' ' D/MID OS II II AIE

Numb er 44 20 F eb ru ary 2006

Scientific Editor: BERTI L NORD ENSTAM Techn ical Editor: GUNNE WI R ENI US NOHLI N _ L h Publi hed and di tri buted b The S edi h Mu eum of Natural H i tor s s y w s s s y, D e artment of Phanero amic Botan p g y,

— 4 kh lm eden PO . Box 50007 SE 1 0 05 Stoc o S , , w

I SSN 0284- 8422

C O NTENTS

- ME EN m n n PELSER P. B . V KAM J F. R . VAN UD : b , , ELD P , DER New co i atio s in J a ba ea MI LL A r a Seneci oneae co . ( ste ace e )

N RDEN TAM A n n MI LL O S J a c ba ea . , B dditio s to the ge us o (Compositae- Seneci oneae)

Nordens ta mia I N m - Seneci oneae LuND rN R . : UN , L D (Co positae ) , a new genu s fiom the Andes of South America

N RDEN TAM B r O S . : mb s No dens ta mia m , New co ination in (Co positae

Seneci oneae r m Ar n na P r E r ) f o ge ti , Bolivia, e u and cuado

ha mn u n n w n NORDEN TAM B . n r N S : Ca a i ot s . o a e u , B , a ge s ofthe m a - Seneci oneae m n r I Co posit e , ende ic to the Ca a y slands

N RD EN B n w O STAM . KALLERSJO M P ELDENASI Ne hrotheca e , , , . . p , a monotypic genus ofthe Compositae - Calenduleae from the southwestern Cape Province

NORDENSTAM G n r r in ri b Calenduleae m , B e e ic evisions the t e (Co positae)

NORDENSTAM r mb n n Seneci oneae , B New gene a and co i atio s in the of the Greater Anti lles

T L i ’ e ficati on n m H i era c zum . H i era i um . : ctot TYLER, yp of a es of sect c Vul a ta b n m r m rn and sect . g ased o ate ial fro southe Sweden

New taxa and combinations pub lished i n this issue

w l 44 2006 C . om . Ne s p ,

M I LL New comb inations in J a coba ea . (Aster aceae Sen eci on eae)

l z Z 2 ’ - V MEMB EN EL ER J F. VELDKAMP R UUD AN PI . P S ETER B , , DER 1 ) Mi mi ni r it De artme t of ota 31 6 P ea rs on H all a U ve s y, p n B ny, ,

O xfor d , O hi o 45056, USA

hi ed u pelserpb@muo o. 2 ) Nati onaal H er ari u m d erla d Univers iteit Leide r an h b Ne n , n b c , 2 A i n Th h r la d P O . Box 95 1 4 300 R d e e , Le , Net e n s iv leidenun . nl veld kamp@nhn .

This a er is dedi a ted to BERTI L NORDENSTAM on the a s i n o his 70th bi r thda p p c occ o f y, F ebrua r 20 2006 y ,

Abstr act

New in sights in the evolutionary relationships between the species of tribe

Seneci oneae show that at least 27 species currently assign ed to Sene ci o sect . a r a r m ne i an d are b r a i n nu J coba ea should b e seg eg ted f o Se c o L . ette pl ced the ge s I n r are and 1 9 new mbin n i n J a ba ea MI L . co L this pape , these species listed co atio s

J a coba ea are published .

I ntr od uction

The resu lts ofrecent molecu lar systematic studies on tribe Senecioneae show that the D RT rm - r a J a ba ea MI UMO . species of Seneci o sect . co ( LL) fo a well suppo ted cl de P 2002 2003 2004 ubl . a n an r r ELSER et al . un ( , , , p ) th t is o ly dist tly elated to othe species

in a r L u ua r bu Sene i n u Sene i vul is . s lly att i ted to c o , i cl d g c o g , the type species of this nu KN P M 1 995 KADEREI T J FF 1 996 AI N G N 2000 PELSER ge s ( OX AL ER , E REY , B OLDE ,

r u . a a m r r a et al . These esults s ggest that sect J cob ea is o e closely el ted to P a ke ra an r r n c d seve al othe main ly New World genera in Seneci oneae . These fi dings are con firm ed in preliminary phylogenetic analyses of nuclear and plastid DNA sequence da ta that are currently carried out as part of ongoin g phylogenetic studies of the tri be focu sing on the intergeneric relation ships b etween its estimated 1 50 n ra r r r ge e and the delimitation ofSene cio (PELSER et al . We the efo e p opose to ran r 27 n i n n DNA t sfe the species of sect . J a coba ea that have bee de tified usi g u n a a r m n S r new mbin n seq e ce d t f o Se eci o to J a coba ea MI LL . eve al co atio s to mm a m i n J a ba ea a r n b n ma r WI 2000 acco od te the co h ve ece tly ee de elsewhe e ( EBE , PELSER VAN DER MEI JD EN The results of our phylogenetic analyses (PELSER et C m w 44 o . Ne s l . 2006 p ,

2 2 r r r r rr WI al . 00 b , howeve , show that th ee species that we e t ansfe ed y EBE

D n m r n L. d . a r ra en i . ar n i . e i u ki r his cus C . S e s s an S s c c s e 2000 S. ( ; g , o , L ) o ly distantly

r J a ba ea and r n are b r r n Sene i . elated to those of co , at p ese t ette etai ed in c o

I n a r umb r ub fi for this p pe , we list all species and a n e ofs speci c taxa which analyses of DNA sequen ce data in dicate that they should be considered members of J a coba ea O n b m m ar and make new comb inations where n ecessary. asis of orphological si il ities b k r urr n n to these species , we elieve that it is li ely that othe taxa c e tly assig ed to

r a in a ea m in n for a Sene ci o are also bette pl ced J coba . New co b atio s these t xa will b e made elsewhere (NORDENSTAM

When n early fin ished with this report we accidentally discovered a publication by GANDOGER ( 1 8 83) where 1 9 new species ofpresumab ly J a coba ea were described an d

m ri r I FNI . r b b are but at the ti e ofw ting we e not listed in The types p o a ly in LY, we have i n m d i r a r r m An not spected the an nvite a curious e de to sort this p oble out . isotype of ‘

ta vi cu N . K . ANK ERSMI T Ba um Bus s um r S. ba s G s s A D ( , ) was expected in L , whe e

’

KOK ANKERSMI T S rb ar m but u not b e u . he iu is deposited, co ld fo nd

D . A r Ed . 4 1 4 : a ba ea vul a r is a o a a MI . . 7 J b e G r b . 5 J c LL , a d ict ( ) Lectotype co g

G N . r . AERT , designated he e

M N H 1 J a cobaea abrotan oli a L . . if ( ) OE C

r L 4 n i J a ba ea a b ota ni oli a . M N H Methodus 1 79 587 . Se ec co f ( ) OE C , ( ) o

PI 1 H r r t i L . Z Aw. UI . a rota n i o i L . . b l us S . 753 869 . e bi chia a b o a ni ol a G f , p ( ) f ( ) in LL ,

— Ar . . 1 1 832 45 1 nom . u erfl . e e i h r i chia S U Nomencl . . . s S n 0 e b . ch Bot ( ) , p c TE D , Bot , ed 2 1 1 840 750 2 1 84 1 557 561 sub H er bichi a a r a ti a , ( ) ; ( ) , ( c p c

i ca r a thi u HERBI CH A . F1 . 1 3 1 43 Sene i Sene c s . G 8 c o p , ddit alic ( ) c o 1 a brota n i oli us s ub s . ca r a thi cus HERBI CH M N Cons . PI . Eur. 879 356 . f p p ( ) NY A , p ( ) R ene i a r t n i v r h I H EK R rt . . S c b o a oli us ar . ca a t i cus HERB C H e e . S o f p ( ) AY , p pec Nov egni 30 2 1 1 6 Ve . . 93 79 g Beih , ( )

ene i m e h r i n 1 1 1 . S n a lus S HU V b . S eb e b . Ver. Naturw. 850 07 c o o oc p C R, e ( )

E b . ene i r i l An l it . . S c ti ol ens is KE NE D in SONKLAR et a . e W o R R ex ALLA TORRE , iss co Al ‘ 1 882 1 33 i . enreis en 2 a Se 1 882 247 in H R I N A Al enfl . p ( nte p ) ; A T GER, tlas p ( ) ( Eur Al r D en fl . 1 890 325 . ene i a rota ni li us var . ti oli ens is op . p p ( ) S c o b fo ( ALLA TORRE)

- KE N HE I I Mi E 2 1 4 2 n e i a brota ni li us sub s . . t. ur. 6 9 1 69 . Se c o R ER ex G , llust Fl . , ( ) o f p

ti r li D l A 1 1 o V r n S u A en fl . r 0 938 ens is G M Jahrb . ( ALLA TORRE) A S , e ei s ch tze p p lpentie e ( ) 22 .

2 m . n . J acobaea adon i di oli a L I S . P ELSER VELDK . c ov. f ( O EL ) , o b

Sene i a donidi li us Lol SEL a 2 A r 1 6 n m n v. ro Sene i o . G . 807 56 o . o c o f . , Fl ll ( p ) , p c o

a brota n i oli n 1 435 id . us . no . Sene i a r temis i i oli us PERs . S n . 2 Se 807 f auct , L c o f , y ( p ) , , C w l 44 2006 m N e s . o . p ,

s u erfl . nom . p

M N H J a oba ea al i n a L . 3 . c p ( ) OE C

lida l L N H M d 1 7 4 . i M etho us 9 587 S a na . S . P1 . J a ba ea a l i na L . co p ( ) OE C , ( ) o g o p , p n n w r l Fl An 1 . Ed . 2 1 778 870 ot e . i nera ia a ina . L . . H s . L 1 753 880 UD . S ( ) ; , g ( ) ( ) C p ( ) , p

ALL. . P m . 1 1 5 20 . . 2 n n w 1 24 var . 7 Pl . . 2 2 1 763 3 . 8 3 38 ot e , ed , ( ) , excl ; , Fl ede ( ) , t , f ( )

2 n — 4 . L Pl . Ed . 4 3 3 1 803 08 m Sene i a i nus . S . . r . . l WI . S LLD , p , , ( ) , iso y c o p ( ) COP , Fl Ca n

1 82 37 1 n . i nera r i a rdi lia U N I ll 2 2 2 1 4 f. u . m 1 77 6 . S 7 G . ed . , ( ) ; L ppl ( ) , iso y C co fo O A , 1 2 1 4 4 1 n n w 1 2 A . 7 1 773 69 J . F . ustr. 77 7 6 ot e . f. Su . 78 Ob s erv. . Bot ( ) ; ACQ , ( ) , t ( ); L , ppl ( )

— v fl . ene i 0 rd li us U N LAI R . Man . H r . 1 1 375 not new nom . su er S i G C b 8 , ( ); p c co fo ( O A ) , e ( l )

H 2 4 n n — 24 1 nom . s u erfl . I NK andb . 1 8 9 7 8 m no . f. Sene i 0 , p L , ( ) , iso y ; L c

— fl . a ba ea r da t I z n am r da tus K H ra 1 7 1 834 61 4 nom . s u er J a O Se co OC , Flo ( ) , p co co P Z , 4 1 852 5 nom. su erfl . ( ) , p

En 1 Me . 7 806 1 09 . Sene i ba ldens is P I R . . c o O , cycl th ( )

n n m a l i na r b b n m n ra- s ecific nam Note This sy o y y of J . p is p o a ly i co plete; the i f p es m m n m n in n b n . S b u u ot b u have ee o itted o e a es have ee cl ded, altho gh new, eca se they

t n as r r r n un r i n I n K ew n i are o e s s . of e cited ifthey we e , we e e co te ed the dex

mbi ua rv P E ER VELDK m . nov 4 a o a ea a B . . . J c b LS . g ( ) , co b

R 1 i B D i nera r i a a m i a BI V . ir ar a m v u S 3 8 1 5 5 Sene i a b us l . C . b S . . u C g , t p icili ( ) c o g ( ) , Pr 1 r 6 838 356 . . i nera i a bi l r WI . ner a r i a bi r odr. i l ( ) , excl syn C co o LLD C co o m i Brv M N n E ub s . 1 s a b ua . Co s . . ur. 879 350 . Sene i i bbos us s ubs . p g ( ) NY A , p Fl ( ) c o g p B A NO . m . . I a . 1 8 2 4 . ambi uus rv. 8 3 9 g ( ) RCA , Co p Fl t l ( )

l H R 1 2 1 i nera r i a i bb s a u . P G . . r . e . 8 7 1 . Sene i C g o ss , Cat o t g Boccadifalco ( ) c o

i D — bbos us GU . C . Prodr. 6 1 8 8 5 ene i0 a m i us s u i us . 3 5 . u U 3 S b bs . bbos G g ( SS ) , ( ) c g p g ( SS ) 1 4 2 4 H . J . nn . Soc . 68 97 7 . C ATER, Bot Li ( )

ene i t I 4 a HE . Dia . Pl . Or . I 6 1 8 6 95 . inera r ia S c t e eus . n o yg BO SS LDR , g ient , ( , C

n br dens is sub s . t et a I HE . M N n E 1 e a e Co s . . ur . 8 79 350 . o p yg (BO SS . LDR ) NY A , p Fl ( ) ene i a m i w H E . GREUTE VVi lldeno ia 33 2003 S b uus sub s . ta eteus oi . R c o g p yg (B ss LDR ) , ( )

5. J a obaea an alo a D VELDK m n v C . . o . o . c g ( ) , c b

Sene i a na lo s D . Pr 1 r n N RDEN TAM u C odr. 6 838 366. S b O S ] . c o g , ( ) elected ove the ext y [F 1 64 1 I ran . ( 989)

Sene i e ili r D Prodr. 1 3 366 . ne i ch s nthem id s s ta b s W . 6 8 e C . 8 S a o e c o p c ALL ex , ( ) c o y

var . ta bilis W . D H K r I n a 1 1 s e C . . f. . . 3 88 339 . p c ( ALL ex ) OO , Fl B it di ( )

Sene i ch th r n moides D . Prodr. 6 1 3 65 n n H NK r 2 1 s a e C 8 8 3 o S . . 789 c o y , ( ) , C RA , Baie Fl ( )

386. w N l . 44 Com . e s 2006 p ,

Pr dr. 6 1 8 366 . ene i ch s a n them v r mbr i rmis D . 8 S r i Sene i s is i o C o 3 o des a . c o y f , ( ) c o y

— t . I n 1 1 9 . n W v r H K f. r i . 3 88 33 Se e 0 la tas ED . ar. s is mbr ii o mis DC . . y f ( ) OO , Fl B i dia ( ) c c GE W W L - 4 haul 1 9 4 n m . r i rmi AS AL in AS AL ME HR . a 8 s is mb i o s DC . S 9 3 o y f ( ) OTRA , Fl piti ( ) , i ri n a . r r o t . i v l , cont a p

D Pr dr. 38 36 a a n n m r R N . 6 i ner a i a de i i ens S C . o 8 6 s b C c p P E G ex , ( l ) , cited sy o y y the m i ta bi li n k wens i in n ub n m S. s e I e s s . dex , is an valid a e , p l shed as a sy ony of p c

4 20 1 8 6 74 . Sene i la etus EDGEw. ran . nn . Soc . . c o , T s Li Bot ( )

i a H I P N. . M EY. H . 6. J a coba ea a u a t . G S q c ( LL) AERT , B C ERB

N MEY R k n W r G . . . S H . O e o . . u 3 1 802 J a ba ea a ua ti a HI P. co q c ( LL) AERT , B C E B , Fl ette a ( )

1 u t u HI Ve . 2 1 1 1 20 H UD s . . An 1 2 1 i a a . 2 0 . Sene i s . S 76 76 3 7 c o q c LL , g yst ( ) ; , Fl gl ( ) , I I 4 n m . Seneci o a coba ea var. a ua ti us HI I BEG . . . 3 1 903 2 1 iso y j q c ( LL) F OR , Fl tal ( ) , m rfl s u e . no . p

R i 1 Seneci o ra ea ltus BE TOL. O u . Sc . 3 1 8 9 1 83 p , p se ( )

MEI JD EN 6b . a ob a ea a ua ti a var er ra ti a BERTO L. P ELSER J c q c . c ( )

ti v r r r t BERT L PEL ER MEI JDEN in HEUKEL . Ne rl . a i . s J a ba ea ua a a . e a a O S de co q c c ( ) , Fl 23 2005 6 ed . ( ) 77 .

i err a ti us BERTOL . Rar. I a . Pl D ec . 1 1 0 62 . en e i a ua ti us var. Sene . 3 8 S c o c , t l ( ) c o q c

rr BER M An 42 — n e a ti us TOL. o . G . W . I n r . . . P r 1 8 77 . Se e i 0 c ( ) C ss in ER EDD , t od Fl al a is ( ) c

i BERT L NEI L R n 1 4 2 1 P I BE . I a . a v err t M 5 . a b ea ar. a us O . . . e 8 6 O 3 j co c ( ) , Fl ( ) ; AOL . Fl t l

b var. rra i I n . . I a . 1 03 4 1 2 n . ene i 1 903 2 1 4 n . s u e t us 9 m S ( ) (i cl c ) ; co Fl t l ( ) , iso y c o u rr RT V - - r 1 2 a a ti us sub s . e a ti us BE OL. TOUR LET . Pl . a . I r et 908 73 q c p c ( ) , Cat sc nd e Loi e ( ) ;

— ti us VA M H W rk 1 1 n m a ea b . erra . u 5 950 50 . Seneci 0 a cob s u s ATT E S , Fl . T ey ( ) , iso y j p c BE 2 4 SU . u u . 1 0 1 1 5 R UY ons . . r 1 9 7 1 3 RTOL. 9 7 C ( ) DRE , Fl To lo s ( ) O , p Fl F ance ( ) ,

n m iso y .

‘

n r 2 2 1 42 1 nom . n . Sene i Se e i ba ba r ea e li is K K . . 7 o . S 90 c o f ROC , Fl iles , ( ) , i val c o

ti r r 1 2 1 1 nom . n a . a ua s v G . . S . 3 8 9 5 u an onba ba a ei oli us WI MM . q c [ ] f RAB , Fl iles ( ) , i v l I ba r ba ra ei l us K . I I in I I P . PI . . 3 1 903 Senecio a coba ea var. o i K j f [ ROC ] F OR F OR AOL , tal ( )

4 ne i er ra ti u s ub s . 2 1 4 I n . I 1 903 1 2 Se s ; co Fl . tal . ( ) c o c p - 2 2 ba r ba ra e i li us WI MM R in H I 111 . . M t Eur. 6 1 9 8 778 o G . f [ AB ] BEGER EG , Fl it , ( )

ba r ba r a e i oli us WI MM G . Seneci o a qua ti cus s ub s p . f [ RAB ]

A P . nn Tids s kr. 57 1 1 1 W J . . . N . 96 73 EDERSE , Bot ( ) ALTERS , Bot Li ‘

Soc . 7 1 1 76 2 r r n m 9 73 ba ba ei oli us . ( , f iso y

Sene i r a ten i K RI H z 1 3 692 sub Sene i s s . . 8 0 c o p LETT C T , Fl Leip ig ( ) cited c o ba r ba r ea i oli u K K R Ex r 1 2 244 s . HB . . G rm . cu s . 83 f ROC ex C , Fl e ( ) nom erfl . s up . 44 006 C w l . om . Ne s 2 p ,

m 7 J acobaea arn autorum VE EN. PELSER c b . nov. . ( L ) , o

t i z un s b r. K ni l s r n ut r VE EN. S e O hm. e . W . M . Seneci a a o um . G o L , g g Bo iss , ath 1 8 8 1 88 9 45 Ciner ar i a ar na utor u V rwis . m Natu s . 8 EN. YM N Cons . 1. Cl ( ) ( EL ) N A , p F

— u r 2 1 8 9 1 63 . en eci al in s bs . a na ut r u E . Eur S . 8 S us o m V EN H YEK Re , uppl ( ) o p p ( L ) A , p

R . 0 2 1 9 1 68 . v V . ert . S . No . e 3 3 0 pec egni g Beih , ( ) ' — P N m . 1 4 r 883 1 . T AI N Senecioe ubescens A cI c E . . W T Seneci o , le Fl Bulg ( ) , non . O i M r 0 9 k 1 1 9 2 2 . anci ci DE EN . p G , agya Bot Lapo ( )

l R a r mb . . 8 . J acob ea au i cu a U . PELSE c (BO RG ) , o nov

in . Pl r . r i la . 1 52 1 9 . Seneci a u cu U o . 8 6 o BO RG C ss , Notes C it ( )

r mb . 9. J acobaea bois s i e i DC. PELSER c . ( ) , o nov

b is s r i DC. Prodr 1 838 00 . en c e . S e io o i , 7 ( ) 3

acobaea cann abi oli a LE . E . Wl EBE 1 0. J f ( SS )

Tu n w na i a . . W E i J ac baea can bi l E E I EB rcz a ino ia 3 2000 62 . Senec o o fo (L SS ) , ( ) i 4 6 1 83 2 2 . cann ab li us E . fo L SS , Linnaea ( l

1 J a ob ae arn i oli I CH N c a c ca W . S K 1 . ( LLD ) RA

S H NK D nks chr. 1 1 1 4 1 J a b ea car n i li a WI . C a e A . M . 1 8 3 8 3 6 co o c ( LLD ) RA , cad unch ( )

— i . Pl . . i n Seneci 0 cam iol cus WI S . 4 3 1 803 1 993 Senec o i anus 3 . LLD p , ed , , ( ) c

i li cus WI . HE H E Fl . E 1 1 1 2 1 2 1 mb . n . i var. carn T T 89 . o ( LLD ) A CO , ngadine ( ) , t , co i val (s ne

- b n e i i n an us u . ar n i li I D UN BLAN . . Se c s bs c us W . asion ) c o p o c ( LL ) BRA Q , Neue

D hr S . N tu . 4 1 0 . a rf. Ges 8 1 9 3 30 enksc . chweiz ( )

m i c . n b r E V . 1 1 b . J acobaea ca oli a s ub s i s u i ca CH NE ARD PELSER c mb nov. p ( ) , o

li s r V . 8 r n r i ns ub i cus CHENE ARD H rb . . 6 1 906 6 . Seneci a i cu va . 3 o c o , Bull e Boiss l l , ( )

- i n u r i s CHENEVARD AN . Denks chr . Seneci o arn i li us s ubs . s b cu UN BL c o c p ( ) BRA Q , Neue

Naturf. es . 48 9 1 S . 0 chweiz G ( 1 3) 30 .

' m 1 2. J a cob aea del hin i oli a V H P ELSER VELDK. c b . nov. p gf ( A L) , o

2 1 9 1 9 1 . 4 Sene i del hin i i oli us S m . V H b . 7 5 c o p f A L, y Bot ( ) , t

Se i R 1 4 n O . ci . 8 1 7 23 eci cr i n tus BE TOL. S o , puse ( l )

1 1 8 2 2 J a ba ea nth r li P P E D . Pr . 3 9 a o i o a J. E . co f R SL C R SL, elic ag ( )

b Y 1 M E . S HE . 3 . J aco aea eru c oli a L. P. G TN. . if ( ) AER , B C RB

J a ea k . W r 1 8 1 c ba eru i oli a L. T MEY CHE B 0 c P. G E N. . . S . Oe on. 3 o f ( ) A R , B R , Fl ette au , l ( ) 20 . M Y. r M r. 1 . 8 M I EB . . C . m A E Ve z . Pfl . G . 83 1 8 1 ; B ex , asp ee ( ) isony

Seneci er u i li u 8 — a ea S . PI . 1 75 69 A 2 . J a b c s L. 3 o fo , p ( ) ddenda ( ) (epithet) co r ubella N H 4 fl . M Methodus 1 79 5 86 nom. s u er OE C , ( ) , p m w N e l . 44 2 0 6 Co . s 0 6 p ,

i ten ui li u . F1 . Aus tri ac . 3 1 775 42 . 2 n Senec s J 78 . Se eci er u i li u s . o fo ACQ , ( ) , t o cfo L i li u CHUBL N m n u MA . bs . te s J S . G Wurte b er 1 834 4 s u 5 1 . p fo ( ACQ) RTE S , Fl g ( ) v r n i T E i eru i li us . a . te u li us CARI O Fl . . 2 1 2 44 Sene 5 87 3 . c o cfo L fo , tude , ed , ( )

Li 1 2 2 J a ba ea s ua ve lens GI I . . tuan . 1 78 01 nom . . co o L B , Fl ( ) inval

R P . P . . 1 8 1 9 1 no r f. J a ba ea tenui li a J . 75 e J UI N " co fo ESL C RESL , Fl Cech ( ) ( to ACQ )

I cono . . Pl . r . 4 1 82 2 Sene i del hi n i i oli us R H . r 6 9 c o p f C B , g Bot C it ( )

m . n n n V H no . o i val , A L

v r b ar. s ub la b a CAR I OT E l . . 2 1 2 44 . Sene i er u i li us . s u 5 87 3 c o cfo L g , tude F , ed , ( )

i a rna s s i N . . n . 1 1 2 Senec ens s G 875 3 . o A D , Fl Lyo ( )

i ha e i N . ARI OT E Pl . 2 1 2 44 N Sene c b rt i G C . 5 87 3 G n 1 875 c o A D ex , tude , ed , ( ) ; A D Fl Lyo ( ) ‘ r 1 23 ( cha be ti Q.

li u . v a utumn N ARI T E F 2 n i i . a li l Se ec eruc s ar s G . C O . . 5 1 872 344 . o fo L A D ex , tude , ed , ( )

ARI OT N . . . 1 5 1 2 . Sene i a utumna lis G N . C G 87 3 c o ( A D ex ) A D , Fl Lyon ( )

m n 1 3h. J a oba ea cr u i oli a s ub s . ar un ens is TURCZ . VELDK . co b . ov. c cf p g ( ) ,

2 2 4 r n Soc . . M o. 0 1 8 7 1 8 ene i Sene i a u ens is U . . S c o g T RCZ , Bull Nat os , ( ) c o

i E I PI . r 1 1 eru i li u . b s . a r une 1 s s u ns s U . W S b . 3 998 66. cfo L p g (T RCZ) EBE, i i ( ,

i o a c a ea . var a n i U in D Pr dr 1 . ene b . r d . o S c o orus . C . 6 838 350 j L g fl T RCZ , ( )

i ra a t var ur D Pr 1 1 ene l u . a h . S c o e s d i cus C odr. 6 838 35 . p , ( )

v n LE . Re ert . 8 1 1 0 1 38 . Se e i blimi H . 9 c o , Fedde, p ( )

4 1 J a coba ea alleran di an a o . D UR I EU PELSER m . nov . g (C ss ) , co b .

i a ller ndi a D r 2 1 Sene a nus . URI EU . Soc . . 855 365 . c o g COSS , Bull Bot F ance ( ) m 1 5 . J a oba ea i an tea D F . P ELSER c . nov. c g g ( ES ) , o b

i Sene i a nteus D F . PI . A . 2 1 800 273 . 234 . c o g g ES , tlant ( ) , t

1 a a n h m n 6. J cob ea a ali oi des SI ex S N . VELDK . c . ov. g p ( EBER PRE G ) , o b

4 z n 1 0 . 1 n a a ha li oides SI E R Kr 2 1 823 322 . Co y g p BER, eise eta ( ) , t , f ,

KUN H Sene i na ha li oides SI E R Kr 1 823 352 nom . non T c o g p BER ( eise eta l ( ) , ‘ ’ in 4 z ha l des S N . S . Ve . 4 1 26 55 . n a s i eberi S U . . na 8 val , g p o ) ex PRE G , yst g ( ) Co y TE D , m 2 1 1 40 4 1 4 r r a ha ide M N on . F1 . Eur. No encl . . li . 8 . i ne a i na o s C S Bot , ed , ( ) C g p NY A , p

‘ 1 1 I l d 879 35 : S s . n . Sene i na ha es ( ) Lectotype EBER (L) , c o g p o ’ I S . S r . EBER tia , designated he e

ene i a r i a B A n I 2 S c s cus E UVERD u . Soc . . G e I 1 1 92 1 1 5 . o , B ll Bot e ve , ( ) C w l 44 2006 N e s . om . p ,

m . nov. 1 7 . J a obaea i n an a L. VELDK . o c c ( ) , c b

PI . 1 753 869 . Sene i i nca nus L. S . c o , p ( )

— k 5 1 8 354 a n m n nu um M R . r d . 08 J a ba ea in a na RAF . e Note co c ( epos New Yo ( ) ) is o e d , 2 2 2 in an r Lituan . 1 1 78 0 ub while J a coba ea i nca na GI LI B . (Fl . ( ) ) was p lished opp essed The mb n n b on n b rk I E A n . V . m wo ( C N, ppe d ) co i atio ased the Li nean asiony is therefore still available for valid publication .

h lla D PELSER om . nov. 1 8 . J a obaea leuco C . c p y ( ) , c b

D a H r . M n . 1 1 1 44 . . 8 3 Sene i leuco h llus C C t . o s c o p y , o t p ( )

1 9 . J a cobaea ma ri ti ma (L .) P ELSER M EI J DEN

L PEL ER MEI EN in H E L F1 N rl 2 2 J a ba ea ma r itima . S JD UKE S . ede . . 3 005 677 . co ( ) , , , ed ( )

r P1 . 1 753 25 . i ner a r i a ma r i tima L . L . S . Pl . . 2 Oth nna ma i tima L . S . 9 o , p ( ) C ( ) , p , ed

— fl 1 1 244 ea m nt a M N H M th d 1 4 n m . er 763 . J a ba t e s e o us 79 5 87 o su . ( ) co o o OE C , ( ) , p 4 r R H . in MOSSLE H andb . . 2 2 1 829 1 79 PI . rm . Excurs . Sene i ma i tima L. R G c o ( ) C B , ed , ( ) ; e 1 2 244 n n L n i n r r i D Pr dr en 83 o . f. Se e Ci e a a C . o . 6 838 355 . S eci ( ) , c o , ( l ) o

r i nera r i a DC . H . . n . oc . 68 1 974 2 3 . bi l sub s . C J S 7 co o p ( ) C ATER, Bot Li n ( )

r r S . PI . 1 753 25 . i nera r ia a na dens is L. . Pl . . 2 1 763 Oth nna Cine a ia L . 9 S o , p ( ) C c , p , ed ( )

1 244 — N n n L PI r r i n . n D . o Sene a a de s is . S . 1 753 8691 Sene i Ci e a i a C . c o c , p ( ) , nec c o

J a ba ea Ci nera r i a I F I . co not in N , see note

n r r r S . Pl . . 4 3 3 1 803 2085 . ene i bi c0 10 r WI LLD . i e a i a bi l WI . S C co o LLD , p , ed , , ( ) c o ( )

I nd . em . H r . Re . P orrn . 1 22 nn 1 n I v TOD . V S an 859 30 30 861 759 on . , o t g ( ) , ; Li aea ( ) ,

n n r r bi l r WI . ARCANG . m . . I a . . 2 1 8 4 672 . Se e i Ci e a i a subs . 9 c o p co o ( LLD ) , Co p Fl t l , ed ( )

i n r r r Pl . H r . R e . a 1 2 1 2 . en i e a i a neb dens is GU . . a 8 7 S ec C o SS , Cat o t g Bocc dif lco ( ) o d H nn 4 bi l r subs . ne r e b ns is GU . . J . . Soc . 68 1 97 273 . Sene i co o p o ( SS ) C ATER, Bot Li ( ) c o i r n r r b dens is U . M . Wa n 1 4 1 3 279 . en Ci e a a s ub s . ne G 98 S eci p o ( SS ) ABB , tso ia ( ) Non o r L RAF neb dens i . n r o s L ec J a coba ea neb odens is ( . ) .

r r P P D Pr 1 22 i ne a i a a ndida . in PRE J . . . . 8 95 . Sene i C c C RESL SL C RESL , elic ag ( ) c o a ndidus PRE D C . Prodr . 6 1 3 355 . . 8 8 c (C SL) , ( )

Sene i i er r 4 n la tti i MEI K K ew u . 1 954 1 5 1 42 . ene i C a i a DC . var . 9 S c o p LE , B ll ( ) c o i l r la tti i MEI K M H R ar E n b var. W o . . G . b r 33 1 975 co o p ( LE) ATT E S , Notes y Bot d di u gh ( ) 4 34.

Oth nna ma r itim n r r a and 0 th0 n a Ci ne a i a . r ub Note o L . L we e p lished m u . We not r m r b n two si ultaneo sly have discove ed who ade the fi st choice etwee the , but r one n ra n un r for Ci nera r i a the fi st epithet is the ge e lly e co te ed, while the epithet in the present context one is always referred to DE CANDOLLE ( 1 83 a nomen novum for w o Ne s l . 44 2 6 C m . 00 p ,

th nna Ci ner ar i a in Seneci b f f O r S. mar i tima o necessitated o ecause o the olde L . . ( 1 78 1)

2 a cobaea maroccan a P. H . D I PELSER m 0. J . . ( AV S) , co b nov

r I H . D R . . r r n i ma canu P. Se e oc s N o G . E b 38 1 980 44 5 c o AV S , otes y Bot a d din u gh ( ) .

ba in uta AV P EL R D m 2 1 J aco ea m C . SE VEL K. c b . nov. . ( ) , o

mi t V I 1 - r r 1 7 1 2 1 . 3 . 3 . S n ci inut AV i ne a i a nu a CA . . 9 f C 3 e e m us C . DC. Pr . , con ( ) , t , o ( ) , od 4 6( 1 838) 3 6.

h r P D cr D . i ner r i a hete lla O TE N V. l 1 798 1 . 1 0 f. 2 . C a O . es . ec 7 op y R GA, ( ) , t ,

h A. ME 22 . J acob aea ot onn ae M . I EB . S EN . ex C . Y ( B ) PR G .

th nna e M N . A rz . P ea o . . J ac ba o S E C . MEY. Ve fl. G . M r 1 3 1 8 1 . . 8 o ( PR G ex , asp ee ( ) n nn I - 2 1 3 r n ci othonna e e eci otho a e M. . . r. . 8 I e o S 80 08 . a o B EB , Fl Tau Caue ( )

BlEB. . . I r . 1 64 9 5 M. N 1 98 8 . ( ) B ORD , Fl . an ( )

n r an la F1 r dr Ci e ar i a ma SM. . G aec . Pro . 2 1 8 1 3 1 79 F1. Graec. 9 1 839 56 . 872 . o , ( ) ; ( ) , t

— li a in t W . E m r r . 1 0 8 0 S i nn a s WI . Caca na a I . H 8 9 5 . enec i tu p LLD , nu o t Be ol ( ) o p ( LLD )

B 28 845 499 . SCH . 1F . r , Flo a ( l )

r ar l r us C i 1 4 ne var rar i or a Seneci i o K . K H 7 1 8 3 47 . Se ci othonnae . o f O , L nnaea ( ) o fl K K C K K CH 2 1 H . 5 ( . O ) O , Linnaea (

MEY. H E . 2 obae alud s a L. P. E TN. B. C B 3 . J ac a o G S p ( ) A R , R 3 1 802 ME C ek n . r ba ea aludos a L. P. G E N . Y. S HE B . O o . W J aco . p ( ) A RT , B R , Fl ette au ( )

a Arr. rr i Y N t. 1 1 Seneci al ud s us L. . PI . 1 75 870 inerar i a s e at lia G 2 o o S 3 . p , p ( ) C fo RA ,

oc . in . P1 2 1 46 nom. s u er D or i al dos a L. O RR . Ann. S r . . 82 9 fl . a u U B it ( l , p p ( ) F , L n Lyon 4 16 1 868 40 . n. s . ( )

— 1 8 L n a ur D Prodr. 6 83 44 . r 2 1 63 12 . Se ec atus C Ci ne ar i a a urea . S . . 7 i , p ed ( ) o , ( )

S. ur s 348 a eu . , non L

WI E E n c ta tar us E S 1 834 1 92 J a c baea ta tar i ca E S E . Se e i i c 9 . o L S , Linnaea ( ) o (L S ) B w Turcz anino ia 3 (2000) 62 .

m . . Li n . 1 82 202 no d rr t li I I B . S li a s e a i a G . tua 7 o go fo L , Fl l ( ) , inval

d . . 1 22 4 2 . Seneci r i ar i us WALLR . Sche r 8 8 o p , C it ( )

— J ac baea k s tel ecka OF1z C 1 2 25 . h mi us H S Pl . . 2 8 8 3 Seneci o bo e c U . TA S , yll Nov ( ) o o , bohemi cus U H i aludos us subs . 4 m. Sen e C Sez nam 1 852 5 no su erfl . ( ) , p c o p p (TA S )

m 1 8 1 242 . E Prodr PI . o 7 C K . . LA , B h en ( )

l D 2 40 40 Comi . P t . 1 8 3 Seneci s ad er i S E . t o A L R, Fl es , ed ( ) 44 00 C N w l . 2 6 om . e s p . 9

t nt s E i o a lud us subs . ome o us C K . Prodr. . m 1 87 242 Senec os 1 . p p LA , Fl Boh en ( )

P R mb . nov. c a a r o ii DE N T. ELSE o 24. J a ob e p e s o n ( O ) , c

r L ust . 44 i r . i Re e t . 1 8 22 n i n Senec e s oomi DE NOT. 9 Se e i nca us o p , p Fl g ( ) c o N N M ons . . Eur. 1 9 . r oo i i DE N . 7 subs . e s n OT C 8 356 p p ( ) Y A , p Fl ( )

i v r. t i u PE S n . 2 1 4 a 8 07 5 S n e i o n an u s ubs . Sen ec o i ncanus a i l c s S Pl . 3 e i s R , y ( ) c c p

m . . I . 1 2 i R . A C N . ital cus PE S PE s O 88 349 . ( R ) ex R A , Co p Fl tal ( )

2 aco aea s ub al in a W. D . J . K CH PELSER VELDK . o nov 5 . J b mb . . p ( O ) , c

i l H 7 4 J . K r 1 1 3 61 4 n m. r r r S nec uba inus W D . C i . ine a i a a na e s 8 o . o l o p O , Flo a ( ) , nov p C p

n . on L. f. . W H EN F1. Ca at . Pr o. 1 1 2 r . B r 8 5 6 . i ner a i l i n . 9 a a a van 13 auct , e g A L , p in ( ) C p [ ] a t L Pl 2 1 76 24 en a l i n u . C 3 1 3 . S eczo s L. s la a S . . . l i n S . ubs . s uba us , p , ed ( ) p ( ) OP p p

J H E R . N v R . K e D . H K ert S . o . Ve . . 1 W. C Y 30 2 93 1 680 . ( O ) A in p pec egni g Beih , ( )

r r r i ur l t . i u a a J . Au iac . 2 4 n i ne a i a c di l a var. a C tr 7 4 1 77 . e eci s 1 7 7 . S C o fo c A Q , Fl ( ) , t o r tu n S . s ubs . u i cula s J . R H . I co . Pl 1 4 2 a i nus L. C C B 2 l a . 82 3 o r . r . p ( ) OP p ( A Q ) C , g Bot C it ( ) , — rd u ur l at 1 35 . Sen eci atus s bs . a i cu us J . J . A N . m . . I . . C C C t o co p ( A Q ) A Q ex R A O , Co p Fl tal 2 4 ( 1 88 ) 3 8 .

ALL. OK . m . n v. 26. J acobaea uni ora V b o fl ( ) EL , co

i L I P — S lida un or a A L. Mé an es . M . S c . Ro r 2 77 1 8 . eci o . 1 3 Sen o go fl , g hilos ath y Tu in ( ) o

— r P m. 1 1 85 00 . . ne un o us ALL ALL. . T i 7 2 1 7 3 RE Z . Se i fi ( . ) , Fl ede ( ) , t , f , non c o h ller i D Y 1 1 97 625 a N 9 0 . A D , Taxon ( )

M P 4 r I . 11 6 1 0 46 G N . CARI OT Seneci chabe tii PET1TM. 9 o , onde , ( ) non A D ex ,

2 1 872 44 . E FL . 5 3 tude , ed , ( )

27 a v ri E J acob ea u l a s G N. . g A RT

— PI . 1 79 44 . 1 0 1 en i o a cobaea a u TN F t . l r . 5 c J a b ea a is G E ruc . Sem 2 1 7 f. S e co v g A R , ( , t . j

P 1 870 . S . 3 L. I . 75 , p ( )

2-3 1 7 2 64 1 — Seneci o ac0 baea S n c o c var nudus E T . i b a W N . 7 e e a o ae . j S O , Bot Univ ( ) j

n T dd ti 1 4 1 98 77 . dus E N JAK as . . Nar. Muz . O . P . 8 0 subs . u W SO C p ( S O ) , Cas , ( )

B i 2 4 n m. L th an . 3 1 78 1 9 o . J ac baea s i cca GI I . . u o L , Fl ( ) , inval

dr 8 8 5 1 Seneci o olios us DC. Pro . 6 3 3 f , ( l )

4 a ea var. r r D 8 48 0 S. c ba Ja d . 8 8 3 Seneci o os culos us J . . G fl ORD , Cat aines ijon l ( l ) , non j o A DC . flos culos us L M.

8 I Hi an. 2 1 65 1 1 9 . Prodr. P . s in WlLLK . N E Seneci o a cob a eoi des WI LLK . j , LA G , p ( ) m w l 44 Ne s . 2006 1 C o . O p ,

2 J a oba ea vu l aris s u b s . d un ens is DUMO RT . P ELSER M EI JD EN 7b . c g p ( )

dunens i DUMORT . PELSER MEI EN i n a b vu l a r is s u . JD J a ea b s s HEUKELS F1 . co g p ( ) , , 2 2 Nederl . ed . 3 005 677 . , ( )

2 — 1 8 7 66. Seneci 0 a coba ea . . un Sene i dunens is DUMO RT . . . s ub s d ens is c o , Fl Belg ( ) j L p D a ni a 1 1 2 RT KADEREI T P. W DUMO . . S 6 986 3 . ( ) ELL , tso ( )

d n n N : MEI ri v . n u W JD Seneci o a coba ea ar us . o EN Gorte a 8 1 976 57 . . j auct ESTO , ( ) , t l

AM v los us L . DC . LAM . F1 . r . . 3 4 1 5 1 63 . Seneci o a coba ea ar. os cu 80 j fl in , F anc ed , ( )

Ann . . nn n n . . 1 4 4 a co a ea c ulos a . 1 J b os OURR Soc . s 6 868 0 Sene i fl F , Li Lyo , ( ) , non c o

flos cul os us JORD .

Acknowled gements

We wou ld like to thank BERTI L NORDENSTAM an d WERNER GRE UTER for valuable

suggestions on earlier drafts of thi s paper. 44 2006 N ews l . Com . p , 1 1

R efer en ces

n r n i n A r L N 2000 . A ofP a cke a Se ec o eae I N J . F. J . . G BA , OLDE phyloge y ( ; ste aceae) based on internal tran scrib ed spacer region sequen ce data and a b road samp

r M le ula r P h lo ene ti cs a nd E l uti n 1 6: 33 1 338 . li ng of outg oups . o c y g vo o

D i dam neci ni r if li i D . ac aea ANDOGE M . 1 883 . e u bus Se o s eruc o . C Jacob e G R, q e g ege L ( )

n i D euts he B ta n is he Mona ts s chr i n s eci ebus adhuc ur i ot s . L . ovis p igit g c o c fi

— — 4 0, 56 59.

A relimi n an f NA ar a n in RE rT W J FF 1 996. ar o K ADE , J . . C . E REY p y alysis c v i tio the m a I HI N D E ri b Seneci oneae . n : . J . . H . J . B ENTJE t e (Co posit e) D , N

‘

tema ti s . P r eedi n s o the I nterna ti na l m i ta Comp os i ta e . Sys c oc g f o Co p os e

1 4 — R a n K w r w 1 v l . G r e UK . C n e en e Ke 994 o 3 9 360 . o f c , , , , oy l Botanic a de s , ,

r i fD r n K N E . . J . D . P ME 1 995 . o end os eneci o i Senecio eae OX, B AL R The o ig n with n the Amer i a urna B ta 2 (Asteraceae) b ased on plastid DNA evidence . c n J o l of o ny 8 :

— 1 567 1 573 .

2 A n h n m NO RDENSTAM . 006 . t e u J a ba ea MI . a , B dditio s to ge s co LL (Co posit e

— m . N le ter 44 : 1 2 1 . Seneci oneae) . Co p e ws t 3

V MEI JD EN 2 2 . a k n n r PELSER P B . G V N . R . AN 00 a: , . , RA E DEEL , B DER T c li g speciose ge e J a ba ea Species composition and phylogenetic position of Seneci o sect . co A ra a a n a an nrDNA u n Amer i a n J urna l o ( ste ce e) b sed o pl stid d seq e ces . c o f

B — ota ny 89: 929 939 .

PE ER B V LS P. . G N . R . AN ME I JD EN 2003 . P r n r , , RAVE DEEL, B DER hylogeny eco st uction in the gap between too little and too much divergence : the closest relatives of

Senecio ja coba ea (Asteraceae) according to DNA sequences and AFLPs . M le ula r P h l n E 2 1 — 2 o c y og e eti cs a nd volution 9: 6 3 6 8 .

PELSE P B VAN DEN F VAN EI E 2 4 H O K . G A N . R . M JD N 00 . R, . . , , , R VE DEEL, B DER The a ba e systematic value of morphological characters in Seneci o sect . J co a A r — S s tema ti B ta n 29: 790 805 . ( ste aceae) . y c o y

PELSE P . R VAN V EN R EL r v R M EI J DEN 2005 . I n: AN MEI JD . HEUK S F l a a n , . B . DER DER , , , o

N r de la nd ed . 2 6 e , 3 77 .

- W I E . I 2 i n 000 . k ri b Senec o eae . EBE, . The annotated chec list of the t e CASS

A r — Tur z a n i n w a 3 : 6 3 . ( ste aceae) in Siberia . c o i 58 C om w l 44 Ne s . 2006 p . ,

A o o e e u J a coba ea MI LL dditi ns t th g n s . (Compositae— Sen eci on eae)

BERTI L NO RDENSTAM Depar tmen t ofPhaner ogami c Bota ny Swed is h M useu m ofNatural His tor y - 4 lm P. 0 . Box 50007 S 1 0 05 S o kh o S ed e , t c , w n

rtil n r n ta m nr m . b e . o de s @ se

Abstr a ct

w m n n in b MI m a - n i ne Eleven ne co bi atio s J a co a ea LL . (Co posit e Se ec o ae)

r b n r n r r m Sene i . p oposed, all ei g t a sfe s f o c o

I ntr od uction

f MI n n r m n i PELSER et al . r n e . The ecogn itio o J a coba ea LL . as a ge us disti ct f o S ec o L (

2006 n a m r r ran r b . ) ecessit tes so e fu the t sfe s , listed elow

New combinations a l a n m n v J cobaea vu ri s G N. s u b s . otl a di ca EUM . . . o . o . g AERT p g (N ) B NORD , c b

r Seneci o a coba ea var . otla ndi c us UM N in UM N AHLFVENGREN Sv. j g NE A NE A , Flo a 2 6.

Sene i a a u t N RN ( 1 I n A c o cob ea s b s . o la ndi cus UMAN S . . O j p g ( E ) TE ER, Fl sel land ( cta Ph Su . 1 62 . yt . ec

m . nov. J a obaea mollis W lL LD . . . o c ( ) B NORD , c b

Sene i m lli WI . Pl 2 1 1 s S . . 3 : 0 c o o LLD , p h b n Prodr. 7 : 0 1 Se e i e es ermus DC . 3 c o p , r Sene i ca l ver ti i I . K H i 1 n o1 . Di a . Pl . Or. . Se . exs cc . 859 no n c o BO SS in OTSC Y , B ss , g Nov 32

Sene i tchiha tche vi i 8 0 18 i n T HI H Min . . 2 : 2 3 8 . C C . A 9 c o , sie Bot

- J a oba ea n udi a ulis H H AM x D N . om . nov. U . . e . D O . c c (B C ) B NORD , c b

- Sene i nudi a uli U H . HAM D D N r dr . . a 1 78 s . . O P o c o c B C ex , Fl Nep l Senec i o a ll n D W . r e s C . P odr. 6: 367 p ALL ex , w l 44 2006 Ne s . Com . p , 1 3

AN H u . . non Soc . Seneci o unna nens is N H Joum . y FRA C , de Bot FR C , B ll r n 39: 303 Bot . F a ce

1 4 1 Sene i bla tta r i i oli us N H Joum . . 0: 5 c o f FRA C , de Bot R VAN e ert . 8 : 359 Sene i ros uli er LEV . c o f , Feddes p 1 0 : 352 r l LEv Re ert . Sene i es ui o i i . c o q , Feddes p

m n W I NK . . . o . ov. J a oba ea ren ardii . c (C L ) B NORD , c b

i n TRAUTV . RE M XI M . WI NK D Pl . . 6 r WI NK . . ecas Sene i rena di i . c o C L , GEL , A L , Nov ( 1 882)

R f. D an kr. 57 z i H . . S Sene i koel i . c o EC , Biol

m . n v. J a obaea kors hi ns k i K H . . . o c y ( RASC ) B NORD , co b i A a I . . A . Sc . R SS. Ser. 1 3 : 352 Sene i k rs hi ns k i K H . c o o y RASC , ct nst Bot cad U

m n J a obaea tib eti a H K . f. . . o . ov. c c ( OO ) B NORD , c b

H K r . I nd . : 40 f. Sene i ti beti cus . . 3 3 c o OO , Fl B it

m r FF H N m n v J a obaea ultib acteol ata . J . . . . . o . c (C E REY Y L C E ) B NORD , co b w 4 lti r te la tus FFR H N Ke u . 02 Sene i mu b a c o . J . . c o C E EY Y L C E , B ll

m . nov. J a oba ea ra h an i oli a W . ex DC . . . o c p f ( ALL ) B NORD , c b

r DC . Prodr. 6: 366 Sen e i a ha n i oli us W . c o p f ALL ex ,

D . Pr dr. 6: 6 non D UM . Sene i di ers i li us W . C o 36 c o v fo ALL ex ,

m . nov. J a obaea ech aeta . . H N K . . PAN . . o c (Y L C E Y ) B NORD , c b H N A x in 1 4 K . Y. PAN Ph ota . S . 9 : 9 Sene i echa etus . . t c o Y L C E , cta y m J a obaea ambracea U . ex DC . . . o . nov. c (T RCZ ) B NORD , c b i r D Pr 4 Sene a mb a ceus U . C . odr. 6: 3 8 c o T RCZ ex ,

r in um . Ja . . 2 1 55 Senecio ma ns hu i cus KI TAM . Jo p Bot

J a obaea ar un ens is U . . om . nov. c g (T RCZ . ) B NORD , c b

i r i M 2 2 1 4 1 . ene i Sene a unens s U . u . Soc . . . 0 8 7 8 S c o g T RCZ , B ll Nat ose , ( ) c o r 1 b s . a nen is UR E . WI E . S b . 1 3 1 998 66. eru i li us . s u r u s cfo L p g (T CZ) BE, Fl i i ( ,

200 . ur r n n m PELSER et al . 6 F the sy o y y, see

Personally I prefer to keep a rg unens is as a species rather than a subspecies under erucifoli a .

R efer en ces

- mb ina n i n PELSER P. V K M V MEI EN 2 J . F. R . AN JD 006 . , , ELD A P , DER New co tio s

44 1 — 1 1 A ra a Seneci oneae . C m . News l . : . J a coba ea MI LL . ( ste ce e ) o p w 4 N l . 4 2 Com . e s 006 1 4 p ,

N r n ta mi a LUND I N Co o e—Sen eci on eae o de s ( mp sita ) , a new genu s fr om the Andes of Sou th Amer ica

* R OGER LUND I N T Swed is h Museu m ofNatur al Hi s tory P Box 50007 -1 04 05 Sto kh olm S ed . O . , S c , w en

Abstr act

- n i n ri The Th e new genu s Nordens ta mi a LUNDI N (Compositae Se ec o eae) is des c b ed .

r n r m 1 856 was r rr G nox s e a nda W . type is y y p EDD dati g f o , which t ansfe ed to w 2 in 1 2 . ne nu b u 0 r b . J FF 99 Aeq ua to i um B . NORD . y C E REY The ge s consists of a o t species distributed along the Andes from northern Argentin a and Bolivia to northern

rn E r M n b n n Nordens ta mi a are Peru and southe cuado . a y of the species elo gi g to

rr n m a in Ae ua tor i um G nox s an d Sene i . cu e tly ispl ced q , y y c o

I ntr od uction

When NORDENSTAM in 1 978 describ ed the genu s Aegua tor i um he restr icted the genu s m r - flowered altem ate- hr b an d tr to co p ise white , leaved s u s ees with a distinct tomen tum of pelta te — stellate trichomes in two layers especially on the abaxial leaf

- P r e no s H e u flowered m mb r Sene i . a x sides . disc ssed the yellow e e s of c o sect gy y

ATRE r n t n n r . CU C . but o as elated, co ge e ic

Later authors su ch as JEFFREY ( 1 992) and ROBI N SON CUATRECASAS ( 1 992) amended P r a e nox s the gen eric concept ofAequa tor i um by includin g the members of sect . gy y — N RDENSTAM as well as adding new yellow flowered Species fr om Bolivia and Peru . O relu ctantly followed su it and provisionally recogniz ed P r a egyn oxys as a subgenus un der Aequa tor i um (NORDENSTAM

r NORDEN STAM has b m D uring my studies of Gynoxys and in discussion s with D . it eco e in creasingly clear that the P ra egynoxys group is generically distin ct from Gynoxys as a rr ra b n n m r t b well as from Aequa tor i um . This is lso co obo ted y o goi g olecula s udies y

b NORDENSTAM r . Seneci oneae n r u PELSER et al . un u the phyloge y g o p ( , p lished; , pe s

m I r r new n r nam Nordens ta mia for co . ) he e p opose the ge e ic e this distinctive taxon An an r an of de sh ubs d trees .

H i s 1 5 2005 at the a e of 50 . ROGER LUNDI N died une ectedl fr om a heart attack on Nov. xp y , , g

manuscript to this arti cle has b een completed and revised by the Editor.

m w l 44 N e s . 2006 C o . l 6 p ,

Er bran n r b or r 2— 1 8 m m n - ub a on ect chi g Sh u s t ees tall , finely to e tose p escent at le st Lea s a rna or r r un a ar and a un r . e yo g veget tive p ts le f de sides v lte te a ely opposite ,

a or r r a narr or - r r petiol te a ely sessile , l nceolate to owly ovate ovate co date , enti e ,

mi r bb and nn a n r a u mar n n a - n or d i ed pi tely vei ed, co i ceo s; gi s shallowly si u te de tate r an or n u u or mu r n b r n or m r a ep d de tic late , apex ac te c o ate , ase t u cate so ewhat co d te; upper Side glab rescent; lower side greyish - white - tomentose with s ub s tellate or

— Ca i tula r am u br n m m b r m . anchi g , so eti es glo ose glochidiate t icho es p hete og o s ,

m r in mi in l m llif nn r u r n fl r n ra r u u te a u be o o b mi s o es ce ces . diate , seve al to n e o s s py a dal y

I n l u r e ub n r a Re e ta le flat or m a n n n . c p c so ewh t co vex , i disti ctly alveolate vo c s cyli d ic l

u a r u u br n u r br 5— 1 0 s ubuni s eriate c psh ped with few, sho t calyc l s acts ; i vol c al acts , ,

- R - r — b n or b n r a abr or n m n . a o ets 3 5 o lo g elliptic o lo g, co i ceous , gl ous fi ely to e tose y fl ,

- — m b b r . D is or ets 5 1 5 rm r r fe ale; ligule yellow; style ilo ed, fe tile c fl , he aph odite; co olla

- - 5 b rr r r b . A r b yellow, lo ed with na owly t iangula ovate lo es nthe s asally sagittate , ecaudate; apical appendage oblong- ovate; en dotheci al tissue polariz ed; filament

- b n a a ar n r a not b a a k n . St bran n ar coll cyli d ic l , s lly thic e ed yle ches li e o lo g , pic lly — s ela s acute {oni cal with a Short cen tral pencil ofhairs; outside distally papillate . Cyp n r P a us br n m r r n m m r b b u a . ho o o phic , o lo g, gla o s , cost te pp istles u e ous , pe siste t

b rb a . white, a ell te

m n I N m : Nordens ta i a re a da . UN o . nov . Type p (W EDD ) L D , c b

A r r A n 1 : 77 S n . : e ua to i um m: G nox s e a nda W . r. Basiony y y p EDD , Chlo ndi a y q ' K w 2 in n i n : r a u J FF e u . 6 co e do a e nd m W . . p ( EDD ) C E REY, B ll , g Type Bolivi , ’

Pr Lareca a ni 1 n P . 130 . 1 . 1 . 1 W . " . a 85 S . ov de j , v llee de Tipua , , EDDELL ( holo , ) Fig

The new genu s comprises about 20 species distributed from northern Ar gentina P E r re a nda r n rn N. th ough Bolivia to orthe eru and s outhem cuado . The type species p r r n ra a u rn P Ar kn be is est icted to ce t l Bolivi and so the eru . eview ofthe own Species will

NORD ENSTAM . published in a forthcomin g paper by B . 44 2006 C N w l . om . e s p , 1 7

R efer en ces

— An an m H F i eldi a n B t . 1 74 ATR ECASAS J . 1 95 1 . S on a . a o . CU , tudies de Co posit e

A new nu and r n in m a . Br i tt n ia A 1 955 . o CUATRECAS S, J . ge s othe ovelties Co posit e

— 8 : 1 5 1 1 63 .

- AT A r R E El én r Ae ua to i um . D iAz P I H I S. J . CU CAS S 1 990 . EDRA TA, g e o q NORD

- Rev. A a d. m . ne . (Seneci oneae Asteraceae) en Colomb ia . c Colo b Cte

— 659 666.

' ’ - A RE A AS 4 i S. J . CU T C S 1 99 . mb en el n r DiAz PI EDRAHI TA . Novedades colo anas ge e o

A t r i m A ra a Seneci oneae . Rev. A a l m i n e ua o u . d. b . C e c . q NORD ( ste ce e , ) c Co o

24 — 7 252 .

i n m a M n J FF . 1 992 . r b Senec o eae r I E REY, C The t i e (Co posit e) in the asca e e slands with

- — n ann k n ra f r b . Kew Bull . 4 4 1 09 a otated chec list ofthe ge e O the t i e 7 : 9 .

AM n mi in n i n m RD E T 1 . NO NS . 978 . tr b Se ec o eae , B Taxo o c studies the i e (Co positae)

— 44 : 1 83 . Op era Bot .

- NO RD ENSTAM . 1 . nu Ae ua tor i um . R . m a en ci nea , B 997 The ge s q B NO D (Co posit e S e o e)

E m 1 — C . News l . 3 : 1 1 6 . in cuador. o p

A n Ae ua tor i um and n s R I N N H . J . CUATRE CASAS 1 992 . G ox OB SO , dditio s to q y y A n P v 2 4 1 1 — 4 r : Seneci o eae in E u r and ru . No on : 1 6. ( ste aceae ) Bolivia, c ado e w m . N l 44 2 C o e s . 006 1 8 p ,

’ " 1 s l T 5 P 21

w u Nor dens ta mi a UND I N . F i . 1 . G nox s re a nda W . ne g y y p EDD , type of the gen s L

W P L r c a a é u 1 85 1 P . n . r . a e a EDDELL s . , Bolivia, ov de j , v ll e de Tip ani , ( holotype) C w l 44 2 06 Ne s . 0 om . p , 1 9

New comb inations in Norden s tamia Co o e-Sen eci on eae r o Ar e ( mp sita ) f m g ntina, B ol v P er u E cu or i ia, and ad

BERTI L NO RD ENSTAM Depar tment ofPhaner ogami c Bota ny Swedi sh M us eum ofNa tu r al History - 4 m d e P. 0 . Box 50007 S 1 0 05 Sto kh ol S , c , we n m nr m nord ens ta . se b er ti l. @

Abstr act

Fourt een new comb inations are made in the genus Nordens ta mi a LUNDI N

m - n i n n r r m Ar n n Se ec o eae . r are P r (Co positae ) The t a sfe s Species f o ge ti a , Bolivia , e u

u rn E u r r u r un r Sene i G nox s or Ae ua tor i um . and so the c ado p evio sly t eated de c o , y y q

I ntrod uction

The new genus Nordens ta mia LUNDI N has been recently established for a gr oup O f An an hr n r n r I N n w b a d b o G nox s e a nda W . UND e de S u s t ees , ased y y p EDD (L The

n and Ae ua tor i um . . rm r nu ge u s is allied to Gynoxys CASS . q B NORD The fo e ge s is

ara r z b n n - flowered mu ch cte i ed y co siste tly opposite leaves , yellow capitula , ch n Ae ua t r i um ORDENSTAM poi ted sty le branches and a tomen tum of Simple hairs . q o (N

1 978 n b a rn r b - floret u ) is disti ct y the lte ate leaves , white flo ets , o tuse disc styles witho t

n - r n ub m n m O f a r m . Nordens ta mi a a ce t al pe cil , and a do le to e tu stell te peltate t icho es

n ra rn - flowered t and m has (ge e lly) alte ate leaves and yellow capi ula , a so ewhat

n - flore A numb r Of u a in poi ted tip to the disc t styles . e species p to now pl ced

P r n Ae ua tor i um n r in r ub en . a e nox s b e G ox s o Sene i . o s q , y y c o ( sect g gy y ) , eed to ran rr r m OfNordens ta mia r m P ru are t sfe ed to No dens ta mia . So e taxa f o Bolivia and e

The nu an m r an still undescribed and will b e tr eated in a fort hcoming paper. ge s is i po t t con stituent in the cloud forest vegetation especially on the Amazonian Side of the An n w e a b e b e un in a P ru . rk i n des , and tax can expected to fo d Bolivi and e Field wo P 2005 2 m i r n n b u in r in D ec . 006 e fii ture e u to Jan . yielded so e nte esti g fi ds to disc ssed i - N I N r . n r A R N pape s The ge e ic position ofthe few Oppos te leaved taxa (cf. BELTR OB SO 200 5) is still uncertai n and will b e further in vestigated . C om N w l 44 2006 2 O . e s . p ,

M ateri al and methods

Herbarium material was made available to me from a number ofherbari a (abbreviations r m i l M P d in a n H M N H M N l 998 . O S U US an cco da ce with OL GRE OL GRE viz a n y , NY, , , , n m n r m m r r ar additio al speci e s f o so e othe he b ia .

New combi n ati on s i n Nordens tami a

N r n m m r i H R B ATRE m o de s ta i a ca a a cens s . O . CU C . . . o . nov . j ( ) B NORD , c b

r r ROB . CUATREC . Novon 2 4 1 1 Basionym : Aequa to i um caja ma cens e H . : P m Pr D r n An : r . ar : . u r Sa r u r P r u Type e u Caja ca ov C te vo , ist ito d es de C te vo , a q e “ ” a na u r a r Pa n am n a a Jaen rr b an 2600 m N cio l de C te vo , c se io jo al , c i o h ci (Cho o l co) , , 2 VI H. 1 987 DiAz O SORES 585 S . M 1 0. U O , ( holo ;

mi a a is h en is UATREC . om . nov. Nordens ta c r . . p s (C ) B NORD , c b

a n m : n x r i h ATRE Bri ni 1 2 1 G o s ca s ens is CU C . tto a : 85 B sio y y y p ,

N v n 2 : 4 1 2 S n m: Ae ua t r i um ca r is hen e ATRE H ROB UATREC . yn o y q o p s (CU C . ) . . C o o

: P r i h n H a M ri 2 m 1 C s b u u 800 0 . . 1 957 H . r . a Type e u p , etwee n co and Tingo a a, , VII , ELLENB ERG 22 1 1 (U

N r n m r B o de s ta i a a b is u . . om . nov. f (CA RERA B NORD , c b

n Ar n . . 332 m: G nox s a br is i i . Soc . Basio y y y f CABRERA, Bol ge t Bot n n m A r w S : e ua to i um a br is i i R . J FF Ke u . 61 y o y q f (CAB ERA) C E REY, B ll : Ar n n a Ju u Va Gr Ri o J r a I I I PETYRI ELLA in BRI Type ge ti , j y, lle ande, o d n, FABR S , CR SC FA S 1 LP K 57 7 ( holo . ;

r m . nov. No dens ta mi a h uan ucon a CUATR EC . . . c ( ) B NORD , o b

n m : n l i an 4 Se e i hua nuconus CUATREC . Fi e d a . 5 Basio y c o , Bot n n Britt n ia 1 5 S m: G no s hua n uc na ATRE ATREC . 8 x o CU C . CU o 8 : y o y y y ( ) , : P ru H an 1 2 W 4 . 9 7 M . S A 5 . U S Type e u uco , , A DA (F holo ;

N rd n ta m n n n o e s i a u i e s is H . R O B. UATR EC . . . om . nov. j ( C ) B NORD , c b

a n m A r : e ua to i um un i nens e H . ROB CUATREC . Novon 2 : 4 1 2 B sio y q j . ,

Jun n : a a a a H a 2 - 2 m 1 2 KI LLI P : P ru . VI ar b ua 700 3 00 7 . . 9 9 Type e i C p p t , ove capistan , , , SMI TH 24434 (US

Norden s tami k n H R B m n a i u . O . CUATREC . . . o . ov. g ( ) B NORD , c b

n m : Ae r A v 4 1 2 ua to i um ki n i i H . ROB . CU TREC . N o on 2 : Basio y q g ,

: b amb a : 1 5 km r m m on r a un r 1 0 600 fi. Typ e Bolivia, Cocha f o Colo i the o d to T a i , 1 K I N 7 . . 978 I H 768 M 0 U S . O II , G B S OP ( holo ; w 44 2 Ne s l . 006 Com . p , 2 1

b n and ra r N. ki n i i r r u u r n z g is a cha acte istic species , s ally easily ecog i ed y the lo g the a r on b r and and and b a br r stiff h i s the anches petioles , the ovate sally oadly co date

a r m n m bran ir a on r i le ves . The Sho t to e tu of ched ha s especi lly the unde s des of the

n i r I t a ar ab r m n a d a r br n n um nt m . leaves is v i le , f o de se l ght g ey to loose ow ish i d e u is

m m a Shrub or s all tree to 3 t ll .

r r n min : a abamba : 26 km r m m r on r Fu the collectio s exa ed Bolivi , Coch f o Co a apa oad to b amba ft KI N I H 7650 MO 1 US OfM n Punku Cocha , , G B S OP ( , N o te ’ on r abamba— mara a S 2850 m S AH SS82A S1 ° oad Coch Co p , , , T L ( ) l NE M n P n r l Pr . rr 5 km a u o e ami n Sihuenca S ov Ca asco, de o te co p c o a , ,

W 27 0 m 1 0 111 1 988 M N NEE 1 8067 NY1 1 Pr arr km E 0 S US . , , , OLO O ( , ); ov C asco, del u b S b r S W 2950 m 1 988 M RI S I 279 NY p e lo de i e ia, , , , , A O ALD AS (

- b a r . rr a ra n r E an A r San a ruz : Pr . a b t C ov C lle o , Loc To esill s T yecto e t e st cia Las as y Lag r a S W 2500- 3000 m 8 V P 1 3 1 6 NY una B av , , , , ARGAS RADO (

b a r Par u a n Amb r a una ra a rr P n ill n n Pr . a o u o r ov C lle o , q e N cio al o o , L g B v y Ce o g e t a do “ ” or Jarr vi cin idad Torredi llas 3 1 00 m p la a desde la de , ,

4 . . 1 96 I M V R R 27 0 NY Pr a r P u 9 J 0 1 . b ar IV , ARD , A GAS , NAVARRO BA RA ); ov Ca lle o , q e ” n Amb r 25 km al San Ju P r r S Nacio al o o , N de an del ot e o , ,

2630 m 1 . 1 I M R V R 2 1 0 . 996 J 88 " , IV , ARD , NAVAR O, A GAS BARRA (NY )

A a n n r m u Pr n rr m r u . Sud u u b devi ti g collectio f o Ch q isaca ( ov Ci ti , Ce o B fete , c e de la ° ’ ° ’ n l n r ri n 2 22 22 - 2 I A r abu ar e e o o e tal O SO S 64 W 50 300 m 9 . V. I 99S pa te t l lado , , , , , RROYO,

S N H 954 NY 1 USZ r in a - a and m n m but n ERRA O OLST , , ) diffe s le f Sh pe to e tu , Si ce it is r n m n n a r u r i . devoid of flowe i g c pitula, the dete i atio is nce ta n

N r n m m n m o de s ta i a li o ens is . . . . o . nov. (B NORD ) B NORD , c b

"

n m . N ew l . 1 1 4 m: Ae ua tor i um li monens e . R . s 3 : Basio y q B NO D , Co p

: E a r M r n - S n : 49 km r m mon r Gu m Type cu do , o o a a tiago f o Li on oad to alaceo , ,

VHI . 1 996 H N 2 2 1 1 6. S A KNU 88 S , T L DSE (

is UATREC . ma b e ons e i fi Ob s . r a n G nox s az ua ens C c c c The el tio ship to y y y , which y p - A r n ur r n n . I n all hr e ua to i um eeds f the i vestigatio fact, t ee Opposite leaved species of q R I N N ( OB SO BELTRAN 2005) are only provisionally placed in Nordens ta mia .

N r n m . nov. o de s ta mi a lon i s t la R EENM CUATREC . . . o g y (G . ) B NORD , c b

a n m Barcinone 3 : 2 2 : Sene i l on i t l us REENM UATREC . . . s G . C 9 B sio y c o g y , Collect Bot , , (1 953) m n n : ATREC . CUATREC . Brittoni a 8 : 1 58 S G nox s lon is z la GREENM . CU y o y y y g y ( ) , ( 1 955)

: P ru Pr M — u Sa la a n ar r m 3600 3700 m A. WEBERBAUER Type e , ov. oq egua, y p , e Ca u as , , 733 1 A(F N w l 44 C m . 2006 2 2 o e s . p ,

H AN H R B Nordens ta mi a as coens is . . O . . R . om . p ( BELTR ) B NO D , c b nov.

H r H . ROB . m . N w l 42 n m: Ae ua to i um as coens e . AN e s . : 5 Basio y q p BELTR , Co p

P ru D t Pa Pr . a am a ra umm r : o . O r Yanacha a Type e , p sco , ov x p p , il to s it ofCo dille a g , via n D 2 m 4 H B a 1 98 D . . SMI H . OTI GER 8 84 M Ri o S 600 1 8 . . 7 U S aniel , , VII , N T ( holo; AMA MO US Z , , m Norden s tami a rimachi an a CUATREC . . . o . . ( ) B NORD , c b nov

n m n r ima hia na UATRE Ph l i 52 : 1 64 : G ox s c C C . to o a Basio y y y , y g

S n m: Ae ua tor i um r ima chi a num UATREC . H . ROB . UATRE v n 2 4 n C C C . No o : 1 3 y o y q ( ) ,

hinca a n M - r : P ru H uanu D to . C o ri H a u E1 M r Typ e e , co , p , highw y Ti go a a u n co , i ado n r i h 2 - 2 m 2 I I I . 1 980 M . RI MACHI 4908 . r Ca s 600 700 1 . US ea p , , , ( holo ; F

r n m m n No de s ta i a s tell ato il os a GREENM . CUATRE C . . . b . ov. p ( ) B NORD , co

n m : ene i tella t il u REENM ATRE S c s o os s G . CU C . . . Barcinone 3 : Basio y o p , Collect Bot , , 264 ( 1

n m A r n ATREC . FF E K w S : e ua to i um s tella to i los um REENM . CU G . J e y o y q p ( ) C E R Y , 47: 62 Bull .

: P r V1 11cabamba Ri o n MA I 4946 Type e u, , Chi chao, CBR DE (F

N rd n m m o e s ta i a ta va ra H . R O B. UATRE C C . . . b . . ( ) B NORD , co nov

a n m: Ae ua tor i um tova r i i H R B ATRE v 2 4 O . CU C . No on : 1 3 B sio y q . , P H n Pr . Ta aca Ani a Mar n H : r . u : b a tr Type e u a cavelica ov y ja, de c valle , e e uachocolpa y n 3300 m 2 1 1 4 I V. 964 78 1 . US Ti tay, , , TOVAR (

Norden m m . n v s ta i a tues ta e CUATREC . . b o . ( ) B NORD. , co

a n m Sene i tues ta e ATRE Fi l i an 4 : CU C . e d a . 6 B sio y c o , Bot S n n m A r to i um tues ta e UATREC . H . ROB . UATRE N v n 2 4 1 3 : e ua C C C . o o : y o y q ( ) ,

D e . H anu 2 : P r . DI OYTKOWSKI 34095 P 700 m D . U W Type e u p u co , illao , , T ESTA AZ (F holo . )

Norden ta mi w ko ws ku UATRE m n s a o t C C . . . o b . ov. y ( ) B NORD , c

a n m : Sene i wo tk w ki i UATRE l i 4 o s C C . Fie d ana . 9 B sio y c o y , Bot S n n m : n w tk w x ATREC . Brittoni a 1 58 G o s o o s ki i CUATREC . CU 8 : y o y y y y ( ) , P H : . fTam o m 4 XI 1 W 4 r u u . o V a YTK b 3500 . . 937 WO O SKI 1 5 Type e u an co, vic de ac s , , , (F holo . )

m w 4 Ne l . 4 20 6 C o . s 0 2 4 p ,

mn B NO RD . a n a ri oth a us . e e u o C , a n w g n s f e Co o e- Sen eci oneae e e c th mp sita , nd mi to the Canary I slands

BERTI L NO RDENSTAM De ar tm e ofPhanero mi otan Sw dis h M u eu m of atur al Hi tor p nt ga c B y, e s N s y B x 0 - 1 4 05 to khol m wed e 0 . o 500 7 S 0 S S P. , c , n m nor d ens ta m nr . se b ertil . @

Abs tr act

RD . r z as a new nu Of ri b Seneci oneae r m Ca na r i otha mnus B . NO is ecogn i ed ge s the t e f o r m i B h n ur i i D ar e r n r Sene . et e co t C . the C an ary I slan ds . Two Species t a sfe s f o c o (sect ) w i l r r m La an d r ne . ru a . . b a thi d Species , C p co B NORD is desc i ed f o the island of

Gomera .

I ntr od uction

The Canary I slan ds are ren owned for their high degr ee of endemism at different ri n On b 500 taxonomic levels of flowe ng pla ts . the Species level a out species out of a

a 1 200 na r n an are n m or 42% D I al . tot l of tive flowe i g pl ts e de ic , ( AV S The numb r n m n ra ar a n mi n n but u u 20 n r e of e de ic ge e v ies with t xo o c Opi io , s ally ge e a r z endemic to the Cana y I slan ds are recogni ed .

an are ar Macar ones i an r R n and numb r ur r The isl ds p t of the Flo istic egio , a e of f the n r r i n r D D N in W in ri b a e Macarones an m e . . P e i a llis . O S ge e a e de ics , g c EET the t e Seneci oneae m a 1 4 1 2 are n m of the Co posit e , with species , of which e de ic to the r I an MW MW 200 1 SW N N M NN 2003 SW N N Cana y sl ds (BRA ELL BRA ELL , E SO A S , E SO

ar n m Accordin g to current taxon omic opin ion four Species ofSeneci o L . e e de ic to the ’

b llei KUNK . SUN . S. ana I an vi z tener i a e CH BI P . o r . S. S . . S C y sl ds , fi ex BOLLE , D , a lm i M r PI r menti o ens s CHR . S . I NK in u and S. he mos a e . p (NEES) ex L B ch , TARD The fi st

l a r i nd a r n b n in . Sene i ned n r S. vu s a is allied to the ge e ic type g cco di gly elo gs sect c o ,

W r a n n m n n i . e . . S. bollei ot n . S he e s is assig ed to a sectio The two last e tio ed Species , ,

a lmens i and her m a e r n t Be thencour ti i DC . AN p s os togethe co sti ute the sect . (C DOLLE 1 838 NORDENSTAM 1 978 J FF , , E REY

a n m a Bethencour tu was at m n b The t xo o ic st tus of sect . discussed so e le gth y NORD ENSTAM I ts isolated and anomalou s position within Senecio was pointed 44 m N ews l . 2006 C o . p , 2 5

E a au a h r b - b r n a illiforrn out . speci lly the c d te ant e s and the o tuse style a ches with p p

- sweeping hairs serve to characteriz e the group .

E na KUNK r h r u nu un r n m Bethencour ti a xceptio lly, EL t eated t is g o p as a ge s de the a e , h u m a n KUNK 1 975 and ma new alt o gh with so e hesit tio ( EL , he de the n B r PI K H n r ti H I wa n t mb . he mos a e UNK . r Bethe cou a s o co inatio ( TARD) EL oweve , C O SY ’ a ub as a nu n CHOI SY S n r r n a ar in v lidly p lished ge s , si ce ge e ic desc iptio ppe ed n m en i a lmen i U H r a m n m viz n S e s s . sy o y y of c o p (B C The e is also ho o y , - in n w i n r H I L . ub 1 833 o n n m a la ti a P . BR . Betencou ti a ST . G (p lished , a sy o y of c the um a Leg inos e) .

Bethen c ur ti i ha n n a r A renewed study of sect . o s led to the co clusio th t it is best t eated A r H I in U H 1 825 r am in n n as a separate genus . l eady C O SY B C ( ) exp essed the s e Op io whe m B th n ur A m n n nam wa n t n r n r na e e co ti a . s s o he i t oduced the ge e ic e e tio ed, this e w nu validly pub lished and cannot no b e adopted for reasons of homonymy. The ge s is here described as new and the proposed name Ca na r i otha mn us needs no further explanation .

Di scussi on

I n several morphological features the new genus Ca na r i otha mn us is reminiscent Of M nu H uber ti a u r bb b narr the alagasy ge s BORY , s ch as the sh u y ha it, ow sessile leaves ,

- H r a um r u ma an d flowered a au a an r . n e o s s ll yellow capitul , c d te the s , etc oweve , close relationship between these two gen era is not suggested and is also not supported by

in r . n n m u ar tu Seneci oneae n Gr u PELSER et al . r o goi g olec l s dies ( phyloge y o p , p og ess) All three taxa of Ca na r i otha mnus are included in a broad study ofthe whole tribe and A rm a kn r n r m r a Sene i . r they fo closely it g oup , which is o ly e otely el ted to c o ve y close relationship between Ca na r i otha mn us and H uber ti a is not indicated by the

l a r is n m u ar a r i in ur S. u but Sene i S s t . u olec l d ta , c o ( ncl d g of co se the type v g ) is eve m r r Ca na r i otha mn us b n a o e distantly positioned in our phylogenetic t ee . elo gs with

A r an m n n m Sene i but un a n n ra in a a few f ic ele e ts ( a ed c o , still ssig ed to ge e ) cl de which a ar a r r um r n m n n r ppe s s Siste to a J a coba ea clade . The ci c sc iptio of o ophyletic ge e ic n r - r tr. a ba ea and r n defin ed o u as Sen e i . s J co cepts s ch c o s , co , othe p ese tly ill r a be u r min a r r a r n a un ecognised t xa will disc ssed in fo thco g p pe s , togethe with p ese t

n tio of the complete phylogenetic tree .

balusterforrn m n ar in n r and m r m a n The fila e t coll s , dist ct a the tails , s all p is atic hex go al ovary wall crystals in taxa now referred to Ca na r i o tha mnus were illu strated in N RD AM 1 B O ENST 978 : . 7 1 OE ( Figs ,

kn Be thenc ur ti i vi z a lmens is and n Sene i . o . S. The two ow Species of c o sect , p m N w l 44 C o . 2006 2 6 e s . p ,

n D ri hermos a e are r ra rr nu Ca na r i otha mn us . , he e t sfe ed to the new ge s u ng the field work within the present study it became apparent that a third taxon exists on the small

I t r rib a na r i otha mn us n G m r . C r u i la isla d ofLa o e a is he e desc ed as a new species , p co

B . NORD .

Taxonomy

a na tha mn us . . en . nov C rio B NORD , g .

B th nc ur tia H I I NK in H n m ille . P B 4 . hr S n : e e o U o . . es c . ar. I ns . 1 8 y C O SY ex L B C , g , hys Can

- n Betencour tia T. H I L. Vo . I n er. ré . 1 : 376 ne i s . S Se . in y , non , y t B sil c o sect B h n rt i D Pr 4 1 et e cou i C . , odr. 6: 1

altema s es s ili a n r - a Fruticuli glabri ramosi erecti vel ascendentes . Folia li ea i lanceolat n ra ar n i nn i i t r r mb i teg vel Sp se de tata vel lobata vel p at part ta . Capi ula pa va co y osa ecal cu lata hetero ama r a flaviflora I u r bracteae i ri ata e 4 y g adiat . nvol c i un s e e pauca (

r f min — — - u 3 5 herrna hroditi r 5 b a . . Floscu li adii e ei 1 3 b revi . Flosc li disci p ; co olla lo at An therae caudata n m r S r m obtus i s ubtruncati e; e dotheciu adiale . tyli a i apice vel

l o l n a br vihir uta papil ati ; areis s tigmatici s fere confluentibu s . Cypselae b o g e e s e vel

a ill a - P a numeros ae b m u b arb ellatae s emi ers i sten p p os e 5 costatae . appi set e al ae in te p tes .

l N a mens is . . Typus : C. p ( EES) B NORD m S br u hr b r an d n bran in . a a rna all gla o s S u s , epeatedly de sely ch g Le ves lte te , sessile , n r b n r or or - b r u m li ea to o la ceolate , enti e , toothed few lo ed to pinnatipa tite . Capit la s all , r mb i n r r n br rm n s n flores cences r a co y ose athe de se acteate te i al y , adi te , yellow

fl w r I n m u ur na ecal culate but subinvolucral o e ed . u r rr a b vol c e na owly c pan late to t i te, y , br r n ma b u r br r 4 Ra - florets fem acts p ese t, s ll, su ulate; invol c al acts unise iate , few ( y

r 1 D - florets rma r 3 r 5 - b b ale , fe tile , few ( isc he ph odite , few ( co olla lo ed; lo es

- n rr midveined . A r u m a owly ovate lanceolate , nthe s tailed; endothecial tiss e ainly

- r m n ar b alusterforrn b n flat b u . adial; fila e t coll s ; apical appendage ovate o lo g , , o t se

S bran b ub r n a a a r a illiforrn tyle ches o tuse to s t u c te , pic lly with few and sho t p p n - r n m n n n a r ar u or ar . sweepi g hai s , i side stig atic ea co ti uo s e ly so Cypsel s te ete , n rr b n r ir u or a b u ir 5 - or a owly O lo g , Sho tly h s te p pillose with o tuse d plex ha s , veined r bb c r o odiurn in P r n er u a r r . b tun o s i ed; p p dist ct of seve al cell laye s appus istles , m nu ar a - b b m r n ba a nn . i tely ell te , white , se i pe siste t, s lly co ate

S ana I n 3 r a . pecies , C y sl ds

r m m n n= o 1 0 . a lmens is O Ch o oso e (C p , RTEGA NAVARRO

K ey to the Speci es of Ca n a ri oth amn us

M - — 1 . n a b n or b n 1 0 mm br a r 3 5 ost leaves la ceol te o lo g o la ceolate , up to o d, Sho tly w l 44 2006 C N e s . om . p , 2 7

- - n 1 — n a lmens is dentate or lobed with ovate oblo g lobes 5 mm lo g 1 . C. p Leaves entire or 2— 5 - lobed with linear- filiform lobes 5— 20 mm long 2

— — - — 1 n 2 n b mm . i 2. a 2 3 cm 3 5 b 5 0 C. ru la Le ves lo g, lo ed; lo es lo g p co

a 3 —7 cm n r or m 3 - b b 1 0— 25 mm Le ves lo g, enti e ostly lo ed; lo es long

3 . C. her mos a e

m . nov. 1 n a riothamn us almens is EEs . . o . Ca p (N ) B NORD , c b

r r m NEE H r h a n m: i ne a i a a l ens is s . r . Tab . 24 B sio y C p , o ae p ys Be ol

r lm H I Bethencou ti a a ens is I NK in U H P . Bes chr . nar . S n . : y p (NEES) C O SY ex L B C , hys Ca

I n 1 4 nom . ille . enecio a lmens is HR M . I NK H 1 S N . s . 8 C S U . c . , g ; p ( EES) ex L in B C ,

Lea es 2 cm 2— 1 0 mm br b n a narr b n - b a v long, oad , o la ceol te to owly o lo g o ov te , m 3— 5 — 7 — b m n ir or b b at n r a u b ostly ( ) lo ed , so e e t e ilo ed , fl te ed, co i ceo s ; lo es ovate to

- — m I — r r b n 1 3 mm n u r . n l u ra l bra ts 4 5 . n t iangula o lo g , lo g , c onate vo c c No disti ct

u ut ma un ar nd s ub inv lu r l ra r n R - l r ets 1 — 3 b a o c a b . a calycul s , s ll ped cul cts p ese t y f o ; b u n m n in n m n 4- in i i n n m in tu e eq alli g la i a le gth; la i a ve ed, with add t o al i co plete ve s

- — D i rets 5 . bran r n t r a a a a . s l 3 St o b pic lly cf o yle ches sho t, exse ted, pic lly o tuse with few rm A r n n l b u ba us terforrn ar. r . t sho t papillifo pili nthe tails lo g, a o t the le g h of the coll

s ela n rr b n — 3 mm n ra - ur 5 m r n Cyp a owly o lo g, lo g , st w colo ed , with i p essed vei s , rm r shortly puberulou s with white papillifo obtuse duplex hai s main ly along the veins .

— - P a us br num r u 3 mm n m n u barb m r n . pp istles e o s , lo g, i tely ellate , white , se i pe siste t

r r 4 a lmens is mm in r P m u 2 00 m s . m . C. p is fai ly co on the Calde a egion ofLa al a p to , also occuni n in b arran at r a 450 m MW MW 1 974 2001 g cos lowe ltitudes ( , BRA ELL BRA ELL , ; 1 65 m LI D On r m r r r un in n u in , Tene ife it is o e a e , fo d a ha df l of localities Las

fiadas ar n 2 m e . R u A m n r Am ar n ar T Ca 000 . auce ou d ( g at oq e del l e d o go , e Boce del , NORDENSTAM 9326 in S) and lower down in the Barranco del Ri o (on steep cliffs at

m a mu r a 1 400 m NORD ENSTAM 9328 in S . r a n r r r , ) The e is si gle eco d f o ch lowe ltitude m ) in Arico in eastern Tenerife .

I n spite of the small flowerheads the plants may b e rather showy when the densely in AMW MW corymbose synflores cences are in full flower (cf. photo BR ELL BRA ELL

‘ ’ a nam Tur a t e S H N K 1 907 MW MW 1 974 The loc l e is g y ( C E C , BRA ELL BRA ELL ,

2 Can ari th mn r n . o a us u i ol a . . s . ov. p c B NORD , p

: r I an La G m r P r u a na Gara na R u la ar Type Cana y sl ds , o e a, a q e N cio l de jo y, oq e de Z cita,

1 020 m honolithi c r k E— NE NORD ENSTAM 9333 S , steep p oc s , aspect, ( K MA 0 holotype; , , isotyp es

Fruti culus ram — l m 2 m n lin r 1 m t et t . a te a c eari a osus glabe c . al us la us Folia lo ga integra vel plerumque 3— 5 - lob ata coriacea; lob i s linearis u squ e ad 1 0 mm longis et m w Ne l . 44 200 C o . s 6 2 8 p ,

m r m r 2 rmn . r nu s ubcor bos a hete o ama ecal l latis Capitula pa va e osa y g ycu ata . I nvolucrum an guste campanulato - cylindraceum; involucri bracteae 5 — 6

- f min i 2— fl vi u — h l n u ra e e 3 a . 3 5 enn a hroditi lineari ob o gae . Flosc li dii Flosc li disci p ;

- l n l n ob o ae ub eru ae . P n mer corolla flava 5 lobata . Cypselae a guste g p appi setae u os ae barb ellatae albae .

- 1 m nd W Le 2 r n n br u r b ca . a . a ves cm m B a chi g gla o s sh u let high ide long, ostly

3— 5 - b m b b or m n ub r u r - r n b lo ed, so e ilo ed si ple , flatte ed , s co iaceo s , g eyish g ee ; leaflo es

— mm n — 2 mm m r n - i r 5 1 0 u br b k . l nea , lo g , wide , apically c o ate with ownish lac tips

a i tula ra num r u in r mb bra a s nflores cences m C p seve l to e o s co y ose cteol te y , s all , n n i l r r m u u u r a d s ub nvo uc al br m . 1 mm ub a hete oga o s; ped c la acts s all , ca long , s ul te .

I n lu re n rr m an a ub ndr n u r br 5 — 6 u ri vo c a owly ca p ul te to s cyli ical; i vol c al acts , nise ate , narr b n - an — 4 mm n and 0 7— 1 2 mm r n 1 — 3 - n owly O lo g l ceolate , lo g wide, g ee , vei ed

r m m n m n r n R - orets 2— 3 u o u rr u mar . a ac te so ewhat ac i ate , so e with a ow sca io s gi s y fl ,

ma r b n r a 2— 3 mm n am n b n — 3 mm fe le , fe tile , light yellow; tu e cyli d ic l , lo g; l i a o lo g ,

- - br n n 1 mm 4 n a 3 . S a r lo g , wide, vei ed , pically toothed tyle nches li ea ,

- — n nu u m r b . D is orets 3 5 rm r . lo g , with conti o s stig atic a ea, apex o tuse c fl , he aph odite r t b u r b r n n b 5 - b b n rr Co olla yellow, u la elow, g adually wide i g a ove , lo ed; lo es a owly

— 1 mm n n mi dv ined a u . An r 1 mm n n u n a e . 8 ov te , lo g , disti ctly , c te the s lo g i cl di g n n m n r r appe dage and tails; distinctly caudate; tails ca . the le gth of fila e t colla o

n r a a a n a narr - b n b n m n lo ge ; pic l ppe d ge owly ovate O lo g , o tuse; e dothecial tissue ai ly D r n lin r m k n n ar n a b alus terforrn . b adial with s all thic e i gs ; coll elo g ted , isc style a ches ear n u n m r b ub r n r with co fl e t stig atic a eas , apically o tuse to s t uncate with o ly sho t and

m air r n 2 mm n a r s elas narr b . few p pillifo h s outside late ally. Cyp owly o lo g, ca lo g,

5 - r bb or - n r a rm a r ma nl a n i ed vei ed, sho tly villous with white p pillifo duplex h i s i y lo g the

n car o odium 1 0 — 1 2 r r r vei s; p p distinct of cell ows; ova y wall c ystals hexa to octagonal , P a u br n m r W m n m - r n flat . s u u b rb b pp istles e ous , hite , i tely a ellate , se i pe siste t, asally

n n an nn u u co nate i to a l s .

This rare species is on ly kn own fr om thr ee small population s at higher altitudes ofLa m r I n I un n n n in r G . t r o e a the type locali y co ted to o ly five pla ts , g owi g pa tly in accessib le rock crevices of the Sheer phonolithi c precipice of Roque de la Zarcita . m n r ar and a un i n These speci e s we e all l ge he lthy. The Species is also fo d two places at

R u A and n m n o h a d m u r . M oq e de g , wit a few so ewhat depa pe ate speci e s y colleague ULF SWENSON in Stockholm (S) in spected one ofthese popu lation s in 1 993 (Roque de A and NE 1 1 m 2 2 n rm a o 50 0 . . 003 SW N N 580 S g , slope , , III , E SO , photo in ) and he co fi s th t

in n i la mu e a it is dw dli g and that its futu re is precar ious . C. rup co st b cl ssified as a n an r highly e d ge ed species .

3 . Can a ri othamn u h rm P1TARD m n s e os ae . . c . ov. ( ) B NORD , o b

a n m: Sene i her mos a e PI i n PI P U I leS an ar 239 B sio y c o TARD TARD RO ST, C ies 44 N ws l . 2006 C om . e p , 2 9

PI K . ar . 25 : 2 n m . : B thencour ti a herm os a e UNKE Cuad . an 8 o Syn . e ( TARD) L Bot C ill eg .

MW MW 1 4 1 N RDEN TAM 1 7 . I ll us tr. : LI D 1 967 P 3 97 . 00 O S 9 8 late ; BRA ELL BRA ELL , Fig ; , Fig 33 G C , F,

Lea ves m or 2— 3 - b m 3- b 3— 7 cm n b r- filiform si ple lo ed, ost leaves lo ed, lo g; lo es linea ,

— — 1 m n I n l u ra l bra ts 6 7 an a mm n . s ela n c . 5 s flatte ed, lo g vo c c , l ceol te , lo g Cyp r n rr b n - ndri mm n r n br n 5 - ri te ete , a owly o lo g cyli cal , lo g , g ee ish to ow ish , st ate,

P a us b r m densely puberu lous with white papilliform duplex trichomes . pp istles se i

mi n ar a — mm n W r n u b b 3 . pe siste t, tely ell te , lo g, hite

hermos a e r a n r m n n r b n r and m r C. is e dily disti guished f o its co ge e s y the lo ge o e The f n hr n rr n are n . t m dissected leaves leaves o e have t ee a ow seg e ts , which flatte ed

not n r a m m a in ra r but as i n r a a m a ( cyli d ic l , as so eti es st ted lite tu e) , the othe t x so e le ves ir r 4 — - may b e ent e o 5 lobed .

her mos a e n mi G m r W r rar and un in a m C. is e de c to La o e a , he e it is e local , fo d few s all

fR u n n n u a n i n Valleherrnos o r n . . on a O o e a d o pop l tio s the egio , e g the steep w lls q Ca o , ° 4 m . ORD ENSTAM 9332 in S . clifis at Presa del Encantadora at 3 0 m S . (N )

Acknowledgements

I wish to thank CECI LI O MASSANET ACOSTA and RI CARDO ME SA COELLO (Tenerife) and J I N LERALTA La G m ra for n m an in ura r AC TO ( o e ) excelle t co p y the field , the C to of the n Gara ona n P rk Dr. AN N N for u r a d rm n j y Natio al a , GEL FER A DEZ , s ppo t pe issio to collect m f r n D r LF W N N f r r na r a n n n a o a d . U S o b o o e s ples studies , E SO pe so l O se v tio s of the

r u i la . popu lation s Of C. p co

Fin ancial support from the Royal Physiographic Society in Lund is gratefully a kn c owledged . m w 44 Ne s l . 2006 C o . 3 0 p ,

R efer en ces

Wi ld F wers o h a na I nd E Z MW E 1 74 . r x MW D . . 9 l t e C s la s . cmo . BRA ELL , BRA LL o f y n Cabildo I nsu lar de Te erife .

nd 1 w r MWE D Z . MW 200 . Wild F l e s o the Ca na r I s la nds 2 . BRA LL , . BRA ELL o f y , ed M E r R r . dito ial ueda, ad id

V P h B hr i un r na r I n ni l U H . O N 1 825 . h s i ca lis c e es e b de Ca is chen s eln . Koe . B C , L y c g g

i n ch . r n . Ak . W s s e s ad , Be li

P r dr mu s tema ti na tura i re n i ve e a bili . A P 1 . l t N . . 838 s s s s s 6 . CA DOLLE , DE o o y g g , vol

Treuttel Wi irtz P r . , a is

N D A. . H MI 1 994 . Centres o P la nt i vers i . D I S. D . H W V. H . t AV S , EY OOD C A LTO f y tr r h i r r t n 1 E r e A i t t n i . u uth W A Guide a nd S a e o e s e a . ca S es t gy f Co v o op , fi , o

mbr . K . A i nd th Middle Eas t WWF/I U N . s a a e . C U Ca idge,

2 n i n a m Ma n I an FF E . 1 99 . ri b Se ec o e e a r J E R Y, C The t e (Co posit e) in the sca e e sl ds with

- — 4 : 49 1 09 . n ann a r k n ra ri b Kew Bull . 7 a ot ted wo ld chec list ofthe ge e ofthe t e .

K 1 n riti c n ra L m r . a a d . B t. G 975 . taxo es c os e a G G o UNK EL . Novedades y la flo de o e a

— 25 : 1 4 . Ca nar . 7 9

r r P w a r K UNK G. 1 980 . D i e Ka na is hen I ns eln und ih e a nz en elt. G EL, c fl ust v Fische , k S ut art r . t tg , New Yo

LI I ) 1 n n r r I n N r k en ka s r bu . Shr. s e Vi d s , J . 967 . Co t i tio s to the flo a ofthe Cana y sla ds o p

- Aka l 1 M t. Na tur v t Kla s n . . 2 . n iv. f rl . d. a 3 Os . e . s e s U O O . o , , slo

m . NO RDENSTAM . 1 a n m in r b Seneci oneae , B 978 . T xo o ic studies the t i e (Co positae)

— 44 : 1 3 . Op era Bot. 8

1 E n la r Ma ar n i a: um r O J . . RR 977 . e c o és c RTEGA, B NAVA O studios flo a algunos n e os

— B t. Ma r n . 3 0 . de cromosomas I I I . o ca o 3 : 7 8 ' I K Bei tr ci z r K enn tn is d r eta ti n der Ca na r is chen ns eln . S H N H . 1 907 . e u e Ve C E C , g g o

- - D 11 r n . E 1 898 1 899 . Bd . . 1 . . G J eutsche Tiefsee xpedition Teil Fische , e a

’

W N N 2 4 M n ien B t. S U . 00 . P er i a llis far starkt Slakte med rOtter akaro es . Sv. o E SO , c ett g i

l i d s/tr 1 — 2 . 98 : 93 06 .

MANN 2 P n fP r A r : Atotal W N N . i a S U. U 003 . o e i ca ll s E SO , S hyloge y ( ste ceae) evidence

2 — 4 r n Ta x n 5 : 533 5 6. app oach reappraising the doub le origin ofwoodi es s . o

N w l 44 C om . e s . 2006 3 2 p ,

N e hrotheca e o o c e u o e p , a n w m n typ i g n s fth Compositae- C alend uleae fr om the southwester n Cape P r ovince

I NO RDENSTAM M I KALLERSJ O PI A E N BERT L , AR LDE AS Swedish M us eu m ofNatur al History

- m PO . Box 50007 S 1 04 05 St kh l Sw d , oc o , e en

kall ers o nr m. s e mari . j @

Ab str act

Ne hr theca KALLERSI O r b as a new nu m a p o B . NORD is desc i ed ge s of the Co posit e l n l n N. ili i lia L. . R . KALLERSJO tr bu in Ca e du eae . The si gle species cfo ( ) B NO D is dis i ted rn Pr n Afri The r u b n the southweste Cape ovi ce of South ca . Species has p evio sly ee m ar r m u n r a i n u in Gi bba r i a . b au a r b t cl ded CASS ec se of si il f uit o phology, is ge e ic lly n r disti ct on mo phological and molecu lar evidence .

I ntr od uction

Ongoing phylogenetic studies ofthe tribe Calenduleae have clear ly demon strated the polyphyletic nature ofthe small genus Gi bba r ia CASS Thi s genus was established by I NI 1 8 1 7 n G bi l r CASS The nu n ot CASS ( ) with a si gle species , co o ge s was well b r r n un r a u e . . H ub de stood y l te a tho s , g ARVEY who listed it as a do tful ge us of the Ar ctotideae .

NORLI ND H 1 4 r n r t r m ( 9 3) ecog iz ed Gi bba ia as a distinct genu s related to Os eosp e um L . and a l ndula H e n a n r t had b n r b especially to C e L . oted th t the ge e ic ype ee desc i ed alr a D E N 1 836 e dy by THUNBERG ( 1 800) as Os teosp ermum s ca br um THUNB . CA DOLLE ( ) r b am a n X r mn l n i D u r a z n desc i ed the s e t xo as e otha us eck o a nus C . witho t e li i g its

n t r r n or Gi ba r i a bi l . AS ide tity with Os eosp e mum s ca br um THUNB . b co o CASS poi ted out by NORLI NDH ( 1 943) the correct name ofthe type of Gi bba r ia is G s ca bra (THUNB . )

T. NORL .

H r N RLI ND H 4 L n O 1 9 3 a n G ili i li a . . . b o oweve , ( ) added seco d species , cfo ( ) T NORL , ased

Os teos er mum ili ci oli um I NN 1 - kn n an r m p f L . (L AEUS 759) . This is a well ow pl t f o the

u rn a r mm n on b M u and u r ar as so thweste C pe, fai ly co o Ta le o ntain, ill st ated as e ly 1 739 by B URMAN .

NORLI ND i li ci oli um H based his taxonomic decision on the generic position of O. f on a n i z l Thi n k - a ra r v a c s e a . s ur a d Si gle cha cte , . the sh pe of the yp is c ved idney sh ped and C N w l 44 2 06 om . e s . 0 p , 3 3

i li ci oli um also provided with a distin ct ventral cavity in Cibba r i a s ca br a as well as O. f

(Fig .

H r are r ff r n in r m r an m r a ar oweve , these two Species ve y di e e t othe i po t t o phologic l ch r in n n ar n n n n acte s b our e ot r . u (see elow) , and opi io they co gene ic This co cl sio is also supported by molecu lar inform ation obtain ed from an ongoin g study of the entire

KALLERSJO et al . i n r r r i n a r b . Gi bba i a u t i e ( , p ep) Both species of we e incl ded n na i b on u n r m nt and tr nL- F n h r phyloge etic a lys s ased seq e ces f o the ge es , toget e with a repres entive sample ofall recogniz ed genera sensu NORD ENSTAM (in press) . The na a i r i a in r f r n a lysis pl ced the two G bba Species ve y di fe e t parts of the tree (Fig . Gi bba r ia ili cifolia was found as sister to all Calenduleae save Ga r uleum and D im r h the a r a Gi bba r i a s a bra n n r u Os teos er o p o c , whe e s c was ested withi a g o p of p

mums Os teos ermum la brum . E . Br. r a . n ur , with p g N as its closest el tive To e s e that it was not a Of n am na n r a DNA r n PCR and case co t i tio we epe ted the ext actio s , n r m u b ut u . An u in ar m n kkn fi n seq enci g, the es lt was the sa e alyses s g p si o y jac i g

1 6 or a an n r n H LSENB E K R N ST 2 1 r u (FARRI S et al . 99 ) B yesi i fe e ce ( UE C O QUI 00 ) es lted m n r n co pletely co g ue t topologies .

A new monotypic genus is here described to accommodate the southwestern Cape

n r nam Ne hr theca n r species Gi bba r i a ili cifoli a . The ge e ic e p o is i spi ed by the r if en orrn cypselas .

- Ne hrotheca . D . KALLERSJ O en . nov. om o i ae Calend u leae p B NOR , g (C p s t )

- l a Frutex erectus vel diflus u s ramosus u sque ad m altus glanduloso hirt el us . Foli altem a s es s ili a - b n an a 2— 6 cm n a 1 — 4 cm a a a ovato o lo ga ad l ceolat lo g l t , pice

- i r mu r n a mar n nua n a a n u . r b eviter c o at , gi e si to de t t vel de tic lata Cap tula solita ia

h r r I n u rum m anu at m n u r bracteae 1 0— 1 3 pedunculata ete ogama adiata . vol c ca p l u ; i vol c i — l n u uni s eri atae vel s ubbi s eri atae lanceolatae 6 1 0 mm o gae mm latae . Flosc li

— - u radii feminei 1 0 1 3; ligulae flavae s ubter plerum que rubro violaceae . Flosc li disci - h m m r ha lvae la r in r r a flava uin elobata . o o o e ca b ae curvae Q ste iles , co oll q q Cyp selae p g

— — reniforrnes 4 5 mm longae 2 3 mm latae cavemam ventralem in cludentes .

L KALLERSJ : l O . i i i lia . . . Type N. cfo ( ) B NORD

Single species :

AL LERSJ O om . nov. Ne hrotheca ili i oli a L . . . K p cf ( ) B NORD , c b

1 1 2 4 S . . . 0: 3 a n m : Os teos ermum ilici oli um L . B sio y p f , yst Nat ed

1 I . a r : H rb . I N . Lectotype , design ted he e e L N (L NN) Fig

i li i li a r i n u h rn a Pr n n u n ab M un N. cfo is widesp ead the so t weste C pe ovi ce i cl di g T le o m u n i n an d r ar a P nin u and on . tai othe p ts of the C pe e s la, also o ta n slopes up to ca

1 n ar n K and r n H k M un in and t ar 000 m S m . orthw ds to Bai s loof F e ch oe o ta s eas w ds N ew l . 44 2 4 C om . s 006 3 p ,

I t m u r r b or r b b n to Swellendam . is a s elly gland la sh u sh u let with flat o lo g to ovate a rm na leaves with a dentate and scabrid margin . The solit ry te i l capitula have r m r or i n n row ra - florets m im r or phylla ies o e less a si gle , yellow y , so et es with a eddish

in n - flor t are r b a d e s . pu ple t ge elow, the disc also yellow

w i a r i a . n k m n r n . r b The ge us G bb CASS is o li ewise o otypic G s ca b a CASS . is a sh u let r r b r ar ubu u n with a ve y diffe ent ha it , with enti e , line to s late and apically p nge t leaves ; n r br are irnbri cate and r are b m r or the i voluc al acts , the ays white a ove and o e less

- orange yellow below.

R efer ences

UR M N 1 Ra r i rum i a na rum l n a r u 1 2 2 B r J . 739 . o a c a t m . 7 tab . 6 . ous s ere B A , fi p , p ,

Amsterdam .

ND A. P. 1 P r dr mus s t ma i r i v ili Tr l 836 . o s e t s e n e eta b s . 5 . eutte CA OLLE , DE o y g g vol

H 1 1 a r i B . 1 1 1 S I NI 7 . u i P Gibb a . ll . Sc Soc. hil . 8 7 : 39 . CA S , . 8 79 .

F I A J . S. V. A. KALLERSJ O M . LI P M D . A. G K U . 1 996 . ARR S, , LBERT, , , , SCO B, , L GE Par irn n a kkni fm r rm n - inin i 1 2 - 1 24 s Cla dis t s : 99 . o yj c g outpe fo s eighbour jo g . c W m . I n : H V . H . . N F l ra a e is HAR VEY . H . 1 865 . W O W. S ns , Co positae AR EY , O DER, o c p H m 3 . S CO . Dub Ju . vol . odges ith , lin; ta , Cape Town

H UE LSENBE K P F R T ON UI S 200 1 . MrBAYES: a i r C , J . . . Q Bayesi n nfe ence of

n r B i n r m i 1 4- i a t s 7 : 75 755 . phyloge etic t ees . o fo c

in r KALLERSJ M . N RD EN T A O O S AM B . E N S P. KE . P. A. R . W . , , , , LDE , BAR R, N OOD p ep Are- evaluation ofgeneric delirnitati ons in the Cal enduleae morphological and m olecular evidence (tentative title) .

LI NN EU . 1 7 t ma n tur 1 I m lvii H lrni ae . 59 . S s e a a e . 0 . . . Sa o A S, C y , ed p L ,

NO RDENSTAM . in r l RE T FF r b Ca enduleae . I n : KADE I J . W. . J , B p ess . T i e , C E REY (eds) , Th F VI I I r cd . K . KUBI TZ KI . e a mili es a nd Genera o Va s ula r P la nts . f c , vol (se ies , ) in n D r rk r A r . r n H b : Flowe g pla ts . icotyledons . ste ales Be li , eidel e g , New Yo r n Sp i ger.

N RL di . M n r O I NDH T. 1 943 . tu n I r S es i the Ca lendulea e . , o og aph of the gene a D im r a r i h the Ca s ta lis Os teos ermum Gi bba i a an d Chr s a nthemo des . o p o c , , p , y

W. K . Gleeru un . C . p, L d

H UNB h F P r dr mu la nta a en i um r r r. o . r. . P . 1 800 . s rum c s J T ERG , C o o p p , pa s poste io E m ali U s ae . d an, p C 44 2006 3 5 News l . omp . ,

n r ofNe hr otheca Os teos ermum ili c i oli um L . p i 1 . t , g F g. Lecto ype of p f e e ic type

I N I NN . KALLERSJO. B NORD . L N (L ) ” ’ i li c ifoli um in LI NNAEUS handwritin g . m N w 44 C . 2006 6 o e s l . 3 p ,

2 a f N hr li i F i g. Cypsel s o ep otheca i cfoli a (L A— r r D— F . . KAL R B NORD LE SJO ( C) and Gi bba i a s ca b a (THUNB . ) T. NORL . ( ) . A D R i E V n r . n t n n , adial side . B , e t al side C , F Lo gi ud al sectio Showing seed X m NORLI NDH and n ra a r . r cavity (left) ve t l c vity ( ight) 6 . F o C w l 44 2 om . Ne s . 006 p , 3 7

n i n f r F ig. 3 . Simplified parsimonyj ackkni fe tree showi g the posit o o Nep h otheca

L- F n in tri b b a on nt and tr n u . the e , sed seq e ce data

r i n Numbers on nodes represent pa s mo y jackknife frequen cies . m . N w l 44 3 8 C o e s . 2006 p ,

Generic r evisions in the tr ib e Calen d uleae (Compositae)

BERTI L NO RDENSTAM D e ar tm ofPhanero ami c ta Sw d i h M u um of atu r al Hi tor p ent g Bo ny, e s se N s y - m wed n P. 0 . Box 50007 S 1 04 05 Sto kh ol S , c , e m n rti l . n r n s e b e o de sta @ rm .

Abs tr act

The generic limits in the tri be Calenduleae are revised with special focus on the

ir um r n te s er mum . r new n r r m u rn Afii v iz Os o ca . c c sc iptio of p L Th ee ge e a f o so the ,

M n ulus . . 2 Nor li ndhi a . . 3 I nul ides . o oc B NORD ( B NORD ( and o B NORD . 1 are b for m n r n i n Os teos ermum or Tr ter is ( Sp . ) esta lished ele e ts p eviously i cluded p ip

w m n a n ar ma an n w r i dhi . n ar n e b e d e No l n a LESS The ecess y co i tio s de , the Species

r r b r m u rn m m b . St . H n m ap te a B . NORD . is desc i ed f o so the ost Na i ia The ele a ende ic t r m n ta - h l n r n rr r Oli oca us . Os eosp e um s a c e e e a e T. NORL . is t a sfe ed to g p LESS as

Oli oca r us burchelli i . m . n v n er mum . b o . Os teoS g p B NORD , co Fi ally, p i n rr ra b k D im r h the a V I . D . p na tum (THUNB . ) T. NORL . is t nsfe ed ac to o p o c A LL as HA i nna ta B . p (THUN ) RV .

I n trod ucti on

Ongoin g phylogenetic studies on the Calenduleae and a review of the tri be for the

F a mi li es a nd Gener a o Vas ula r P lants NORDENSTAM in r n f c ( , p ess) have ecessitated r E nu evisions ofgeneric limits within this small but in triguin g tri be . specially the ge s

t r r Os eos e mum . in r r m r r p L is need ofa efined ci cu sc iption, a p ocess which was sta ted

Bla xi m D m r t I an r urr the r r . u i h V d with the t ansfe ofsect to o p o heca A LL . the es ection of n r Tr t r 4 RDEN TAM a Oli oca r us . ORDENSTAM 1 99 a b NO S i e is . N ge e p LESS and g p LESS ( , , I n thi s paper some furt her elements are segregated from Os teosp er mum and

r r n w r b e r . A r nu Ne h otheca . . desc i ed as th ee gene a fou th new ge s , p B NORD KALLERSJO ra r m i r n r b ar r G bba ia . a d , is sepa ted f o CASS desc i ed in a sep ate pape

AM l r r a N RDEN T . Am u r LERSJO et al . O S et a . KAL ( olec la study ( , in p ep ) is unde w y to

u n ri E a a n m b el cidate the phyloge y of the t be . speci lly the ffi ities and li its etween

hr a n th m FAB R Os teos ermum are u r and b e C ys e oides . and p still n esolved will W NORDENSTAM discussed in forthcoming papers (cf. also OOD

m w 4 Ne s l . 4 2006 C o . 4 0 p ,

Tr t r is l nd s ti na S nn 6: 97 Os teos er mum cla nd s ti n um n . : i e c a e e Sy p LES , Li aea p R NORLI ND H S i n Calenduleae I : 328 H t : (LESS T. NO L . ; , tudies the olo ype ” ”

H rb . HUN . 307 1 3 a lendula h br ida . 3 S HUNB . UP T ERG S loco , e T BERG no C y , fol ( ” ann a Tr i r i a nde ti na I N te s l s b . THUNB) . The holotype is ot ted p c y LESS G (cf NORLI NDH 1 943:

4 2 2 I ll tr. . M ra S. A r 1 9 9 . 275 MARLOTH r S. Afi. 1 3 us : J . K 9 BLAC , Flo ust alia , , Fig ; , Flo a ,

1 4 1 — h P a 67 . A NORLI ND H 9 3 . 33 a . l te Fig ; , Figs f,

— — Er nnua rb 2 5 dm m m or br n n . rb u 3 6 ect a l he high; ste si ple a chi g Leaves he aceo s , (

— m m i in n a r r a a 1 0 cm n 1 3 c b r o a . ) lo g , wide , o tuse , a g ns s uately de t te ep nd Capitul l xly

— r mb or m m ar 2 5 cm n r br . co y ose so eti es solit y; pedicels lo g , spa sely acteolate

— — — - fl r — 1 n I n u ra br 8 1 0 5 7 mm 2 4 mm . Ra o ets 8 0 r u vol c l acts , long, wide y , sho t, eq alli g in u r br i n n or n r or u ur the vol c al acts le gth slightly lo ge , yellow, with witho t a p plish

— a mm n . black basal Spot . Cypsel 8 9 lo g

V 1 6 U N r m m n mb r : 2n 1 6 NORLI N DH 1 963 CHOUK SANO A et al . 9 8 Ch o oso e u e ( , , T R ER

D istri bution : Westem Cape from the Cape Pen in sula north to C alvin ia and L .

ama u an n n n rnm am b a a r S n m N q al d and exte di g i to southe ost N i i , e stwa ds to welle da

an d Oudtshoom .

NORLI ND H r z n n m a lendula mons tros a and Tr i ter is la ndes tina eali ed the sy o y y of C p c , r r m n H e a but nevertheless chose the latte epithet because of the bette typ e Speci e . lso

k m tr n r r H r r mark NORLI ND H ons os a . b disli ed the epithet as i app op iate oweve , as e ed y ,

f m n tros a un m ak b am a n as T la ndes ti na . the type O C. o s is ist a ly the s e t xo c

M l h r m 2. onocu us os e oides D C . . b . . y ( ) B NORD , co nov

n r r Prodr. 6: 458 m : Tr i te is h os e oides DC . Basio y p y ,

in Os teos er mum h s er ides D . . NORLI ND H S S n . : o o C y p y ( ) T NORL ; , tudies the

Calenduleae I : 333 n r : D REGE 2737 a Pr . . Lectotype (desig ated he e) , C pe ov , L

- m m a : b n K u and r D NORLI ND H l . o . Na aqu land etwee o se Ga ip (G C) . cited this speci en as

r n n type without furthe i dicatio of its status .

ll RL 4 i — I ustr. : NO IND H 1 9 3 F . 33 k. , g j

Er or n n ann ua rb 2— 5 — 7 dm r m r rn ect asce di g l he ( ) high , with a o atic odou ; ste m ra r r n n r n n in a rb u ode tely to ichly b a chi g; lower b a ches asce d g . Le ves he aceo s to s ubcam os e u 1 0 cm an d 2 cm a bt r mar n , p to long wide , apic lly o use to ounded , gi s

in u - n a r r an a - m r na a r mb o u . s ate de t te ep d with c llo s uc o te teeth C pitula laxly co y ose , 1 7— 8 mm m n 1 m n r I n r ra ca . 3 a u 0 c b . u b few to y, pedicels p to lo g , acteolate vol c al cts ,

n — 4 - - — m n r R a floret ca . 1 3 or ran r 2 3 3 mm . s lo g , wide y , yellow o ge colou ed, ti es lo ge

— mm n n u ra bra a 7 8 . tha invol c l cts . Cypsel s lo g 44 2006 C m N ews l . o . p , 4 1

= Chr omosome number: 2n 1 6 (NORLI NDH

D r bu : r rn a Pr n : V n Rh n r a s do a n and . ist i tion No thweste C pe ovi ce y p , C lvi ia L ama ua an r N q l d dist icts .

r a n u r m M mons tros us b n ra ar e This Species is e dily disti g ished f o y the lo g ys , which

n n u r br or n r and n r n - r u . twice the le gth of the i vol c al acts lo ge , ofte o a ge colo ed

N rli n dhi a . . n nov. o B NORD , ge .

t o ermum E n b . n a Os e s . e es tr a ta . . u u p sect f T NORL s sect Co f s T. NORL .

N. a m le ten HAR : . Type p c s ( V) B NORD .

H er ae ann uae an u - u n l n i a altem a b gl d loso p besce tes g a d bus ad api cem atris . Foli caulina herbacea in tegra vel sinuato - dentata vel repan da petiolata vel s es s ili a bas aliter

m r emi am lexi caulibus . a u r I u r bracteae s ub un i s eri atae s p C pit la co y bosa adiata . nvol c i - l n l u r feminei flavi a ranti ci u villoso g a du os ae . Flosc li adii vel u a t bo villoso et

a tri alatae r x l l r f glan duloso . Cypsel e vel fe e e a atae g ab ae e enestratae interdum

u i - r r landulos i flavi lob i s atr urei n ros tratae . Flosc li d sci Q ste iles spa se g opurp s si e ven i in clerench mati u nthera l t u ifi S dist cte s y b s ; a e ca cara ae ca datae; stylus apice b dus .

— - L rn Annua rb an u ar ub n b a k n . ea es l he s , gl d l p esce t with l c tipped gla ds v alte ate ,

in - n - or u n or u b in . Ca i tula petiolate sessile , s ato de tate de tic late , ase half clasp g p

I n l u ra l bra ts s ub uni s eriate or n ar b r a ar r mb ra . co y ose , diate vo c c e ly ise i te , p tly

r n m r 1 - n ar R - r m n 8 . l ets r u u . a ove lappi g, e o s (ca villous gla dul y f o fe ale , fe tile , yellow to orange; tube glandular and villou s with mu lticellular two - tiered (apically 1 s elas m or m r m m m or tiered) trichomes . Cyp ho o di o phic with apical cavity (so eti es s all

- D is - r e ts ab n u n ra 3 n or W in m n r ra . l se t) , witho t fe est , wi ged gless , so e Ofte ost te c f o n m r - b a k r a W u s clerench mati c fu ctionally ale , yellow with pu plish l c tips ; co oll itho t y ran an u ar u r a b a a abr u m r n n st ds , gl d l o tside; co oll lo es pic lly gl o s , with thin a gi s with thi lateral vein s but without s clerenchymati c strands; marginal resin canals present; tube

r an ar An r a ara - au b u ua in m n r in sho t, gl dul . the s c lc te c date with tails a o t eq ll g fila e t colla length; endothecial ti ssu e polariz ed with distinct thicken ings; filament collar straight n r a un rm u k n a n a r an ar - a cyli d ic l , ifo (witho t thic e ed cells); apic l appe d ge t i gul ov te

- l k r m r i one in row b a k a u mid ined r u . a g ned with dist ct cell , l c ish , c te , with a da esin d ct

S a a r n m nu bifid or r b b mam rm a tyle pex Sho t co e , i tely sho tly ilo ed, with illifo papill e , r h a u ar ran b ase of cone with a ring or collar of ob tuse hai s; s aft with two v sc l st ds .

- O r abr u 2 n . va y gl o s , Vei ed

m n r not k . Ch o osome no . now

r NW Pr n u rn m b a . Th ee species , Cape ovi ce , so the Na i i

This new genu s differs from Tr ip ter is by the cypselas with small or even lackin g apical

a u r and - floret r a u n s clerench mati c c vity witho t fenest ae , disc co oll witho t disti ct y w 44 N l . 2 Com . e s 006 4 2 p ,

d b b nn a b n Nor li ndhia in r m I nuloi es cf. vei s . is dist ct f o ( elow) y the a u l ha it with u n rna a r mb s nflores cence r n non - u r r ca li e alte te le ves , co y ose y , p ese ce of gland la hai s

D n r m M n cul us are a n f - floret r etc . istinctio s f o o o found in the ven tio o the disc co ollas

- (no s clerenchymati c strands) and cyp s elar morphology (non fenestrate) .

K ey to the species of Nor li ndhia :

1 3 - i 2 . Cyp selas w nged Cypselas without wings (or with rudimentary in complete wings)

Ra - florets 2— 3 m r n n u ra bra 2. y ti es longe tha i vol c l cts

R ay- florets equalling involu cral bracts in len gth or Slightly longer

N. bre vi ra di a ta . . . . 2 . (T NORL ) B NORD

m . nov. 1 Norli n dhia am l e tens H . . . co . p c ( ARV ) B NORD , b

H V ND . 3 : W. H . H E S . m : Tr i ter is a m le tens HA . Basiony p p c RV ; ARV Y in AR EY O ER, Fl Cap N ORLI ND H S in 42 teos er mum a m le tens H R . 7 . Os p p p p c ( A V) T , tudies the 1 ama ua alenduleae I : 339 a r : B E 586 . C Lectotype (design ted he e) R GE , L N q land

4 Prodr. 6: 57 . Tr ter is a m lex i ca ulis n u DC . ur r . : i F the syn p p se s , p p non '

Afi. P . T con s a BOL . H . U ran . S. T a m lexi a ulis HUN . S r p c (T B ) LES ; fi ; BOL S , T s hil 4 Mus . : 1 7 Ann Afr. 9 HUT H . J . HUTCHI N SON . S. 394 T ra ilis J . g c C , ,

— I ll N RLI ND H 1 43 Fi . 35 a l . ustr. : O 9 , g

m an d a - N. a m le tens nnu rb p c is an a al he to tall , with petiolate to sessile h lf clasping leaves and with richly branchin g s ynflores cences of radiate showy capitula . The

- - R a - florets or vegetative parts are glan du lar hairy with black tipped glan ds . y yellow

- - A an r rk ra florets ar e b ut rk . o nge , while the disc yellow da tipped lso the the s have da

a a n . tips , especially the pical ppe dage

D istri bution : Western Cape Province from Van Rhyn s dorp an d Calvinia throughout L .

Namaqualand .

m 2 . Norli n dhi a brevi ra di ata . . . . o b . n ov. (T NORL ) B NORD , c

NORLI ND H St in a m : Os teos ermum brevi ra di a tum . . B siony p T NORL , , udies the

— m 1 50 m Calenduleae I : 345 : S H H 84 1 5 Pr . an Type C LEC TER , Cape ov , Cl willia , ,

1 896 S fide NORLI NDH 1 943 in BOL G GRA K P PRE W Z . ( holotype; isotypes B , , , , , , , , )

I ll RL H 4 m— NO I ND 1 3 . 35 O . ustr. : 9 , Fig C 4 om . N e ] . 4 2006 p ws , 4 3

Annu rb m an u ar— b . a rna or al he to high , gl d l pu escent Leaves lte te , petiolate sessile

a - n rb a - b b n - 3— 8 cm n 1 and h lf claspi g, he ceous , elliptic o long o o lo g lanceolate, lo g, m I n m c wide . co pari son with the preceding species the capitula are fewer and less

u u mu r r r r r conspic o s , with ch sho te yellow ays , ha dly exceeding the phylla ies in len gth .

D ri bu n : S a r rr n r m am n ist tio c tte ed occu e ces f o Clanwilli orthwards in Calvini a and L . Namaqua lan d to southernmost Nami b ia (MERXMULLER

3 . N orli n dhi a a tera . . S . . p B NORD , p nov

: NORDENSTAM N 1 2 mi H n UN 3 b Warrnb ad D r. : a Type L DGRE , Na ia, ist ochstei , isol ted kopje 2 1 km SE of Ai - Ai s 62 km on Ai - Ai s road from road junction 4 1 km N of

1 74 NORDENSTAM : N Vi ols dri fi 25 . . 9 8 UN 1 23 S PRE . ) , VI , L DGRE ( holotype , isotype)

: . . I ll us tr . Fig l

a m l ecten te H R . . ffini N. p ( A V) B NORD a s sed indumento b revi ori et mi nus glan duloso landibus u u brevioribus et r r m raeci ue ex alati s interdum g q oq e , p aese ti cypselis p p vel dim ntalibus a i c li u t r irn l li allis ru e p a b s e aro s u bas a bus in s tructi s .

ar ann ua rb 1 m br r r u u r - ub n Co se l he to high , anching , sho tly hi s te and gland la p esce t,

m n Lea e u u m br br n m 1 cm k. s so ewhat gl ti o s , old ste s and anches gla esce t; ste to thic v a rn and am au or r b n - b lte ate , sessile plexic l sho tly petiolate , o lo g o lanceolate to

rr - b 3— 6 cm n 1 — 2 cm n ir or r b un na owly elliptic o long, lo g , wide, e t e Sho tly and l tly few

b or - n a s ub cam os e rrri dvein ed ain r a ra ins a u or bt lo ed de t te , , with f te l te l ve , c te o use

mu r . Ca i tula num r u rmi and r r mb r and c onate p e o s in te nal axilla y lax co y s , adiate,

- — fl w r P narr n ar filiform 2 4 mm n bra . yellow o e ed . edicels with owly li e to lo g cts

I nv l u r e m r u r br 1 8 — 2 1 n ar b r in r- an a o c he isphe ical; invol c al acts , e ly ise iate , l ea l ceol te , 6— mm n mm W a umi n a r r u u ar and r 9 . lo g , ca ide , c te , sho tly hi s te and gland l spa sely

- 2 1 u 1 — 1 mm n r b . 5 R o ets ca . rr ar u m r . a setose , with na ow sc io s a gins y fl ; t e lo g , villous

2 - r m ar bt ri m ami n rr b n — ob a with tie ed ulticellul o use t cho es ; l na a owly o lo g l nceolate ,

— - - bran narr i r . 2 mm n 9 1 2 mm n 2 mm 4 . S lo g, wide, veined tyle ches owly l nea , ca lo g,

b l - a 4 mm n — 2 mm r and s ua . s e as b o tuse Cyp ellipsoid o ov te , lo g , wide , te ete u lly

n n u rr br u for m a n lo gitudi ally s lcate with a few fu ows , gla o s except so e set e alo g the

rr a r abr n or m r n or m 3 r u n fu ows , l te gl esce t u icate , without wi gs so e with ed ced wi gs a and m m b a a a u n ra a na apic lly so eti es also s lly, with apical c vity witho t fe est e (occ sio l - r n D i - l r e ts n m r or 3 m n r r b . s cypselas with l s all fe est ae) , light g eyish ow ish cf o u e ous , mm n 4 n b r . u mb ma r a ca . mm functionally le . Co oll lo g; tu e sho t, ca lo g, villo s; li rr am ar u al roxirn all r b ri a ar na owly c panulate , sp sely villo s especi ly p y; co olla lo es t ngul

u br u n m n a a a in a ra in ovate , ac te , gla o s (o ly i utely p pillate outside pic lly) , with th l te l ve s r n m s m n b i n an d n in wn r with thin scle e chy a, ana to osi g elow s uses exte d g do the co olla n n n as thin vein s (no distin ct s clerenchym ati c strands exte di g dow the corolla) . m w 4 N e l . 4 2 4 4 C o . s 006 p ,

Ant r 2 mm n u rr r an ar he s long , disti ctly ca date , apical appendage na owly t i gul ,

u m a k n i n m n r ar z ac te , so ewh t thic e ed the iddle; e dothecial cells sho t, pol i ed (few n r z n n m n S a k n n o r rt un rm . a thic e i gs ho i o tal walls o ly); fila e t colla sho , ifo tyle pic lly

ifi r a narr b n abr u 2 - n n an d b d . O r . co ical , ste ile v y owly O lo g , gl o s , vei ed

D ri bu n : O n kn n r m n in u rnm m b r ist tio ly ow f o the type collectio so the ost Na i ia , whe e it was foun d gr owi ng i n schist and gravel on an isolated kopje .

N a t ra r r m rm r b W in a ru m n r . p e diffe s f o the fo e two species y the gless cypsel s ( di e ta y n m m r n r r ub n r and rt r an wi gs so eti es p ese t) , the Sho te p esce ce with fewe Sho e gl ds , the n r narr r r r narr m u b r m r . ostly e ti e leaves , and the owe invol c al acts with ow sca ious a gins

I n ul oides . . en . nov. B NORD , g

E ene tra ta ub . T men t a . . Os teosp er mum sect . f s s sect o os T NORL

rr i n r l H rb r u s ubte aneus l os us . os u ata u e a pe ennis , ca dex g Folia petiolata spath lata vel rotundata integra sub cori acea utrinque albido- tomentosa apice acuta ad rotundata

a a ar a s ca os a ra a a . I n u r bracteae in terdum cuspidata . C pitul solit i p di t vol c i

- subuni s eriatae s ubbi s eri atae i n r r mar n r fimbri atae . u r ad , te io es gi e sca ioso Flosc li adii f min fl v i n l r e ei a i u s cco c anes ce tes . r r tri a atae vel l tei , in y Cyp selae t iangula es , late a

u rugosa et api cem versu s glan du lifera; cavem a apicalis magna efenestrata . Flosc li

- i r v S u r a m n u bilobus . disci Q ste iles fla . ty l s ste ilis pice i te

I : nuloides t ment s a . fil . . D Type o o (L ) B NOR .

Mon otypic :

l m n I n u oides to e tos a L . fil . . . om . nov. ( ) B NORD , c b

fil . I NNE fil . u . Pl . z 3 4 n m : Ca lendula t men t s a . S 8 Basio y o o L ; L , ppl

Tr r fil E n m S n . : i t i m n L n e s t e t s a . . s s I N S e . : 90 y p o o (L ) ; LESS G , y opsis g Co p

Os teos er mum t m n t m fil RL H in alenduleae I : e s u . . . . NO I N D S C p o o (L ) T NORL , , tudies the 3 r d K atTerku ls r r Oct b . 3 6 H : HUN S . n . Ca . . S a olotype T BERG , e p B pel y ivie , , he “ ” 2 4 - t HUNB HUNB no . 07 0 UPS H B : Ca lendula t men s a E T ERG ( T UN ); isotypes o o , T RG , “ M NTI N rb . O S rb wARTz D HL Dr . S S a he ( ) , he ( ) , A

r n Os teos ermu u t D Pr . 4 1 83 ur s . : a u odr 6: 63 F the y p m c sp id m C . (

I ll RLI ND 4 tr. : NO H 1 3 Fi 34 us 9 . , g .

m n b r m L ma n r u a r nn a r o b r b . ea es To e tose pe e i l he s all su sh u v i ly os l te , petiolate

a u a rb n ir a a mu r n or r a sp th l te to o iculate , flat, e t e , pic lly c o ate sho tly cuspid te; petiole m i t um a a ra a . I n l u r a l bra ts b a n . Ca ul r ase so ewhat cl spi g p solit y, sc pose , di te vo c c subun i s eri ate nn r n - fi in R - r t ma - r r u mbri ate mar . a o e s , i e o es with sca io s g s y fl fe le fe tile;

mina but urn n b u u n r n rm n n . la yellow t i g l e po d yi g , epide is with elo gate co vex cells s elas 3 - n ar a a a but a ki n n ra ru b n Cyp wi ged, with l ge pic l c vity l c g fe est , gose etwee the C w 44 2 Ne s l . 006 om . p , 4 5

n and a i a m a a u ar- a r D is - orets n n m wi gs p c lly so ewh t gl nd l h i y. c fl fu ctio ally ale , yellow; r a b ri an ar- a ran u n br a mar in ma a a a a co oll lo es t gul ov te , with t sl ce t o d g s , s ll p pill e pic lly, an d ub a r n n i h k n r n ma no n u ar r n s l te al thi vei s w t thic e ed scle e chy ; egla d l hai s , o ly i r An r n mu u on r a an d a . a b k lt cell la glands co oll cypsel s the apical appe d ge ovate, lac ish , d n a n m n uba u an r . S b mam a a a s c te co c ve , with disti ct a gi s tyle with o tuse ill te p pill e ,

n r n - style apex bilobed dow to the i g of short sweeping hairs .

D istribution : A sin gle species with scattered localities in the southwestern Cape

Pr n . I t r b m n rar n r r r n n are r . ovi ce is pe haps eco i g e , Si ce the e athe few ece t collectio s (cf RLI NDH 1 943 The i r bu r n r m u b a and Paar in n r NO p . d st i tion a ges f o T l gh l the o th to

a P n n u and n in u an d a r M Ba n C pe e i s la Caledo the so th , e stwa ds to ossel y. The si gle r r r m a P ni n r m 1 2 eco d f o the C pe e su la dates f o 89 .

S m n : a n Kl nri vi ersber e am r b r Au Z EYHER 3068 elected Speci e s C ledo , y g Zwa te e g , gust,

S ND r S n D iv. emkloof a r R r H errnanus 1 50 m ( O ER sc ipsit, ); Caledo , F N tu e ese ve , , ,

BURMANN 1 1 22 B L r a Div ar n a P r 1 1 . NW . 983 O r . IV. , C ( ); B ed sdo p , h d sto y fl ts ofThe oo t R 2 1 r r iv 2 f a n E m 5 . D( . 1 933 J . P. H . A K 768 S s D . mi W o lo g the li oad, , COC ( ); B eda do p , les

I X. 1 2 NORD EN TAM 1 3 PRE 420 D e H r a 1 7 . 96 S 95 M S 3 Napie , s ndy veld, , ( , , ); ( BC) , oop ,

Potb er r R r 1 km NW Potb er u b m un in 1 90 m g Natu e ese ve , of g ho ses at ase of o ta ,

2 1 1 PRE nb onkers hoek r R r 5 . . 979 . U 996 S J VII , C B RGERS ( ); telle osch , Natu e ese ve , ’ ’ ° ° 4 1 2 1 D D n m r 1 9 A. R . W 0 S . 33 S . 33 59 S I 8 S7 E 1 9 . . 99 , , VIII , OOD ( ) ( ) , welle da dist , N 2 1 B L I X. 1 5 ESTERHU YSE 78 7 O Leeuri vi erber 1 . 9 8 foothills of g , , ( ); N foothills of K P 42 I X. 1 83 J . VLO 720 RE . 3 1 b r m a n n r Bonn iedale 1 300 ft 3O. 9 Lange e g ount i s ea , , , ( ) ( ’ 4 P HN N RE 1X. 98 . 8509 . AB Ri r a a r 2 1 5 m 28 . ), ve sd le i field, , l , BO E G )

t men t s um in ub n own n . NORLI ND H ( 1 943) placed 0 . o o a s sectio of its withi his sect As n m m a a k n ra u an E enes tr a ta . f the a e i plies , the cypsel s l c fe est e , altho gh they have apical cavity which is very large .

t ment s um r r m all or m Os teos er mum an d The hab it of O . o o diffe s f o ost species of p Tr i t r I t r nn a r u m n and un p e is . is a low pe e i l with os late to e tose leaves scapose ped cles m m O f m n ff r n r m Tr i ter is are bearing a Single capitulu . So e the ai di e e ces f o p the

b n c s elar n r non - s clerench mati c na n a se ce of yp fe est ae , the thin y ve tio of the disc

floret r mam a m a a . co olla , and the ill te stig atic p pill e ofthe styles

- to Oli oca r us Tr a nsfer of Os teosp erm um s an ctae h el en ae T. NORL . g p

m . n ov. Oli ocar us b urchelli i H K . f. . . o g p ( OO ) B NORD , c b

H na: 290 H : in MELLI SS St . f. n m : Tr i ter is burchelli i HOO K . Basio y p , , ele olotype An r m NORLI ND H 1 939: 787 1 943: 7 K . t . H na 1 86 MELLI SS . 6 S No , ele , ( ) isotype cited f o B ( ,

1 84) may not be extan t .

- Noti s er 1 939: 787 n ta helena e . . . S n . : Os teos er mum s a e y p c T NORL , Bot , N w l 44 200 C om . 6 e s . 4 6 p ,

Typ e as above .

H na . I wa r z new H K This species is endemic to St . ele t s ecogni ed as a species by OO ER fil a in Tr i ter is b u n ar r m . , who pl ced it p eca se of the a gul cypselas , with th ee al ost

- NORLI ND H ank win g like primary angles and thr ee less protruding secon dary angles . s Tr ip ter is in Os teosp er mum and re- n amed the species becau se of homonymy

NORLI ND H 1 939 H e not a in ub nu Tr i ter is r ( , did pl ce the species s ge s p , howeve ,

a ca nthos ermum and mur i a tum . Xenis mi a but grouped it with O . p c in sect of subgenu s Os teosp ermum

a ca nthos ermum Althou gh NORLI ND H mentioned 0 . p as the closest ally of

- l endula m ta e helena e r mb fini 0 . ca ceu O . s a n c , he pointed out its ese lance and af ty to

li a r u a ub nu . r r z in . O oc s n sect g p , u ssigned as to s ge s The latte is ecogni ed as a

r u b r u r and St . H n a n ar b r sepa ate gen s y the p esent a tho , the ele a t xo cle ly elongs the e

on m r a n b n u ra ra r r . o phologic l evide ce such as ha it, i vol c l and flo l cha acte s

Molecu lar evidence fr om I TS and chloroplast sequ ences also demonstrate a close

s a n ta e - helena e Oli oca r us ca lendula ceus relationship between 0 . c and g p

KALLE I l in r . RS O et a . ( , p ep )

P r r rr li r 0 . a ca nthos e mum DC . . b e r O oca us e haps also p ( ) T NORL has to t ansfe ed to g p , I f where it was placed by BOLUS and this issue will be further investigated . this 1 836 b m n n m um r rr Xen is mi a DC . ass ption p oves co ect, ( ) will eco e a sy o y of

i r Ol g oca p us Less .

Tr an sfer of Os teosp erm ump i n n atum to Dimorp hotheca

r m u s te s ermum lr Os teos e mum i nna tum HUN . . O o p p (T B ) T NORL . is ano alo s in p a eady b nnu b and r i r r i n u r and y its a al ha it p ofoundly p nnatipa tite leaves , unise iate vol c e

- b non ra are m r or r b . especially y the white ( yellow) ys , which o e less pu plish elow

NORLI NDH 1 943 : 1 9 1 in n Of own Aca nthotheca DC . . . ( ) placed it a sectio its , ( ) T NORL , b n a n b t r in ased o the genus Aca nthotheca D C . The t xon fits defi itely et e

D im r h the a r n n D nn HUN . HA t . b . i a a o p o c , whe e it o ce elo ged, as p (T B ) RV This species D r Aca nthotheca C . 1 838 will b e furthe studied in our molecu lar phylogeny group . ( )

b m n m fD m 1 4 RE ER et al . n i r h V 75 cf. G UT eco es a sy o y o o p otheca AI LL . ( ;

N ews l . 44 2006 4 8 C omp .

RD N T 2 4 An r n t r W A. R . . NO E S AM 00 . Os eos e mum OOD, B inte esti g new species of p A r - Calendu leae r m rn a Pr n S u A r ( ste aceae ) f o the weste C pe ovi ce , o th f ica ,

r S. A r Bot. 69 2 . r n n k u Ch s a nthemo ide . 7 s . : 5 578 p ovidi g a li to the gen s y f J . C N w l 44 m . 2 o e s . 006 p , 4 9

Fi . 1 N r li ndhi a a tera . g . o p B

X - X - A: fl r 4 . D D i r r a : r : Ra o et : flo et 8 . Lowe le ves , B Uppe C y , sc , x

E - i f - X f i fl r : 1 6 . G : T o di s c floret 1 : r a o d s c o et 8 . 6 . Co oll , laid F p style, X X H I na n 8 . NORDENSTAM : 8 . J : UN N , Cypselas , Cyp sela, longitudi l sectio , L DGRE 1 2 D el . . NORDENSTAM . 3 (S) . B w Ne l . 44 200 C om . s 6 5 0 p ,

New gener a and comb in ations in the Sen eci on eae ofthe G r eater Antilles

BERTI L NO RD ENSTAM D artm t ofPhanero ami c ota n Swed i h M u eu m of tu ral Hi tor ep en g B y, s s Na s y - 1 4 t kh lm w d n P 0 . 50007 S 0 05 S S . Box , oc o , e e

nor d ensta m nrm . se b ertil. @

Ab str act

n n w Generic limits in the tri be Seneci oneae ofthe Gr eater Antilles are revised . Seve e

n r are r b : Antilla nthus . . H err era nthus . . Lundi n ia ge e a desc i ed B NORD , B NORD , E i L Z lekma n a . . Nes a m elos . D . e nis . . and emis ia B . NORD . , B NORD , p B NOR , o B NORD ,

A r w m ina n m . As n n r ar n . 33 ne b are e o B NORD . ltogethe co tio s ade a co seque ce the e

d P enta ca li a . in ub H in n u Of Sene i . an dige o s species c o L CASS C a , ispaniola and

Jamai ca .

I ntr od ucti on

The present study is part of an ongoin g revision of generic limits in the Seneci oneae

I t b n Gr r An r u r ub and H an . r of the eate tilles , pa tic la ly C a isp iola has ee necessa y to n n mb r r m are n uffi k n in disti guish a u e ofnew gene a, so e ofwhich still i s ciently now all

r n r neci n ae in details and deserving a closer study. The t eatme t of the t ibe Se o e the

Gr r A u n ow or r n b n r n r m eate ntilles has p to ece tly ee ve y co se vative , with ost species m ni n - r r r OfSeneci o L . u umb r ra e ai g in an ove expanded concept , pl s a n e oft nsfe s to the

r un r an - fin n P n lia poo ly de stood d ill de ed ge us e ta ca CASS .

The Jamaican Seneci oneae were sorted out in a preliminary way by NORDENSTAM

dontocli ne . . r z new n m n r viz . O who ecogni ed two e de ic ge e a , B NORD and J a cma i a N h r I N N in P . RD T es e b H . R B O . we e accepted y OB SO ROCTOR who however added the genu s P en ta ca li a with three Jamaican species (two described as on ran rr new and e t sfe ed fr om Seneci o) .

I n r 5 A I N 1 63 ri b eneci oneae r r n b r the Flo a de Cuba vol . ( LA 9 ) the t e S was ep ese ted y fou n r Thr ee are m n m Er t n u e e a . a e h i tes o e g of these quite s all , ely c with species (pl s one I n ar t Emili a m t n m Sha era . v ie y) , with two species , and the ono ypic and e de ic f n 2 1 l a r is Se e i r r z in u n S. vu c o Species we e ecogni ed, cl di g the type species , g , which is mm n in u r 20 r nnu u u b a . a widesp ead a al weed, altho gh nco o C The othe species listed un r Sene i are all n n u r b or in m m m b a but de c o i dige o s sh u s v es , ost ofthe ende ic to Cu , m l umbeu S. tr i neurus . On e two of the also occurrin g on H ispan iola (S. p s and ) ofthe w N e s l . 44 2006 Com . p , 5 1

lmi r nci ll wa n ub Sene i s vi z . S. a o o s r rr nu P enta ca li a 1 982 C an c o , , t a sfe ed to the ge s in b P an r r n r b a m n s Old elti a . . y ROCTOR, and othe was ece tly desc i ed the o otypic f B NORD LUND I N (NORDENSTAM LUND I N The remain der are now disposed in four new

n ra and n l r r na S. u a is m in nu in ub an ge e , o ly the occasio l weed v g e ains the ge s the C

flora .

I n H ispaniola (Haiti and the Domin ican R epub lic) three small en demic gen era have n H er i a b r z vi z . d t Ma tt eldi a . and Ekma n a B RH I D I o o . i o us O . ee ecogni ed , URB f URB p pp r b m r or n a b n r n b Twelve species of Sh u s , s all t ees vi es h ve ee t aditio ally een placed in Senecio (LI OGI ER One particu larly deviatin g taxon is described elsewhere as the w u I ur bia R N RD N ne n . O E STAM r r n r ri in gen s g B NO D . ( Fu the ge e a of the t be

H an a are m r or Ere hti te Emili r s a C as s oce ha lum. isp iol the o e less weedy c , and p

S m ten ar ft r R I N N P BORHI D I r ra k o e ye s a e OB SO and ROCTOR, followed thei t c and transferred another eleven Cub an species an d one from the D ominican Republic to H e a r n w P en ta ca li a . lso desc ibed a e species fr om Cuba as P en ta ca li a a cuna e B RHI D I r O P. a lmi ronci llo BORHI DI n r rr , elated to ( No e ofthese t ansfe ed species is

in P enta ca lia i n r n R I N b a N CUATRECASAS . . well placed , which the o d se se of OB SO , a o , is r An nu r rn m i n a Ri a r n r rnm a la ge dean ge s with a no the li it Cost c , whe e the o the ost u of ara mo a n n W n br n ofP enta ca lia r o tpost p veget tio is fou d . ithi the oad co cept the e are two n r u tr ub r b CUATRECASAS R I N N (at least) disti ct g o ps , eated as s gene a y , OB SO and P enta ca lia r but b r r a n r as b J FF ME m f. othe s , ette t e ted as ge e a , y E REY, BRE R and ysel

n u r m r i n or r a Monti ca li a . J FF nu se s st icto co p ises v es epiphytes , whe e s C E REY is a ge s of erect shr ubs with closely set leaves and termi n al s yn flores cences . These two ge nera both have an An dean distribution and are not represented in the Caribbean .

The purpose of the present paper is to b riefly describe the new genera and make the n Afirller r m n k and u n ar new mb . ecess y co inations t eat e t with eys disc ssio s will follow, and a global overview of the Seneci oneae with a generic key will b e published in the

F a mili es a nd Genera o Vas ula r P la n ts NORDENSTAM r . f c ( , in p ess)

New gener a and combin ati ons

An ill an th n n t e . ov. us B . NORD. , g

ru r i n a caulin a s es s ili a breviter a lineari a F tices e ect vel s cande tes . Foli vel petiolat vel

- a a au a lura lanceolata ad ob longo ovata supra glabra s ubtus tomentosa . C pitul p c vel p

— - r r a I n r r corymb osa cymos o pani cu lata discoidea vel inte dum adi ta . voluc i b acteae

feminei u . u uni s ri Rece taculurn alveolaturn brum . u r e atae . p gla Flosc li adii vel n lli Flosc li An therae b s a ittatae br herrna hr oditi r r in ra lobis ovati s . disci p ; co olla es ife , asi g vel evi

r S ram n r n r s ti mati ci s confluentibu s obtus i te caudatae . tyli i li ea es i tus a eis g apice pilis

- — - Pa r elli ti co oblon ae 8 1 0 cos tatae ub n . b evibu s spars i s . Cypselae p g p esce tes ppi n seta e numeros ae basi conn atae alb ae persiste tes . N w l 44 2 6 C om . 00 2 e s . 5 p ,

Typ us : An ti lla nthus ekma n i i (ALAI N ) B . NORD .

— - Lea es au n Er or n n r b 2 m b r n m m n . ect sca de t sh u s ; a ches ostly white to e tose v c li e ,

or h rt in r or an a b n - 3— 1 3 cm n 1 — 4 sessile s o ly petiolate , (l ea ) l ceol te to O lo g ovate , lo g , cm n r or m m a nu - n br u or a wide , e ti e so eti es sh llowly si ate de tate , gla o s ( at le st

br n b n or r - m n b n m r n u u gla esce t) a ove , de sely white g ey to e tose e eath; a gi s s ally

a i t ula r i n rm n r mb or m - an u a revolute . C p few to seve al te i al co y ose cy ose p ic l te s n flores cences or r r r a - flowered or r r W — r m y , discoid a ely adi te , yellow a ely hite c ea

I n l u r e n r narr m anu ur n b ra . a colo ed; pedu cles cteate vo c cyli d ical to owly c p late ,

— — — a u n u r br un r 5 1 3 b n n b u . c lyc late; i vol c al acts ise iate , O lo g la ceolate , o tuse ac te

R r a D - r t — 1 r a u ar narr e e ta le b . is o e s 5 3 b b c p c gla ous , alveol te c fl ; co oll tu l elow, owly am anu a or nn - b b n rr midlined c p l te fu el shaped a ove; lo es deltoid to a owly ovate , ,

ll n R n n r n i n r n ft a d . apically s ubcucu ate a d papillate . esi canals o e p ese t co olla style An r b a a a a or r au a a a a n a bt the s s lly s gitt te sho tly c d te; pic l ppe dage ov te , o use; n l r S endothecial tissue radial with elongated cells; filame t collar b a uste forrn . ty le branches apically obtuse with rather few and short lateral sweeping- hairs; in side with

- — - la n 8 1 r rt ir u . ma ar a n nu u . s e s b 0 bb stig tic e s co ti o s Cyp elliptic o lo g , i ed, sho ly h s te P a us bri num r u n r u u r r r b nn pp stles e o s , Sle de , s ally Sho te than the co olla, asally co ate ,

r n white , pe siste t .

r nu r b ub 1 7 r r n z . This is the la gest ge s of the t i e in C a, with species p esently ecog i ed

1 An i n h n B RHI D I m n t ll a t us acu a e O . . o . ov. . ( ) B NORD , c b

m: P en ta li un B RHI D1 A t H un 1 2 ca a a c a e O c a . . 37 : 88 99 Basiony , Bot g i A UN 1 34 1 2 SV ub Pr . Vrllas rr Es cambra in s axos s : J . S Type C A , C a, ov Las , ie a de y, a ar m n P P r — m HA r 900 930 V. 1 939 C . c lc eis o tis ico ot e illo, , ( holo

2 An mi r n il M m . till an th us al MA o c lo G . . o b . . ( ZA O EZ) B NORD , c nov m n : Sene i a lmi ronci llo MA G M An H . . 1 9: 277 Basio y c o ZA O EZ, ales ist Nat

S n . : P enta ca lia a lmi r nc ll MA M R P Ar n Ar r o i o G G. . J . b . y ( ZA O EZ) ROCTOR, old o

M . ur r s n . : a li r E n n D r REEN C a a dis l GRI S B . o C . Sene i b i tt n i i G F the y c co o , ; c o o , Field

Mu . Nat . H 1 64 . Ser. 2 s . 3 3 ist , Bot

WRI H 2 u : . 870 b . Type C G T , C a

An tillan th u n 3 . s az ule s is A I N . . om . . ( LA ) B NORD , c b nov

n m : en i n S e az ule s is A I N n r. a . M H N at . D e O us . . Basio y c o LA , Co t c s ist Colegio La ” a n 1 8 : 1 0 S lle o .

O I arr : A AI N M . U I 7352 ub Or n S rr A a Type L L PEZ F G E RAS , C a , ie te , ie a zul , B aco , L V ” ” S . S U0 ( holo , , 44 2006 C m N w l . o . e s p , 5 3

n 4. An till a th us bis eri atus A I N . . m n ov. ( LA ) B NORD , co b .

n m Sene i bis er i a tus A I N n r. a . Mu H N a : t . O s . . a D e La B sio y c o LA , Co t c s ist Colegio ” 1 8 : 1 1 Salle no .

: A I N 3429 u a n rr M 2 - L VI I . 1 53 . b Or S oa 5 26 . 9 S Type LA , C , ie te, ie a de , ( holo , NY

An till an th us ca ri n a tus R EENM . MB V 5 . G . . . NO . ( ) B NORD , CO

r M i 4 2 a n m : Sene i a i na tus REENM . H G us . . . 1 6 . Ser. 3 3 B sio y c o c , Field st Nat , Bot (1 91 2)

S n . : P enta ca li a a r i na ta GREENM . BORHI D I A Hun 1 2 a . 37 : 88 99 y c ( ) , ct Bot g .

A H F 4 . S 079 ub Or n S rra Moa am an n 9 0 m : J . S 0 Type A ER , C a, ie te, ie de , C p Be ito , ,

An till an th u u be RE ENM m n v . s o . o . 6 c ns is (G . ) B . NORD. , c b

a n m n i n i REENM M H i 1 4 r 24 : Se e cube s s G . us . . . 6 . Se . 3 B sio y c o , Field st Nat Bot (1 912)

P n t u REENM B RHI D I H un 7 1 2 n li n A . : S . : e a ca a c be s is G . O . 3 88 99 y ( ) , cta Bot g

4 i A. H F 084 ub a n rr Moa m San n 900 m : J . S Or S Type A ER , C , ie te, e a de , Ca p Be ito , ,

(F, NY) .

m n 7 . An ti ll an th us ekm an u A AI N . . o . ov. ( L ) B NORD , c b

M H . D La n O . us . . e a n m : Sene i ekma n i i A I N r. B sio y c o LA , Co t cas ist Nat Colegio ” 1 8 : 1 1 Salle no . : A I N A UN O I U I R 5606 uba Or n S rra ri a en Type LA , C A L PEZ F G E AS , C , ie te , ie de C st l , ” ” - h 2 L NY V O . c arras cos 7 . I V. 1 956 S . S U , ( holo , , , iso)

m . nov. 8 . An ti lla n th us eri oca h us GREENM . . . o rp ( ) B NORD , c b

M rr 1 3 : 257 a n m : Sene i er i oca r hus GREEN . B sio y c o p , To eya n 1 2 B RH I D I A a . H u . 37 : 88 99 P n r a r ha REENM . O S n . : e ta ca lia e i oc G y p ( ) , ct Bot g

r n r m m Toa am La ar a 22 A. H F 4 1 49 ub O S nt es : J . S y yp A ER , C a, ie te , f o Ca p to C p B g , A SH F 8257 m G r S S rr MO ra m n J . . (F, NY, f g e t); A ER , Ca p La lo ia, of ie a de

- M 24 M ra m n . XI I . 1 91 0 O oa 30 . , (F, NY, f g e t) m w 44 Ne s l . 20 C o . 06 5 4 p ,

m Antill a nth us le u cole is GREENM . . . . nov. 9. p ( ) B NORD , co b

4 Mus . H . . 1 6 . Ser. 324 m : Sen e i le ucole is GREENM . Basiony c o p , Field ist Nat Bot ( 1 912)

BORHI D I A . H n . 37 : 89 1 992 P n t li a l c le is REENM . S n . : e a ca eu o G y p ( ) , cta Bot u g

A H F 4 1 46 ub m m ar 400- 450 m : J . . S Ori Type A ER , C a, ente , Ca p Toa to Ca p La B ga, ,

- 1 1 0 22 26. II . 9

m . nov h m A I N . o 1 0 . Antilla nt us oaens is . . ( LA ) B NORD , c b

M L 1 n O . us . H . . a S 1 8 : a n m : Sene i moa ens is A I N ri b . B sio y c o LA , Co t cas ist Nat alle l

P enta ca li a moa ens is A I N BORHI D I A . Hu . 37 : 89 1 992 S n . : y ( LA ) , cta Bot ng 2 2 M n n 2 1 4 L M . VI I 9 7 S . : ON M N 3 03 S rr oa J 0 . Type LE CLE E TE , ie a de , i a oh son, ( holo ,

A I N . . om . nov. 1 1 Anti ll a n th us molden kei GRE ENM . ex . ( LA ) B NORD , c b

H A I N n ri b . O . Mus . . . s n m: Sene i moldenkei GREENM . Ba io y c o LA , Co t cas ist Nat “ ” l H n 1 1 2 De S ban o . 8 : Colegi o La a le , a a, 2 1 4 HA rr 9 5 C . 1 : A UN 1 775 b Or Gr Moa 1 6. . Type C A , Cu a, iente, an Tie a, , IV ( holo )

Anti h h m 1 2. lla nt us ac le is G REENM . . . b . nov. p y p ( ) B NORD , co

2 Mus . H . . 1 64 . Ser. 3 5 a n m : Seneci o a ch le is GREENM . B sio y p y p , Field ist Nat , Bot (1 912)

- A. H F 4 u n Ri ani am 22 26 . H. 1 1 0 : J . S 008 b a Or o m 9 Type A ER , C , ie te, Ya guey to C p Toa, F NY ( , )

1 An illa nth h n 3 . t us ac odus GREENM . . . om . ov. p yp ( ) B NORD , c b

n m . Pub . Mus . . H . i a : Sene i a ch odus GREENM . B sio y c o p yp , l Field Nat ist Ch cago , Bot 2: 325 P 1 . : 2 n n BORHI D I A a . H u . 37 : 89 99 S e ta ca li a a ch oda GREENM . y p yp ( ) , ct Bot ng

A H F 1 r 24 : J . . S 8 86 b Ori n S S rr Moa m G Type A ER , Cu a, e te , of ie a de , Ca p La lo ia

30 . XI I . 1 9 1 0 fia t NY men . ( holotype, F g )

1 4 An n h AL I N m n . ti lla t us s a u etii ov. g ( A ) B . NORD . , co b .

n 1 2 m: Sene i s a u eti i A I N n ri . M H . Nat . S 1 8 : b . O us . Basio y c o g LA , Co t cas ist La alle

m N w l 44 2006 C o . e s . 5 6 p ,

EB . om . nov . Leon is tri n ca ra (GRI S ) B . NORD , c b

4 Mem . Am. A . . S. 8 : 5 1 a n m: Sene i tr i neurus GRI E . B sio y c o SE , cad N H un 89 1 992 P enta ca li a tr ineur a GRI E BORHI D 1 Acta . n . : Sy ( SE ) , Bot g prob . I H 327 ub M n r Guan anam S " : . W Type C R G T , C a, o teve de , t o ( isotype )

“ b 50 : i le n is Le nis RI T N WI L N u . rr . u r r s n . : Sene Fu the y c o o ( o B T O SO , B ll To ey Bot Cl

m An . 7 : 43 1 S b . Sene i d min ens is . c o o g URB , y till

4 D m n r n anz a . M 33 7 Sto . S n t es : H . . UR KH I y yp v T C E , o i go p ope Co st

: . 1 . I llus tr. Fig

S an n r b br u in m ar but un br and c de t sh u , gla o s ost p ts , yo g anches , peduncles petioles ru u - r u and un m m n an u ar- a a b fo s aspe ulo s , ped cles so eti es fi ely gl d l p pill te elow the rn n r a ar r n m m ri a . Lea ves o capitulu ; ste s st te alte ate sho t xill y flowe i g shoots , ovate to

cm n cm n r or ri ba or r 1 elliptic, lo g , wide , e ti e shallowly t lo te g ossly

4 - n a rba u r - n r r m n ar b a a —b a b s de tate , petiol te , he ceo s , th ee e ved f o e the se; le f l de a ally

— n r n a or n n r 1 3 cm n . cu eate to t u c te occasio ally hastate; petiole Sle de , lo g P edun l es S n o res cences r mb rm a in a ra r . y fl co y ose , te in t g the l te l axilla y leafy shoots c n r 1 — 4 cm n i r bra ru u ru u r ri m Sle de , lo g , with a few l nea cts , aspe lo s with fo s sho t t cho es

- Ca itula m an a r am u and m im a a a an u r . so et es apic lly p pill te gl d la p ca p ul te , hete og o s ,

- w I n lu ra l bra ts un r a 8 — 1 2 r or di s ciforrn flo ered . adiate , yellow vo c c ise i te , ( narr b n an — 7 mm n 1 — 2 mm d n 3— 5 - n owly o lo g to l ceolate, lo g , wide , isti ctly vei ed i b k h n in rr r n r u n u and ub ru u w th lac is (Ofte te uptedly) esi ife o s vei s , apically ac te p e lo s ,

br 2— 5 r abr u or rar m ub r u r u u . a u u othe wise gl o s ely so ewhat p e ulo s th o gho t C lyc l s acts ,

n ra r r an ri but r n in z 5— 8 mm n — 1 mm ge e lly sho te th the phylla es va yi g si e, lo g, wide ,

n r and umi ar ub n or r ru u ri m or li ea ac nate , sp sely p esce t with whitish sho t fo s t cho es in n R e ta le r ub abr u m m m an r . e h s gl o s , so eti es with a edi es vei c p c Slig tly convex , sho tly

R - l rets ma 1 — 5 firnbrilliferous but not n a or i a . a , de ticul te dist nctly lveolate y f o fe le ,

abr u ub in r a 2— 5 mm n ami b n or narr - b n gl o s ; t e cyl d ic l , lo g; l na O lo g owly elliptic o lo g ,

— mm n - - r n in ar abr u b ara 1 5 3 5 n . S b lo g , vei ed tyle a ches l e , gl o s , o tuse , with sep ted

- — r ba a bu r i ma n D is l rets 1 6 24 rma hr . stig tic li es . c f o , he p oditic Co olla s lly tu la , widen ng b — 6 mm br u r 5 - b b — o mm a ove , long , gla o s , sho tly lo ed; lo es deltoid vate ,

n a m an ub a a ark r n an s ub cucullate and i : a lo g , with edi s pic l d esi c al , apically papill te A r n n n a b a a but au a u . mm . o tside nthe s lo g i cl appe d ge , s lly sagittate, ec d te; ‘ ’ apical appen dage narrowly ovate- lan ceolate; filament collar b alusterform (with n r a t ark r n n br n r a b . S e l ged cells sally) yle with two d esi ca als , anches li ea with continuou s stigmatic area inside; base in durated; stylopodium distin ct; style

- and r r b ran n ar b n ca . 1 mm n b ches li e o lo g , lo g , apically o tuse with few ve y sho t

- la s - b n br u m m a ra n a r or ub ab r u . s e l te l sweepi g h i s s gl o s Cyp elliptic o lo g , gla o s (so eti es w N e s l . 44 2 0 C om . 0 6 p , 5 7

F i 1 Leon i s tri n ca ra G I E . . g. . ( R SE ) B NORD

A x - X - X - D florets 6. D r a ofdi fl P r n 1 R fl r 6 . sc ore a . a o et t o tio ofplant, B y , C isc , Co oll , l id

- X — out E ofdis c floret 1 2 . 6 . S m n 1 2 . S br G 6. A F t . , x ta e s , x F yle anches , x Cypselas , ( L

MI H 4 in . D el . . NORDENSTAM . 3238 in G EKM N 1 504 S . B S T al . S; A ) B m w 44 Ne s l . 2006 Co . 5 8 p ,

n - ur r r P a us a a 1 0 r b rw b . few pical setae) , with disti ct st aw colo ed i s , othe ise light own pp m r u r ri a m nu b rb r n nn a br u ba . istles n e o s , plu ise te , i tely a ellate , white, pe siste t, sally co te

Leonis tri neura is a vin e which occur s in Cuba and H ispani ola (both H aiti and n ri nam m r m Dominican Republic) . The ge e c e is si ply adopted f o the epithet of the

Le n is Anti lla n thu an d syn onym Seneci o leon is BRI TTON WI LSON . o agrees with s n m - The some other genera in havi g continuous stig atic areas on disc floret s tyles . nfl n ar n r r n r n t m The an s y ores ce ces are axill y alo g the athe sle de scande t s e s . pl t is

r n r n r u ofien b ki r u in r - floret st o gly esi ife o s with distinct, lac sh esin d cts phylla ies , disc corolla and style .

When URB AN described Seneci o domi ngens is he remarked that it has no related “ ni f mi r n n r m Antillean taxa ( Nulli alii ex antilla s a f s ) . The e is nothi g to disti guish it f o

r nd ran a N r k i r a S. tr i neura a , howeve , it is st ge th t URBA ove loo ed th s widesp e d species .

u r ab r not r r n i n The species is q ite va i le , which is pe haps su p isi g View of its wide - r distr ibution on the two largest islan ds of the Greater Antilles . The leaf Shape va ies n r m r n from e ti e and ovate to ore or less distinctly trilobate o grossly de tate . The br m r n i n n r m r rt m calyculus acts Show uch va iatio le gth , f o ve y Sho to al ost the length

m r and z fra - floret r b r m or n u ra br . nu b O s ofthe i vol c l acts The e si e y is also va ia le , f o one two or r r and m t are b r b as d rm to at least five a ely eight, so e capi ula est desc i ed iscifo au r mi ra - fl r t bec se ofthe educed la na of the y o e s .

Leon is tr i neura is rather similar in vegetative morphology to Ma ttfeldi a tr ip li ner vi a

m n nu n m H a an n r b r U RB . . a , a o otypic ge s e de ic to iti This is lso a sc de t sh u with alte nate - Ma t eldi a r m are n r r . f petiolate leaves , which ovate, e ti e and th ee veined d is distinct o Le on is by the b ilabiate ray- florets and the narrow and few- flowered capitula with onl y

five phyllaries .

m Nes a elos . . n n v p B NORD , ge . o .

ru s candent r m n i l r nt a altem a a elli tico F tices es amis to e tos s g ab es ce es . Foli petiol ta p ovata ad obovata integra margin e dentata vel denticu lata coriacea costata basi rotundata vel obtusa interdum cuneata apice obtusa mucron ata supra glabres centia n a r h ter ama r a s n flores centii s itida s ubtus dense tomentosa . Capitul pa va e og adiat in y axillaribus r m i i I u r m cal culatum n r bracteae co y bose d s pos ta . nvol c u y ; i voluc i s ubuni s eri atae vel fere b i s eri atae linear i - lanceolatae vel anguste oblongae glabrae vel

- los culi ra m r fl r m a . los culi pubescentes . F dii pauci la ina b evi ava vel c e ea lba F disci herma hrodita r s u eme am u a uin uelob ata lobi s ovati s p ; co olla p c pan lat q q , vel l n l i m a An th ra a a ittatae . r ceo at s apice papillati s . e e b si obtus ae vel s g Styli a i apice

subtruncati eferrentibus numeros i s r s ti m ati ci s s e arati s . pilis , a eis g p Cypselae stri atae- costatae api cem versu s Sparse s etos ae aliter glabrae; carpopodium C N ws l 44 2006 om . e . p , 5 9

m P i di stin ctu . cO os ae b a b ar ellata nn appi setae p al e b e basi co atae .

Lianas or scandent shrub lets with tomentose glabrescent stems; trichomes soft and

bran n L - — flexuous . ea ves r a a b 3 7 cm , chi g alte nate , petiol te , elliptic ov te to o ovate , 2— 4 cm n r n n a m r rrri dri bb ed long, wide , e ti e with de tate to de ticul te a gins , and with late ral in r u b r un — ob or b mu r n ve s , co iaceo s , ase o ded tuse cuneate; apex o tuse c o ate; u r n a ar or m abr n an d a ppe side i iti lly aneose loosely to entose , gl esce t glossy with r u n n rn r br n or r n m n etic late ve atio patte , lowe side ow ish g eyish de sely to e tose; petiole

- Ca i tula in rmina or a r r 3 1 0 mm . r r mb m r m u Sho t, p te l axilla y l te al co y s , s all , hete oga o s ,

R e ta l flat n r . ec e r u um n a . I n l u r e adiate p c , sho tly de tic late with ac i ate sc les vo c calyculate with few small subulate calycu lus bracts; i nvolucral bracts subuni s eri ate to

m b r a 5 — 8 n ar - an a n rr b n br u or m n al ost ise i te , , li e l ceol te to a owly o lo g , gla o s to e tose;

r r m m r m n R - or ets 2 min r n r b r u o b u ar . a i ne acts with sca io s e ano s gi s y fl few ( la a sho t,

— - D is - re t 5 mm n b n r am or a a 2 3 fid . l s to lo g , o lo g, yellow, c e y white , pic lly c f o

rm r 5— 1 3 r 4 — 5 mm n t b ar b m a b he aph odite , ; co olla lo g , u ul elow, ca panul te a ove ,

- n - An r 5 b b u r or a . lo ed; lo es ovate to la ceolate , apically callo s inc assate p pillate the s

g b b or a au a a a a n a narr ovate lanceolate . asally o tuse s gittate , ec d te; pic l ppe d ge owly Sty le bran ches apically subtruncate with numerous short sweeping- hair s; stigmatic

la ri a or r bb in u r r br u r ar . s e s a eas sep ated Cyp st te i ed , ciliate ppe half, othe wise gla o s;

P a us br num r u — c u r r a b arb a car o odium . p p distinct pp istles e o s opio s , plu ise i te , ell te , H b nn r n . r . . white , asally co ate , pe siste t Th ee spp , ispaniola

S m r r nu are n n ab n ir o e cha acte istics of the ge s the sca de t h it, the petiolate e t e leaves n or n u mar n n r u u r urfa an d m n with de tate de tic late gi s , the shi y etic late ppe s ce to e tose r ma and r - r - flowered u r r t lowe Side , the s ll sho t ayed few capitula , pl s cha acte s ofs yles and an thers .

m n v Nes am elos l ucens P I R . in LAM . . . o . o . 1 . p ( O ) B NORD , c b

2 : 34 1 1 8 1 . in LAM . En . Su . n m : C n z a l ucens P I . Basio y o y O R , cycl ppl ( l ) 4 m An . 3 : 1 3 P I N S b . Sene i lu ens . S n . : y c o c ( O R ) URBA , y till

H a . : N ECTOUX s . n . P Type , iti ( ) Sene i n z ides S . 3 : 508 mi n ens is SFR . r r s n . : C n z a d Fu the y o y o g , yst c o co y o 4 1 2 Prodr. 6:

m nov. 2 Nes am elos h ottean a U . EKM N . . . . p ( RB A ) B NORD , co b

23A 1 1 : 93 EKM N Arki v . . a n m : Sene i hottea nus . B sio y c o URB A , f Bot ( ) E EKM N H 7430 H an H i : Ma H rn r u Type : . L . A , isp iola, ait ssif de la otte , weste g o p ,

— I XI . 1 926 Torb ec M rn rm n m n r in k 1 400 1 500 m 27 . , o e Fo o , li esto e idge , thic ets , ,

S . 1 ( holo ,

I ll 2 : . . ustr. Fig m N ] 44 2006 Co . e . 6 0 p ws ,

F i . 2. Ne a m l h tt an a E KM N . . g s p e os o e (URB . A ) B NORD

- X - X x m n 1 2 . R fl r 6 . D floret 6 . D S A P r n . a o ets o tio of l B y , C isc , ta e s ,

- x E r n fl r 2 1 2 EKM N H7430 i n S . S b o et 1 . . tyle a ches of disc , F Cypsela , x ( A , typus) D l N RD EN O STAM . e . B . C m N w 44 o . e s l . 2006 p , 6 1

m a l ai n . I M NE ALM . . F i . 3 . N a l u g es p e os (J J E Z ) B NORD.

P r n X - x - X I n u r 6 . Ra floret 6 . D D 1 . fl t . ore 6 . o tio of B vol c e , C y , isc ,

E r a — X - m n 1 2 . G S bran i ofdi s c floret a d 6 . S a ofd s c floret Co oll , l i out, x F t e s , tyle ches x 1 2 . LI D el . . NORDENSTAM . OGI ER 22276 in JBSD i sot us . ( , yp ) B m w Ne l . 44 2 0 Co . s 0 6 6 2 p ,

ALM. c m n 3 . Nes a m el os alai n i i J . J I MEN . . b . ov. p ( EZ ) B NORD , o

r a i n i i I MEN AL l i on n A a m: H e odoti a a l J . J M Co ecc f. . Sci . Re . D om . Basiony EZ , Co c d p 2 ( l 1 5

MEN ALM . BORHI D I A a km us a i n i . I E a ni o a a l i J J . H . 37 : 1 1 1 S n . : y p pp ( EZ ) , ct Bot ung

I ER 222 r V a n anz a l n n : LI OG 76 P . e H rb . Type , ov de La eg , Co st , Co ve to , ( e D JI MENE . JBS Z holo , iso ll : . 3 . I ustr. Fig

H rr ran th us . en . nov. e e B . NORD , g

m m h m in i n - n r rb r r 3 a r o ot ara eos o m labres centes . F utex vel a o pa va ad alt , a i to e tosi g Folia altem a petiolata oblanceolata vel lan ceolata ad anguste elliptico- oblonga usque 20 cm longa et 4 cm lata integerrima acuta supra glabra costata s ubtus dense

n — m n l m r — mb m u 1 3 c . ura u m c r to e tosa, petiol s lo gus Capitula p vel n e osa cy osa o y osa

a I n u r m an m an laturn al latum n r r discoide . vol c u guste ca p u c ycu ; i voluc i b acteae bi s eri atae lerum ue 8 b n — lan ceolatae tomentos a o tus e p q o lo go e b a . Flosculi henn aphr oditi ; antherae caudatae; styli rami apice truncati vel s ubobtus i tantum m nu ill ti i m i i nfl n i l r li r r ill i a a r st at c s co ue t bus et ate a te a at s . i te p p a eis g extensis , do so p p

- ir r P numeros ae b Cypselae 1 0 costatae h s utae; ca popodium di s tinctum . appi setae al ae b ar ell atae r n a in ann l nn ta b pe siste tes b si u o co a e .

r v Type : Seneci o i a lis GREENM .

Monotypic :

H rr r h e e ant us ri val is GREENM . . . com . nov. ( ) B NORD , b

n m n Se e i r iva lis REENM . Mus . H . 1 64 . er. 326 : G . S Basio y c o , Field ist Nat , Bot A H F 4 4 r r 4 - m : J . . S 3 5 b O S rr r W 50 550 Type A ER , Cu a, iente , ie a de Nipe , nea oodfo d, ,

I ll : . 4 . ustr. Fig

Er r b or ma — m m r n r 3 . S r r b ect Sh u s ll t ee high te s te ete, with la ge pith, gla esce t; young

r - Le a rn or b r m n . a ves a a b anches a achnoid to e tose lte te , petiol te , o lanceolate n rr - b ir 7— 20 cm n — 4 cm lanceolate to a owly elliptic o long, ent e , lo g , wide , acute; u r br u midveined and a n r a r n r ppe side gla o s , with f i te l te al vei s ; lowe side densely

r - m n r mu - br n r r m m r b app essed to e tose with light g ey ch a ched cu led t icho es , id i prominent and glabrous; margins somewhat revolute; leaf- blade tapering in to a

— l rmin 1 3 cm n . a i tu a r n m or r mb petiole lo g C p te al , fai ly de sely cy ose co y ose , discoid, u I nv l u r e n dr n rr am anu r m n calyc late . o c cyli ical to a owly c p late , sho tly to e tose; n u r r - — — 3 mm b 8 b r a b n an 8 1 0 mm n 1 . 5 i vol c al acts , ise i te , o lo g l ceolate , lo g , ( wide ,

r m n b r a nn r m r n bt bra 3— 5 sho tly to e tose , su co i ceous with thi e a gi s , o use; calyculus cts ,

w Ne s l . 44 2 C om . 006 6 4 p ,

— m n - n an a 1 3 mm n r r . Re e ta le l ceol te , lo g , app essed , sho tly to e tose la ate c p c flat to D i - or 2 — 2 ub n a a and m nu fimbri llate not n u a . s ets 0 5 s co vex , lveol te i tely ( de tic l te) c fl , i r mm n r rm r r b b . 8 b b rr he aph od te; co olla p o a ly yellow, c lo g , tu ula elow, na owly

m anu a . mm n b n rr mm n b b 0 3 . ca p l te a ove , tu e lo g , lo es a owly ovate , c lo g ,

in m na in a u llat midve ed ar s bcucu e u . and with gi l ve s , acute, apic lly with papillate o tside

An r 3 mm n r i u a n a narr the s lo g , sho tly to dist nctly ca d te , apical appe d ge owly 2 mm n n n a b . S br n . arr b r ovate , o tuse tyle a ches c lo g , owly o lo g, apic lly t uncate to

b r r a k n n - r n slightly o tuse , with ve y sho t p pillae and lac i g sweepi g hai s , i side

n inu u m m ar n in r u co t o sly stig atic with stig atic eas exte d g late ally to the o tside , non

la narr n r m r b ma a u a a . s e s b stig atic do sal side eset with s ll c te p pill e Cyp owly o lo g , te ete ,

1 0- r bb n ir u br r — 6 mm n — 1 mm i ed , de sely h s te with light own hai s , lo g, wide;

r i n P a us br num r r r e r ca o od um . p p disti ct pp istles e ous , plu ise iat , st aight, glossy white ,

— r n 6 7 mm n b a a nn . pe siste t , lo g , s lly co ate

I nam new n for P P HE A of CI TMA H ab an a for m n n e this ge us EDRO ABLO RRER , , his e i e t knowledge and interest in Cuban Compositae .

n n w nu r u n m en r v i REENM The si gle species ofthis e ge s was p evio sly a ed S ecio i a l s G .

I t is a large shr ub or small tree with big leaves an d some un usual floral features . The u r br are u u two r u invol c al acts s ally eight and disposed in se ies , a condition not uniq e in u in m n r m r r ri a r b ri are o . the t i e , altho gh ost ge e a the phylla es o e less unise te The styles of the disc- florets are special by having a continuou s stigmatic area in side the bran an d a n r a b style ch which lso exte ds to the outside , which is do s lly eset with

4 E . ur rm r br are m n . t poi ted papillae (Fig ) F the o e , the apex ofthe s yle anches so ewhat n k n - r bt a d r r . o use lac i g sweeping hai s , only p ovided with few and Sho t papillae The

n m n m on a a n m - bra ur r de se to e tu ad xial le f sides co sists of uch nched c led hai s , resembling those ofsome species ofthe Andean genu s Nordens ta mi a LUND I N (2006) a m lthough they are ore tightly packed .

Lun di n i a n n . e . ov. B NORD. , g

Frutex erectus vel arbor parva ad 5 m alta glabra sed partibu sj uvenilis albo- tomentos i s

labres centi u a lt m a a a n — an a n r g b s . Foli a e petiolata lanceol t ad oblo go l ceol ta i teg a vel m r i u min I n m lura mb r a . ru a g ne pl s us ve denticu lata . Capitula p cory osa adiat voluc campanulatum calyculatum; involucri bracteae uni s eri atae plerumque 8 oblongae mar n h lin m u a R a u m c nvexum fimbrillatum u . gi e y ae . ecept c lu o vel dentic latu Flosc li r 5— 8 fenri nei a am n n 8— 20 henn a hroditi r flava adii fl vi l i a oblo ga . Flosculi disci p ; co olla

b bu a l i i ll ll i th r audatae . ob s ovat s s ub cucu ati s a i at s . An e ae c asi tu los , anguste apice p p S ram oblon i r s ti mati ci s di s creti s a trun cato r b tyli i g a eis g , pice vel levite o tuso pilis lateralibus brevi u i nter um n n m elli tico- oblon ae b s d pe icillo ce trali o ato . Cypselae p g breviter vill s a 1 - - P a numeros ae barbellatae a b a o e 0 cos tatae vel nervatae . appi set e l e

r a r vi r co oll b e o es . C m N w l 44 o . e s . 2006 p , 6 5

F i . 5 . Lu n di n i a l umbea I R . . g p (G R SE ) B . NORD

A r br X Ra D - floret D r ofdi s c- flor X 1 2 et . Flowe ing anch, l B y C isc , Co olla E am n 2 G x 1 2 ORDEN TAM S S bran ofdi s c 1 . N S t e s , F tyle ches Cypsela , ( UN I N 3 1 7 in . D l . NORD ENSTAM . L D S) e . B m w 4 Ne s l . 4 2006 Co . 6 6 p ,

E 1 I Mem . A . . S. 8 : 5 5 : Senecio l umbeus G . Type p R SE , cad N

The genus is named for my fri end and colleagu e ROGER LUNDI N ( 1 955 who was m an n n r ri in E u r ar also a good co p io o seve al field t ps c ado and the C ibbean .

Monotypic :

m n v Lun di ni a l umbea G I S E . . c b . o . p ( R E ) B NORD , o

Mem . Am r. A . . . 2 8 : 5 1 5 a n m : Sene i l umbeus GRI SEB . S B sio y c o p , e cad N , 2 a r n V r ad r ri l m : . W I H 3 8 b M vu oru . Type C R G T , Cu , p ope o te e de ipas

: . 5 . I llus tr. Fig

Er r b or m n r r 2— 5 m a br u or m m ect sh u s all and sle de t ee , t ll , gla o s so ewhat to entose

- un r . Lea ves altemate or b especially in yo g pa ts , petiolate, lanceolate o long lanceolate ,

5— 1 5 cm n 1 — 4 cm n ir or i n u a mar i s rb lo g, wide, e t e with de tic l te g n , he aceous , apically

i tula ra mb r u or m a umina b un . Ca r ac te so ewh t ac te, asally c eate p seve l , co y ose, adiate , - w r flo ered . I nv l u r e m u u i nv lu a l bra ts 5 yellow o c ca pan late , calyc late; o c c (

i r rr b n m m r n . Re e ta le n un se iate , na owly o lo g, acute , so e with hyaline a gi s c p c co vex ,

R - r t — a n — narr fimbrillate or denticulate . ay flo e s 5 8 ; lamin oblo g owly obovate; style

- — D i r ets 20 rm r . b or m m n ft r . s o 8 o tuse so eti es poi ted with a tu of hai s c fl , he aph odite r b rr or u m s ub cucullate Co olla lo es na owly ovate , with witho t a faint idline, apically ,

An r u a r u m a . t apex and o tside so ewh t papillate the s ca date; tails at le st colla leng h ,

n r S br r- b n m m u r in t o r. so eti es eq alli g colla leng h longe tyle anches linea o lo g, apically r or m bt rt r m m m an r ft t uncate so ewhat o use with sho late al pili , so eti es with a edi hai tu ;

r a r r n u m a el stigmatic a e s sepa ated; style b a ch o tside so ewhat p pillate . Cyp s as elliptic n - b r u . 1 0 n . P a us br m r u r o lo g , sho tly villo s , ca vei ed pp istles nu e o s , slende , white ,

b arb rt r r . ellate , sho e than the co olla

Vernacular name (Cuba) : Retama arbol (ALAI N

I r - hrub r r This species is foun d on Cuba and on H ispaniola . t is a la ge leaved s o slende n 2 m m r r r are tree betwee and 5 meters tall . So e of the ost cha acte istic featu es the

r ma r a - floret a a r sepa ated stig tic a e s of the disc style , the c udate nthe s , and the

n de ticulate receptacle .

A r u one ar ab rri on b an . ppa ently j st v i le species , occu ng oth isl ds

E kman i a le . . en . nov. B NORD , g

rutices u r lerum u r n r a altema i r mar n F vel s ff utices p q e esi ife i . Foli petiolata nteg a vel gi e n - bu s ubtu m n r r l r nti a Ca itula lura de tato lo lata s to e tosa sup a glab a vel g ab es ce . p p vel numerosa i n s yn flores centi am terrrrin alem di spos ita radiata vel interdum discoidea . I n u r r Fl s culi r au f minei flavi . vol c i b acteae subun i s eri atae . o adii p ci e Flosculi disci n m r herma hr di i n h r l u a ittatae S ram u e osi p o t ; a t e ae ecaudatae sed p erumq e s g . tyli i 44 2006 C N ews l . om . p , 6 7

ma n i a b u h u URB . . . i 6. E lek F g. c ( ) B NORD

TAM D l . . NORDENS . r n bran EKM N H 1 395 in S . e Flowe i g ch , ( A ) B m w 4 . 4 2006 C o Ne s l . 6 8 p ,

i v i r ti m i i r i l truncati s ubobtus i bre b us s at c s s e a at s . abrae vel pil s , a eis g p Cypselae g vel

— - ll P a numer s ae b arb e atae ba . hirs utae 8 1 O cos tatae . appi set e o al e

r Lea ve rna S r b or ub r b m m tr n n r u . s a h u s s sh u s , so eti es s o gly esi ife o s alte te , petiol te n r or n - b u mar n ua m n b b r u or e ti e with de tate lo late gi s , us lly to e tose elow, gla o s

r n a a i tula r num r u in r or rmin r m or glab esce t bove . C p seve al to e o s late al te al co y bose

n r m m - fl w r R le m s n flores ce ces r o o e ed . e e ta cy ose y , adiate so eti es discoid, yellow c p c I n lu r e m anu a -c u s n u ra abr u m nu a a . flat, gl o s , i tely lveol te vo c ca p l te p haped; i vol c l bra s ubuni s eri ate or sub - b r 5— 1 3 m n and m r bb r cts ise iate , , to e tose id i ed, inne with

- R r t 1 ra m . r u mar in narr b n . a l e s sca io s g s , owly o lo g y f o to seve l , fe ale , yellow D is - ore ts num r u rma r r bu r n n b 5 - b c fl e o s , he ph odite; co olla tu la , wide i g a ove , lo ed; r a b n rr n a midveined ft n r n co oll lo es a owly ovate to la ceol te , , o e with esi , apically

An r a a r au a a a n a - n papillate . the s s gitt te to Sho tly c d te; apic l ppe d ge ovate la ceolate , bran r n or m bt r - ir ob tuse . Style ches t u cate so ewhat o use with sho t sweeping ha s ;

las narr b n abr or r 8 — 1 0 m a ar r . s e stig tic eas sepa ated Cyp owly o lo g , gl ous hi sute ,

P a s b r num r u n r. r 1 0 . r bb car o odi um in . u i ed; p p dist ct pp istles e o s , sle de Ci ca spp ,

Hispan iola .

E i r h n n RB . . Type : lekma n a ba a o e s is (U B . NORD

m m n r r r u for E . . EKM N b a and The gen s is na ed L A , e ine t explo e of the flo a of Cu

brin numb r r n m in n r H ispan iola i n the last century. This gs the e of gene ic a es ho ou of EKM N Ekma ni a Ekma ni a nthe Ekma n i ocha r is Ekma n io a us A to nine ( , , , p pp , A F Ekma nochloa Ekma nom ces Elekma ni a Ma neki a M r tekma n i a . On , y , , , y ) ly DOL

ENGLER can match this record in generic eponymy.

Elekma n ia r n nu i n H an a n n u is the la gest se ecioid ge s isp iol , with i e species , pl s r iz . . a ma nens is er tes i i r a one or un r b . v S s r pe h ps two desc i ed Two species , andfi we e r b b N as m m u but m r am n desc i ed y URBA having ho oga o s capitula, the ate ial ex i ed of n n r A k E ha iti ens is are r these species is co siste tly adiate . few species , li e . t uly discoid however.

m m . nov. 1 . Elek an i a ba rahon en s is . . . o (URB ) B NORD , c b

n m: ene i a ra h n n i Arkiv . 23A l : 1 S b o e s s . 9 Basio y c o URB , Bot ( l ) n P nta r DI 1 2 . BORHI A Hu . 37 : 88 99 S : e ca li a ba a honens is U . y ( RB ) , cta Bot, ng r ub h in p ob . p lis ed

: EKM N H 6795 S D m : r r a ru S rr Type A , anto o ingo Co dille a de B ho co , ie a de los

VHI . 1 26 . 1 . Corrri s ari os in n an mm n b 1 800 m 29 . 9 S , pi el ds , co o a ove , ( holo iso C w l 44 2 om . N e s . 006 p , 6 9

m n h lek a i a b a i t . . D m 2 . E U . o . nov. c ( RB ) B NOR , c b

n m : ene i R 4 S buchi i . 1 e ert. 7 : 07 Basio y c o URB , Fedde p

: U H 203 1 H a M m ran an in r " 1 800 m alt . r Type B C , iti , o e T ch t, sylva ap ica, (B dest oyed ) ll : . 6 . I ustr. Fig

r ki n a ara r b r n r n r in m n r This st i g species with ch cte istic ha it is st o gly esi ife ous a y pa ts , even in the florets .

man i a uerte 3. El ek u m . n ov s . . . o b . f (URB ) B NORD , c

a n m : enec i u r S mb . An . : S o e tes i i . 7 558 B sio y f URB , y till : U 1 743 S D mi n : Pr D e V in rn R 2500 m Type F ERTES , anto o go ov. la ega Lo a osilla (B " destroyed )

This was described as having homogamous capitula (also mainta ined in LI OGI ER but all material later collected (in Cordillera Central) are shrub s with heterogamou s radiate capitula .

4 m n h n K m n El ek a i a aiti e s is U . . . o . ov. . ( R G URB ) B NORD , c b

n m n i h K RB in N m An 4 : Se e a i ti ens is U U . S b . : 69 Basio y c o R G , URBA , y till l

: PI CARDA 1 065 H a r P r - au - Pr n in al es tri bus r " Type , iti , p ope o t i ce p (B dest oyed )

has a n b i b n u fr n an r b This species disti ctive ha t, ei g a s f utesce t li a, desc i ed as “ ” “ ” i N S rei z klimm er or r a ra n ar a and m . p t ep do , with li e le ves s all disco d capitula URBA “ ” ma be r but 1 900 r mark : S di stincti s s irna n a arctii s affi ni . ( ) e ed pecies , ulli lii s This y t ue, at least provisionally the Species is grouped here with the majority oi span i olan taxa I n r m r r h r in in the new genu s Elekma n ia . flo al o phology the species ag ees with ot e s

kuekentha li i n i i n n are not n k E . this ge u s . The leaves u li e those of , o ly d st ctly n arrower.

m n 5 . E l ekman i a kueken tha l u . E KM . . . co . ov. (URB ) B NORD , b

23A 1 1 : 92 EKM Ar ki v . a n m : Sene i kueken tha li i . B sio y c o URB , Bot ( ) ’ : EKM N H 949 1 H : Ma Ma u l Ar cahai e a i i Type A , aiti ssifdes the x , , C ye Nicolas , h lls de

1 . 1 . 600 m 1 1 928 S . , ( holo iso )

m . n ov. 6. Elekman i a ma rci an a U . E KM . . . o ( RB ) B NORD , c b

2 A 1 1 : 93 . EKM Ar ki v . 3 n m : Sene i ma r ia nus . Basio y c o c URB , Bot ( ) - Mar u h rn M m : EKM N H 8078 H Ma Ma u St . Type A , aiti , ssif de the x , c , so t e slope of o e C w l 44 2006 Ne s . om . 7 0 p ,

Z emis i a dis olor Sw. . . F i g 7 . c ( ) B NORD

’ P 2647 1 826 as inera r i a dis l r . Ma . r m r . F o Cu tis s Bot g late ( , C co o )

m . N w l 44 2 Co e s . 006 7 2 p ,

herma hr oditi r cremea uin uelobata lobi s n a m ana instructi s p ; co olla q q , li e edi apice S ram s ubtruncati a i cali u r i An therae caudatae flavae . b s i nc as s at s . tyli i pilis p paucis

r i 8 - nervia a illato- c o odi um brev ib us ar s timati ci s s e a at s . ar , eis p Cypsela p p villosa; p p m a n m r ae a a arb ellatae r n P u e os b b . di s tinctu . appi set e l e pe siste tes

Er or r r b 1 — 4 m m n r but ect , diffuse diva icate sh u high , to e tose th oughout, leaves

i ll L e a rna a n rr n a — 1 br n adax a . eav s 5 3 gla esce t y lte te , petiol te , a owly ovate to la ceol te , cm n — 4 cm ub r u n u ub n r m r in lo g , wide , s co iaceo s , with de tic late to s e ti e a g s , m r bb nn u ub b b r u or id i ed and pi ately veined , apically ac te to s o tuse , ase o nded ub r a u r b m n br u and in r r n s co d te; ppe Side eco i g gla o s sh y, lowe side pe sistently de sely

- m n i tul n m r u in rm na r m s n fl r cen r Ca a b o es ce . white to e tose . p u e o s te i l co y ose y , adiate

- m and u n u ra I n lu r e a am anu . c vo c cupsh ped c p late , ca wide high , calyc late; i vol c l

— - — Re e ta le br u m u . R orets 3 6 br un r 8 1 3 . a acts ise iate , c p c gla o s , in tely alveolate y fl

— - r h i — D is o ets enn a hrodit c . 1 0 1 5 ft n min b 2 3 mm . (o e la a o long , long , white c fl p , ca ; mi lin k n An r r m or b n d ed . r co olla c ea yellowish; lo es disti ctly , apically thic e ed the s

r m r n n f u . S b b r distinctly ca date , yellow tyle c ea y white; style a ches su t u cate with ew

x i ll ma a ara la n 2 mm n - n r r ab a a r . C s e b 8 e sho t pili y; stig tic a e s sep ted yp o lo g , lo g, r a rm b mu n u a r car o odi um ved, sho tly villous with p pillifo white o tuse cilagi o s h i s; p p

— P us r num r u ca . r a 3 4 mm rb in . a b b b dist ct pp istles e o s , ise i te , long , white , a ellate ,

m a . r n . One s . Ja pe siste t p , aic

A r A M 1 2 fl w rh r r cco ding to DA S ( 97 ) the o e eads are f ag ant .

m W m . nov. Z e is i a dis ol or S . . o c ( ) B NORD , c b

a n m r r r W r V I n 1 1 4 : i ne a i a dis l S P odr. e . d . O cc . : B sio y C co o ARTZ, g i i l r W D 4 1 2 r w . : en P i l S n S e d s S C . Prodr. 6: enta ca l a dis c S . y c o co o ( ) , o o ( )

H . ROB . . Amol Arb r 1 1 d . , J o 3

”

W R am dis l r WA r S . r : S J . S Lectotype (he e designated) A TZ, aica, C co o [ RTZ sc ipsit] ( )

’

I llustr. : B M 2 4 a . . 6 7 . 7 . Curtis s ot . g t Fig

‘ ’ E n m : r n m r r m z emi s m m b po y y The gene ic a e is de ived f o , the i ages ade y the Taino ‘ ’ in r r n m m G A r r n one r . people the eate ntilles , as ep ese tatio s of ofthei gods , na ed Ze i

fl i ni i A M 1 2 Sene i The a t es ofthis distin ctive taxon are still un clear. DA S ( 97 ) pointed to c o a lmi r n i ll n w m r o c o o in Antilla nthus cf. b r but ( , a ove) as the closest elative , o e

r b b m r n r are r u Ma tt eldi a Nes a m elos p o a ly so e othe ge e a close , s ch as f , p and Ekm i m r ama an r r rr P enta ca li a b H . a n op app us . Two o e J ic species we e efe ed to y

R I N N in P r H R B I f v P. i n rna ta H . ROB . P. s ubdis colo . O . OB SO ROCTOR iz . o and they b n Z emis i a I ann m m n n are not elo g in c ot tell at this o e t, o ly that they well placed in

P en ta ca li a m r b e r r n and u r m n . These atte s will fu the i vestigated disc ssed in fo thco i g a r p pe s . C w 44 2 06 Ne s l . 0 om . p , 7 3

Acknowledgemen t

f r r n me J HN PRUSKI D r. H M A. N NI o Many thanks to Mr . O and T O AS ZA O p ovidi g with referen ces an d copies of protologues of a number of species .

R efer en ces

i I n M na J m . n W. F l wer i n la nts o J a ma i ca . . A M . D . 1 972 . DA S , C o g p f U v dies , o , a aica

i ni Pu r R Ed t . U v. D E E LI I E R 1 F l ra de u a 5 . A AI N HE M N R . . . OG 963 . C b L , R A O ( ) o de e to ico ,

Ri o Piedras .

1 2 r in n m new in r RHI D I A. 99 b . ub BO , (p o p lished New a es and Species the flo a of

— 5 0 . H n . 9 Acta Bot. u 7 Cuba and Antilles . g

i R D . P r M r . La F l r a de la H is a n i la 8 . San R A H . 1 96 . LI O GI E , . 9 o p o ed o de aco s ,

m in r b Seneci oneae m . NO RD ENSTAM . 1 978 . n , B Taxo o ic studies the t i e (Co positae)

— 44 : 1 83 . Op era Bot .

I n ur bi a new nu A r a - Seneci oneae r m NO R DENSTAM . 2006 . , B g , a ge s of the ste ace e f o

Wi lldenowi a 36 S u . H ispan iola . ( pecial iss e)

W. . FF E I KAD EREI T J . J r r b Seneci oneae . n : RD EN TAM . in . NO S , B p ess T i e , C E R Y (eds)

K KUBI TZ KI . VI I I ri cd . . The F a mi li es a nd Genera o Vas ula r P la nts . f c , vol (se es , ) n H b r rk: D i n . A ra . r r n an . t Flowe i g pl ts co yledo s ste les Be li , eidel e g, New Yo

Sprin ger. 3 ma a . Arnold Ar bor . 6 M r n ra J J . PR G R . 1 982 . OCTOR, o e additio s to the Flo of a ic

L 1 99 31 5 .

— 4 1 4 70 . Antill . : 69 S mb . i ha i ti ens is . N I 1 900 . Sene URBA , . c o y m N w 44 . 2006 4 Co e s l . 7 p ,

L ectot ificati on o e o H i era ci um ec yp fnam s f s t. r m Vul a ta b r l H i e a ci u and sect. g ased on mate ia fr om souther n Sweden

TORBJ ORN TYLER

De . ofE lo Plant E lo and S emati c pt co gy, co gy yst s SOlve ata n 37 S—22362 u d Sw d n g , L n , e e tor bj

D edi a ted to P r I L NORDENSTAM wh s e ea r l i nteres t i n b ta n a nd the c of BERT , o y o y m s i ta e was unded b his un le the H i era i um- ex er t S N NORD ENSTAM on Co p o fo y c , c p TE , 7 r F 2 the as i n o his 0th bi thda on ebrua r 0 2006. occ o f y y ,

Abstract

Lectotypes for 94 combination s in H i era ci um based on materi al from the Swedish r p ovinces of GOtaland and eastern Svealand are selected and presented .

I ntrod uction

The aim of the present study has been to select lectotypes for all n omen clatur al

H i era i um H . . combination s of taxa currently placed in H i era ci um sect . c and sect ’ ’ Vulga ta (in cluding the bifi dum- group ) described fr om material (at least in part) collected in the GOtaland provin ce or the eastern part of the Svealand provin ce in rn All u kn r m r r southe Sweden . species of these gr o ps own f o this a ea have ecently b n r and r b in r im r r n m n r Y 1 998 2002a b ee t eated desc i ed p el ina y egio al o og aphs (T LER , , , 2003a b 2004a S m r an mbina n r b , , , o e ofthe elev t co tio s have al eady een typified b SENNI K OV 2003 YL 2000 2004b but r mb r mbi n y ( ) and T ER ( , ) a la ge nu e of co ations

Of m ina n rank f ar or r m . b O b e ain these , all co tio s at the v iety a ove, except those r b HN an E I desc ibed y ZA d a few ofthe oldest combinations in the group made by . FR ES

J . I N u . I n m C . b are and b and L DEBERG (as disc ssed elow) , analysed typ ified elow ost a in t for mb n b r r f r ar c ses , select g lecto ypes these co inatio s has een elatively st aight o w d n r m r ar n a in rotolo ues ar r si ce the o iginal ate ial is cle ly i dic ted the p g , l ge pa ts of it has been refoun d in Swedish pub lic herbaria and the material has been found to con form m rr u mb in n n rn . H r with the cu ent sage ofthe co atio s co ce ed (ifany) oweve , a few so e m what more complicated cases are discussed and explained in some detail below. So e particu larities relevant for this study concerning the nomenclature and typificati on of Scandinavian H i era ci a have previously been described and discu ssed by TYLER New l . 44 2006 C om . s p , 7 5

(2000) and SENNI K OV Some ofthe original material have been distri buted in

r u r ex s i ccata ri n ab ft n m a m n va io s se ies of , and p ted l els , o e with so ewh t e e ded text, n b b ut all m n are r u b r have the een added, speci e s he e q oted y thei

i n a u u - r n a r b n a . o gi l , s ally hand w itte l els (whe ext nt)

Am n mbi n r u a r main lectot if m o g the co ations in this g o p th t will still e to yp y, those ade by ZAHN ( 1 92 1 - 23) Shou ld b e lectotypified with material in the ex s i ccata ofDAHLSTEDT r or in ir n m r b b rotolo ues . n r u as di ectly d ectly i dicated in the p g U fo t nately, ost p o a ly none of the duplicates of these exs i ccata actually seen by ZAHN survived the second w H r in u a I BN 20 a for r ar. C Ar wo ld oweve , s ch c ses the ( ticle Note ) llows the selection of lectotyp es from the remaining duplicates (i s osyntypes ) ofthese exs i ccata

r S . Si exs i ccata D H b u b n (e . g . those p esent in ) nce the of A LSTEDT O vio sly have ee compiled with great care an d are highly homogeneous this shou ld not meet any

r m H r m r m ina n ma b r u r b . r r u b b pa tic la p o le s oweve , since the e is a la ge n e ofco tio s de y ZAHN b ased on material from throughout Scandinavia distri b uted in the ex s i ccata of

DAHLSTEDT I consider it appropriate to treat these together in a futur e separate study.

I an d . J . I N E Concern in g the few old and important combinations of E . FR ES C L DEB RG FR l FR and H tenole is t . . t H . H . di a ha num . vu a um s a ar e f unt ified . . th t still le yp (e g p , g p r r n m a a r no b u r n ma r a LI NDEE . ) they all ep ese t co plic ted c ses whe e o vio s o igi al te i l has been refound and where the extant material ofthese authors for vari ou s reasons has to H r I am rr n rk n on m b e in r r r r . te p eted with g eat ca e oweve , cu e tly wo i g ost ofthese cases and they will b e the topic of separate pub lications in the n ear fu ture .

Lectotypifications

- HAN 1 900 . I : 1 8 20 n s s . s ematole is JO ss . 1 2b . acidotum DAHLST . 89 H . ( exs i dex) p p ( p

’ H r s zn Vid r nar Gillers ho den 1 898 r : Vs m . Lectotype selected he e allefo s , ett to p a j ; ;

’

K J H N N in S. leg . . O A SSO

ca es iti um NORRL NN Note : This combin ation is a taxonomic syn onym of H . ( ) BRE ER

AM 1 2 — 2 S . 9 3 6 JOHANSS. T STENSTR . ( 1 892) and H chr is ti a nens e (DAHLS . ex ) ( ) and this also became the view of its author as i ndicated by a determi nation stri p from 1 924 attached to the lectotype .

1 920 . H . acroma dar u m SAM JOHANSS. , in ( p

’ GUNN udenn anni a S ra na Elds und 1 7 ni 1 904 . Lectotype selected here: S , t ng s , ; ju ; leg AR

’ ’ m has r b n as H s i lva ti cum SAMUELSSON in S. The speci en fi st ee identified ma da rum b m but a r H . a cr o T no . sp ilop hyll um DAHLS . [ l te changed to y

SAMUELSSON . m New l 44 . 2006 Co s . 7 6 p ,

HANS 1 7 a lbi n otum DAHLST . JO S. 90 . H . ex ( p

’ ’

l . Dann m ra 1 . K H N N Ex J . Lectotype selected here : Upp e o ; 900; Leg . O A SSO ( 4 n Hierac . an . XI V: 8 m ri b in S. D H H rb . S A LSTEDT , e c d ; with e e ded p nted la el)

1 894 . a n ractum Ss . e u r c b e DAHLST . H . f p y y ( p

’ Lectotype selected here : Smaland; H eljaryds angar pa in s krankt lokal (torr mark)

’

H N N in S. 1 884; K . JO A SSO

a crochr is tum DAH LST . Note : This combin ation is a taxonomic synonym of H . ex ANSS JOH .

AM . 1 920 . H a n o mum OHANSS. S . is to J ( p

’

r : Vastmanl and : Va an r a r a an 1 9 1 7 . Lectotype selected he e st fo s , F ge st , lov g; ; leg ’ GUNNAR SAMUELSSON in S.

in OHANSS. AM . 1 20 H . a rros i orme DAHLST . J S 9 f , ( p

’ h n 1 EU NI Stren as D om rost a e . 899 t r : SOderm1 . Lecto ype selected he e gn , p g ; GE E

’ EX HL H r H i ra n XI r n b KOH . D b . e c . S . : 50 LER ( A STEDT, e ca d ; with p i ted la el with the site ” ” emended to Langberget and the date changed to in S.

m E M 1 badi ell u . G A . 955 . H . L O ( p

’ r l D m r 1 H D H H a di ellum : 890 . b Lectotyp e selected he e : Up . anne o a; ; A LSTEDT;

ER . A M v. Tids kr . 1 55 r . in . L O. ; Vide S Bot . 9 ; O ig S

m m D AH L D 1 H . cacu i na tu ST. AHLST. 899 . XI : ( ) ( , exs

4 a n ra ctum s s . ca cumi na tum DAHLST . 1 89 a . Bas H . f p ( p

’ r : Sma r u k n en tradb evaxt b r an Lectotype selected he e land, Fage h lts soc e i och e gig g

’

1 83 K . NNROTH S. invid r u rr ar . LO Fage h lts he g d 8 ; J . in

H . ca es i r um A M N RRL 1 8 i o . O 88 . fl L O ex . ( p

’ D es cr r A M 1 1 r v u 88 1 . 5 o H . s i l a ti cum s b s . 5 . o ig . L O. ( p ) p p

’ ’

t r l h lm k r 1 A M in S. V S 879 S. UI Lecto ype selected he e : Up . ax o a po; ; L Q ST

’ E r D H 1 872 S. A M UI . H . t : bb . pi ype selected he e Upland , Ti le ; L Q ST (det A LSTEDT m n on i n the left speci e the sheet S.

nam wa r b NORRLI N Note : This case is somewhat complicated . The e s fi st validated y “ ” but ALM V u r and u b e n r r n r he quoted Q . as a tho this sho ld i te p eted as an i di ect ’ r r n r n i l va ti cum u s . 5 i n A M UI u H . s s b efe e ce to p L Q ST Th s , the desc iptio given r and by ALMQUI ST ( 1 8 8 1 ) is the original description and protologue . Seve al localities gatherings are mentioned in this protologue but the one here selected as lectotype is N ws l . 44 2006 C om . e p , 7 7

h n one an a a f t e s ar u be a r ain . H r o ly ext t tod y ( as co ld sce t ed) oweve , the lectotype gatherin g consists of two Specimens apparently collected late in the season and thus r u f r rm i i r not o ina n . r r n an t s r ve y usef l dete tio The efo e , the selectio of ep ype app op iate and the one selected here was collected by the author at a Site mentioned in the

rotol o ue u r u r m n was a ar n not n p g , altho gh this pa tic la speci e pp e tly ide tified with this

’ un a r H r in DAHL TEDT an n . n ab S S a d species til l te ( oweve , the exta t l el ofthe epitype is h d Since the origin al label is missing it is indeed possible that the specimen was origin ally referred to this species by its collector and author ")

n r n DAHLST . 1 2 I H . caes i o i es ce s FR . STENSTR . : g ( ex ) ( 89 b exs .

ilva ti cum s s . ca es i on i es cens FR . TENSTR 1 . Bas : H . s r 889 p g ex S . ( p

’ r : D als land S n b Skn m an an a r na Lectotype selected he e , te e y , ell L ged och B lde s s ’

E . N OM . 1 888; K . O . STE STR in S

Note : The epithet ca es i on ig r es cens on this plant appeared already in FRI ES ( 1 847) but was not validly published sin ce the species was not definitely accepted there .

1 2 1 . ca n ed orme DAHLST . 9 H . ip if ( p

’

l . ra ar ins . Rafs ten 8 1 9 1 5 . r : n s . U : G Lectotype selected he e . p ; pp so p , , ; Leg ’

E h K HAN N in . RI N I D et . H . DAHL x . rb . . JO SSO S H . GSE LE; STEDT ( e )

2 chi ridotum OHAN SS. 1 9 0 . H . J ( p

’

k r k l van 1 1 8 . H D als lan d . Eds les o n r O 9 r : ab . Lectotype selected he e g, B ac e a ga , g; ’ n n 1 6 J H N N SAMUELSSON H i erac . . . o . GUNN SAMUELSSON Ex . leg . AR ( O A SSO , sca d exs ; with emen ded printed label) in S.

1 905 . hl r ma u r um JOHANSS. H . c o o ( p

’ Lectotype selected here : OstergOtland : Aby Vid lands vagen till Gravers fors ; ’

in S. K . J H N N 1 900; Leg . O A SSO

1 925 . com lex um JOHANSS. H . p ( p

' ’ ’ HA M D t 0 . e . 1 896 Le . r : Sdm Ostertal e Skarfs ta 6 . Lectotype selected he e j , , ; g GSTRO ; ’

1 24 in . K . JOH . 9 S

4 7 1 925 . 8 malmea n um JOHANSS. 1 1 1 var. 1 889 . con tri n n s NORRL . H . s g e ( p ) ( p , ’ Ma lmea num

’ ’

ANSSON in S. Rens atra 1 925 K . JOH t r : U l . : Lecto ype selected he e p Bo , ;

Note : This var iety differs fr om the typical form of the species by very finely denticulate leaves an d more acute phyllar ies with almost no stellate hair s and is thu s H r n kn n r m n arb ra r n ma m r r k. the disti ct and y e it specific an oweve , it is o ly ow f o two e y cons tr i n ens localities and its relation to the very widespread an d rather variable H . g is m w 44 Ne s l . 2006 Co . 7 8 p ,

thus diffi cu lt to determi ne .

2 u rimontan um DAHLST . JOHANSS. JOHANSS. 1 90 . H . c p in ( p

’ Lectotype selected here : Suecia : Ves tm anl and in colle lapidoso prati s ilvati ci ad 1 K ’ E K arber ar . u n r b r 898 . . J H N N x . D H H rb . opp g p Lj s a s e g , ; leg O A SSO ( A LSTEDT, e

XI ] in . r S a . : 67 S Hi e ac . c nd )

m HAN 1 dali cu JO SS. 900 . H . ( p

’ ’

D als ko s a 1 8 . K . H N N . in t r : D als l . Lecto ype selected he e g st tion 99; leg JO A SSO S.

AN SS. 1 27 . n i ra DAHLST . H 9 H . de g ns ( ) JO ( p

D : H . tenebr i cos um . den i ra ns AHLST. 1 893a . Bas . ssp g ( p

’ n r Es keback et s acellrun aroeciae r u Lectotype selected here : H ab . i te p Fage h lt ’

1 883 . K . J . LONNROTH S. Smolandiae . Legi in

ha n I N E . 1 2 . H . di ap oides L DE ( 88 p

’ r m E - AVALL D S . et . OGoth . G . HYLTEN C I US et K . . U EN Lectotype selected he e : Suec . ol F S

H r . ex i cca m n r r ar x . I N t E . S an s ta f a ( L DEBERG , ie c d the le speci e , p ese ved sep tely in

LD .

Note : The specim en moun ted to the right ofthe lectotype belongs to a rather different

chlor odes DAHLST . DAHLST . but m r species (possibly H . ( ) the ate ial is not convincing) an ff r m di a ha noides r m r n on d r n H . di e s f o the diag oses of p , as well as f o the desc iptio b a m an in in m r s im r i n the la el cco p y g the exsiccate , in hav g nu e ou s ple hai s the

r H r r a s nflo es cence . b i y oweve , the lectotype fits well oth the o ig nal di gnoses and the n curre t usage of the name .

H . dia ha n i de LI ND EB 1 2 1 1 var m L NNR AHL T 1 4 . o 88 . . atri ca ill u O . D S . 89 a p s . ( p ) p ex ( p

’ r : Sma a r k n n ra Lectotype selected he e l nd, Fage hults soc e vid vage f n as t i fv 4 1 /7 1 883 . g g aregarden till Es keback nara gam la badhus et bland gras och buskar. K ’ J . LONNR OTH in . S.

Note : This is a very pecu liar plant only known from the type gathering (consisting of I t ha r m l l u in 5 m s . s b subam exi cau a speci en ) oad, sessile and al ost p ca l e le ves , an extremely long and dense in dument Of glan dular hairs with few intermixed b lackish simple hairs in the s yn flores cence and an indistinct b and of stellate hairs along the m r n r I a ar n n t r a H . di a ha noides a gi s ofthe oute phyllaries . t is pp e tly o closely el ted to p LI ND EB d b e r r r r . an I t ma b e can not ega ded as a va iety ofthat species . y closely elated

H . rna tum DAH L T D n but r b not on s ecific . H r S . AHLST . b c to o ( ) is p o a ly p oweve , si ce this an n b n n m r an n ur r no ur n r pl t has o ly een fou d o ce o e th a ce t y ago , the e is ge t p actical n r a n m r are r r a r n eed to esolve its t xo o ic position and rank. The e seve al late g the i gs

N w l 44 m . 0 0 Co e s . 2 06 8 p ,

n label) i S.

O HAN . AM . OHANSS. AM . 1 2 2 H . h aboens e (J SS S ) J S ( 9 3 6 exs .

4 ha boens e OHANSS . SAM . 1 92 . r i ns var. J : H . s a s de Bas . p ( p

’ H a Domm eb r 1 r V . e GuNNAR t : b et 908 . Lecto ype selected he e g o , g ; leg

’

SAMUELSSON in S.

AM . 1 24 . hi HANSS. S 9 H . ans JO ( p

’ ’ k A k r t rk k 1 2 1 K N r : S o b ac 9 . J H N Ex . Lectotype selected he e : Nar e . s e ; ; O A SSO (

i ccata no . 1 5 m n r n J H N N SAMUELSSON H i erac . an . exs 7 O A SSO , sc d ; with e e ded p i ted label) in S.

Note : This plant is on ly known from a few specimens which are very Similar to the

m r r r A M n r r n s ubc as s um . NN a d a e ea lier described a d o e widesp ead H . ( L O ) BRE ER they most prob ably conspecific .

rr n va r i n cu e s SAELAN NORRL . 1 889 . 1 50 . ol eta tum JOHANSS. SAM . 1 920 . H . ex ( p ) ( p

’ k k 2 / r : Vastm an land: L us narsber . Sun b or bac e 5 6 Lectotype selected he e j g ds Naset, j ;

’ 1 1 L N GUNN SAMUE SSO S. 9 0; leg . AR in

Note : This tax on differs fiom the typ ical form of H i ncurr ens (main ly known from

— marm ra a and rt r n um n in Finland) by e . g . Spotted o ted le ves sho e i d e t the

I t u b e r r r rm s ynflores cence . sho ld ega ded as a species of its own ight and the fo al r m n n m eco bi atio will be made by me elsewhere ( s c) .

H . i n te rati ron s OHANS 1 900 . g f J S. ( p

’ r m Gr h e s zn r n rr om Torrvar s und : Vs . t tt Lectotype selected he e y y , vid ett to p o p ; ’ 1 m n K . OHANSSON EX. D H H r . H i rac . an . XI I : 57 898 . J b e S ; leg ( A LSTEDT , e c d ; with e e ded r n a i n p i ted l bel) S.

H . n ter v n DAHLST . 1 9 b . I V: i e ie ns DAHLST . 8 3 ( ) ( , exs

B H . r mu a ra m Ss . i n er v n i n DAH LST 1 1 . I V as . : t : e e s . 89 p ( a, exs

Lectotype selected here : [description] Suecia : Ves tmanlan d prope Charlottenb org ’

Kun r D H Hierac . Le . . . . P T . EX. o s ubumbros i s 1 890 . O ad gs locis , g C V ORA ( A LSTEDT, Ex i a 4 in s cc ta I V: 8 ) S.

h r h n H . i t a o to JOHANSS. 1 00 p y ( 9 p .

’ Lectotype selected here : Suecia : Ves tmanland i n prato solo calcareo orientem ’

r u ad katviken H N N Ex D H H rb . h e 1 898 . K . J . S ar. Gr t tt ve s s p y y , ; leg O A SSO ( A LSTEDT, e Hi 4 erac . an XH : 1 in S. Sc d . ) N m . w 44 2 Co e s l . 006 p , 8 1

- H . oh a n s s omi D AHLST . AN JOH SS. 1 2 SAM . 9 3 26 j ( ) ( exs .

’

H . s r Bas . : a s iden ha s s s . o ns s on i i D 1 AHL ST . 1 4 p p j ( 893a p . 6; J oha ns s on i i

t r : R Srnl . : o ber a s zn Vrs malen aen mos s bevaxt kl a arr Lecto ype selected he e g g , p ipp i b ’

k . 1 888 K . J H N N s og ; O A SSO in S.

- H . uelii DAH L ST . OHAN J SS. SAM . 1 2 2 j ( ) ( 9 3 6 exs .

’ H s ilva ti . : cum s s . ueli i D 1 AH LST . 57 Bas p j ( 893a p . ; J ueli i

’ ’ r : Stockh . : U leviken 1 886 O . UEL in Lectotype selected he e gg ; ; J S.

H . koe i n en s e JOHANSS. 1 p g ( 929 p .

’ ’ t r : Vas tmanl K in : . J nn a 1 928 K . J H N N Lecto ype selected he e op g oha esd l , O A SSO

Ex . J H N N SAMUELSSON H i r e ac . an . exs iccata n 477 m n o . r n ( O A SSO , sc d ; with e e ded p i ted a l bel) in S.

H . lam ocran um JOHANSS . AM 1 92 . p S . ( 0 p

’ ’ t r : Vs m: Ski n ka b r norrut 1 K . 9 1 8 le . J H N N in Lecto ype selected he e n s tte e g, ; g O A SSO

S.

H . l a rs s omt JOHANS SAM in OHANS 2 S. S J . 1 9 0 . , ( p

’ An Lectotype selected here : D s l . imskog: Lilla B racke paMurgroneb erget 1 9 1 8 ’ GUNN L . SAMUE SSON Ex . J H N N SAMU ELSSON H i r e ac . an . no . 36 leg AR ( O A SSO , sc d exs . ; m n ri n ab in with e e ded p ted l el) S.

H . l e idi ce s DAH LST DAHL T 1 . 900 . XI I : 60 var v S . . i etuli oli um JOHAN SAM . p p ( ) ( exs ) f SS. 4 ( 1 92 p .

’ ’

r bruk. 1 E : Sodml . nn a 899 U NE K H in Lectotype selected he e La ; GE O LER S. Mounted on this sheath is also a label with the name vi etulifoli um and a description ’ in OHAN N of it J SSO S han d .

Note : This is a dubiou s taxon ; as poin ted out by its author typ ical forms of it have longer and more poin ted leaves and narrower phyllaries with more conspicuous

a r n in ma as m ar a r stellate h i s a d a dist ct apical co co p ed to the typic l ace ofthe Species . I n a n ar ft n n a m r num r u m a r in dditio , this v iety o e has violesce t le ves and o e e o s si ple h i s

flore c nc H r man m n xhibi n m h ara r s n s e e . the y oweve , y speci e s e t o ly so e oft ese ch cte s an d even the original material (from two different sites an d collected duri ng several years) is variable in many respects making it diffi cu lt to evaluate this variety. The n in a in narr n an d ma ar lectotyp e closely fits the diag oses h v g ow, poi ted co te phyll ies with conspicuou s stellate ha irs an d it is mounted together with a han d- written r n JOHANSSON but k n a an d ha r h ar un desc iptio of , it lac s the violesce t le ves i y p yll ies fo d in man y other specimens referred to this variety (including some collected at the same m w Ne s l . 44 200 Co . 6 8 2 p ,

locality) .

JOHANSS. 1 927 . l e idi orme STENSTR . H . p f ( ) ( p

m TEN le idi or e S STR . 1 889 . Bas H murorum s sp . p f ( p

’ i S b s zn r n l tet b r . 1 8 88 t r : D als l . : a Lecto ype selected he e tene y , Balde s as p ett e g , ’ M in E . S N O S. K . O . TE STR

m n m AHL T l 2 . f n m A M D S . 9o b . ecti atul u . H . l on i a u NORRL . g ( ) ( , exs p ( L O ex

AN 1 2 . JOH SS. 9 7 DAHLST. ) ( p

A M AH L T 1 4 l D . a . a es i um . ecti na tu um . S 89 Bas H . c ssp p L O ex ( p

’ r : Su : S kh m i n s ilvati ci s umbros i s ad D r ar n Lectotype selected he e ecia toc ol ju g de ,

’

Ex iccata I V: 67 . Ex D Hi erac . s 1 890. . HL S Legi ipse ( A STEDT, ) in

rm lon ima num b Note : This form differs fr om the typical fo ofthe variable species H . g y ta ller grt and more deeply cut leaves with numerou s leaf- like appen dages on the I n ar u r u n ar e br r ar u i and petiole . p tic la the ca li e leaves oadly t iangul in o tl ne deeply m i JOHANSSON pinnatisect with large basal teeth maki ng the truncate hastate at base . ( 1 927) considered the taxon as merely a modification ofthe species in Shaded habitats but since then many more specimen s have been collected and it appaers to me as ifthis morphology has a genetic backgr ound and may b e worthy of taxonorrri c recogn ition .

H r u r m n are fi a r n . oweve , c ltivation expe i e ts needed to get a de nite nswe to this questio

ma ur h m I N 1 oce al u 936 . H . p FOL ( p

’ t r : D als land Eds les ko s s n H n n an sbacke Lecto ype selected he e g , oghede , oppe g , ’ 1 P A N H D H 09 . R 9 . . . . S. , leg LA SSO ; det A LSTEDT in

m dii rm 2 e o e M M DAHLST. 1 896 . X: 7 . H . f ( AL E) ( exs )

: H s ilva ti cum . medi i orme MA M AN N 1 890 . Bas . ssp f L E [ DERSSO ] ( p

’ ’ N in m . t . Malrn r Ols s an 1 890 . G O . A: . r : Sr S So tu . . Lectotype selected he e , g g d ; Leg

S.

m al tr h m 1 925 . H . eg o ac el u JOHANSS. ( p

’ b J HAN N Ex r . r : rrnl k : k k 1 1 924 K . Lectotype selected he e S . Nac a Nyc elvi en ; O SSO ( he

J HAN N . K . O SSO ) in S

H . membros um OHANS AM 1 24 S S . 9 . J . ( p

’ ’ l n K 4 2 K OHANSSON Ex r : n r tri /7 1 9 3 . J Lectotype selected he e : Up . Lidi go a a a nelund ; (

K . H N N i n . herb . JO A SSO ) S

H . meti ce s ALM DAHL T AH L T l 2 I 4 1 var eu dometi ce s . D S . 89 b . : . s p ( Q ex S . ) ( exs ) p p C m w l 44 o . Ne s . 2006 p , 8 3

AHL T 1 4 89 a . D S . ( p

’ r : Su a: Sma in b m r A k Lectotype selected he e eci land etulo ac o ad Bordsjo par . s eryd 1 884 ’ Ex . D H . H b . Hi era 2 r c . S . V: 6a in S. , legi ipse ( A LSTEDT, e cand )

: Thi n n m n u n Note s taxo is o ly a odificatio of the species witho t taxo omic value (cf. ’ ’ JOHANSSON 1 927) and in my Opin ion the species H meti cep s is in turn a taxonomic n n m for r r r b H ca n i ALM STENSTR . DAH LST . sy o y the ea lie desc i ed p es ( O. ex )

m n i 1 2 1 H . ol bdi o des DAHLST . y ( 9 p .

’ t r : o rml Stren as a n Ki n n 1 S de . n 898 Lecto ype selected he e g vid v ge le Lu da . ; ’

EU NEKOH D et . DAHL T S. GE LER; S . in

H . m un dul orme AHLST DAHL i D . 1 900 . I I I : f ( ) ST. ( exs X

H i nte ra tum . mun r . : i 1 893a . Bas g ssp dulfo me DAHLST . ( p

’ r : n l n 1 bb H as o 886 S. A M UI . Lectotype selected he e Upla d , Ti le , j ; L Q ST; (det H D H in S. . A LSTEDT

Note : This combin ation has long been considered a taxonomic syn onym ofH p a ta le

NORRL.

m muror um s s . n i o l an dul er u . . WESTERL. 1 890 . 1 H . p g r g if C G ( p

’ ’

1 . . R nn H l n . W r : b e sobrunn e 890 . G UN Lectotype selected he e o e y, Leg C ESTERL D in S.

h n m D AHL T HAN n JO SS. a d Note : The lectotype clearly belongs to H p s eudodi ap a u ( S . ) ’ ’ the name n ig rog la nduliferum has never been in wider u s e .

m 1 0 . n as to h ll u JOHANSS. 90 H . p y ( p

’ ANSSON n Vast otetor 1 898 K . JOH t r : Vs m . Gr th tte s z Lecto ype selected he e y y , g p; ; m n r n a in XI I : 5 b S. Ex H r Hi erac . Sacnd . 9 D H b . ( . A LSTEDT, e ; with e e ded p i ted l el)

1 2 . n r t n HANSS. SAM . 9 0 H . e i odo JO ( p

’ r : Vastman land: nar b r r b r b orkb acke Lectotype selected he e Ljus s e g , B att e g ( j );

’ N i r SAMUELSSO H e ac . . SAMUELSSON Ex . J H N N 1 9 1 0 le . GUNN ; g AR ( O A SSO , scand

ex i t n 46 m n ri ab S. s cca a o . ; with e e ded p nted l el) in

n n VUK . 1 882 . o n i o la n dulos um LONNR . H . gr g ( p

’ as cui s r Flenvik aroeci ae H an var Lectotype selected here : H ab . in p p ope p g ’ in NNROTH S. K . J . LO Gotlandiae 1 853 Legi .

Note : The lectotype consists of badly developed Specimen s of the ear lier described d H n i r o la ndulos um has n n b n r ar H ell u idum LAEST . an p c , g g i deed lo g ee eg ded as a m w Ne s l . 44 2 4 Co . 006 8 p ,

mi n m at taxono c syno y of th species .

todon m TENSTR . DAHLST 1 . I X: 9 va r r o ea tu S . 895 3 . oc i odes JOHANSS. H . p ( ) ( exs ) ( )

1 2 - 2 in JOHANSS. SAM . 9 3 6 . 3 1 7 JOHANSS. , ( exs )

1 0 . Bas H ocr i odes JOHANSS. ( 90 p

’ h e s zn Elfvesto s bruk 1 r : Vs m . r t tt r G 898 . Lectotype selected he e y y , p ; ; leg ’ Ex DAHLST r i r m n JOHANSSON . . H b . H e ac . S . XI I : 50 r ab in K . ( e cand ; with e e ded p inted l el)

S.

Note : This variety is not mu ch different from the type of the species and does in my

Opin ion not merit taxonomic recognition .

H a mi n um OHANSS. AM 1 2 e os J S . 9 0 . . p ( p

’ r : Vastmanland: u r b r r b r b orkb acke Lectotype selected he e Lj sna s e g, B att e g , j ;

’ in 1 . . UNN R L SON 909 leg G A SAMUE S S.

lm r DAHL T 4 H . a e i S . JOHANSS. SAM . 1 92 . 1 38 p ex ( p ,

’ m a Vid llmar rd n 1 r : . Sa o a Gu s f o e 908 . Lectotype selected he e Boh st d, l v ng j ; leg ’

J H N N SAMUELSSON H ierac . an . exs i ccata no . 1 3 GUNN SAMUELSSON Ex . 8 AR ( O A SSO , sc d ; m n r in with e e ded p inted label) S.

Note : This comb in ation is a taxon omic synonym Of the ear lier described H a cuden tulum OMANG as was recogn ised by JOHAN SSON already in 1 926 (as evident r m f o determination strips in S) .

H . a rama urum OHANSS. 1 p J ( 900 p .

’

G h e s zn Vast oteto 1 898 . r : Vs m . r t tt r Lectotype selected he e y y , g p ; ; leg ’

J HAN N Ex . D H r H i r n XI m r n K . T H I n b . e ac . S . : 56 O SSO ( A LSTED , e ca d ; with e e ded p i ted a in l bel) S.

m e n 1 2 1 . H . p cti atu DAHLST . ( 9 p

’

l . A RI NGSELLE V l . : G Det. r : U a Orta und . Lectotype selected he e pp ddo , ; Leg ;

’

H . D H n s . ori inalex in . A LSTEDT; . p g S

H . erl a xu m 1 02 . 1 va r. obtus i d n DAH LST . OHAN S. SAM . p JOHANSS. ( 9 p 3 ) e s ( ex J S )

JOHANSS. SAM . 1 92 ( 3 p .

H . tus idens DAH n . : L JOHANSS. AM . 1 920 . no DAHLST . Bas ob ST. ex S ( p ex NOTO

’ r : Vastman land : nar b r Brattber s hall b orkbacke 2/7 Lectotype selected he e Ljus s e g, g ( j ) ’ 1 9 1 GUNN R AMUELSSON in 0 . S ; leg A S.

H . h a i DAHLST . JOHANs s AM 2 eo s s . 1 9 0 . p p ex S . ( p C N w l 44 2006 om . e s . p , 8 5

’ Lectotype selected here : Suecia : Vestmanland solo caespitos o humidi us culo i n prato ’

m ar. G h e 1 8 7 l . K H N N Ex D H r u r t tt 9 . e . J H rb . p ope sacell p y y , , g O A SSO ( A LSTEDT , e

S . XI : 8 1 in . H ierac . cand ) S m hr i on i u JOHANSS. 1 05 . 1 H . p yg ( 9 p

h ’ m . H elms e d zn H 1 t r : S r . s s r 893 . Lecto ype selected e e j y , N etse yd; ; leg ’

K H N N in S. . JO A SSO

m l an ta i n oli u JOHANSS AM . 1 20 . H . p g if S ( 9 p

’ Vas tmanlan d: Va an r r a k n 1 r : r 1 9 7 . Lectotype selected he e st fo s , Fage st i pa e ; ; leg

’ AMUELSS N EX HAN N L i n GUNN S O . JO SSO SAMUE SSON H erac . . exs i ccata no . 52 AR ( , sca d ; m n r with e e ded p inted label) in S.

um u r m DAHL T HAN l li e u S . JO SS. 1 927 . H . p g ( ) ( p

: H s a r co h llum Ss . lumuli erum DAH LST . 1 893a Bas . [ p y ] p p g ( p

’ r D l l n R r Gun r n r Lectotype selected he e : Suecia : a s a d ad ostock pa . na s as loco ape to

’

A. FRYXELL Ex . D H H r . Hi erac . an . 11 : 1 in S. a r 1 890 . b S p ico , , leg ( A LSTEDT, e c d )

1 27 . l um uli er um var. li a den i u m JOHANSS. 9 H . p g p ( p

’

L n b r . 1 926 . R N t r : Bohus lan n b s n . Lecto ype selected he e Teg e y e , e g AG AR

’ OHLSEN 111 8 .

Note : This combin ation is a taxonomic synonym ofH ca lligla ucum OMANG .

AM 24 1 . o eli um OHANSS. S . 9 H . p lyp J ( p

’ r Nrk K br : S ar a bruk b rkan 1 92 1 : Lectotype selected he e : . vist o v t ( jo g) ’

i r n x i ta no . 1 5 Ex OHANSSON SAMU ELSSON H e ac . . e s cca 9 K J H N N . J . O A SSO ( , sca d ; with in emended prin ted label) S.

in OHANSS 1 9 2 J . 0 . H . orri en ti orme B BI . p g f A ST , ( p

’ Brannk rka Lectotype selected here : Su ecia: Soderrn anlan d ad Jakob sb erg par. y loco ’

H r . Ex . D H b humidius culo ramin os o in r s ilvati co 1 890 . g p ato , , legi ipse ( A LSTEDT , e 4 in 11 : 7 S. Hierac . Scand . )

4 AM . 1 92 . ar v r ns OHANSS. S H . p if o J ( p

’ ’ EX A. RI NGSELLE . l S n : r ik 1 922 le . G Lectotype selected here : Up . i go T anv ; g ( i 1 m n r n n x ta no . 97 J H N N AMUELSSON Hi erac . . e s cca O A SSO S , sca d ; with e e ded p i ted

a in l bel) S.

24 AN AM . 1 9 . roli n u a tum JOH SS. S H . p g ( p m w l 4 Ne s . 4 2006 8 6 Co . p ,

’

l . : Varmdon r r : U fb 1 906 . G O . M M Lectotype selected he e p , Lo e ga , ; Leg AL E ; ’ 2 in K . OH . 1 9 3 S. detenu . J

1 4 f HAN rolix um NORRL . 888 . 9 . occos i ce s JO SS. 1 929 H . p ( p ) fl p ( p

’

U l . R ds s zn U lln a s teni b orkba k 1 r : c e 90 1 . Lectotype selected he e pp y , , g j ; ; Leg ’

H N N S. K . JO A SSO in

Note : This form differs from the typical form of the species only by somewhat more abundant stellate tomentum on the phyllaries and it is probably merely a modification of the species and does not merit taxonoa recognition .

r m n M LME DAHLST 1 X11 e a ens . 900 . : H . ( A ) ( exs

AN : H s i lva ti cum . r ema nens M M N 1 89 1 . 1 79 Bas . ssp AL E ([ DERSSO ] p )

’ ’

t . M n Mar tu an 1 A N in t r : rml . S S ks 888 G : M ME S. Lecto ype selected he e , ah , g ; ; AL

m n n rm D T HAN 2 r a i JO SS. 1 9 7 . H . e e tfo e ( AHLS . ) ( p

: H s erra t ons . r ema nenti rme DAHL ST . 1 3a . Bas . [ ifi ] ssp fo ( 89 p

’ r i TEN TR Ki nn l k kl : H d s ti nctum S S . V . : eku l e Mor e ef9 . 7 . 1 890 Lectotype selected he e ; g , ; ’ ’

A. FRYXELL in . r a a a H r m n n i rm DAHL T Bidr . I I S On the sheet the e is lso l bel e a e tfo e S . 2 ’ K . JOH . 1 5 Determ. 9

Note : This species is only known from the type collection preserved on two sheets in H r S. r and ri r r . A r n oweve , the o igin status of this gathe ng is athe confused cco di g to

r t l n k n ri n n kl o o o ue b A. FRYXE LL MOrke ef the p g , the o ly now gathe g was do e y at on

Mt . H unneb er H r n an m r m a . b g oweve , the o ly ext t ate ial is acco panied with l els ’ ’ i n i MOrkeklef a on M Ki nn ekulle . b are dicat ng that is instead situ ted t. These la els ’ t n i n D DT r AHLSTE S a r b ar e m i . u r w it e h nd (the o iginal la els iss ng) Th s , it is clea that D H has n m n r n rotolo ue r A LSTEDT see these speci e s , they fit the desc iptio in the p g ve y ’ ’

and r are no r kn n ri n b FRYXELL r m MOrkeklef . S well the e othe ow gathe gs y f o till , r b r b D H f m b r r howeve , the la els w itten y A LSTEDT identi y the as elonging to a athe ff r n i tin TR DAHL T w fir in i . e . H d s ctum STENS . S . r as di e e t Species , ( ) , and it appea s as ifit st 1 925 that JOHANSSON put them i n conn ection with the protologue of I m r n H r ema nenti or me . f t is difficu lt to tell for sure how this co plexity has a ise . H r m n are n a ma r oweve , the lectotype speci e s the o ly ext nt te ial of this Species and beyon d doubt they belong to the same gathering as the ones mention ed in the protologue and are thu s available as lectotypes even though it appears likely that it r u a DAH r r n a was othe d plicates th t LSTEDT had at hand when w iting the desc iptio , th t these original specimens have vanished and that the now extant material has been

S r n D H r added to late whe A LSTEDT had fo gotten their status .

N w l 44 m . 2006 C o e s . 8 8 p ,

m HANSS DAHL T l a s a rs i den s s s . s ubci n erell u JO . S . 893 . H . p p ex ( p

’ Lectotype selected here : Suecia : Smalan d paroeciae Forserum ad Lattarp i n pratis ’

m r i 1 9 1 . K . H N N Ex . D H H rb . Hierac . S . I 2 in u b os s 8 J : 7 S. , , leg O A SSO ( A LSTEDT, e cand )

i n rme STENSTR . DAHL T . Note : This combin ation is a taxonomic synonym ofH . fo ( ) S

‘

h OHAN AM . 1 92 . s teen o ii J SS. S 0 H . fi ( p

’ r tm lan L u nar ar n D ammr nin n D : Vas an d: s sb s s . o e at . Lectotyp e selected he e j g , j g ; ’

1 1 I n arrrl are: K TEENHOFF . S k m Ex . J H N N AMUELSSON Hi erac . VI I . 0 . . 6 s S S ; toc hol ( O A SSO ,

m ri n rr n n exs i ccata no . 59 n b u scand . ; with e e ded p ted la el with the locality e o eo sly give ” ” as Fjallb erget and the date as i n S.

h m AM 1 24 JOHANSS. S . 9 . H s tibeop yll u DAHLST . ex ( p

’ ’ 2 Ex N r : r d m b r 1 9 0 . J H N Lectotyp e selected here : H all . Ulla ed F i he s e g ( O A SSO

i i n 2 m n r n a in . SAMUELSSON H erac . . exs ccata o . 90 b S , scand ; with e e ded p i ted l el)

DAHLST . 1 . XI I I : 56 var. con turbans JOHANSS. H s ubatron i tens (DAHLST. ) ( 908 exs X )

1 24 . SAM . ( 9 p

’

m : Ekomahult 1 922 . K . JOHANSSON Ex . Lectotype selected here : Smal . Fe sjo ; leg ( 2 m n r n J H N N SAMUELSSON H i erac . n . exs i ccata . 93 O A SSO , sca d no ; with e e ded p i ted label) in S.

ff r n r a r ir um r b : n r r t n str. as b Note This taxo is athe di e e t f om H s ub oni te s S . c c sc i ed y H r n r a r au r and r b b i n own r . l te tho s it is p o a ly a species its ight oweve , si ce the o iginal material of H s uba tr on itens this far has been sought for in vain it has yet not been possible to fix the u sage of this name and as long as this has not been done the status

’ ’ contur n r ma n in of ba s will e i uncerta .

r m D u DAHLST . 1 89 . XI : H . s bi ri u u AHLST . 9 g ( ) ( , exs

4 . r i r r i um DAHLST . in P 1 89 . : H mu orum s s . s ub u Bas p g , ORAT ( p

’ t r : Su : Ves tmanland ad Ar b a in r Lecto yp e selected he e ecia og Silva loco ape to , ’

1 2 . . V in 89 . O . ON P Ex . D H H rb . Hierac . S . I V: 39 S. leg C ORAT ( A LSTEDT, e cand )

H . u o r n DAHLST . OHANSS. 1 27 . s b bs cu a s (DAHLST . ) ex J ( 9 p

' H rr 4 ti ons . u ura ns DAH LST . i n P 1 89 . . : . s e a s bobs c Bas fi ssp , ORAT ( p

’ Lectotype selected here : Suecia : Ves tmanland in s ilvati ci s ad Sands kogshagen prope ’ K 4 in 1 8 1 . V N P H r Hier c . I : 0 un or 9 . x . C . O O E . D H b . a S gs , , leg ORAT ( A LSTEDT, e cand )

S.

Note : I n my opinion the type belongs to the earlier described H s uder ma nn i cum

DAHLST M M r n b r . but mu ma r r rr H s ubobs cu a s AL E , ch of the te ial efe ed to y late C w 44 2 N e s l . 006 om . p , 8 9

authors are prob ab ly better referred to H s tens tr oemi i D AH LST .

r m AN s ubte s cis s u JOH SS. 1 900 . H . ( p

’ Lectotype selected here : Suecia : Vestmanland loco humidius culo in prato s i lvati co ad h ’ rrv r und ar. r K . J H N N EX. D HL H rb . To a s G t tte 1 898 . p p y y , , leg O A SSO ( A STEDT, e n XI I 4 S . : 9 in . Hi erac . ca d ) S

s ubtri an ul are STENSTR . DAHLST . 1 892 . I H . g ( ) ( b exs :

v m r : H s il a ti cu . s ubt i a n la re TENSTR . 1 88 . Bas . ssp g u S ( 9 p

’ ’ r FRYXELL in S. t : Dls l : D als ko s s zn H n 1 8 89 A. Lecto ype selected he e g , ede ; ;

m r H r r M n r n r t i a n ula e A . NORRL . a d a e b Note : This taxo is ve y Si ila to . g ( L O ) they est i treated as conspec fic .

H . s uderman n i cum DAHLST . M M in M M 1 896 . AL E , AL E p

’ ’ A N M M r : Smi . St. Malrn Brannkarr . . 1 890 . . G O . : . Lectotype selected he e , d Leg AL E in S.

n r m AN in ARWI D N 1 2 H . ta a odei u JOH SS. SSO 9 8 . , ( p

’

EU NI K H nu . r rml ann k 2 / 1 0 . Lectotype selected he e : Sode . L a bru . 9 6 90 GE E O LER; dete ’

K OH . 1 27 in S. . J 9

toerens e OHANSS. 1 925 . H . J ( p

’ ’ k 1 2 K OHANSSON in m: R nni barrs o 30/7 9 7 . . J r : Srm . Sa Lectotype selected he e le o nge, i g

4 turbi n i ce s DAHLST . 1 89 a . H . p ( p

’ Lectotype selected here : Suecia : Upland ad D ann emora in collibus et s copuli s locis ’

i r an . H rb . H e ac . S Ex . D H s icci oribus in r s ilvatico 1 890 . p ato , , legi ipse ( A LSTEDT, e c d

11 : 56) in S.

XI ] urti cae ons AHLST . DAHLST . 1 900 . : H . fr (D ) ( exs

r LST 1 l . : H i l va ti cum . ur i ca e ons DAH . 89 b Bas . s ssp t f ( p

’ ’

D H in S. 1 885 H . r : r : K r n Onn e en 7 . Lectotype selected he e No ge istia ia, ; ; A LSTEDT

AM 1 2 . ven et oli um JOHANSS. S . 9 0 H . if ( p

’ 1 5 I n s amlare 5 . . Lectotype selected here : D al arne : Granges b arg; D at . VII ; ’

AM 1 1 7 in S. K . OH . . S . 9 K . G STEENHOFF . S k m J . toc hol ; det

vill a n n e HAN S. 1 1 4 . H . tti g e s JO S ( 9 p

’

Ex . HANSSON . K . JO M a s zn r 1 899 . r : Sdm. Lectotype selected he e ellos , Nyto p; ; leg ( w N l . 44 200 C om . e s 6 9 0 p ,

1 m ri n in i i a ri Ex iccata no . b JOHANSSON H erac Rario a s 00 S. , ; with e ended p ted la el)

di v m OHANSS DAHLST . 1 4 vi olas cen s var. ba o i olaceu J . 89 a . H . ex ( p

’ m zn La rr an 1 r ma . rs e ttar : S o ru s s 888 . Lectotype selected he e l F , p (to lov g) ’

K J H N N in S. . O A SSO

m n a n a n mi n m H u i c us is OHAN SS Note : This co bi tio is a t xo o c syno y of p b p J .

h r m HAN AM 1 2 . s tO o u JO SS. S . 9 0 30 H . xy p ( p )

’ 2 ml n n r 6. . 1 9 I ns a are Lectotype selected here : Vs m: Ljus arstorp s s ; Fjallbe get; Dat . VI ; ’

K . STEENH OFF in S.

” Note : The parish Ljus nars torp given on the label does not exist and is certainly an ” ” rr r f r n r m n n in rotol e o o Ljus arsbe g which is the site e tio ed the p ogue .

Acknowledgement

This study is part of the project The H awkweeds of Sweden foun ded by Th e

Swedish Species I n itiative and Gyllens ti ems ka Krapperups stiftels en . 44 2006 C m New l . o . s p , 9 1

R efer en ces

i dn in i l i - A M UI E . 1 955 . S r sb o o s k k mm n r ar H i r k e a i um artor. L Q ST, p g g o e ta till ett p c en kB t — Sv s o . Tids kr: 49 : 1 70 1 80 .

' ' 8 1 d r z r A M UI S . i 8 Stu e O e s lci tet H i er a ci um. k lm S . L Q ST, l fi g toc ho

ARWI DSSO N T 1 2 Hi r i k — . . B t. N i r 2 9 8 e ac olo is a bidra . o t 2 2 , g g o s e 1 9 0 .

i d i - ENNE M . 1 892 . S r da b dra kannedom af n an H r i e a i um rm r I . BR R, p g till Fi l ds c fo e

M dd — l . Soc . F a una F l . F enn . 1 : 7 1 1 e e 8 5 3 .

M 1 ri i - NNE . 895 . S dda b dra kann edom om n an H r rm i e a i um r I V. BRE R, p g till Fi l ds c fo e A F a una F l . F enn . 1 3 n . 1 . cta Soc .

D HL H . 1 89 1 . H iera ci a icca ta Ex i exs I V. s ccata A STEDT, a ( )

m n a Ber i an ka Tr l D H H . l 89 l b . O a r s ad arden od ade n a r m r A LSTEDT, g i g g y elle ind e k n hi r i r A H r ta ti Ber . 1 . 7 . a da e ac e . c o g n

D H T H 1 2 bidra kannedomen m k n H i ra i mfl r . . 89 a . a r o S a e c u o a A LS EDT, N g a g till es

— B t. Not er 1 4 1 o i s ( l 5 66.

D HLS H 1 2 H r r i um H i r i m n - H Ex i 89 b . e ba e a c oru Sca dina via e I I . s ccata A TEDT, . ( )

H H Bi r i - D . 1 893a . dra s dostra S H eraciumflora I I . Sv. Ve t. Ak. A LSTEDT, g till y ve iges

25 . 3 . H a ndl . n

- D H H . 1 9 H er r i um H i era i o m c ndina via e I V V. E e 8 3b . ba c ru S a A LSTEDT, ( xsiccat )

- D HL H 1 4 Bi d stra ri Hi raciumfl ora I I I . v. Vet. Alc H a ndl . . 89 . dra s O S e S A STEDT, g till y ve ges

3 . 26 n .

Exs iccata D HL H 1 H r a r i um H i er i rum ca ndi na vi a e I X. 895 . e b a c o S A STEDT, . ( )

D H S H 1 H r a r m H i r i m ca ndina via e . Exs iccata 896 . e b i u e a c oru S A L TEDT, . X ( )

Exs iccata DAHL H 1 r r r m n i n ia XI . 899 . H e ba i um H i e a ci oru Sca d av e STEDT, . ( )

— Ex i ccata D H H 1 H r r H r m ndin ia e XI I I I I . s 900 . e ba i um ie a ci oru Sca a v A LSTEDT, . X ( )

Ex i ccata D HLS H 1 2 H r r i r m ca ndina via e XI V. s 90 . e ba i umH e a ci oru S A TEDT, . ( )

- Exs iccata D H r r r m n i n i a e I I )Q GI I . HL . 1 908 . H e ba i umH ie a ci oru Sca d av A STEDT, XX ( )

D H H 2 mellan s ven s ka H i er a c i a Silva ti c i or mia . 1 9 1 . A LSTEDT, . Nya syd och f r A k. Bot. 1 7 n . 2 f. . 2 2 n . . Ark. Bot. 8 N . li n Dr H . E . H N Hi eracia Venn landica e . J F T. 1 936 . o , a O A SSO collecta f i E r m N rma le XI I . Exs ccata FRI E . 1 847 . H er ba i u S , o ( )

Archi eraci er ran D m Vastman lan d D als land . J H N N K . 1 900 . O A SSO , Nya f ala e , och

- 2 I I I 11 . 7 . Bih t Aka d. H a ndl . 5 : . Sv. Ve N w l . 44 9 2 C om . e s 2006 p ,

N K 1 2 Archieraci umfloran inom Dalarnes s ilur mr éi il an tr k H N . 90 . o de S s a ten . J O A SSO , i j

- n l 2 11 V . v. et. Aka d. H a d 8 : 1 . 7 . Bih. S n

N a r bidra kanned men om Hiera iumfl r r S . O HANSSO K . 1 905 . o c o an o r J , N g a g till i s d a ve ige

— B N ti er 97 1 28 . ot. o s

Bidr a kannedomen om as trikl ands Archiera iumfl r B O HAN N K 1 07 . ot. J SSO , . 9 g till G c o a .

- Notis er 1 1 9 .

H N N K 1 1 4 D ia os er an rnarknin ar n a a s all ntare v n k J . 9 . n r s s e s a O A SSO , g och g till g y N r - Hieraciumfonn er. Bot. otis e 65 87 .

H i era ci a Silva ti ci ormi a r n S r la land . Bot. N ti er O HANSSO N K . 1 920 . a J , Nya f f ve iges g o s

- 1 65 00.

°

Na1 a Hieraci d tockholm strakten . ve kB t. Ti ds 1 O HANSSO N 1 2 . r umf n 9: J , K . 9 5 g y i S S ns o kr.

4 -4 85 89.

AN SSO N K 1 2 Enum erantur H i era ci a vul a ta ueci cae . Ark B t 2 1 An 1 . O H . 7 . . . 5 J , 9 g S f o .

- ANSSO N K . 1 2 N a mellan v nska Hi er cier. ven kBot. I i 2 2 O H e a s d skr. J , 9 9 . y s S 3 : 3 9 335 .

L N 2 J HAN N K . G SAMUE SSO 1 9 0 . H ieraciumfl oran Vas tmanlan d . Ark. Bot. O SSO , i f 1 n 1 4 6 . .

N — 2 H i era cia s ca ndi na vi ca exs i cca ta . J O HANSSO K . G SAMUELSSO N 1 923 6. , Uppsala

kh m Exs iccata Stoc ol . ( )

D a la r na H i ra i lva t i rmi a . z J O HANSSO N K . AMUE LSSO N 1 23 . e a i i , G S 9 s c S c fo Leip ig .

O HANSS N L N 4 v n Ar hi i r J O K . G SAMUE SSO 1 92 . s e ska c erac er. Bot. Notis e , Nya

— 1 35 1 60.

I ND . 1 2 H r ka r I r J 88 . i e a ci olo is bid a . G b . L EBERG , C . g g ote o g

LO NNR TH Berattels e m en b otani s k r O r Sm . O K . J . 1 882 . a a , o esa i st a land och p Gotland

-A r r v. ka d. O e s . 39 . S Vet. fi

MA M N i mfl r AND S 1 890 . Bidra oderman lands Hi erac u o a . G O . S L E , ( ER SO g till

— Bot. Notis er 4 88 9 .

M A M AN r G O . S N 1 89 1 N a bidra Soderm anlands H ieraci umflo a . L E, ( DER SO . y g till

— Bot. N l is er 1 1 o 78 8 1 .

AN N 1 i r i mfl r I I MA ME O . l G 896 . b idra Soderman an ds H e ac u o a . L , ( DERSSO Nya g till

B r - Notis e 1 57 1 1 . ot. 7

N R R LI N Bidra H i er a i u - fl r k in i k f n m r O J . P. 1 888 . m o an s and av s a a o , g till c i h l s elle sta ar A a F t Soc . a una F l . F enn . 111 n . 4 . del . c

N RR L I N H ra I H r r 1 2 H in r . O J . P. 1 8 89 . i u i e i m . n : e ba i um Mus ei F enn ci . . , [ c ] , ed els gfo s

P V — O . O N 1 894 Kun n H i r i r B r 4 4 . . . s ortrakte s e ac e . ot. Notis e 33 ORAT, C g N ews l . 44 2006 C om . p , 9 3

ENNI K O V A. . 2 T ifi ti on m H i era c i um A r S 003 . ca n , N yp ofso e ( ste aceae) ames pub lished

i An B t F i 4 2 — 2 b S au r . 1 n o . enn ci 0: 9 3 1 . y wed sh tho s .

N T OM K . . E 1 r S O . 889 . Va mla nds ka Archi era c i er . a TE S R , Ups la . ' H a h r r T . 1 998 . s s och kli bbl o i s od a GOta la nd a lla ilda a r ter s om YLER, T g og ppfi v ‘ u i vi ts dn Shri ne H a lla nd Bleki n e Ola nd och Smcila nd s a mt en lis ta ppg fi , , g ,

Over atr a a de a dven ti v a r t 4 a er. 6 . ub as u m n LBF p fi ( pp , p lished s pple e t to

Medlems la d 1 2 b 998 z ) .

m n . 2000 . r and lectot ificati on H i ra i u S e m . TYLER, T No e clatu e yp of wedish c species

— r J . B ta n 20 3 1 . No d. o y 9 03

2 2 Os ter Otlan d s k fi l r H i r r . 00 a . s s i u H o bb o e a m . i e a i um in TYLER, T g g [ c sect c the

r n Of s ter tl n n B r — O O a d S ot. Notis e 1 52 . p ovi ce g , wede ]

2 2 . n k . H r 00 b G a S o s fibblor H i e a i um . H i era i um and TYLER, T otla ds g och g [ c sect c

Vul a ta in r in an n Ri 22 2— . di G S . n . 3 . sect g the p ov ce of otl d, wede ] No

2 ter otland ha fi l r H i era m a in th r n T. 003a . s i V t O s b o . u . ul a e TYLER, g g [ c sect g p ovi ce of

r n B r — Notis e 1 36 . O o S n ot. ste g tla d, wede ]

2 3 Va t r l k H fi l r r H r 00 b . s e Ot an ds S o s a bb o H i e a i um . i e a i um TYLER, T. g g och g ( c sect c

- Ca ll una 20 no . 2 3 u l . Vulga ta ) . S pp

ko ha fi l r te or s omr adet B hu s lan D als lan d L . 2004a . S s bb o GO b o TY ER, T g och g i g , och r r n r r n d H i e a i um and . Vul a ta in G bu an [H i e a ci um sect . c sect g the othe g egio ‘ — r n D l l n and B hu lan Vr i vrci n e 22 : 1 6 9 . the p ovi ces a s a d o s ] . g

2 4 Lecto ificati ons n m H iera ci um kn n r m . 00 b . t TYLER, T yp of a es ofspecies of ow f o the

F 4 1 — 3 Ann . Bot. enn . : 79 8 . Swedish provinces Gotland and Ostergotland .

— 2 H kfi l r M l l ka en D a hne 1 6 2 1 03 . 005 . o bb o a ar ands . TYLER, T. i p p

W UN 1 B r nn m n m Ronneb tra ktens a una och G 890 . id a ti ll kd edo e o ESTERL D, C . g y f

F l ra . tockh hn o S o .

- 2 z . P nz nrei ch I V: 80 . I n : A. EN D as a e Z HN K . H . 1 92 1 1 2 H iera ci um . 9 3 . A , GLER, fl Leip ig

1 - 2 — 4 1 2 1 1 1 4 — 1 705 1 . 7 576 9 1 . 577 1 1 6 9 (pp . , pp , pp m New l 44 2006 Co . s . 9 4 p ,

New taxa and combinations published in this is sue

Antillanthus . . en. . : . 5 1 B NORD , g nov p

mb . . : . 2 Antillan thus acun ae BORHI DI . N . c 5 ( ) B ORD , o nov p

l ll M m An lanth us a mironci o MAZ G E . N . c b . nov . 2 til . : 5 ( A O Z) B ORD , o p

A N m . 2 Antillan thus a ulens is I . . c . : . 5 z ( LA ) B NORD , o b nov p

mb . nov. . 53 Antillan thus bi seria tus A I N . . c : ( LA ) B NORD , o p

mb . v. : . 3 Antill anthus carinatus GREENM. . . co no 5 ( ) B NORD , p

m . Antillan thus cub ens is GREENM. . . b . . : 53 ( ) B NORD , co nov p

m . Antillanthus ekmami A I N . ORD. b . : . 53 ( LA ) B N , co nov p

An anth us er r hu REENM. . . c m . n v : . till i oca s G b o . 53 p ( ) B NORD , o p

le M m v Antillanthus ucole is GREEN . . N . c b . no . : . 54 p ( ) B ORD , o p

mb . . : . 54 Antillanthus moaens i s A I N . . c ( LA ) B NORD , o nov p

m . n v. : . Antillan thus moldenkei GREENM. A I N . N . c o ( LA ) B ORD , o b p

m n v 54 An tillan thuS ach le is GR EENM. . . co . o . : . p y p ( ) B NORD , b p

m . . : . 4 Antillantlzus ach odus GREENM. . ORD. c b 5 p yp ( ) B N , o nov p

A I B RD c mb . . : . 4 An an thu s u ii N . till s a et O . 5 g ( LA ) N , o nov p

mb . n v. : . 55 Antillanthus s ha eri GREENM. . . c o f ( ) B NORD , o p

mb . nov. : . 5 Antillanthus s ubs uarrosus GREENM. . N . c 5 q ( ) B ORD , o p

m . nov. . Antillanth us tri chotomus GREENM. . N . c b : 55 ( ) B ORD , o p

mnu . en . . : . 26 Can ari otha s . N B ORD , g nov p

m . . : . 28 mn a T R B . c b Canariotha us h ermos e PI . N ( A D) ORD , o nov p

m . : . 27 Canariothamn us almens i s NEES . N . co b . nov p ( ) B ORD , p

m r c la . S . : . 27 Can ari oth a nus u i o . . p B NORD , p nov p

66 Elekmani a . N . en. . : . B ORD , g nov p

mb . . : . 68 Elekmania barahonens is B. . . (UR ) B NORD , co nov p

m . . : . 69 B . Elekmania ba chi t . . b (UR ) B NORD , co nov p

69 m . nov. : . B . Elekmania uertes ii U . B. f ( R ) NORD , co b p

N w l 44 C om . 2006 9 e s . 6 p ,

V DK m . nov. : . 8 a a a mi n uta A P ELSER VEL . o J cob e (C ) , c b p

m 1 o . nov. : . 2 J a coba ea moll i s (WI LLD. ) B . NORD. , c b p

m : . 1 J a coba ea m ultibracteolata . J FF . . HE N . . o . . 3 (C E REY Y L C ) B NORD , c b nov p

- m J a cobaea n udi a ulis U H HAM . ex D . DO N . . o . nov. : . 1 2 c (B C ) B NORD , c b p

m . nov. . J acobaea ers oon u D E . PELSER o : 9 p ( NOT ) , c b p

m 1 r h n li a W . D . J a obaea a a i o C . . . . : . 3 c p f ( ALL ex ) B NORD , co b nov p

n W m n J a oba ea re a rdii . I NK . . . o . ov. : . 1 3 c (C L ) B NORD , c b p

m . nov. : . 9 J a oba ea s ubal ina K H PELSER VELDK . o c p OC ) , c b p

m 1 J a oba ea tibeti a H K . f. . . b . . : . 3 c c ( OO ) B NORD , co nov p

AL ELDK m n v J a oba ea un i lora L. V . . o . : . 9 c f ( ) , co b p

m . nov. : 1 2 J a oba ea vul a ris s ub s . otl an di ca EDM . . . co . c g p g (N ) B NORD , b p

Leon is . . en . . : . 55 B NORD , g nov p

Leon is tri n ca ra GRI SEB . . om . nov. : . 56 ( ) B NORD , c b p

Lun di n ia . . en . nov. : . 64 B NORD , g p

m . . : 66 Lun di n i a l umbea GRI SEB . . o . p ( ) B NORD , c b nov p

M n l n n v o o u us . . e . o . : . 38 c B NORD , g p

M n r m . nov. : . 40 o ocul us h os e oides D . . co y ( C) B NORD , b p

Mon o ul us mo u U M f . m ns tros s . . . b . . : . 39 c (B R ) B NORD , co nov p

Ne hrotheca KALLE n n RSJ O e . ov. : . 33 p B . NORD. , g p

N hr h m . nov. : . 33 e ot eca ili i oli a L . . . KAL LERSJ O o p cf ( ) B NORD , c b p

N m l es a e os . . en . nov. : . 58 p B NORD , g p

N m l os al ain i i I MENE ALM m n 2 es a e J . . . b . ov. : . 6 p (J . Z ) B NORD co p

N h n m n es am elos ottea a . E K M 9 o . ov. : . 5 p (URB .) B . NORD. , c b p

Nes am elo s l u ens P I . in LAM m o . nov. : . 59 p c ( O R . ) B . NORD . , c b p

N orden s ta mi a LUN I N en . . : . 1 5 D , g nov p

N r mi a ca amar n is R B TR E m 20 o dens ta H . ce s . O . CUA C . . . o b . . : j ( ) B NORD , c nov p

Nor m m 2 dens ta i a car is hen s is CUATREC . . . . . : . 0 p ( ) B NORD , co b nov p

Nord n m m . n v. 20 e s ta i a abris u . . o : . f (CABRERA) B NORD , co b p News l . 44 2006 C om . p , 9 7

m dens ta i a h uan ucona CUATREC . . c m n v : 20 r o . o . . No ( ) B NORD. , b p

mi n n n Nordens ta a u i e s is H . R O B. CUATREC . . . om . nov. : . 20 j ( ) B NORD , c b p

de ta mi a ki n R B ATR E m 2 Nor ns u H . O . C U C . . . o . nov. : . 0 g ( ) B NORD , c b p

m m i a li on ens is . . . . m . . 21 Nordens ta : . (B NORD ) B NORD , co b nov p

m n r m . nov. . 21 Nordens ta i a lo i s l a GR EENM . CUATREC . . . : g g ( ) B NORD , co b p

r n m a n H A R B v 22 No de s ta i as coe s i s . N H . O . . . om . no . : . p ( BELTR ) B NORD , c b p

m r n m n v Nordens ta i a e a da W . UND I N o . o . : . 1 6 p ( EDD ) L , c b p

m ma hi a n ATRE m n v . 22 Nor den s ta i a ri c a CU C . . . . o . : ( ) B NORD , co b p

m . n ov. : . 22 n mi a il R EENM ATR E . . Norde s ta a s tell to os a G . CU C . p ( ) B NORD , co b p

m . nov. : . 22 r n t mi a va ra H . R B. UATR EC . . . o No de s a ta ( O C ) B NORD , c b p

m ATRE . m nov. . 22 Nordens ta i a tues ta e CU C . . . : ( ) B NORD , co b p

m . nov. : . 22 mi a w k w ki i UATREC . . . o Norden s ta oyt o s (C ) B NORD , c b p

41 en . nov. : . Norli n dhi a B . NORD. , g p

m n v 42 Norlin dhi a am l e ten s HAR . . . o . o . : . p c ( V ) B NORD , c b p

r n r . . S . nov. : . 43 No li dhi a ap te a B NORD , p p

m n . 42 Norli n dhi a brevi r adi ata T. . . . o . ov. : ( NORL ) B NORD , c b p

m n : . 45 Oli ocar us b urchelli i H K . f. . . o . ov. g p ( OO ) B NORD , c b p

Z emi s i a . . en . nov. : . 71 B NORD , g p

2 W m . nov. : . 7 Z emi s i a dis ol or S . . b c ( ) B NORD , co p