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Western North American Naturalist 81(1), © 2021, pp. 54–62 Cold tolerance of mountain stoneflies (Plecoptera: Nemouridae) from the high Rocky Mountains SCOTT HOTALING1,*,†, ALISHA A. SHAH2,†, MICHAEL E. DILLON3, J. JOSEPH GIERSCH4, LUSHA M. TRONSTAD5, DEBRA S. FINN6, H. ARTHUR WOODS2, AND JOANNA L. KELLEY1 1School of Biological Sciences, Washington State University, Pullman, WA 2Division of Biological Sciences, University of Montana, Missoula, MT 3Department of Zoology and Physiology and Program in Ecology, University of Wyoming, Laramie, WY 4Northern Rocky Mountain Science Center, U.S. Geological Survey, West Glacier, MT 5Wyoming Natural Diversity Database, University of Wyoming, Laramie, WY 6Department of Biology, Missouri State University, Springfield, MO ABSTRACT.—How aquatic insects cope with cold temperatures is poorly understood. This is particularly true for high-elevation species, which often experience a seasonal risk of freezing. In the Rocky Mountains, nemourid stoneflies (Plecoptera: Nemouridae) are a major component of mountain stream biodiversity and are typically found in streams fed by glaciers and snowfields, which are rapidly receding due to climate change. Predicting the effects of climate change on mountain stoneflies is difficult because their thermal physiology is largely unknown. We investigated cold tolerance of several alpine stoneflies (Lednia tumana, Lednia tetonica, and Zapada spp.) from the Rocky Mountains, USA. We mea- sured the supercooling point (SCP) and tolerance to ice enclosure of late-instar nymphs collected from a range of thermal regimes. SCPs varied among species and populations, with the lowest SCP measured for nymphs from an alpine pond, which was much more likely to freeze solid in winter than flowing streams. -
Ent21 3 229 256 Sharova.Pmd
Russian Entomol. J. 21(3): 229256 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2012 Æèçíåííûå ôîðìû è àäàïòèâíàÿ ðàäèàöèÿ ëè÷èíîê æóæåëèö (Coleoptera: Carabidae) ìèðîâîé ôàóíû Life forms and adaptive radiation of ground beetle larvae (Coleoptera: Carabidae) of the World fauna È.Õ. Øàðîâà, Ê.Â. Ìàêàðîâ I.Kh. Sharova, K.V. Makarov Ìîñêîâñêèé ïåäàãîãè÷åñêèé ãîñóäàðñòâåííûé óíèâåðñèòåò, êàôåäðà çîîëîãèè è ýêîëîãèè, óë.Êèáàëü÷è÷à, ä. 6, êîðï. 5, Ìîñêâà 129278, Ðîññèÿ. E-mail: [email protected]; [email protected] Moscow State Pedagogical University, Department of Zoology & Ecology, Kibalchicha str. 6, build. 5, Moscow, 129278 Russia ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: ëè÷èíêè, æóæåëèöû, ìîðôîëîãèô, æèçíåííûå ôîðìû, àäàïòàöèè, ñïåêòðû æèçíåííûõ ôîðì, ïàðàëëåëèçì. KEY WORDS: larvae, Carabidae, morphology, life forms, adaptations, life form spectra, parallelism. ÐÅÇÞÌÅ. Ïðåäëîæåíà èåðàðõè÷åñêàÿ ñèñòåìà âî÷íû è ñâîäèëèñü ê îïèñàíèþ îòäåëüíûõ âèäîâ. æèçíåííûõ ôîðì ëè÷èíîê Carabidae ìèðîâîé ôàóíû Îáîáùåíèå ýòèõ ñâåäåíèé ñòàëî âîçìîæíûì ëèøü ñ íà îñíîâå ñèíòåçà îðèãèíàëüíûõ è îïóáëèêîâàííûõ ïîÿâëåíèåì ïåðâûõ ñâîäîê ïî ìîðôîëîãèè è ñèñòå- äàííûõ ïî èõ ìîðôîëîãèè, ñèñòåìàòèêå è ýêîëîãèè. ìàòèêå ëè÷èíîê Carabidae [van Emden, 1942; Jeannel, Íîâàÿ ñèñòåìà æèçíåííûõ ôîðì ëè÷èíîê Carabidae 1941, 1942; Larsson, 1941].  Ðîññèè ïåðâûå îïðåäå- îòëè÷àåòñÿ îò ïðåæíåé [Øàðîâà, 1981] ââåäåíèåì ëèòåëè ëè÷èíîê æóæåëèö áûëè ñîñòàâëåíû Çíîéêî äîïîëíèòåëüíûõ êàòåãîðèé.  ñèñòåìó âêëþ÷åíî 6 [1929], Ãèëÿðîâûì, Øàðîâîé [1954] è Øàðîâîé êëàññîâ è 45 ãðóïï æèçíåííûõ ôîðì, õàðàêåòðèçóþ- [1958, 1964]. Íàêîïëåííûå ñâåäåíèÿ î ëè÷èíêàõ ùèõñÿ êîìïëåêñîì ìîðôîëîãè÷åñêèõ è ïîâåäåí÷åñ- æóæåëèö ïîçâîëèëè èçó÷àòü èõ àäàïòèâíîå ìíîãî- êèõ àäàïòàöèé. Íà îñíîâå íîâîé ñèñòåìû æèçíåííûõ îáðàçèå. Âïåðâûå íåñêîëüêî ìîðôî-ýêîëîãè÷åñêèõ ôîðì ëè÷èíîê Carabidae ïðåäëîæåíà ãèïîòåçà èõ àäàï- òèïîâ ïî÷âîîáèòàþùèõ ëè÷èíîê æóæåëèö âûäåëèë òèâíîé ðàäèàöèè îò ìàëîñïåöèàëèçèðîâàííûõ çîîôà- Ì.Ñ. -
Volume 36, No 1 Summer 2017
Newsletter of the Biological Survey of Canada Vol. 36(1) Summer 2017 The Newsletter of the BSC is published twice a year by the Biological Survey of Canada, an incorporated not-for-profit In this issue group devoted to promoting biodiversity science in Canada. From the editor’s desk......2 Information on Student Corner: Membership ....................3 The Application of President’s Report ...........4 Soil Mesostigmata as Bioindicators and a Summer Update ...............6 Description of Common BSC on facebook & twit- Groups Found in the ter....................................5 Boreal Forest in Northern Alberta..........................9 BSC Student Corner ..........8 Soil Mesostigmata..........9 Matthew Meehan, MSc student, University of Alberta, Department of Biological Sciences Bioblitz 2017..................13 Book announcements: BSC BioBlitz 2017 - A Handbook to the Bioblitzing the Cypress Ticks of Canada (Ixo- Hills dida: Ixodidae, Argasi- Contact: Cory Sheffield.........13 dae)..............................15 -The Biological Survey of Canada: A Personal History..........................16 BSC Symposium 2017 Canadian Journal of Canada 150: Canada’s Insect Diversity in Arthropod Identification: Expected and Unexpected Places recent papers..................17 Contact: Cory Sheffield .....................................14 Wild Species 2015 Report available ........................17 Book Announcements: Handbook to the Ticks of Canada..................15 Check out the BSC The Biological Survey of Canada: A personal Website: Publications -
Current Views on Chelicerate Phylogeny —A Tribute to Peter Weygoldt
Current views on chelicerate phylogeny —A tribute to Peter Weygoldt The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Giribet, Gonzalo. 2018. “Current Views on Chelicerate phylogeny— A Tribute to Peter Weygoldt.” Zoologischer Anzeiger 273 (March): 7– 13. doi:10.1016/j.jcz.2018.01.004. Published Version doi:10.1016/j.jcz.2018.01.004 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:37308630 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Open Access Policy Articles, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#OAP 1 Current views on chelicerate phylogeny—a tribute to Peter Weygoldt 2 3 Gonzalo Giribet 4 5 Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard 6 University, 26 Oxford Street, CamBridge, MA 02138, USA 7 8 Keywords: Arachnida, Chelicerata, Arthropoda, evolution, systematics, phylogeny 9 10 11 ABSTRACT 12 13 Peter Weygoldt pioneered studies of arachnid phylogeny by providing the first synapomorphy 14 scheme to underpin inter-ordinal relationships. Since this seminal worK, arachnid relationships 15 have been evaluated using morphological characters of extant and fossil taxa as well as multiple 16 generations of molecular sequence data. While nearly all datasets agree on the monophyly of 17 Tetrapulmonata, and modern analyses of molecules and novel morphological and genomic data 18 support Arachnopulmonata (a sister group relationship of Scorpiones to Tetrapulmonata), the 19 relationships of the apulmonate arachnid orders remain largely unresolved. -
Acari Scientific Classification Kingdom
Acari - Wikipedia https://en.wikipedia.org/wiki/Acari From Wikipedia, the free encyclopedia Acari (or Acarina) are a taxon of arachnids that contains mites and ticks. The diversity of the Acari is extraordinary and its fossil history Acari goes back to at least the early Devonian period.[1] As a result, Temporal range: acarologists (the people who study mites and ticks) have proposed a Early Devonian–Recent complex set of taxonomic ranks to classify mites. In most modern treatments, the Acari is considered a subclass of Arachnida and is PreЄ Є O S D C P T J K Pg N composed of two or three superorders or orders: Acariformes (or Actinotrichida), Parasitiformes (or Anactinotrichida), and Opilioacariformes; the latter is often considered a subgroup within the Parasitiformes. The monophyly of the Acari is open to debate, and the relationships of the acarines to other arachnids is not at all clear.[2] In older treatments, the subgroups of the Acarina were placed at order rank, but as their own subdivisions have become better-understood, it is more usual to treat them at superorder rank. Most acarines are minute to small (e.g., 0.08–1.00 millimetre or 0.003–0.039 inches), but the largest Acari (some ticks and red velvet mites) may reach lengths of 10–20 millimetres (0.4–0.8 in). Over 50,000 species have been described (as of 1999) and it is estimated that a million or more species may exist. The study of mites and ticks is called acarology (from Greek ἀκαρί/ἄκαρι, akari, a type of mite; and -λογία, -logia),[3] and the leading scientific journals for acarology include Acarologia, Experimental and Applied Acarology and the Peacock mite (Tuckerella sp.), International Journal of Acarology. -
Geological History and Phylogeny of Chelicerata
Arthropod Structure & Development 39 (2010) 124–142 Contents lists available at ScienceDirect Arthropod Structure & Development journal homepage: www.elsevier.com/locate/asd Review Article Geological history and phylogeny of Chelicerata Jason A. Dunlop* Museum fu¨r Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity at the Humboldt University Berlin, Invalidenstraße 43, D-10115 Berlin, Germany article info abstract Article history: Chelicerata probably appeared during the Cambrian period. Their precise origins remain unclear, but may Received 1 December 2009 lie among the so-called great appendage arthropods. By the late Cambrian there is evidence for both Accepted 13 January 2010 Pycnogonida and Euchelicerata. Relationships between the principal euchelicerate lineages are unre- solved, but Xiphosura, Eurypterida and Chasmataspidida (the last two extinct), are all known as body Keywords: fossils from the Ordovician. The fourth group, Arachnida, was found monophyletic in most recent studies. Arachnida Arachnids are known unequivocally from the Silurian (a putative Ordovician mite remains controversial), Fossil record and the balance of evidence favours a common, terrestrial ancestor. Recent work recognises four prin- Phylogeny Evolutionary tree cipal arachnid clades: Stethostomata, Haplocnemata, Acaromorpha and Pantetrapulmonata, of which the pantetrapulmonates (spiders and their relatives) are probably the most robust grouping. Stethostomata includes Scorpiones (Silurian–Recent) and Opiliones (Devonian–Recent), while -
Powell Mountain Karst Preserve: Biological Inventory of Vegetation Communities, Vascular Plants, and Selected Animal Groups
Powell Mountain Karst Preserve: Biological Inventory of Vegetation Communities, Vascular Plants, and Selected Animal Groups Final Report Prepared by: Christopher S. Hobson For: The Cave Conservancy of the Virginias Date: 15 April 2010 This report may be cited as follows: Hobson, C.S. 2010. Powell Mountain Karst Preserve: Biological Inventory of Vegetation Communities, Vascular Plants, and Selected Animal Groups. Natural Heritage Technical Report 10-12. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Richmond, Virginia. Unpublished report submitted to The Cave Conservancy of the Virginias. April 2010. 30 pages plus appendices. COMMONWEALTH of VIRGINIA Biological Inventory of Vegetation Communities, Vascular Plants, and Selected Animal Groups Virginia Department of Conservation and Recreation Division of Natural Heritage Natural Heritage Technical Report 10-12 April 2010 Contents List of Tables......................................................................................................................... ii List of Figures........................................................................................................................ iii Introduction............................................................................................................................ 1 Geology.................................................................................................................................. 2 Explanation of the Natural Heritage Ranking System.......................................................... -
(Insecta, Coleoptera) В Фауне Арктики. Сообщение 1
ЗООЛОГИЧЕСКИЙ ЖУРНАЛ, 2014, том 93, № 1, с. 7–44 ЭКОЛОГИЯ И ЗООГЕОГРАФИЯ УДК 595.76 ОТРЯД ЖЕСТКОКРЫЛЫХ (INSECTA, COLEOPTERA) В ФАУНЕ АРКТИКИ. СООБЩЕНИЕ 1. СОСТАВ ФАУНЫ © 2014 г. Ю. И. Чернов1, О. Л. Макарова1, Л. Д. Пенев2, О. А. Хрулёва1 1 Институт проблем экологии и эволюции им. А.Н. Северцова РАН, Москва 119071, Россия e(mail: [email protected] e(mail: oa([email protected] 2 Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, София, Болгария e(mail: [email protected] Поступила в редакцию 01.09.2013 г. Жесткокрылые, крупнейший отряд насекомых, в условиях Арктики уступают двукрылым первен ство в полноте освоения среды. На долю Coleoptera приходится около 13% энтомофауны тундровой зоны, однако несколько семейств жуков сохраняют в высоких широтах значительное видовое раз нообразие и существенную ценотическую роль. В этом сообщении мы даем обзор циркумполярной колеоптерофауны Арктики. На основе оригинальных данных, литературных сведений и фондовых коллекционных материалов с использованием экстраполяций и аналогий отмечены особенности таксономического и экологического разнообразия подотрядов, серий и семейств Coleoptera, про анализированы широтнозональное распределение и северные пределы распространения видов, специфика адаптаций и ценотических связей. Ключевые слова: Арктика, жуки, природная зональность, видовое разнообразие, ареал, адаптации. DOI: 10.7868/S004451341401005X Жесткокрылые (Coleoptera) – самый крупный основных широтных трендов параметров их раз отряд насекомых, включающий почти 386500 ви нообразия. дов (Slipinski et al., 2011). На его долю приходится Накопленные к настоящему времени данные почти 40% видов класса. Доминирование этого свидетельствуют о том, что в фауне Арктики на отряда в энтомофауне наиболее отчетливо в теп долю жесткокрылых приходится около 13% видо лых поясах, в тропиках и субтропиках. В умерен вого богатства насекомых, в северной полосе ном поясе жесткокрылые составляют треть видо тундровой зоны их доля снижается до 4%, а в по вого богатства насекомых. -
Vol 4 Part 2. Coleoptera. Carabidae
Royal Entomological Society HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS To purchase current handbooks and to download out-of-print parts visit: http://www.royensoc.co.uk/publications/index.htm This work is licensed under a Creative Commons Attribution-NonCommercial-ShareAlike 2.0 UK: England & Wales License. Copyright © Royal Entomological Society 2012 ROYAL ENTOMOLOGICAL SOCIETY OF LONDON . Vol. IV. Part 2 -HANDBOOKS FOR THE IDENTIFICATION / OF BRITISH INSECT-s COLEOPTERA CARABIDAE By CARL H. LINDROTH LONDON Published by the Society and Sold at its Rooms .p, Queen's Gate, S.W. 7 August I 974- HANDBOOKS FOR THE IDENTIFICATION OF BRITISH INSECTS The aim of this series of publications is to provide illustrated keys to the whole of the British Insects (in so far as this is possible), in ten volumes, as follows: I. Part 1. General Introduction. Part 9. Ephemeroptera. , 2. Thysanura. , 10. Odonata. , 3. Protura. , 11. Thysanoptera. , 4. Collembola. , 12. Neuroptera. , 5. Dermaptera and , 13. Mecoptera. Orthoptera. , 14. Trichoptera. , 6. Plecoptera. , 15. Strepsiptera. , 7. Psocoptera. , 16. Siphonaptera. , 8. Anoplura. II. Hemiptera. III. Lepidoptera. IV. and V. Coleoptera. VI. Hymenoptera : Symphyta and Aculeata. VII. Hymenoptera : lchneumonoidea. VIII. Hymenoptera : Cynipoidea, Chalcidoidea, and Serphoidea. IX. Diptera: Nematocera and Brachycera. X. Diptera : Cyclorrhapha. Volumes II to X will be divided into parts of convenient size, but it is not possible to specifyin advance the taxonomic content of each part. Conciseness and cheapness are main objectives in this series, and each part is the work of a specialist, or of a group of specialists. Although much of the work is based on existing published keys, suitably adapted, much new and original matter is also included. -
A New Genus and Three New Species of the Mite Family Arctacaridae (Parasitiformes, Mesostigmata) from North America O
Entomological Review, Vol. 83, No. 7, 2003, pp. 868–886. Translated from Zoologicheskii Zhurnal, Vol. 82, No. 8, 2003. Original Russian Text Copyright © 2003 by Makarova. English Translation Copyright © 2003 by MAIK “Nauka/Interperiodica” (Russia). A New Genus and Three New Species of the Mite Family Arctacaridae (Parasitiformes, Mesostigmata) from North America O. L. Makarova Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow 119071, Russia e-mail: [email protected] Received March 20, 2002 Abstract—A new genus Proarctacarus and three new species, P. canadensis (Rocky Mountains, Alberta, Can- ada), and P. johnstoni (Rocky Mountains, Idaho, Utah, USA), and P. oregonensis (Coast Ranges, Oregon, USA), are described and the diagnosis of the family Arctacaridae is refined. The majority of records of species of the ge- nus Proarctacarus is associated with mountain coniferous forests. The regular arrangement of setae (transverse rows) and sigillae on the large soft opisthosoma suggests the presence of 10 abdominal segments (besides telson), including pregenital segment VII. The setae of this segment (2–3 pairs) are situated dorsolaterally at the border between podosoma and opisthosoma and are distinctly marked by the last row of podosomal setae (segment VI) and first complete row of opisthosomal setae associated with sigillae sg (genital segment VIII). The setae on pregenital segment VII are described in Mesostigmata for the first time. The taxonomic value of the character “presence/absence of ambulacrum I” and the taxonomic position of the cohort Arctacarina are discussed. Until recently, the family Arctacaridae has been of different countries. An analysis of the type series of represented by a single genus Arctacarus Evans in- A. -
Ecology of Alpine Carabid Beetles (Coleoptera, Carabidae) in a Norwegian Glacier Foreland, with a Special Focus on Claw Wearing to Indicate Relative Age
© Norwegian Journal of Entomology. 12 December 2017 Ecology of alpine carabid beetles (Coleoptera, Carabidae) in a Norwegian glacier foreland, with a special focus on claw wearing to indicate relative age SIGMUND HÅGVAR, RONNY STEEN & DANIEL FLØ Hågvar, S., Steen, R. & Flø, D. 2017. Ecology of alpine carabid beetles (Coleoptera, Carabidae) in a Norwegian glacier foreland, with a special focus on claw wearing to indicate relative age. Norwegian Journal of Entomology 64, 82–111. Nine species of carabid beetles (Coleoptera, Carabidae) were pitfall-trapped during two years in an alpine glacier foreland of southern Norway. A two-year (biennial) life cycle was documented for Nebria nivalis (Paykull, 1790), N. rufescens (Ström, 1768), and Patrobus septentrionis Dejean, 1828. This was based on the simultaneous hibernation of larvae and adults. In P. septentrionis, both larvae and adults showed a considerable activity beneath snow. A limited larval material of Amara alpina (Paykull, 1790) and A. quenseli (Schönherr, 1806) from the snow-free period indicated larval hibernation. A. quenseli was, however, not synchronized with respect to developmental stages, and its life cycle was difficult to interpret. Measurements of claw lengths in eight species showed a considerable wearing during adult life. The data indicated that some individuals might hibernate a second time and thus experience two egg-laying seasons. Wearing of mandible tooth was not suited as age indictor in Nebria nivalis and N. rufescens, since some individuals hatched with a small tooth. Supplied with literature data, a “niche profile” is presented for each species. The most important ecological factors that contributed to niche segregation were: preference for vegetation-free ground, occurrence along the successional and time gradient, humidity or temperature preference, nocturnal versus diurnal activity, flight ability, food habits, phenology, and the ability to be active under snow. -
A Key to the Genera of the Ground-Beetle Larvae (Coleoptera, Carabidae) of the Paleartic Region
ISSN 0392-758 X MUSEO REGIONALE DI SCIENZE NATURALI A key to the genera of the Ground-beetle larvae (Coleoptera, Carabidae) of the Paleartic region Kirill V. Makarow ESTRATTO dal Bollettino del Museo Regionale di Scienze Naturali - Torino Volume 12 - N. 1 - 1994 Boll. Mus. reg. Sci. nat. Torino Vol. 12 - N. 1 pp. 221-254 4-5-1994 K.irill V. MAKAR.ow* A key to the genera of the Ground-beetle larvae (Coleoptera, Carabidae) of the Palearctic region INTRODUCTION It is necessary to use the data about the preimaginal stages, especially lar vae, forthe successful decision of many taxonomic and ecological questions, but it's rather difficult, because of the little knowledge on the larvae biology and morphology as well as the absence of modern identification keys. So, van Emden's monograph (1942) appears hitherto the most complete generalizing work. The available keys for the USSR (Sharova, 1958, 1964) and Middle Europe carabid larvae (Hurka, 1978; Arndt, 1991) do not contain a number of larval descriptions, published from 1942-1991. Besides, the finemorphologi cal investigations ofY. Bousquet and H. Goulet (Goulet, 1979; Bousquet and Goulet, 1984) produced on new opportunities forlarval descriptions. Unfor tunately, this method is currently poorly used (Arndt, 1989a, b; Arndt and Hurka, 1990; Bousquet, 1985a, b; 1986; Hurka, 1986; Makarow and Shilenkov, 1991 etc.) and can not be considered as a basis of the practical keys. The present work contains a key for determination of the main Palearctic carabid groups and takes into account most of the works that followed van Emden's monograph.