A Geoconservation perspective on the trace fossil record associated with the end – Ordovician mass extinction and glaciation in the Welsh Basin
Item Type Thesis or dissertation
Authors Nicholls, Keith H.
Citation Nicholls, K. (2019). A Geoconservation perspective on the trace fossil record associated with the end – Ordovician mass extinction and glaciation in the Welsh Basin. (Doctoral dissertation). University of Chester, United Kingdom.
Publisher University of Chester
Rights Attribution-NonCommercial-NoDerivatives 4.0 International
Download date 26/09/2021 02:37:15
Item License http://creativecommons.org/licenses/by-nc-nd/4.0/
Link to Item http://hdl.handle.net/10034/622234 International Chronostratigraphic Chart v2013/01 Erathem / Era System / Period
Quaternary
Neogene
C e n o z o i c Paleogene
Cretaceous
M e s o z o i c Jurassic M e s o z o i c Jurassic
Triassic
Permian
Carboniferous
P a l Devonian e o z o i c P a l Devonian e o z o i c
Silurian
Ordovician s a n u a F y
r Cambrian a n o i t u l o v E s ' i k s w
o Ichnogeneric Diversity k p e 0 10 20 30 40 50 60 70 S 1 3 5 7 9 11 13 15 17 19 21
n 23 r
e 25
d 27 o 29 M 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Cambrian Palaeozoic Modern e m i T 55 53 51 49 47 45 43 41 39 37 35 33 31 29 27 25 23 21 19 17 15 13 11 93 91 89 87 85 83 81 79 77 75 73 71 69 67 65 63 61 59 57 9 7 5 3 1 0 10 20 Number of(Treatise Ichnogenera Part W) 30 Ichnogeneric Diversity Ichnogeneric 40 50 60 70 Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) International Chronostratigraphic Chart v2013/01 Time Europe Series / Epoch Stage / Age Ma BP Erathem
r Holocene e n p le a p d ri U id b n la ia M a s Pleistocene la C e 2.588 G Piacenzian Pliocene Zanclean
n ia n n ia n o h ia rt g n o n a T a it L u n n q Miocene ia a A in lli s a n s v lia e a a rr g M e i C rd S u 23.03 B a
Chattian i n Oligocene z Rupelian o
Priabonian i c Bartonian
Eocene Lutetian
Ypresian
Thanetian
Paleocene Selandian Danian 66.00 Maastrichtian
Campanian
Santonian Upper Coniacian
Turonian
Cenomanian
Albian
Aptian
Barremian Lower Hauterivian
Valangian M Berriasian 145.00 e Tithonian s Upper o z o i c M e s Upper Kimmeridgian o Oxfordian z
n ia n o o n th ia a n B le i Middle n a ia A v n llo ia c a c jo C a B Toarcian
Pliensbachian Lower Sinemurian Hettangian 201.30 Rhaetian
Norian
Upper Carnian
Ladinian
Middle Anisian
Lower Olenekian / Induan 252.17
Lopingian Changsingian / Wuchiapingian
Guadalupian Capitanian/Wordian/Roadian
Kungurian
Artinskian Cisuralian Sakmarian
Asselian 289.90 Gzhelian
Kasimovian
Moscovian
Pennsylvanian Bashkinian Serpukhovian
Visean
Tournasian Mississipian 358.90
Famennian Upper
Frasnian P Givetian a Middle l Eifelian e o Emsian z Lower Pragian o i c P a l e o z Lower o Lochovian 419.20 i Pridoli c Ludlow Ludfordian/ Gorstian Wenlock Homerian / Sheinwoodian Llandovery Telychian/Aeronian/Rhuddanian 443.40 Hirnantian
Katian Upper
Sandbian
Darriwilian Middle
Dapingian
Floian Lower Tremadocian 485.40 Furongian Stage 10 Jiangshanian / Paibian Series 3 Guzhangian / Drumian / Stage 5 Series 2 Stage 4 / Stage 3
Stage 2 Terreneuvian Fortunian 541.00 Pre-Cambrian
Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Europe System Series Stage Holocene Quaternary Pleistocene Astian Pliocene Pontian Sarmatian Tortonian Miocene Helvetian Burdigalian Aquitanian Chattian Oligocene Rupelian Tertiary Lattorfian Ludian Bartonian Eocene Auversian Lutetian Ypresian Sparnacian Paleocene Thanetian Montian/Danian Maastrichtian Campanian Santonian Upper Cretaceous Coniacian Turonian Cenomanian Cretaceous Albian Aptian Barremian Lower Cretaceous Hauterivian Valangian Berriasian Tithonian Upper Jurassic
Jurassic Upper Jurassic Kimmeridgian Oxfordian Callovian Middle Jurassic Bathonian Jurassic Bajocian Toarcian Pliensbachian Lower Jurassic (Liassic) Sinemurian Hettangian Rhaetian Upper Triassic Norian Carnian Triassic Ladinian Middle Triassic Anisian Lower Triassic Scythian Tartarian Upper Permian Kazanian Kungurian Permian Artinsikian Lower Permian Sakmarian Asselian
Stephanian
Upper Carboniferous Westphalian
Namurian Carboniferous
Visean Lower Carboniferous Tournasian
Fammenian Upper Devonian Frasnian
Devonian Givetian Middle Devonian Couvinian Emsian Lower Devonian Siegenian Devonian
Lower Devonian Gedinnian Pridoli Upper Silurian Ludlovian Silurian Wenlockian Lower Silurian Llandoverian
Ashgillian
Upper Ordovician
Caradocian Ordovician
Llandeilian Llanvirnian Lower Ordovician Arenigian Tremadocian
Merioneth Cambrian St David Comley
Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) North America Stage Series System Holocene Quaternary Pleistocene
Foley Pliocene
Clovelly Duck Lake Miocene Napoleonville Anahuac
Chakasaway Oligocene Vicksburgian Tertiary
Jacksonian
Claibornian Eocene
Sabinian
Midwayan Paleocene
Navarroan Tayloran
Austinian Gulfian
Eaglefordian Woodbinian Cretaceous Washitan Fredricksburgian Comanchean Trinitian Nuevoleonian Coahuilan Durangoan
Portlandian Upper Jurassic
Jurassic Kimmeridgian Upper Jurassic Oxfordian Callovian Bathonian Middle Jurassic Jurassic Bajocian Toarcian Pliensbachian Lower Jurassic (Liassic) Sinemurian Hettangian Rhaetian Norian Upper Triassic Carnian Triassic Ladinian Middle Triassic Anisian Scythian Lower Triassic Ochoan Upper Permian Guadalupian Permian Leonardian Lower Permian Wolfcampian
Virgillian Upper Pennsylvanian Missourian Desmoinesian Pennsylvanian Middle Pennsylvanian Atokan Morrowan Lower Pennsylvanian Chesteran Upper Mississippian Meramecian Mississippian Osagian Lower Mississippian Kinderhookian Bradfordian Chutauquan Cassadagan Chemungian Senecan Fingerlakesian Taghanican Devonian Tioghniogan Erian Cazenovian Onequethawan Deerparkian Ulsterian Devonian
Ulsterian Helderbergian Cayugan
Niagaran Silurian
Alexandrian Richmondian Maysvillian Cincinattian Edenian Mohawkian Chazyan Champlanian Ordovician Whiterockian
Canadian
Trempealeuan Franconian Dresbachian Cambrian Albertan Waucoban
Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) A A A A n A c g A m n r A A a la l e c r c n A lo p li h c a t s g h d a h n h p c i e a t o id ri o o n i e h r i c ti ri c c o ic h c h c c h h n h a h n h h n n a n p n i n n i i s u h u u u u u u s s e s m s s m m a North America Ichogeneric Diversity % Erathem Diversity of previous
4 r e n 2 0 p le a p d ri U id b n la ia 1 M a s 5 la 8 C e G 2 1 0 8 0 2 9 8 0 1 3 n 0 ia n 1 n ia n 0 o h ia 1 rt g n 3 o n a 0 T a it L u 1 0 n n q ia a A in lli 3 9 s a n s v lia 1 4 e a a rr g 1 M e i C rd 3 S u 1 B 3 4 e 2 1 0 n 9 0 2 1 0 o 9 0 2 1 z 0 9 0 o 2 6 9 7 4 1 i 0 3 0 c 4 1 0 3 0 4 9 3 8 4 9 4 6 4 9 1 6 0 5 1 0 1 0 5 9 1 6 5 8 3 3 6 1 0 4 2 6 1 0 3 0 6 1 0 3 0 6 1 0 3 0 6 9 3 8 6 1 1 4 4 5 1 0 1 6 0 5 1 0 1 6 0 5 1 0 1 6 0 5 9 1 6 8 5 1 0 1 7 2 M 5 1 9 1 6 3 e 2 1 0 s 9 0 o z o i c M e s 2 1 0 o 9 0 2 8 z 9 3 o 3 9 5 7 3 1 i 0 6 3 c 3 9 5 5 3 9 7 7 3 1 0 8 3 3 9 7 7 3 9 8 0 4 1 0 1 2 0 4 1 0 1 2 0 4 1 1 1 2 1 3 9 1 8 7 3 1 1 1 9 5 3 1 3 1 4 6 2 1 0 5 0 2 1 0 5 0 2 7 5 1 3 1 0 5 0 3 1 0 5 0 3 9 5 0 3 1 0 9 0 3 1 0 9 3 3 1 0 8 0 3 9 8 7 3 1 0 9 0 3 1 0 1 9 0 3 1 0 1 9 3 3 1 0 1 8 0 3 1 1 1 8 9 3 1 0 2 0 3 1 0 2 0 3 1 0 2 0 3 1 P 0 2 7 a 3 1 0 0 0 l 3 1 0 a 0 0 3 8 e 0 1 3 1 o 0 7 0 3 1 z 0 7 o 0 i c P a l a e o z 3 1 o 3 7 2 i 2 1 0 c 8 0 2 1 0 8 0 2 1 0 8 4 2 5 7 1 5 9 1 3 8 5 9 1 4 6 5 1 1 1 1 6 4 4 1 0 1 9 2 4 1 0 1 8 0 4 1 0 1 8 2 4 1 0 1 7 2 4 9 1 6 8 4 1 0 1 7 2 4 1 3 1 6 5 3 1 0 1 4 0 3 1 0 1 4 0 3 1 1 1 4 3 3 7 0 5 4 2 2 0 2 1 Pre-Cambrian 8 39 Mean 11.9427984485467 Standard Deviation Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) Ichnogeneric Diversity
0 10 20 30 40 50 60 70 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 T 41 43 i 45 47 m 49 e 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91 93
Number of Ichnogenera (Treatise Part W) A A A B B A A A n A rt s a a c g A m n r A h a A l r A a la l e c r A r p s a o c n s A lo p li h c A r o h A A A t A A n c B a t p g h d a h r t p o s s s e s o c B B n h r c o i e a e A h o t t t r t u g o e e t o id i o n i e n o d id e e e o e li l i a e r h r i c ti ri c c o A ic rt p i i ri ri ri p ro c o c c lo g ic h c h c c h h n r o h h c c a c c h h s h o r a h a h n h h n n a d r y h h c n h y s n s n n a u n p n i n n i i s e li a c n n i i n c o i i i i p e u h u u u u u u s l te r u u u te te u u m te te te te h r s e s m s s m m a ia s ia s s s s s s s a s s s s e ia
r e n p le a p d ri U id b n la ia M a s 1 la C e G 1
1
n 1 ia n n ia n o h ia rt g n o n a 1 T a it L u n n q ia a A in lli s a n 1 s v lia e a a rr g M e i rd S u 1 B 1
1
1
1
1
1
1
1
1
1 1
1 1
1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1
1 1
1 1
1 1
1 1
1 1
1 1
1
1 1
1 1
1 1
1
1
1
1
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1 1 1
1 1 1
1 1 1
1 1 1
1
1 1
1 1
1 1
1
1
1
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1 1 1
1 1 1
1 1 1
1 1 1
1 1
1 1
1 1
1 1
1
1
1
1
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1 1 1
1 1 1
1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1
1 1
1 1
1 1
1 1
1 1
1 1
1
1
Ichnogeneric Diversity Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 0 10 20 30 40 50 60 70 1 1 3 3 5 5 7 7 9 9 11 11 13 13 15 15 17 17 19 19 21 21 23 23 25 25 27 27 29 29 31 31 33 33 35 35 37 37 39 39 T 41 41 43 43 e i 45 45 m
47 i 47 m 49 T 49 e 51 51 53 53 55 55 57 57 59 59 61 61 63 63 65 65 67 67 69 69 71 71 73 73 75 75 77 77 79 79 81 81 83 83 85 85 87 87 89 89 91 91 93
Number of Ichnogenera (Treatise Part W) Retention of ichnogenera (%) Ichnogeneric Diversity Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 0 10 20 30 40 50 60 70 1 1 3 3 5 5 7 7 9 9 11 11 13 13 15 15 17 17 19 19 21 21 23 23 25 25 27 27 29 29 31 31 33 33 35 35 37 37 39 39 T 41 41 43 43 e i 45 45 m
47 i 47 m 49 T 49 e 51 51 53 53 55 55 57 57 59 59 61 61 63 63 65 65 67 67 69 69 71 71 73 73 75 75 77 77 79 79 81 81 83 83 85 85 87 87 89 89 91 91 93
Number of Ichnogenera (Treatise Part W) Retention of ichnogenera (%) Ichnogeneric Diversity Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 0 10 20 30 40 50 60 70 1 1 3 3 5 5 7 7 9 9 11 11 13 13 15 15 17 17 19 19 21 21 23 23 25 25 27 27 29 29 31 31 33 33 35 35 37 37 39 39 T 41 41 43 43 e i 45 45 m
47 i 47 m 49 T 49 e 51 51 53 53 55 55 57 57 59 59 61 61 63 63 65 65 67 67 69 69 71 71 73 73 75 75 77 77 79 79 81 81 83 83 85 85 87 87 89 89 91 91 93
Number of Ichnogenera (Treatise Part W) Retention of ichnogenera (%) B ro o A A k A A n r A B B B B h C A n A A t s a a r o C C c g A m r h a A l r o B o l B C h li A a la l e c r A A r p A A s A a o B B s r o z u h C m c n s A lo p li h c r o h A t A n c B B i t a b n a C C o C o i a a t p g c h d a h r th p o s s s e s u o c B if B B fu ri n k r y ly a a m n r c n h r o o in e a e A o t t te r te g o e e e a i i c c v u e c p u h d c t o id i i e n o d id e e o li l i a r s fo f rc o h a n r o a o u ti h r i c ti ri c c o A ic rt p i i ri ri ri p ro c o c c lo g c u u p i li n ri a le ti n ri li c ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s
r e n p le a p d ri U id b n la ia M a s la C e G 1
1
n 1 ia n n ia n o h ia rt g n o n a 1 T a it L u n n q ia a A in lli s a n 1 s v lia e a a rr g M e i rd S u 1 B 1
1
1
1
1 1
1 1
1 1
1 1
1 1
1 1
1 1
1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1 1 1
1 1
1
1
1
1
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1 1 1
1 1 1
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1 1 1
1 1
1
1
1
1
1
1
1
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1 1 1
1 1 1
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1 1 1 1 1 1
1
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1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1 1
1 1 1
Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) B ro o A A A B k A A n A r B B B h C C g A n A t s a a r o C C l C D c A m e r h a A l r o B o l B C h i C o C C a A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y n h i r o o in e a A o t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s
r e n p le a p d ri U id b n la ia M a s la C e G 1 1 1
1 1 1
n 1 1 1 ia n n ia n o h ia rt g n o n a 1 1 1 T a it L u n n q ia a A in lli s a n 1 1 1 s v lia e a a rr g M e i rd S u 1 1 1 1 B 1 1
1 1
1 1
1 1
1 1 1
1 1 1
1 1 1
1 1 1
1 1 1
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1 1 1
1 1 1 1 1
1 1 1 1
1 1 1 1
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1 1 1 1
1 1 1 1 1
1 1 1 1
1 1 1
1 1 1
1 1 1
1 1
1 1 1
1 1
Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) B ro o A A A B k A A n A r B B B h C C D D D D g A n A t s a a r o C C l C D e D id D F c A m e r h a A l r o B o l B C h i C o C C a n e im D ip a A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c a t p g c h d a re th p o s s s e s u o c e B f B B fu i n v r y ly a a m n rc c c n in r m o n u c in y y a l o a r ip o l d h g a i n h i r o o in e a A o t te te r te g o e e a i i c c u e c p u a h d t o o o o s C o r l d li l D e t o ic p c o o i y s s t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v o r n o a im s i g t u h li r d m n r c i F h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n r n p d h h n e h r p h d n x a n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a e i i t o n n i r n p o n i i e u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu l te u o u u te io u h u in u u n s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s ix s m n ra s s s n s a ra s a m m a
r e n p le a p d ri U id b n la ia M a s la C e G
n 1 1 ia n n ia n o h ia rt g n o n a 1 1 T a it L u n n q ia a A in lli s a n 1 1 s v lia e a a rr g M e i rd S u 1 1 B
1
1
1
1 1 1 1 1
1 1 1 1 1
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1 1
1 1
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1 1
Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) B ro o A A A B k A A n A r B B B h C C D D D D G g A n A t s a a r o C C l C D e D id D F G H c A m e r h a A l r o B o l B C h i C o C C a n e im D ip a F o G G a H A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s lu n e c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c F u c i G G y y n a a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y a l o a r ip o l d h g a i u s a G y ro G ro p n h i r o o in e a A o t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D e t o ic p c o o i y s s F c t G k d r r y y tz H lo t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a im s i g t u h li r d m n r c i F u u ip l s i a i ro p r p c H h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F c s l o ta c G n c h o h h a a ti ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r o o y c d h u h c y l y e l m c h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n r n p d h h n e h r p h d n x a p k n o l n h l it l o h n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a e i i t o n n i r n p o n i i e id p h e te i r a i o li h li li p ip n u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu l te u o u u te io u h u in u u n e s u r l te d r te r te e te n o e u s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s ix s m n ra s s s n s a ra s a m m a s is s ia la s ia ia s te s s s ia a s s
r e n p le a p d ri U id b n la ia M a s la C e G 1 1
1 1
n 1 1 ia n n ia n o h ia rt g n o n a 1 1 T a it L u n n q ia a A in lli s a n 1 1 s v lia e a a rr g M e i rd S u 1 1 B 1 1
1 1
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1 1
1 1
1
1
Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) Ichnogenera - Retention (% of previous Stage)
0 50 100 150 200 250 1 3 5 7 9 11 13 15 17 19 21 23 25 27 29 31 33 35 37 39 41 43 e 45 m i 47
T 49 51 53 55 57 59 61 63 65 67 69 71 73 75 77 79 81 83 85 87 89 91
Retention of ichnogenera (%) B ro o A A A B k Ic A A n A r B B B h C C D D D D G H H h g A n A t s a a o r o C C l C C D e D id D i F G H H e H I t c A m e r h a A l r B o l B C h i C o C a n e im D p a F o G G a H e H H e x H o c h A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s lu n e li H i H r H h y c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c F u c i G G y y n a c e H e lm a m m y n I o I a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y a l o a r ip o l d h g a i u s a G y ro G ro p o e li e lm p a is d o c i m n h i r o o in e a A o t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D e t o ic p c o o i y s s F c t G k d r r y y tz H lo H d li c l in o n t o h d b t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a im s i g t u h li r d m n r c i F u u ip l s i a i ro p r p c H a r c o i in d io s ra I c n i r h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F c s l o ta c G n c h o h h a a ti r o o r o th i th i n c u y c i ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r o o y c d h u h c y l y e l m c p m l h c th o c a d ro c h m s h c h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n r n p d h h n e h r p h d n x a p k n o l n h l it l o h e it a h o h e n u n h n p n i n n i iu s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a e i i t o n n i ri n p o n i i e id p h e te i r a i o li h li li p ip n p i h p o p n li r id y i p i n u h u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu l te u o u u te o u h u in u u n e s u r l te d r te r te e te n o e u u te u h n id s u te m e lu te lu ic te u s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s ix s m n ra s s s n s a ra s a m m a s is s ia la s ia ia s te s s s ia a s s s s s e e a is s s a a s s s a s s
r e n p le a p d ri U id b n la ia M a s la C e G 1 1
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n 1 1 ia n n ia n o h ia rt g n o n a 1 1 T a it L u n n q ia a A in lli s a n 1 1 s v lia e a a rr g M e i rd S u 1 1 B 1 1 1
1 1 1
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1 1 B ro o A A A B k Ic A A n A r B B B h C C D D D D G H H h I g A n A t s a a o r o C C l C C D e D id D i F G H H e H I t m K L L c A m e r h a A l r B o l B C h i C o C a n e im D p a F o G G a H e H H e x H o c h p Is K L A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s lu n e li H i H r H h y o o o u L a e i c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c F u c i G G y y n a c e H e lm a m m y n I o I I l n L n th a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y a l o a r ip o l d h g a i u s a G y ro G ro p o e li e lm p a is d o c i m n r p Ix u in a L ic a t n h i r o o in e a A o i t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D e t o ic p c o o i y s s F c t G k d r r y y tz H lo H d li c l in o n t o h d b o r o p d e o p L ic o t o d i r i e n o d d e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a im s i g t u h li r d m n r c i F u u ip l s i a i ro p r p c H a r c o i in d io s ra I c n i r g In e d a K h r v a i a r g h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F c s l o ta c G n c h o h h a a ti r o o r o th i th i n c u y c i l c d i li K i i i ic m id s p L a r ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r o o y c d h u h c y l y e l m c p m l h c th o c a d ro c h m s h c y i ic c c n c c y i i p i e t a h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n r n p d h h n e h r p h d n x a p k n o l n h l it l o h e it a h o h e n u n h p s t h h e g n h n c e s n e p n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a e i i t o n n i r n p o n i i e id p h e te i r a i o li h li li p ip n p i h p o p n li r id y i p i n h i y n n c e i n c i h c p n r h u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu l te u o u u te io u h u in u u n e s u r l te d r te r te e te n o e u u te u h n id s u te m e lu te lu ic te u u fe o u u k l u u lu te n u i e io u s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s ix s m n ra s s s n s a ra s a m m a s is s ia la s ia ia s te s s s ia a s s s s s e e a is s s a a s s s a s s s x n s s ia la m s s s a s ra a n s
r e n p le a p d ri U id b n la ia M a s la C e G 1
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n 1 1 ia n n ia n o h ia rt g n o n a 1 1 T a it L u n n q ia a A in lli s a n 1 1 s v lia e a a rr g M e i rd S u 1 1 B 1
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1 1 B ro o M M A A A B k Ic e M o A A n A r B B B h C C D D D D G H H h I n g A n A t s a a r o C C l C C D e D id D i F G H H e H I t m K L L L M ro ix M o c A m e r h a A l r o B o l B C h i C o C a n e im D p a F o G G a H e H H e x H o c h p Is K L o M M s o m M A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s lu n e li H i H r H h y o o o u L a e i p M a a M o c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c F u c i G G y y n a c e H e lm a m m y n I o I I l n L n th m r e to M M te n o M y a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y l o a r ip o l d h g a i u s k a G y ro G ro p o e li e lm p a is d o c i m n r p Ix u in a L ic a t L h a m g g m e e i i o r r n h i r o o in e a A o t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D a e t o ic p c o o i y s s F c ti G d r r y y tz H lo H d li c l in o n t o h d b o r o p d e o p L ic o o o c r a s M n c c p u i t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a im s i g t u h li r d m n r c i F u u p l s i a i ro p r p c H a r c o i in d io s ra I c n i r g In e d a K h r v a i a r g b L c a i i g i i s i h r h e a h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F c s l o ta c G n c h o h h a a ti r o o r o th i th i n c u y c i l c d i li K i i i ic m id s p L a r i o t n ll ti r c c o ic c i a i n p ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r o o y c d h u h c y l y e l m c p m l h c th o c a d ro c h m s h c y i ic c c n c c y i i p i e t a c e o ic c a h h n a h c t c s o h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n r n p d h h n e h r p h d n x a p k n o l n h l it l o h e it a h o h e n u n h p s t h h e g n h n c e s n e p h c n h h p n n e t n h e h t d n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a e i i t o n n i r n p o n i i e id p h e te i r a i o li h li li p ip n p i h p o p n li r id y i p i n h i y n n c e i n c i h c p n r h n k i p n t i i r u i n r n e i u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu l te u o u u te io u h u in u u n e s u r l te d r te r te e te n o e u u te u h n id s u te m e lu te lu ic te u u fe o u u k l u u lu te n u i e io u u e u s n u o te u e b u u io u r te s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s ix s m n ra s s s n s a ra s a m m a s is s ia la s ia ia s te s s s ia a s s s s s e e a is s s a a s s s a s s s x n s s ia la m s s s a s ra a n s s ia m is is s n s m is a m s n s ia s
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1 1 1 B r P P S o M a P t o M l a P S e T A A A B k Ic e M o a ra r p i r A A n A r B B B h C C D D D D G H H h I n O O e h o P P R o g S T a g A n A t s a a o r o C C l C C D e D id D F G H H e H I t m K L L L M ro ix M o n O o P P P h P a to P s s Q h R e S y c c A m e r h a A l r B o l B C h i C o C a n e im D ip a F o G G a H e H H e x H o c h p Is K L o M M s o m M N c O h P a P a P e g P P P P P p r e P R a h S n r S S S S S u r T h T h A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s lu n e li H i H r H h y o o o u L a e i p M a a M o N to is p r a a l e P e o h P P P o a u t u a R b i R S S p S S g w t t t u s i T T T a o c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c F u c i G G y y n a c e H e lm a m m y n I o I I l n L n th m r e to M M te n o M y e c h m e la la a e a n e P ta p h h y la o o a t m d P e e R d a z u S S a a S p p S e a e e r u b n a e e la o a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y a l o a r ip o l d h g a i u s a G y ro G ro p o e li e lm p a is d o c i m n r p Ix u in a L ic a t L h a m g g m e e i i o r r e o p o a lm e e le o lm t n rm e l h o P y l P la n P l ly ro le o m o ri b a i d d o s a a e g c S ir i i l l l ll S o b l te g T i e rt s rr m n h i r o o in e a A o t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D e t o ic p c o o i y s s F c t G k d r r y y tz H lo H d li c l in o n t o h d b o r o p d e o p L ic o o o c r a s M n c c p u i o s o O io t o o m z P t o e c lo g g y k o v te d e d o o R o c b b r i a S ip S o r r p io d la o b g o r o a s c ra a o t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a im s i g t u h li r d m n r c i F u u ip l s i a i ro p r p c H a r c o i in d io s ra I c n i r g In e d a K h r v a i a r g b L c a i i g i i s i h r h e a n k N d l id m h in o i c a n i a g y b P h o h P la i a ti ir i b r i c n m g o R o e e i t S l S c k d o o ir m i s g ti il l r g m e m T h i s T w a h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F c s l o ta c G n c h o h h a a ti r o o r o th i th i n c u y c i l c d i li K i i i ic m id s p L a r i o t n ll ti r c c o ic c i a i n p e o e i d i i p d h e c h r a c li l ti i h y s d y il io ti n s c d g c ib i c i ii e a l r o p l l c ti a a c o h o e p p o c c l p o o e i d T a a i ti c in ig ti ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r o o y c d h u h c y l y e l m c p m l h c th o c a d ro c h m s h c y i ic c c n c c y i i p i e t a c e o ic c a h h n a h c t c s o r l r c h c o c th h ic e n h e a tu h c y c c o c i o to ic g c o th m h ic u h o c h c c r o u y a s h la la h c p r o y o l s h h r o h o y s r c n c o i a n s c c h o i T a c h a h n h h n n a d r y h h c n h y o n s n n a u u g a y h h c h e is p h r h n h h p p a o a o z it h h ic y n r n p d h h n e h r p h d n x a p k n o l n h l it l o h e it a h o h e n u n h p s t h h e g n h n c e s n e p h c n h h p n n e t n h e h t d i it e h a h r h o y t lc d n l t n h p h h o p c h m h m p n t l n h n h h e r f p ll s y r r n h o it n it m y y a r n p e a k z h r n m s h h n l n h n p n i n n i i s e li a c n n i i n c i i i i p e i i p t n n h n r t i n i id n i n n ir id e d p ia le h n n c a e i i t o n n i r n p o n i i e id p h e te i r a i o li h li li p ip n p i h p o p n li r id y i p i n h i y n n c e i n c i h c p n r h n k i p n t i i r u i n r n e i e h i n m n p n c y u r iu ia is li i n h n n le d h i p n li t i y a i li n i n n i p fe h i e c i i i n r u li e i h c t p p i li h l p e i n p i a e n n i id li is g n u h u u u u u u s l te r u u u te te u u m te te te te h r lu te te e o u u u u io r te u te e u te u u o e z e e h d n u u u u u u lu l te u o u u te io u h u in u u n e s u r l te d r te r te e te n o e u u te u h n id s u te m e lu te lu ic te u u fe o u u k l u u lu te n u i e io u u e u s n u o te u e b u u io u r te te o te u u h u id u o o n o te u u u u u u e o te s n u u te u o te o r te te u te u u te h l u u l u fe te te u t b c n te o o u o h h u te u lu h r n u h u b d l u u te e te o e u s e s m s s m m a ia s ia s s s s s s s a s s s s e ia s s s s n s s s s n ia s s s s s s s s n s a ra s e ia a s s s s m s s ix s m n ra s s s n s a ra s a m m a s is s ia la s ia ia s te s s s ia a s s s s s e e a is s s a a s s s a s s s x n s s ia la m s s s a s ra a n s s ia m is is s n s m is a m s n s ia s s s s s ia s a s a s n ra n m a a s m s s s s s s n s is ia s s s s n s n ia s s s s s s s a la s m ia s x s s s ia a ia ia s s s s n e a m s s s e ia ia s a m ia ia ia s s s s s a a s
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1 B r P P S o M a P t o M l a P S e T A A A B k Ic e M o a ra r p i r U A A n A r B B B h C C D D D D G H H h I n O O e h o P P R o g S T a T r g A n A t s a a o r o C C l C C D e D id D i F G H H e H I t m K L L L M ro ix M o n O o P P P h P a to P s s Q h R e S y c u o c A m e r h a A l r B o l B C h i C o C a n e im D p a F o G G a H e H H e x H o c h p Is K L o M M s o m M N c O h P a P a P e g P P P P P p r e P R a h S n r S S S S S u r T h T h T h A a la l c rc A A ro p A A A s A a o B B s r o z u a h C m C p C o C t C y C c D d s y D ip l E E s lu n e li H i H r H h y o o o u L a e i p M a a M o N to is p r a a l e P e o h P P P o a u t u a R b i R S S p S S g w t t t u s i T T T a o T T r a V c n s A lo p li h h r h t A n c B B i i tr a k b n C C o C o i a o C C o r o s r e C y l t D e r m D m o l ip o c u F c F u c i G G y y n a c e H e lm a m m y n I o I I l n L n th m r e to M M te n o M y e c h m e la la a e a n e P ta p h h y la o o a t m d P e e R d a z u S S a a S p p S e a e e r u b n a e e la o T ic p e o Z a t p g c h d a re th p o s s s e s u o c e B f B B fu i n r y ly a a m n rc c c n in r m o n u c in y y a l o a r ip o l d h g a i u s a G y ro G ro p o e li e lm p a is d o c i m n r p Ix u in a L ic a t L h a m g g m e e i i o r r e o p o a lm e e le o lm t n rm e l h o P y l P la n P l ly ro le o m o ri b a i d d o s a a e g c S ir i i l l l ll S o b l te g T i e rt s rr m re r ri h s T U lm l o n h i r o o in e a A o t te te r te g o e e a i i c c v u e c p u a h d t o o o o s C o r d li l D e t o ic p c o o i y s s F c t G k d r r y y tz H lo H d li c l in o n t o h d b o r o p d e o p L ic o o o c r a s M n c c p u i o s o O io t o o m z P n t o e c lo g g y k t o v te d e d o o R o c b b r i a S ip S o r r p io d la o b g o r o a s c ra a o p ia c T e U k Y t o d i r i e n o d id e r o r li l i a l r s fo f rc o h a n r r o o r u i h n n p is p o s p v lo r n o a im s i g t u h li r d m n r c i F u u ip l s i a i ro p r p c H a r c o i in d io s ra I c n i r g In e d a K h r v a i a r g b L c a i i g i i s i h r h e a n k N d l id m h in o i c a i a g y b P h o h P la i a i ir i b r i c n m g o R o e e i t S l S c k d o o ir m i s g ti il l r g m e m T h i s T w a v h o r i y m U in i a o h r i c ti i c c o A ic rt p i i ri ri i p o c o c c o g c u u p i li n i a le ti n i li c li i i s C p h r o ru h o i i d p e s c r y li i c a i o i i c e F c s l o ta c G n c h o h h a a ti r o o r o th i th i n c u y c i l c d i li K i i i ic m id s p L a r i o t n ll ti r c c o ic c i a i n p e o e i d i i p d h e c h r a c li l ti i h y s d y il io ti n s c d g c ib i c i ii e a l r o p l l c ti a a c o h o e p p o c c l p o o e i d T a a i ti c in ig ti ti e i p is c li c f n c W k p ic h c h c c h h n r o h h c c a c c h s h s h o r a ic rm n l h c c i c t d r c d to c h c c s o o h i h p o l c c r h d o e h a o c c h t c o s c fo h li r o o y c d h u h c y l y e l m c p m l h c th o c a d ro c h m s h c y i ic c c n c c y i i p 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