Permophiles Issue #35 1999 EXECUTIVE NOTES Notes from the SPS Secretary Notes from the SPS Chair
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A Geoconservation perspective on the trace fossil record associated with the end – Ordovician mass extinction and glaciation in the Welsh Basin Item Type Thesis or dissertation Authors Nicholls, Keith H. Citation Nicholls, K. (2019). A Geoconservation perspective on the trace fossil record associated with the end – Ordovician mass extinction and glaciation in the Welsh Basin. (Doctoral dissertation). University of Chester, United Kingdom. Publisher University of Chester Rights Attribution-NonCommercial-NoDerivatives 4.0 International Download date 26/09/2021 02:37:15 Item License http://creativecommons.org/licenses/by-nc-nd/4.0/ Link to Item http://hdl.handle.net/10034/622234 International Chronostratigraphic Chart v2013/01 Erathem / Era System / Period Quaternary Neogene C e n o z o i c Paleogene Cretaceous M e s o z o i c Jurassic M e s o z o i c Jurassic Triassic Permian Carboniferous P a l Devonian e o z o i c P a l Devonian e o z o i c Silurian Ordovician s a n u a F y r Cambrian a n o i t u l o v E s ' i k s w o Ichnogeneric Diversity k p e 0 10 20 30 40 50 60 70 S 1 3 5 7 9 11 13 15 17 19 21 n 23 r e 25 d 27 o 29 M 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93 Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity 0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 n 23 r e 25 d 27 o 29 M 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 c i o 63 z 65 o e 67 a l 69 a 71 P 73 75 77 79 81 83 n 85 a i r 87 b 89 m 91 a 93 C Number of Ichnogenera (Treatise Part W) -
Guadalupian, Middle Permian) Mass Extinction in NW Pangea (Borup Fiord, Arctic Canada): a Global Crisis Driven by Volcanism and Anoxia
The Capitanian (Guadalupian, Middle Permian) mass extinction in NW Pangea (Borup Fiord, Arctic Canada): A global crisis driven by volcanism and anoxia David P.G. Bond1†, Paul B. Wignall2, and Stephen E. Grasby3,4 1Department of Geography, Geology and Environment, University of Hull, Hull, HU6 7RX, UK 2School of Earth and Environment, University of Leeds, Leeds, LS2 9JT, UK 3Geological Survey of Canada, 3303 33rd Street N.W., Calgary, Alberta, T2L 2A7, Canada 4Department of Geoscience, University of Calgary, 2500 University Drive N.W., Calgary Alberta, T2N 1N4, Canada ABSTRACT ing gun of eruptions in the distant Emeishan 2009; Wignall et al., 2009a, 2009b; Bond et al., large igneous province, which drove high- 2010a, 2010b), making this a mid-Capitanian Until recently, the biotic crisis that oc- latitude anoxia via global warming. Although crisis of short duration, fulfilling the second cri- curred within the Capitanian Stage (Middle the global Capitanian extinction might have terion. Several other marine groups were badly Permian, ca. 262 Ma) was known only from had different regional mechanisms, like the affected in equatorial eastern Tethys Ocean, in- equatorial (Tethyan) latitudes, and its global more famous extinction at the end of the cluding corals, bryozoans, and giant alatocon- extent was poorly resolved. The discovery of Permian, each had its roots in large igneous chid bivalves (e.g., Wang and Sugiyama, 2000; a Boreal Capitanian crisis in Spitsbergen, province volcanism. Weidlich, 2002; Bond et al., 2010a; Chen et al., with losses of similar magnitude to those in 2018). In contrast, pelagic elements of the fauna low latitudes, indicated that the event was INTRODUCTION (ammonoids and conodonts) suffered a later, geographically widespread, but further non- ecologically distinct, extinction crisis in the ear- Tethyan records are needed to confirm this as The Capitanian (Guadalupian Series, Middle liest Lopingian (Huang et al., 2019). -
Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships
438 Article 438 by Saswati Bandyopadhyay1* and Sanghamitra Ray2 Gondwana Vertebrate Faunas of India: Their Diversity and Intercontinental Relationships 1Geological Studies Unit, Indian Statistical Institute, 203 B. T. Road, Kolkata 700108, India; email: [email protected] 2Department of Geology and Geophysics, Indian Institute of Technology, Kharagpur 721302, India; email: [email protected] *Corresponding author (Received : 23/12/2018; Revised accepted : 11/09/2019) https://doi.org/10.18814/epiiugs/2020/020028 The twelve Gondwanan stratigraphic horizons of many extant lineages, producing highly diverse terrestrial vertebrates India have yielded varied vertebrate fossils. The oldest in the vacant niches created throughout the world due to the end- Permian extinction event. Diapsids diversified rapidly by the Middle fossil record is the Endothiodon-dominated multitaxic Triassic in to many communities of continental tetrapods, whereas Kundaram fauna, which correlates the Kundaram the non-mammalian synapsids became a minor components for the Formation with several other coeval Late Permian remainder of the Mesozoic Era. The Gondwana basins of peninsular horizons of South Africa, Zambia, Tanzania, India (Fig. 1A) aptly exemplify the diverse vertebrate faunas found Mozambique, Malawi, Madagascar and Brazil. The from the Late Palaeozoic and Mesozoic. During the last few decades much emphasis was given on explorations and excavations of Permian-Triassic transition in India is marked by vertebrate fossils in these basins which have yielded many new fossil distinct taxonomic shift and faunal characteristics and vertebrates, significant both in numbers and diversity of genera, and represented by small-sized holdover fauna of the providing information on their taphonomy, taxonomy, phylogeny, Early Triassic Panchet and Kamthi fauna. -
Carbon and Strontium Isotope Stratigraphy of the Permian from Nevada and China: Implications from an Icehouse to Greenhouse Transition
Carbon and strontium isotope stratigraphy of the Permian from Nevada and China: Implications from an icehouse to greenhouse transition Dissertation Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By Kate E. Tierney, M.S. Graduate Program in the School of Earth Sciences The Ohio State University 2010 Dissertation Committee: Matthew R. Saltzman, Advisor William I. Ausich Loren Babcock Stig M. Bergström Ola Ahlqvist Copyright by Kate Elizabeth Tierney 2010 Abstract The Permian is one of the most important intervals of earth history to help us understand the way our climate system works. It is an analog to modern climate because during this interval climate transitioned from an icehouse state (when glaciers existed extending to middle latitudes), to a greenhouse state (when there were no glaciers). This climatic amelioration occurred under conditions very similar to those that exist in modern times, including atmospheric CO2 levels and the presence of plants thriving in the terrestrial system. This analog to the modern system allows us to investigate the mechanisms that cause global warming. Scientist have learned that the distribution of carbon between the oceans, atmosphere and lithosphere plays a large role in determining climate and changes in this distribution can be studied by chemical proxies preserved in the rock record. There are two main ways to change the distribution of carbon between these reservoirs. Organic carbon can be buried or silicate minerals in the terrestrial realm can be weathered. These two mechanisms account for the long term changes in carbon concentrations in the atmosphere, particularly important to climate. -
A New Late Permian Burnetiamorph from Zambia Confirms Exceptional
fevo-09-685244 June 19, 2021 Time: 17:19 # 1 ORIGINAL RESEARCH published: 24 June 2021 doi: 10.3389/fevo.2021.685244 A New Late Permian Burnetiamorph From Zambia Confirms Exceptional Levels of Endemism in Burnetiamorpha (Therapsida: Biarmosuchia) and an Updated Paleoenvironmental Interpretation of the Upper Madumabisa Mudstone Formation Edited by: 1 † 2 3,4† Mark Joseph MacDougall, Christian A. Sidor * , Neil J. Tabor and Roger M. H. Smith Museum of Natural History Berlin 1 Burke Museum and Department of Biology, University of Washington, Seattle, WA, United States, 2 Roy M. Huffington (MfN), Germany Department of Earth Sciences, Southern Methodist University, Dallas, TX, United States, 3 Evolutionary Studies Institute, Reviewed by: University of the Witwatersrand, Johannesburg, South Africa, 4 Iziko South African Museum, Cape Town, South Africa Sean P. Modesto, Cape Breton University, Canada Michael Oliver Day, A new burnetiamorph therapsid, Isengops luangwensis, gen. et sp. nov., is described Natural History Museum, on the basis of a partial skull from the upper Madumabisa Mudstone Formation of the United Kingdom Luangwa Basin of northeastern Zambia. Isengops is diagnosed by reduced palatal *Correspondence: Christian A. Sidor dentition, a ridge-like palatine-pterygoid boss, a palatal exposure of the jugal that [email protected] extends far anteriorly, a tall trigonal pyramid-shaped supraorbital boss, and a recess †ORCID: along the dorsal margin of the lateral temporal fenestra. The upper Madumabisa Christian A. Sidor Mudstone Formation was deposited in a rift basin with lithofacies characterized orcid.org/0000-0003-0742-4829 Roger M. H. Smith by unchannelized flow, periods of subaerial desiccation and non-deposition, and orcid.org/0000-0001-6806-1983 pedogenesis, and can be biostratigraphically tied to the upper Cistecephalus Assemblage Zone of South Africa, suggesting a Wuchiapingian age. -
GEOLOGIC TIME SCALE V
GSA GEOLOGIC TIME SCALE v. 4.0 CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST HIST. ANOM. (Ma) ANOM. CHRON. CHRO HOLOCENE 1 C1 QUATER- 0.01 30 C30 66.0 541 CALABRIAN NARY PLEISTOCENE* 1.8 31 C31 MAASTRICHTIAN 252 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 Lopin- 254 32 C32 72.1 635 2A C2A PIACENZIAN WUCHIAPINGIAN PLIOCENE 3.6 gian 33 260 260 3 ZANCLEAN CAPITANIAN NEOPRO- 5 C3 CAMPANIAN Guada- 265 750 CRYOGENIAN 5.3 80 C33 WORDIAN TEROZOIC 3A MESSINIAN LATE lupian 269 C3A 83.6 ROADIAN 272 850 7.2 SANTONIAN 4 KUNGURIAN C4 86.3 279 TONIAN CONIACIAN 280 4A Cisura- C4A TORTONIAN 90 89.8 1000 1000 PERMIAN ARTINSKIAN 10 5 TURONIAN lian C5 93.9 290 SAKMARIAN STENIAN 11.6 CENOMANIAN 296 SERRAVALLIAN 34 C34 ASSELIAN 299 5A 100 100 300 GZHELIAN 1200 C5A 13.8 LATE 304 KASIMOVIAN 307 1250 MESOPRO- 15 LANGHIAN ECTASIAN 5B C5B ALBIAN MIDDLE MOSCOVIAN 16.0 TEROZOIC 5C C5C 110 VANIAN 315 PENNSYL- 1400 EARLY 5D C5D MIOCENE 113 320 BASHKIRIAN 323 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 1500 CALYMMIAN 6 C6 APTIAN LATE 20 120 331 6A C6A 20.4 EARLY 1600 M0r 126 6B C6B AQUITANIAN M1 340 MIDDLE VISEAN MISSIS- M3 BARREMIAN SIPPIAN STATHERIAN C6C 23.0 6C 130 M5 CRETACEOUS 131 347 1750 HAUTERIVIAN 7 C7 CARBONIFEROUS EARLY TOURNAISIAN 1800 M10 134 25 7A C7A 359 8 C8 CHATTIAN VALANGINIAN M12 360 140 M14 139 FAMENNIAN OROSIRIAN 9 C9 M16 28.1 M18 BERRIASIAN 2000 PROTEROZOIC 10 C10 LATE -
Globivalvulininae, Foraminifera) from the Wordian (Middle Permian) of Nw Iran
Rivista Italiana di Paleontologia e Stratigrafia (Research in Paleontology and Stratigraphy) vol. 125(1): 1-11. March 2019 GLOBIGAETANIA ANGULATA GEN. N. SP. N. (GLOBIVALVULININAE, FORAMINIFERA) FROM THE WORDIAN (MIDDLE PERMIAN) OF NW IRAN VALERIO GENNARI* & ROBERTO RETTORI *Corresponding author. University of Perugia, Department of Physics and Geology, Via Pascoli, 06123 Perugia, Italy. E-mail: [email protected] To cite this article: Gennari V. & Rettori R. (2019) - Globigaetania angulata gen. n. sp. n. (Globivalvulininae, Foraminifera) from the Wordian (Middle Permian) of NW Iran. Riv. It. Paleontol. Strat., 125(1): 1-11. Keywords: Systematics; New taxon; Small foraminifera; Globivalvulinidae; Biseriamminoids; Palaeozoic. Abstract. A new small biseriamminoid foraminifer, Globigaetania angulata gen. n. sp. n., is here described from the Wordian of a Permian–Triassic sedimentary succession of NW Iran (Zal and Poldasht stratigraphic sections). The new taxon, dedicated to Prof. Maurizio Gaetani, is characterised by peculiar morphology, coiling, and structures that are characteristic of the subfamily Globivalvulininae Reitlinger, 1950, family Globivalvulinidae Reitlinger, 1950, superfamily Biseriamminoidea Chernysheva, 1941. The introduction of the new taxon contributes to the knowledge of the systematics and evolution of the Palaeozoic biserial microgranular foraminifera. INTRODUCTION subfamilies (Globivalvulininae Reitlinger, 1950; Paraglobivalvulininae Gaillot & Vachard, 2007; Although the Permian Period was character- Dagmaritinae Bozorgnia, 1973 and Paradagmariti- ised by two major biological crises (end-Guadalupi- nae Gaillot & Vachard, 2007). This family due to its an and end-Permian) (Jin et al. 1994; Erwin 2006; richness and abundance in shallow-water carbonate Clapham et al. 2009; Shen & Bowring 2014), it was environments became a useful biostratigraphic tool also an important phase of marine biotic diversifi- for regional correlations in the Middle–Late Permi- cation (Vachard et al. -
Permian (Artinskian to Wuchapingian) Conodont Biostratigraphy in the Tieqiao Section, Laibin Area, South China
Permian (Artinskian to Wuchapingian) conodont biostratigraphy in the Tieqiao section, Laibin area, South China Y.D. Suna, b*, X.T. Liuc, J.X. Yana, B. Lid, B. Chene, D.P.G. Bondf, M.M. Joachimskib, P.B. Wignallg, X.L. Laia a State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Wuhan, 430074, China b GeoZentrum Nordbayern, Universität Erlangen-Nürnberg, Schlossgarten 5, 91054 Erlangen, Germany c Key Laboratory of Marine Geology and Environment, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, 266071, China d Key Laboratory of Marine Mineral Resources, Guangzhou Marine Geological Survey, Ministry of Land and Resources, Guangzhou, 510075, China e State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, 39 East Beijing Road, Nanjing, 210008, R.P. China f School of Environmental Sciences, University of Hull, Hull HU6 7RX, UK g School of Earth and Environment, University of Leeds, Leeds LS2 9JT, UK *Corresponding authors Email: [email protected] (Y.D. Sun) © 2017, Elsevier. Licensed under the Creative Commons Attribution- NonCommercial-NoDerivatives 4.0 International http://creativecommons.org/ licenses/by-nc-nd/4.0/ 1 Abstract Permian strata from the Tieqiao section (Jiangnan Basin, South China) contain several distinctive conodont assemblages. Early Permian (Cisuralian) assemblages are dominated by the genera Sweetognathus, Pseudosweetognathus and Hindeodus with rare Neostreptognathodus and Gullodus. Gondolellids are absent until the end of the Kungurian stage—in contrast to many parts of the world where gondolellids and Neostreptognathodus are the dominant Kungurian conodonts. A conodont changeover is seen at Tieqiao and coincided with a rise of sea level in the late Kungurian to the early Roadian: the previously dominant sweetognathids were replaced by mesogondolellids. -
Middle Permian (Wordian) Foraminifers of the Furen Limestone, Oita Prefecture, Japan
人と自然 Humans and Nature 22: 21−25 (2011) Report Middle Permian (Wordian) foraminifers of the Furen Limestone, Oita Prefecture, Japan Fumio KOBAYASHI Division of Earth Sciences, Institute of Natural and Environmental Sciences, University of Hyogo / Division of Natural History, Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda, Hyogo, 669-1546 Japan Abstract The Furen Limestone exposed in Yatsu-cho, Usuki City, Oita Prefecture is contained as an exotic block in the Jurassic siliciclastic rocks of the Okugawachi Formation in the Southern Chichibu Belt. In addition to the known Early Permian (Asselian) fusulines, Middle Permian (Wordian) foraminifers of eight species of fusulines and seven species of non-fusuline foraminifers were newly found from the limestone. Among them, Neoschwagerina craticulifera, Verbeekina verbeeki, Chusenella cf. conicocylindrica, and Pseudodoliolina sp. are systematically described. Key words: Middle Permian foraminifers, Furen Limestone, Chichibu Belt Introduction (Kambe and Teraoka, 1968). However, paleontologic description of Paleozoic foraminifers is very few in The Furen Limestone exposed at north of Tomari, this area and almost confined to that of Huzimoto Yatsu-cho, Usuki City, Oita Prefecture is famous (1937). of the germination of an imposing limestone cave locally called as “Furendo” or “Furendokutsu” (Figure 1). The limestone corresponds to the western extension of the Permian Tsukumi Limestone (Kambe and Teraoka, 1968) and is contained as an exotic limestone block in the Jurassic siliciclastic rocks of the Okugawachi Formation (Teraoka et al., 1992; Sakai et al., 1993) in the Southern Chichibu Belt in Kyushu. Huzimoto (1937) described seven species of Early Permian fusulines from the Furen Limestone. The most important species among them is Pseudoschwagerina orientale described as a new species. -
Proposal of Guadalupian and Component Roadian, Wordian And
Permophiles Issue #34 1999 REPORTS Proposal of Guadalupian and Component morphoclines, absolute dates, and paleomagnetics. Roadian, Wordian and Capitanian Stages as International Standards for the Middle Permian Historic Preamble Prolonged deliberation of SPS members culminated in the man- Series dated formal postal vote by Titular (voting) Members that approved subdivision of the Permian System into three series, in ascending Brian F. Glenister order Cisuralian, Guadalupian and Lopingian (Report of Presi- University of Iowa dent Jin Yugan, Permophiles #29, p. 2). The “——usage of the Department of Geology Guadalupian Series and constituent stages, i.e. the Roadian, the Iowa City, IA 52242, USA Wordian and the Capitanian Stage for the middle part of the Per- mian.” was approved unanimously by 15 voting members. Pro- Bruce R. Wardlaw posal of the Guadalupian as a chronostratigraphic unit of series U. S. Geological Survey rank (Girty, 1902) predates any potential competitors by decades 926A National Center (Glenister et al., 1992). Of the three component stages currently Reston, VA 22092-0001, USA recognized, the upper two (Wordian and Capitanian) enjoy com- parable priority. Capitanian was first employed in a Lance L. Lambert lithostratigraphic sense by Richardson (1904) for the massive reef Department of Physics limestones of the Guadalupe Mountains of New Mexico and West Southwest Texas State University Texas, and the Word was used by Udden et al. (1916) for the San Marcos, TX 78666-4616, USA interbedded clastic/carbonate sequence in the adjacent Glass Moun- tains. Both were used in a strictly chronostratigraphic sense first Claude Spinosa by Glenister and Furnish (1961) as substages of the Guadalupian Stage, referenced by their nominal formations and recognized else- Permian Research Institute where through “ammonoid and fusuline faunas”. -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2014/02 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age Erathem / Era System / Period GSSP GSSP age (Ma) GSSP GSSA EonothemErathem / Eon System / Era / Period EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon GSSP age (Ma) present ~ 145.0 358.9 ± 0.4 ~ 541.0 ±1.0 Holocene Ediacaran 0.0117 Tithonian Upper 152.1 ±0.9 Famennian ~ 635 0.126 Upper Kimmeridgian Neo- Cryogenian Middle 157.3 ±1.0 Upper proterozoic Pleistocene 0.781 372.2 ±1.6 850 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian 1000 Callovian 166.1 ±1.2 Quaternary Gelasian 2.58 382.7 ±1.6 Stenian Bathonian 168.3 ±1.3 Piacenzian Middle Bajocian Givetian 1200 Pliocene 3.600 170.3 ±1.4 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.62 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Langhian Sinemurian Proterozoic Neogene 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- Hettangian 2050 Burdigalian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 28.1 Gorstian Oligocene Upper Norian 427.4 ±0.5 2500 Rupelian Wenlock Homerian -
Lopingian, Permian) of North China
Foss. Rec., 23, 205–213, 2020 https://doi.org/10.5194/fr-23-205-2020 © Author(s) 2020. This work is distributed under the Creative Commons Attribution 4.0 License. The youngest occurrence of embolomeres (Tetrapoda: Anthracosauria) from the Sunjiagou Formation (Lopingian, Permian) of North China Jianye Chen1 and Jun Liu1,2,3 1Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 2Chinese Academy of Sciences Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 3College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing 100049, China Correspondence: Jianye Chen ([email protected]) Received: 7 August 2020 – Revised: 2 November 2020 – Accepted: 16 November 2020 – Published: 1 December 2020 Abstract. Embolomeri were semiaquatic predators preva- 1 Introduction lent in the Carboniferous, with only two species from the early Permian (Cisuralian). A new embolomere, Seroher- Embolomeri are a monophyletic group of large crocodile- peton yangquanensis gen. et sp. nov. (Zoobank Registration like, semiaquatic predators, prevalent in the Carboniferous number: urn:lsid:zoobank.org:act:790BEB94-C2CC-4EA4- and early Permian (Cisuralian) (Panchen, 1970; Smithson, BE96-2A1BC4AED748, registration: 23 November 2020), is 2000; Carroll, 2009; Clack, 2012). The clade is generally named based on a partial right upper jaw and palate from the considered to be a stem member of the Reptiliomorpha, taxa Sunjiagou Formation of Yangquan, Shanxi, China, and is late that are more closely related to amniotes than to lissamphib- Wuchiapingian (late Permian) in age. It is the youngest em- ians (Ruta et al., 2003; Vallin and Laurin, 2004; Ruta and bolomere known to date and the only embolomere reported Coates, 2007; Clack and Klembara, 2009; Klembara et al., from North China Block.