d’Alemanya Universidad i alde Cantabria, Consejería de Cultura del Gobiernode Bèlgica. de Cantabria Universität Tübingen, Ulmer Museum Institut fur Naturwissen- schafliche Archäeologie (Universität Tübingen) Royal Belgian Institute of Natural Sciences 11 UNIVERSITAT ROVIRA I VIRGILI HOMININ-CARNIVORE INTERACTION DURING THE PLEISTOCENE: DEVELOPMENT OF NEW METHODS FOR THE STUDY OF HUMAN BEHAVIOR THROUGH A TRANSDISCIPLINARY AND EVOLUTIONARY APPROACH Edgard Camarós Pérez Agraeixo al (Cantàbria) haver-me permès realitzar elsE. Camarós experiments vinculats amb aquesta Tesi en el marc d’un conveni amb l’IPHES. A les persones que m’han ajudat en algun moment de la meva recerca. Gràcies al Dr. San- Parque de la Naturaleza de Cabárceno tiago Borragán ( ), a la Prof. Katerina Harvati, Prof. Hervé Bocherens i Dra. Dorotheé Drucker ( ), Prof. Valentín Villaverde (Universitat de València), Prof. Carlos Díez ( ), Prof. Jesús Jordá Pardo (UNED), Dr. AlfredParque Sanchis de (Museu la Naturaleza de Prehistòria de Cabárceno de València), Prof. Pilar Utrilla i Prof. Lour- des Montes ( ) i Dr.Universität Fréderic Plassard. Tübingen Segur que m’oblido d’algú. Universidad de Burgos Però no em puc quedar aquí. Vull agrair-liUniversidad als Drs. Florent de Zaragoza Rivals i Jordi Rosell que m’acollissin aquell estiu a Teixoneres amb els braços oberts i m’ajudessin quan estaba literalment penjat, i establissin les bases reals del que és aquesta Tesi Doctoral. Ells saben que els hi dec molt.
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