Information Sheet on Ramsar Wetlands (RIS) Categories Approved by Recommendation 4.7, As Amended by Resolution VIII.13 of the Conference of the Contracting Parties
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Descent of Man
THE DESCENT OF MAN, AND SELECTION IN RELATION TO SEX, BY CHARLES DARWIN, M.A., F.R.S., &c. IN TWO VOLUMES.—VOL. II. WITH ILLUSTRATIONS. LONDON: JOHN MURRAY, ALBEMARLE STREET. 1871. [The right of Translation is reserved.] ERRATA. CONTENTS. PART II. SEXUAL SELECTION-continued. CHAPTER XII. SECONDARY SEXUAL CHARACTERS OF FISHES, AMPHIBIANS, AND REPTILES. FISHES : Courtship and battles of the males — Larger size of the females — Males, bright colours and ornamental appendages; other strange characters — Colours and appendages acquired by the males during the breeding-season alone — Fishes with both sexes brilliantly coloured — Protective colours — The less con- spicuous colours of the female cannot be accounted for on the principle of protection — Male fishes building nests, and taking charge of the ova and young. AMPHIBIANS : Differences in structure and colour between the sexes — Vocal organs. REP- TILES : Chelonians — Crocodiles — Snakes, colours in some cases protective — Lizards, battles of — Ornamental appendages — Strange differences in structure between the sexes — Colours — Sexual differences almost as great as with birds .. Page 1-37 CHAPTER XIII. SECONDARY SEXUAL CHARACTERS OF BIRDS. Sexual differences — Law of battle — Special weapons — Vocal organs—Instrumental music — Love-antics and dances — Deco- rations, permanent and seasonal — Double and single annual moults—Display of ornaments by the males .. .. .. 38-98 vi CONTENTS OF VOL. II. CHAPTER XIV. BIRDS—continued. Choice exerted by the female — Length of courtship — Unpaired birds — Mental qualities and taste for the beautiful — Preference or antipathy shewn by the female for particular males — Vari- ability of birds — Variations sometimes abrupt—Laws of varia- tion — Formation of ocelli — Gradations of character — Case of Peacock, Argus pheasant, and Urosticte . -
00 WNAN 2007 New PAGE Template
Western North American Naturalist 81(1), © 2021, pp. 54–62 Cold tolerance of mountain stoneflies (Plecoptera: Nemouridae) from the high Rocky Mountains SCOTT HOTALING1,*,†, ALISHA A. SHAH2,†, MICHAEL E. DILLON3, J. JOSEPH GIERSCH4, LUSHA M. TRONSTAD5, DEBRA S. FINN6, H. ARTHUR WOODS2, AND JOANNA L. KELLEY1 1School of Biological Sciences, Washington State University, Pullman, WA 2Division of Biological Sciences, University of Montana, Missoula, MT 3Department of Zoology and Physiology and Program in Ecology, University of Wyoming, Laramie, WY 4Northern Rocky Mountain Science Center, U.S. Geological Survey, West Glacier, MT 5Wyoming Natural Diversity Database, University of Wyoming, Laramie, WY 6Department of Biology, Missouri State University, Springfield, MO ABSTRACT.—How aquatic insects cope with cold temperatures is poorly understood. This is particularly true for high-elevation species, which often experience a seasonal risk of freezing. In the Rocky Mountains, nemourid stoneflies (Plecoptera: Nemouridae) are a major component of mountain stream biodiversity and are typically found in streams fed by glaciers and snowfields, which are rapidly receding due to climate change. Predicting the effects of climate change on mountain stoneflies is difficult because their thermal physiology is largely unknown. We investigated cold tolerance of several alpine stoneflies (Lednia tumana, Lednia tetonica, and Zapada spp.) from the Rocky Mountains, USA. We mea- sured the supercooling point (SCP) and tolerance to ice enclosure of late-instar nymphs collected from a range of thermal regimes. SCPs varied among species and populations, with the lowest SCP measured for nymphs from an alpine pond, which was much more likely to freeze solid in winter than flowing streams. -
From Characters of the Female Reproductive Tract
Phylogeny and Classification of Caraboidea Mus. reg. Sci. nat. Torino, 1998: XX LCE. (1996, Firenze, Italy) 107-170 James K. LIEBHERR and Kipling W. WILL* Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract ABSTRACT Characters of the female reproductive tract, ovipositor, and abdomen are analyzed using cladi stic parsimony for a comprehensive representation of carabid beetle tribes. The resulting cladogram is rooted at the family Trachypachidae. No characters of the female reproductive tract define the Carabidae as monophyletic. The Carabidac exhibit a fundamental dichotomy, with the isochaete tri bes Metriini and Paussini forming the adelphotaxon to the Anisochaeta, which includes Gehringiini and Rhysodini, along with the other groups considered member taxa in Jeannel's classification. Monophyly of Isochaeta is supported by the groundplan presence of a securiform helminthoid scle rite at the spermathecal base, and a rod-like, elongate laterotergite IX leading to the explosion cham ber of the pygidial defense glands. Monophyly of the Anisochaeta is supported by the derived divi sion of gonocoxa IX into a basal and apical portion. Within Anisochaeta, the evolution of a secon dary spermatheca-2, and loss ofthe primary spermathcca-I has occurred in one lineage including the Gehringiini, Notiokasiini, Elaphrini, Nebriini, Opisthiini, Notiophilini, and Omophronini. This evo lutionary replacement is demonstrated by the possession of both spermatheca-like structures in Gehringia olympica Darlington and Omophron variegatum (Olivier). The adelphotaxon to this sper matheca-2 clade comprises a basal rhysodine grade consisting of Clivinini, Promecognathini, Amarotypini, Apotomini, Melaenini, Cymbionotini, and Rhysodini. The Rhysodini and Clivinini both exhibit a highly modified laterotergite IX; long and thin, with or without a clavate lateral region. -
Ent21 3 229 256 Sharova.Pmd
Russian Entomol. J. 21(3): 229256 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2012 Æèçíåííûå ôîðìû è àäàïòèâíàÿ ðàäèàöèÿ ëè÷èíîê æóæåëèö (Coleoptera: Carabidae) ìèðîâîé ôàóíû Life forms and adaptive radiation of ground beetle larvae (Coleoptera: Carabidae) of the World fauna È.Õ. Øàðîâà, Ê.Â. Ìàêàðîâ I.Kh. Sharova, K.V. Makarov Ìîñêîâñêèé ïåäàãîãè÷åñêèé ãîñóäàðñòâåííûé óíèâåðñèòåò, êàôåäðà çîîëîãèè è ýêîëîãèè, óë.Êèáàëü÷è÷à, ä. 6, êîðï. 5, Ìîñêâà 129278, Ðîññèÿ. E-mail: [email protected]; [email protected] Moscow State Pedagogical University, Department of Zoology & Ecology, Kibalchicha str. 6, build. 5, Moscow, 129278 Russia ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: ëè÷èíêè, æóæåëèöû, ìîðôîëîãèô, æèçíåííûå ôîðìû, àäàïòàöèè, ñïåêòðû æèçíåííûõ ôîðì, ïàðàëëåëèçì. KEY WORDS: larvae, Carabidae, morphology, life forms, adaptations, life form spectra, parallelism. ÐÅÇÞÌÅ. Ïðåäëîæåíà èåðàðõè÷åñêàÿ ñèñòåìà âî÷íû è ñâîäèëèñü ê îïèñàíèþ îòäåëüíûõ âèäîâ. æèçíåííûõ ôîðì ëè÷èíîê Carabidae ìèðîâîé ôàóíû Îáîáùåíèå ýòèõ ñâåäåíèé ñòàëî âîçìîæíûì ëèøü ñ íà îñíîâå ñèíòåçà îðèãèíàëüíûõ è îïóáëèêîâàííûõ ïîÿâëåíèåì ïåðâûõ ñâîäîê ïî ìîðôîëîãèè è ñèñòå- äàííûõ ïî èõ ìîðôîëîãèè, ñèñòåìàòèêå è ýêîëîãèè. ìàòèêå ëè÷èíîê Carabidae [van Emden, 1942; Jeannel, Íîâàÿ ñèñòåìà æèçíåííûõ ôîðì ëè÷èíîê Carabidae 1941, 1942; Larsson, 1941].  Ðîññèè ïåðâûå îïðåäå- îòëè÷àåòñÿ îò ïðåæíåé [Øàðîâà, 1981] ââåäåíèåì ëèòåëè ëè÷èíîê æóæåëèö áûëè ñîñòàâëåíû Çíîéêî äîïîëíèòåëüíûõ êàòåãîðèé.  ñèñòåìó âêëþ÷åíî 6 [1929], Ãèëÿðîâûì, Øàðîâîé [1954] è Øàðîâîé êëàññîâ è 45 ãðóïï æèçíåííûõ ôîðì, õàðàêåòðèçóþ- [1958, 1964]. Íàêîïëåííûå ñâåäåíèÿ î ëè÷èíêàõ ùèõñÿ êîìïëåêñîì ìîðôîëîãè÷åñêèõ è ïîâåäåí÷åñ- æóæåëèö ïîçâîëèëè èçó÷àòü èõ àäàïòèâíîå ìíîãî- êèõ àäàïòàöèé. Íà îñíîâå íîâîé ñèñòåìû æèçíåííûõ îáðàçèå. Âïåðâûå íåñêîëüêî ìîðôî-ýêîëîãè÷åñêèõ ôîðì ëè÷èíîê Carabidae ïðåäëîæåíà ãèïîòåçà èõ àäàï- òèïîâ ïî÷âîîáèòàþùèõ ëè÷èíîê æóæåëèö âûäåëèë òèâíîé ðàäèàöèè îò ìàëîñïåöèàëèçèðîâàííûõ çîîôà- Ì.Ñ. -
Coleoptera: Introduction and Key to Families
Royal Entomological Society HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS To purchase current handbooks and to download out-of-print parts visit: http://www.royensoc.co.uk/publications/index.htm This work is licensed under a Creative Commons Attribution-NonCommercial-ShareAlike 2.0 UK: England & Wales License. Copyright © Royal Entomological Society 2012 ROYAL ENTOMOLOGICAL SOCIETY OF LONDON Vol. IV. Part 1. HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS COLEOPTERA INTRODUCTION AND KEYS TO FAMILIES By R. A. CROWSON LONDON Published by the Society and Sold at its Rooms 41, Queen's Gate, S.W. 7 31st December, 1956 Price-res. c~ . HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS The aim of this series of publications is to provide illustrated keys to the whole of the British Insects (in so far as this is possible), in ten volumes, as follows : I. Part 1. General Introduction. Part 9. Ephemeroptera. , 2. Thysanura. 10. Odonata. , 3. Protura. , 11. Thysanoptera. 4. Collembola. , 12. Neuroptera. , 5. Dermaptera and , 13. Mecoptera. Orthoptera. , 14. Trichoptera. , 6. Plecoptera. , 15. Strepsiptera. , 7. Psocoptera. , 16. Siphonaptera. , 8. Anoplura. 11. Hemiptera. Ill. Lepidoptera. IV. and V. Coleoptera. VI. Hymenoptera : Symphyta and Aculeata. VII. Hymenoptera: Ichneumonoidea. VIII. Hymenoptera : Cynipoidea, Chalcidoidea, and Serphoidea. IX. Diptera: Nematocera and Brachycera. X. Diptera: Cyclorrhapha. Volumes 11 to X will be divided into parts of convenient size, but it is not possible to specify in advance the taxonomic content of each part. Conciseness and cheapness are main objectives in this new series, and each part will be the work of a specialist, or of a group of specialists. -
Appendix O19749
Oikos o19749 Gerisch, M., Agostinelli, V., Henle, K. and Dziock, F. 2011. More species, but all do the same: contrasting effects of flood disturbance on ground beetle functional and species diversity. – Oikos 121: 508–515. Appendix A1 Tabelle1 Table A1. Full species list representing the standardized number of individuals per species for the study sites Steckby, Woerlitz, and Sandau. Density expresses the proportion of species standardized abundances to total abundance. Macropterous = winged, brachypterous = wingless, dimorphic = both forms can appear with a species. Body size is the average of maximum and minimum values found in the literature (for references see below). Wing Reproduction Body size Species names Steckby Woerlitz Sandau Density Morphology Season In mm Acupalpus dubius 0.032 0 0.016 0 macropterous spring 2.6 Acupalpus exiguus 1.838 1.019 0.71 0.005 macropterous spring 2.7 Acupalpus parvulus 0.081 0.038 0.032 0 macropterous spring 3.6 Agonum dolens 0.032 0.038 0.081 0 dimorph spring 8.8 Agonum duftschmidi 14.966 2.755 0.016 0.025 macropterous spring 8.2 Agonum emarginatum 116.659 4.472 25.194 0.208 macropterous spring 7.2 Agonum fuliginosum 0.097 0.038 0 0 dimorph spring 6.7 Agonum lugens 0.177 0 0.081 0 macropterous spring 9 Agonum marginatum 0.371 0.075 0.113 0.001 macropterous spring 9.2 Agonum micans 19.502 4.208 23.71 0.067 macropterous spring 6.6 Agonum muelleri 0 0.019 0 0 macropterous spring 8.2 Agonum piceum 0.468 0 0.016 0.001 macropterous spring 6.4 Agonum sexpunctatum 0.032 0 0.016 0 macropterous spring 8.2 Agonum -
Holocene Palaeoenvironmental Reconstruction Based on Fossil Beetle Faunas from the Altai-Xinjiang Region, China
Holocene palaeoenvironmental reconstruction based on fossil beetle faunas from the Altai-Xinjiang region, China Thesis submitted for the degree of Doctor of Philosophy at the University of London By Tianshu Zhang February 2018 Department of Geography, Royal Holloway, University of London Declaration of Authorship I Tianshu Zhang hereby declare that this thesis and the work presented in it is entirely my own. Where I have consulted the work of others, this is always clearly stated. Signed: Date: 25/02/2018 1 Abstract This project presents the results of the analysis of fossil beetle assemblages extracted from 71 samples from two peat profiles from the Halashazi Wetland in the southern Altai region of northwest China. The fossil assemblages allowed the reconstruction of local environments of the early (10,424 to 9500 cal. yr BP) and middle Holocene (6374 to 4378 cal. yr BP). In total, 54 Coleoptera taxa representing 44 genera and 14 families have been found, and 37 species have been identified, including a new species, Helophorus sinoglacialis. The majority of the fossil beetle species identified are today part of the Siberian fauna, and indicate cold steppe or tundra ecosystems. Based on the biogeographic affinities of the fossil faunas, it appears that the Altai Mountains served as dispersal corridor for cold-adapted (northern) beetle species during the Holocene. Quantified temperature estimates were made using the Mutual Climate Range (MCR) method. In addition, indicator beetle species (cold adapted species and bark beetles) have helped to identify both cold and warm intervals, and moisture conditions have been estimated on the basis of water associated species. -
Coleoptera: Carabidae) Peter W
30 THE GREAT LAKES ENTOMOLOGIST Vol. 42, Nos. 1 & 2 An Annotated Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae) Peter W. Messer1 Abstract A survey of Carabidae in the state of Wisconsin, U.S.A. yielded 87 species new to the state and incorporated 34 species previously reported from the state but that were not included in an earlier catalogue, bringing the total number of species to 489 in an annotated checklist. Collection data are provided in full for the 87 species new to Wisconsin but are limited to county occurrences for 187 rare species previously known in the state. Recent changes in nomenclature pertinent to the Wisconsin fauna are cited. ____________________ The Carabidae, commonly known as ‘ground beetles’, with 34, 275 described species worldwide is one of the three most species-rich families of extant beetles (Lorenz 2005). Ground beetles are often chosen for study because they are abun- dant in most terrestrial habitats, diverse, taxonomically well known, serve as sensitive bioindicators of habitat change, easy to capture, and morphologically pleasing to the collector. North America north of Mexico accounts for 2635 species which were listed with their geographic distributions (states and provinces) in the catalogue by Bousquet and Larochelle (1993). In Table 4 of the latter refer- ence, the state of Wisconsin was associated with 374 ground beetle species. That is more than the surrounding states of Iowa (327) and Minnesota (323), but less than states of Illinois (452) and Michigan (466). The total count for Minnesota was subsequently increased to 433 species (Gandhi et al. 2005). Wisconsin county distributions are known for 15 species of tiger beetles (subfamily Cicindelinae) (Brust 2003) with collection records documented for Tetracha virginica (Grimek 2009). -
(Insecta, Coleoptera) В Фауне Арктики. Сообщение 1
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ, 2014, том 93, № 1, с. 7–44 ЭКОЛОГИЯ И ЗООГЕОГРАФИЯ УДК 595.76 ОТРЯД ЖЕСТКОКРЫЛЫХ (INSECTA, COLEOPTERA) В ФАУНЕ АРКТИКИ. СООБЩЕНИЕ 1. СОСТАВ ФАУНЫ © 2014 г. Ю. И. Чернов1, О. Л. Макарова1, Л. Д. Пенев2, О. А. Хрулёва1 1 Институт проблем экологии и эволюции им. А.Н. Северцова РАН, Москва 119071, Россия e(mail: [email protected] e(mail: oa([email protected] 2 Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, София, Болгария e(mail: [email protected] Поступила в редакцию 01.09.2013 г. Жесткокрылые, крупнейший отряд насекомых, в условиях Арктики уступают двукрылым первен ство в полноте освоения среды. На долю Coleoptera приходится около 13% энтомофауны тундровой зоны, однако несколько семейств жуков сохраняют в высоких широтах значительное видовое раз нообразие и существенную ценотическую роль. В этом сообщении мы даем обзор циркумполярной колеоптерофауны Арктики. На основе оригинальных данных, литературных сведений и фондовых коллекционных материалов с использованием экстраполяций и аналогий отмечены особенности таксономического и экологического разнообразия подотрядов, серий и семейств Coleoptera, про анализированы широтнозональное распределение и северные пределы распространения видов, специфика адаптаций и ценотических связей. Ключевые слова: Арктика, жуки, природная зональность, видовое разнообразие, ареал, адаптации. DOI: 10.7868/S004451341401005X Жесткокрылые (Coleoptera) – самый крупный основных широтных трендов параметров их раз отряд насекомых, включающий почти 386500 ви нообразия. дов (Slipinski et al., 2011). На его долю приходится Накопленные к настоящему времени данные почти 40% видов класса. Доминирование этого свидетельствуют о том, что в фауне Арктики на отряда в энтомофауне наиболее отчетливо в теп долю жесткокрылых приходится около 13% видо лых поясах, в тропиках и субтропиках. В умерен вого богатства насекомых, в северной полосе ном поясе жесткокрылые составляют треть видо тундровой зоны их доля снижается до 4%, а в по вого богатства насекомых. -
Vol 4 Part 2. Coleoptera. Carabidae
Royal Entomological Society HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS To purchase current handbooks and to download out-of-print parts visit: http://www.royensoc.co.uk/publications/index.htm This work is licensed under a Creative Commons Attribution-NonCommercial-ShareAlike 2.0 UK: England & Wales License. Copyright © Royal Entomological Society 2012 ROYAL ENTOMOLOGICAL SOCIETY OF LONDON . Vol. IV. Part 2 -HANDBOOKS FOR THE IDENTIFICATION / OF BRITISH INSECT-s COLEOPTERA CARABIDAE By CARL H. LINDROTH LONDON Published by the Society and Sold at its Rooms .p, Queen's Gate, S.W. 7 August I 974- HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS The aim of this series of publications is to provide illustrated keys to the whole of the British Insects (in so far as this is possible), in ten volumes, as follows: I. Part 1. General Introduction. Part 9. Ephemeroptera. , 2. Thysanura. , 10. Odonata. , 3. Protura. , 11. Thysanoptera. , 4. Collembola. , 12. Neuroptera. , 5. Dermaptera and , 13. Mecoptera. Orthoptera. , 14. Trichoptera. , 6. Plecoptera. , 15. Strepsiptera. , 7. Psocoptera. , 16. Siphonaptera. , 8. Anoplura. II. Hemiptera. III. Lepidoptera. IV. and V. Coleoptera. VI. Hymenoptera : Symphyta and Aculeata. VII. Hymenoptera : lchneumonoidea. VIII. Hymenoptera : Cynipoidea, Chalcidoidea, and Serphoidea. IX. Diptera: Nematocera and Brachycera. X. Diptera : Cyclorrhapha. Volumes II to X will be divided into parts of convenient size, but it is not possible to specifyin advance the taxonomic content of each part. Conciseness and cheapness are main objectives in this series, and each part is the work of a specialist, or of a group of specialists. Although much of the work is based on existing published keys, suitably adapted, much new and original matter is also included. -
An Annotated Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae)
The Great Lakes Entomologist Volume 42 Numbers 1 & 2 - Spring/Summer 2009 Numbers Article 3 1 & 2 - Spring/Summer 2009 April 2009 An Annotated Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae) Peter W. Messer Follow this and additional works at: https://scholar.valpo.edu/tgle Part of the Entomology Commons Recommended Citation Messer, Peter W. 2009. "An Annotated Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae)," The Great Lakes Entomologist, vol 42 (1) Available at: https://scholar.valpo.edu/tgle/vol42/iss1/3 This Peer-Review Article is brought to you for free and open access by the Department of Biology at ValpoScholar. It has been accepted for inclusion in The Great Lakes Entomologist by an authorized administrator of ValpoScholar. For more information, please contact a ValpoScholar staff member at [email protected]. Messer: An Annotated Checklist of Wisconsin Ground Beetles (Coleoptera: C 30 THE GREAT LAKES ENTOMOLOGIST Vol. 42, Nos. 1 & 2 An Annotated Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae) Peter W. Messer1 Abstract A survey of Carabidae in the state of Wisconsin, U.S.A. yielded 87 species new to the state and incorporated 34 species previously reported from the state but that were not included in an earlier catalogue, bringing the total number of species to 489 in an annotated checklist. Collection data are provided in full for the 87 species new to Wisconsin but are limited to county occurrences for 187 rare species previously known in the state. Recent changes in nomenclature pertinent to the Wisconsin fauna are cited. ____________________ The Carabidae, commonly known as ‘ground beetles’, with 34, 275 described species worldwide is one of the three most species-rich families of extant beetles (Lorenz 2005). -
Ecology of Alpine Carabid Beetles (Coleoptera, Carabidae) in a Norwegian Glacier Foreland, with a Special Focus on Claw Wearing to Indicate Relative Age
© Norwegian Journal of Entomology. 12 December 2017 Ecology of alpine carabid beetles (Coleoptera, Carabidae) in a Norwegian glacier foreland, with a special focus on claw wearing to indicate relative age SIGMUND HÅGVAR, RONNY STEEN & DANIEL FLØ Hågvar, S., Steen, R. & Flø, D. 2017. Ecology of alpine carabid beetles (Coleoptera, Carabidae) in a Norwegian glacier foreland, with a special focus on claw wearing to indicate relative age. Norwegian Journal of Entomology 64, 82–111. Nine species of carabid beetles (Coleoptera, Carabidae) were pitfall-trapped during two years in an alpine glacier foreland of southern Norway. A two-year (biennial) life cycle was documented for Nebria nivalis (Paykull, 1790), N. rufescens (Ström, 1768), and Patrobus septentrionis Dejean, 1828. This was based on the simultaneous hibernation of larvae and adults. In P. septentrionis, both larvae and adults showed a considerable activity beneath snow. A limited larval material of Amara alpina (Paykull, 1790) and A. quenseli (Schönherr, 1806) from the snow-free period indicated larval hibernation. A. quenseli was, however, not synchronized with respect to developmental stages, and its life cycle was difficult to interpret. Measurements of claw lengths in eight species showed a considerable wearing during adult life. The data indicated that some individuals might hibernate a second time and thus experience two egg-laying seasons. Wearing of mandible tooth was not suited as age indictor in Nebria nivalis and N. rufescens, since some individuals hatched with a small tooth. Supplied with literature data, a “niche profile” is presented for each species. The most important ecological factors that contributed to niche segregation were: preference for vegetation-free ground, occurrence along the successional and time gradient, humidity or temperature preference, nocturnal versus diurnal activity, flight ability, food habits, phenology, and the ability to be active under snow.