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University of Florida Thesis Or Dissertation Formatting
SYSTEMATICS OF TRIBE TRICHOCEREEAE AND POPULATION GENETICS OF Haageocereus (CACTACEAE) By MÓNICA ARAKAKI MAKISHI A DISSERTATION PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORIDA IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY UNIVERSITY OF FLORIDA 2008 1 © 2008 Mónica Arakaki Makishi 2 To my parents, Bunzo and Cristina, and to my sisters and brother. 3 ACKNOWLEDGMENTS I want to express my deepest appreciation to my advisors, Douglas Soltis and Pamela Soltis, for their consistent support, encouragement and generosity of time. I would also like to thank Norris Williams and Michael Miyamoto, members of my committee, for their guidance, good disposition and positive feedback. Special thanks go to Carlos Ostolaza and Fátima Cáceres, for sharing their knowledge on Peruvian Cactaceae, and for providing essential plant material, confirmation of identifications, and their detailed observations of cacti in the field. I am indebted to the many individuals that have directly or indirectly supported me during the fieldwork: Carlos Ostolaza, Fátima Cáceres, Asunción Cano, Blanca León, José Roque, María La Torre, Richard Aguilar, Nestor Cieza, Olivier Klopfenstein, Martha Vargas, Natalia Calderón, Freddy Peláez, Yammil Ramírez, Eric Rodríguez, Percy Sandoval, and Kenneth Young (Peru); Stephan Beck, Noemí Quispe, Lorena Rey, Rosa Meneses, Alejandro Apaza, Esther Valenzuela, Mónica Zeballos, Freddy Centeno, Alfredo Fuentes, and Ramiro Lopez (Bolivia); María E. Ramírez, Mélica Muñoz, and Raquel Pinto (Chile). I thank the curators and staff of the herbaria B, F, FLAS, LPB, MO, USM, U, TEX, UNSA and ZSS, who kindly loaned specimens or made information available through electronic means. Thanks to Carlos Ostolaza for providing seeds of Haageocereus tenuis, to Graham Charles for seeds of Blossfeldia sucrensis and Acanthocalycium spiniflorum, to Donald Henne for specimens of Haageocereus lanugispinus; and to Bernard Hauser and Kent Vliet for aid with microscopy. -
South American Cacti in Time and Space: Studies on the Diversification of the Tribe Cereeae, with Particular Focus on Subtribe Trichocereinae (Cactaceae)
Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2013 South American Cacti in time and space: studies on the diversification of the tribe Cereeae, with particular focus on subtribe Trichocereinae (Cactaceae) Lendel, Anita Posted at the Zurich Open Repository and Archive, University of Zurich ZORA URL: https://doi.org/10.5167/uzh-93287 Dissertation Published Version Originally published at: Lendel, Anita. South American Cacti in time and space: studies on the diversification of the tribe Cereeae, with particular focus on subtribe Trichocereinae (Cactaceae). 2013, University of Zurich, Faculty of Science. South American Cacti in Time and Space: Studies on the Diversification of the Tribe Cereeae, with Particular Focus on Subtribe Trichocereinae (Cactaceae) _________________________________________________________________________________ Dissertation zur Erlangung der naturwissenschaftlichen Doktorwürde (Dr.sc.nat.) vorgelegt der Mathematisch-naturwissenschaftlichen Fakultät der Universität Zürich von Anita Lendel aus Kroatien Promotionskomitee: Prof. Dr. H. Peter Linder (Vorsitz) PD. Dr. Reto Nyffeler Prof. Dr. Elena Conti Zürich, 2013 Table of Contents Acknowledgments 1 Introduction 3 Chapter 1. Phylogenetics and taxonomy of the tribe Cereeae s.l., with particular focus 15 on the subtribe Trichocereinae (Cactaceae – Cactoideae) Chapter 2. Floral evolution in the South American tribe Cereeae s.l. (Cactaceae: 53 Cactoideae): Pollination syndromes in a comparative phylogenetic context Chapter 3. Contemporaneous and recent radiations of the world’s major succulent 86 plant lineages Chapter 4. Tackling the molecular dating paradox: underestimated pitfalls and best 121 strategies when fossils are scarce Outlook and Future Research 207 Curriculum Vitae 209 Summary 211 Zusammenfassung 213 Acknowledgments I really believe that no one can go through the process of doing a PhD and come out without being changed at a very profound level. -
Espostoa Br.& R
2021/09/25 08:03 1/10 Espostoa Br.& R. Espostoa Br.& R. Cet article est basé sur l'original écrit par Graham Charles et publié dans le British Cactus and Succulent Journal 17(2): 69-79 (1999). Vous en trouverez une traduction sur le Cactus Francophone : Espostoa Br.& R. Il est ici modifié et corrigé à la lumière de nouvelles découvertes. Notez que dans l'article original, la fig 1. à la page 68 a été incorrectement légendée. On doit lire “Espostoa melanostele dans le Canyon Tinajas, près de Lima GC157.04” Although cerei are not the most popular cacti for growers restricted by cultivation in glasshouses, almost all collections have at least one representative of this beautiful genus. Most species are easy to cultivate and the hairy stems of many species make them an attractive addition to any collection, even in a small glasshouse. In pots, they are slower growing than many cerei, a factor which also makes them good show plants, frequently seen winning the Cereus class. For exhibitors, those species with stems covered with white hair catch the judge's eye. The hair can also hide minor blemishes on the stem which would more likely get spotted and count against other less covered species! In keeping with the fashionable trend towards the broader concept of genera, Espostoa has been expanded in recent years to include species previously included in the separate genera Pseudoespostoa Backbg., Thrixanthocereus Backbg. and Vatricania Backbg. Recent molecular studies have shown that Vatricania is not an Espostoa and so it will be excluded from this revised article. -
Connoisseurs' Cacti
ThCe actus Explorer The first free on-line Journal for Cactus and Succulent Enthusiasts 1 Siccobaccatus 2 Morangaya pensilis Number 17 3 Espostoa in Tenerife ISSN 2048-0482 4 Barranco Rambla de Ruiz December 2016 5 Juab and Utah County The Cactus Explorer ISSN 2048-0482 Number 17 December2016 IN THIS EDITION Regular Features Articles Introduction 3 A naturalised population of News and Events 4 Espostoa melanostele on Tenerife 21 In the Glasshouse 9 Travel with the cactus expert (16) 25 Journal Roundup 14 Where lizards dare: an excursion to Barranco On-line Journals 15 Rambla de Ruiz (Tenerife) 29 The Love of Books 18 Juab and Utah County, Utah, throughout the Society Pages 51 year 2015 36 Plants and Seeds for Sale 55 A Happy Medium? Morangaya pensilis . 41 Books for Sale 62 Cover Picture: Siccobaccatus dolichospermaticus See page 9 The No.1 source for on-line information about cacti and succulents is http://www.cactus-mall.com The best on-line library of succulent literature can be found at: https://www.cactuspro.com/biblio/en:accueil Invitation to Contributors Please consider the Cactus Explorer as the place to publish your articles. We welcome contributions for any of the regular features or a longer article with pictures on any aspect of cacti and succulents. The editorial team is happy to help you with preparing your work. Please send your submissions as plain text in a ‘Word’ document together with jpeg or tiff images with the maximum resolution available. A major advantage of this on-line format is the possibility of publishing contributions quickly and any issue is never full! We aim to publish your article quickly and the copy deadline is just a few days before the publication date. -
Cactus Explorers Journal
Bradleya 34/2016 pages 100–124 What is a cephalium? Root Gorelick Department of Biology and School of Mathematics & Statistics and Institute of Interdisciplinary Studies, Carleton University, 1125 Raven Road, Ottawa, Ontario K1S 5B6 Canada (e-mail: [email protected]) Photographs by the author unless otherwise stated. Summary : There are problems with previous at - gibt meist einen abgrenzbaren Übergang vom tempts to define ‘cephalium’, such as via produc - photosynthetisch aktiven Gewebe zum nicht pho - tion of more hairs and spines, confluence of tosynthetisch aktiven und blütentragenden areoles, or periderm development at or under - Cephalium, die beide vom gleichen Triebspitzen - neath each areole after flowering. I propose using meristem abstammen. Cephalien haben eine an - the term ‘cephalium’ only for a combination of dere Phyllotaxis als die vegetativen these criteria, i.e. flowering parts of cacti that Sprossabschnitte und sitzen der vorhandenen have confluent hairy or spiny areoles exterior to a vegetativen Phyllotaxis auf. Wenn blühende Ab - thick periderm, where these hairs, spines, and schnitte nur einen Teil der oben genannten Merk - periderms arise almost immediately below the male aufweisen, schlage ich vor, diese Strukturen shoot apical meristem, and with more hairs and als „Pseudocephalien“ zu bezeichnen. spines on reproductive parts than on photosyn - thetic parts of the shoot. Periderm development Introduction and confluent areoles preclude photosynthesis of Most cacti (Cactaceae) are peculiar plants, cephalia, which therefore lack or mostly lack even for angiosperms, with highly succulent stomata. There is almost always a discrete tran - stems, numerous highly lignified leaves aka sition from photosynthetic vegetative tissues to a spines, lack of functional photosynthetic leaves, non-photosynthetic flower-bearing cephalium, CAM photosynthesis, huge sunken shoot apical both of which arise from the same shoot apical meristems, and fantastic stem architectures meristem. -
Bradleya 31/2013 Pages 142-149
Bradleya 31/2013 pages 142-149 Coleocephalocereus purpureus has a cephalium; Micrantho - cereus streckeri has a pseudocephalium (Cereeae, Cactoideae, Cactaceae) Root Gorelick Department of Biology, School of Mathematics & Statistics, and Institute of Interdisciplinary Studies Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6 Canada. (email: [email protected]). Photographs by the author Summary : The putatively closely related cactus Introduction genera of Coleocephalocereus , Micranthocereus , Cactaceae (cacti) in the tribe Cactoideae have Cereus , Monvillea , and Stetsonia have a wide a wide range of reproductive anatomies ranging range in specialization of reproductive portions of from cephalia to pseudocephalia to forms where the shoot, from cephalium to pseudocephalium to reproductive and vegetative structures are indis - no specialization. After briefly summarizing the tinguishable (Buxbaum, 1964, 1975; Mauseth, shifting uses of the terms ‘cephalium’ and ‘pseudo - 2006). cephalium’, I provide gross morphological evi - For instance, Melocactus Link & Otto, Disco - dence that Coleocephalocereus purpureus has a cactus Pfeiffer, and Espostoa Britton & Rose have true cephalium that is formed of a continuous true cephalia in which the flowering parts are not swath of bristles and hairs, with its underlying photosynthetic because every epidermal cell con - thick cortex of parenchyma replaced by a narrow tains a modified leaf that is a hair, bristle, or layer of cork. By contrast, Micranthocereus streck - spine, with no stomata amongst the epidermal eri has a pseudocephalium composed of nothing cells (Mauseth, 2006). Furthermore, there are more than larger hairier areoles in which the un - changes to the internal anatomy of cephalia, derlying epidermis is still photosynthetic and the where the underlying cortex is not a wide swath of underlying cortex is still a thick layer of highly succulent parenchyma, but instead a thin parenchyma without any noticeable cork. -
Chapters 13-14
13. ROCKERY IX This feature runs in a North-South direction from near the Western end of Rockery VIII. It is a small feature, 21 meter long and about 11 meters wide at the middle. On this feature are grown plants belonging to the collection of the late SHRI O. P. BANSAL, who was a keen plant grower. For several years he was associated with the Rose Society of India. In his later years he took to growing cacti. He expired in 1994, when be was actively involved in the activities of the National Cactus and Succulent Society of India, Chandigarh. After his sudden death in 1994 the family expressed the desire to donate the plants to this Garden. So after building Rockery VIII for MR. G. C. JAIN, this rockery came into being. MR BANSAL had a small collection, but quite a large number were big 175 specimen plants. Most remarkable were three mature plants of Melocactus maxonii with large cephalium, big groups of Coryphantha elephantidens, and several large Ferocactus herrerae. Out of the three big Melocacti, one still survives near the Northern end. The others did not survive long. Now several younger plants have been planted alongside this specimen. Starting from the Northern end there is an Agave americana variegated plant. In the background of this rockery there are several large Ritterocereus pruinosis, Yucca aloefolia, Kalanchoe behariensis, Aloe arborescens and some Agaves. There are three Opuntia mamillata fa. monstrose plants on this feature. A large Espostoa lanata is near the Northern end. There is a larger plant of Napolea cochenillifera. -
Connoisseurs' Cacti
TheCactus Explorer The first free on-line Journal for Cactus and Succulent Enthusiasts Quebrada Botija, Chile Parodia turbinata Number 6 Copiapoa taltalensis ISSN 2048-0482 Echeveria trianthina November 2012 Navajoa fickeisenii The Cactus Explorer ISSN 2048-0482 Number 6 November 2012 IN THIS EDITION Regular Features Articles Introduction 3 Parodia turbinata : a confused taxon 26 News and Events 4 Discocactus horstii in Habitat 34 Recent New Descriptions 15 A Day Trip to the Peak of Botija 37 In the Glasshouse 17 A must see site - Suripujio 43 Journal Roundup 20 Echeveria trianthina and E. halbingeri The Love of Books 22 var. sanchez-mejoradae 46 Cactus People Histories 24 Matucana aurantiaca in the Cordillera Blanca 49 Society Page 62 Copiapoa taltalensis in the vicinity of Esmeralda 52 Plants and Seeds for Sale 65 Travel with the Cactus Expert (5) 57 Books for Sale 68 Last minute additions Ferocactus cylindraceus or F. acanthodes ? 25 iSpot South Africa 48 The No.1 source for on-line information about cacti and succulents is http://www.cactus-mall.com Cover Picture Copiapoa atacamensis and Eulychnia iquiquensis at the top of Pico Botija. Photo: Philippe Corman Invitation to Contributors Please consider the Cactus Explorer as the place to publish your articles. We welcome contributions for any of the regular features or a longer article with pictures on any aspect of cacti and succulents. The editorial team is happy to help you with preparing your work. Please send your submissions as plain text in a ‘Word’ document together with jpeg or tiff images with the maximum resolution available. -
Espostoa(Vatricania)
Haseltonia58 26: 58–67. 2019 Haseltonia 26: 58–67. 201958 Espostoa (Vatricania) guentheri have unusual cephalia or pseudocephalia Root Gorelick Department of Biology and School of Mathematics & Statistics and Institute of Interdisciplinary Stud- ies Carleton University, 1125 Raven Road (unceded Algonquin territory),Ottawa, Ontario K1S 5B6 Canada. Email: [email protected] Manuscript received 19th August 2019 Abstract: Espostoa (Vatricania) guentheri is a peculiar putative Espostoa, having some traits in common with Espostoa sensu stricto (cephalia in which many or most epidermal cells produces a spine, bristle, or trichome), some traits in common with Thrixanthocereus (cephalia with bristles instead of hairs; patches of epidermal cells in the cephalium that lack spines, bristles, hairs or trichomes), and some unique traits (gradual transition to cephalium formation; disjunct habitat). Gradual cephalium formation in E. guentheri is common, but not universal, which might be either because Vatricania is not closely related to Espostoa or that Vatricania is of hybrid origin. E. guentheri may have a pseudocephalium, rather than a true cephalium, with the reproductive part of shoots in subgenus Vatricania being morphologically intermediate between those of subgenus Espostoa and subgenus Thrixanthocereus. Keywords: cephalium, Espostoa, disjunct species, hybrid origin, pseudocephalium, Thrixanthocereus, Vatricania INTRODUCTION Previously (Gorelick, 2016: 118), I defined a cephalium as: part of a cactus shoot arising directly from the shoot apical meristem, with the cephalium composed of confluent areoles from which flow- ers originate, bearing copious spines and trichomes, and underlain by a thick periderm in lieu of an even thicker cortex. Bristles and hairs on flowering parts are longer than those on non-flowering parts. -
(Cactaceae Juss.) Species
Acta Agrobotanica DOI: 10.5586/aa.1697 ORIGINAL RESEARCH PAPER Publication history Received: 2016-05-24 Accepted: 2016-10-03 Anatomical and morphological features Published: 2016-12-20 of seedlings of some Cactoideae Eaton Handling editor Barbara Łotocka, Faculty of Agriculture and Biology, Warsaw (Cactaceae Juss.) species University of Life Sciences – SGGW, Poland Halyna Kalashnyk1*, Nataliia Nuzhyna2, Maryna Gaidarzhy2 Authors’ contributions 1 HK: carried out the experiments Department of Botany, Educational and Scientific Center “Institute of Biology and Medicine”, and wrote the manuscript; Taras Shevchenko National University of Kyiv, S. Petlyury 1, Kyiv 01032, Ukraine 2 NN: designed the anatomical Scientific laboratory “Introduced and natural phytodiversity”, Educational and Scientific Center experiment and contributed “Institute of Biology and Medicine”, Taras Shevchenko National University of Kyiv, S. Petlyury 1, to data interpretation; MG: Kyiv 01032, Ukraine designed the experiment, * Corresponding author. Email: [email protected] critically read the manuscript and contributed to data interpretation Abstract Funding Three-month-old seedlings of 11 species of the subfamily Cactoideae Melocac( - This study was financed from the research project tus bahiensis, Melocactus curvispinus, Echinopsis eyriesii, E. mirablis, E. peruviana, No. 14БП036-01 at the Taras Oreocereus celsianus, Rebutia flavistyla, Rebutia minuscula, Astrophytum myrios- Shevchenko National University tigma, Mamillaria columbiana, and M. prolifera) have been studied. These plants of Kyiv. exhibit a uniseriate epidermis, covered by a thin cuticle. Except for E. peruviana Competing interests and A. myriostigma, no hypodermis could be detected. The shoots of all studied No competing interests have specimens consist mainly of cortex parenchyma with large thin-walled cells. The been declared. -
Handbook of Shows
Handbook of Shows 11th edition 2021 British Cactus and Succulent Society www.bcss.org.uk Contents Contents Page Preface....................................................................................................................2 1.0 Introduction ...................................................................................................4 2.0 Cactus Classes in the Schedule...................................................................4 2.3 Cactus Groups..............................................................................................5 2.4 Abbreviations used for Groups and Subgroups of Cacti ..............................9 2.5 List of Cactus genera with details of Group eligibility.................................10 3.0 Succulent classes in the Schedule.............................................................15 3.3 Succulent Groups .......................................................................................16 3.4 Abbreviations used for Groups and Subgroups of Succulents...................20 3.5 List of eligible Succulent genera, with details of Group eligibility...............21 4.0 List of Taxa of a Controversial Nature ........................................................28 5.0 Succulent plant families reference listing ...................................................30 6.0 Notes for Exhibitors ....................................................................................37 7.0 Notes for Judges ........................................................................................40 -
A Taxonomic Backbone for the Global Synthesis of Species Diversity in the Angiosperm Order Caryophyllales
Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2015 A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales Hernández-Ledesma, Patricia; Berendsohn, Walter G; Borsch, Thomas; Mering, Sabine Von; Akhani, Hossein; Arias, Salvador; Castañeda-Noa, Idelfonso; Eggli, Urs; Eriksson, Roger; Flores-Olvera, Hilda; Fuentes-Bazán, Susy; Kadereit, Gudrun; Klak, Cornelia; Korotkova, Nadja; Nyffeler, Reto; Ocampo, Gilberto; Ochoterena, Helga; Oxelman, Bengt; Rabeler, Richard K; Sanchez, Adriana; Schlumpberger, Boris O; Uotila, Pertti Abstract: The Caryophyllales constitute a major lineage of flowering plants with approximately 12500 species in 39 families. A taxonomic backbone at the genus level is provided that reflects the current state of knowledge and accepts 749 genera for the order. A detailed review of the literature of the past two decades shows that enormous progress has been made in understanding overall phylogenetic relationships in Caryophyllales. The process of re-circumscribing families in order to be monophyletic appears to be largely complete and has led to the recognition of eight new families (Anacampserotaceae, Kewaceae, Limeaceae, Lophiocarpaceae, Macarthuriaceae, Microteaceae, Montiaceae and Talinaceae), while the phylogenetic evaluation of generic concepts is still well underway. As a result of this, the number of genera has increased by more than ten percent in comparison to the last complete treatments in the Families and genera of vascular plants” series. A checklist with all currently accepted genus names in Caryophyllales, as well as nomenclatural references, type names and synonymy is presented. Notes indicate how extensively the respective genera have been studied in a phylogenetic context.