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Logunov 2003 a Revision of the Genus Yllenus.Pdf INSTITUTE FOR SYSTEMATICS AND THE MANCHESTER MUSEUM, ECOLOGY OF ANIMALS, THE UNIVERSITY OF MANCHESTER, SIBERIAN BRANCH OF THE RUSSIAN MANCHESTER, UK ACADEMY OF SCIENCES, NOVOSIBIRSK, RUSSIA A revision of the genus Yllenus Simon, 1868 (Arachnida, Araneae, Salticidae) by Dmitri V. Logunov & Yuri M. Marusik Edited by Kirill G. Mikhailov Illustrated by the authors KMK Scientific Press Ltd. Moscow v 2003 D. V. Logunov & Y. M. Marusik. A revision of the genus Yllenus Simon, 1868 (Arachnida, Araneae, Salticidae). Moscow: KMK Scienetific Press Ltd. 2003. 167 pp. In English. Ä. Â. Ëîãóíîâ, Þ. Ì. Ìàðóñèê. Ðåâèçèÿ ðîäà Yllenus Simon, 1868 (Arachnida, Araneae, Salticidae). Ìîñêâà: èçäàòåëüñòâî ÊÌÊ. 2003. 167 ñòð. Íà àíãëèéñêîì ÿçûêå. Kirill G. Mikhailov (the editor) This is a worldwide taxonomic revision of the genus Yllenus, with 65 valid species being diagnosed, figured and (re)described. A key to three species groups and distribution maps for all revised species are also provided. The # neotype is designated for Yllenus albifrons (Lucas, 1846). Lectotypes are designated for the four following species: Attus albocinctus Kroneberg, 1885 ($); Attus vittatus Thorell, 1875 (#); Yllenus hamifer Simon, 1895 ($); Attulus validus Simon, 1889 (#). Three species names are newly synonymized: Yllenus somonensis Prószyñski, 1982 with Y. validus Simon, 1889; Yllenus israelensis Logunov, 1996 with Y. tschoni (Caporiacco, 1936); Yllenus probatus Weso³owska, 1996 with Y. mirandus Weso³owska, 1996. Three new combinations are proposed: Yllenus auriceps (Denis, 1966) comb.n. (ex Attulus), Yllenus maoni- uensis (Liu, Wang et Peng, 1991) comb.n. (ex Philaeus) and Euophrys skalanicus (Dobroruka, 2003) comb.n. (ex Yllenus). Three specific names are treated as nomina dubia: Attus pusio Simon, 1871, Euophrys skalan- icus (Dobroruka, 2003) and Yllenus kronebergi Roewer, 1951. Twenty nine new species are described: Ylle- nus algarvensis sp.n. (#; from Portugal); Y. aralicus sp.n. (#$; from the E. Caucasus and Middle Asia); Y. bakanas sp.n. (#$; from Kazakhstan); Y. bucharaensis sp.n. ($; from Uzbekistan); Y. charynensis sp.n. ($; from Kazakhstan); Y. dalaensis sp.n. (#$; from Kazakhstan); Y. dunini sp.n. (#$; from Azerbaijan and Kazakhstan); Y. erzinenis sp.n. (#$; from S. Siberia and Mongolia); Y. gavdos sp.n. (#$; from W. Mediter- ranean); Y. guseinovi sp.n. (#$; from the E. Caucasus and Middle Asia); Y. halugim sp.n. (#; from Israel); Y. karakumensis sp.n. (#$; from Turkmenistan); Y. karnai sp.n. (#$; from NW India); Y. kononenkoi sp.n. (#$; from Kyrghyzstan); Y. kotchevnik sp.n. (#; from Turkmenistan); Y. mirabilis sp.n. (#; from Uzbeki- stan and Turkmenistan); Y. murgabicus sp.n. (#$; from Tajikistan); Y. nigritarsis sp.n. (#; from Turkmen- istan); Y. nurataus sp.n. ($; from Uzbekistan); Y. pamiricus sp.n. (#$; from Tajikistan); Y. pseudobajan sp.n. (#$; from SW China); Y. pseudovalidus sp.n. (#$; from Turkmenistan and Kazakhstan); Y. shakhsenem sp.n. (#; from Turkmenistan); Y. tamdybulak sp.n. ($; from Uzbekistan); Y. turkestanicus sp.n. (#$; from Middle Asia); Y. uiguricus sp.n. (#$; from Kazakhstan); Y. uzbekistanicus sp.n. (#$; from Uzbekistan and Turkmenistan); Y. zhilgaensis sp.n. (#$; from Kalmykia and Kazakhstan); Y. zyuzini sp.n. (#$; from Kaza- khstan and Turkmenistan). This work is intended for arachnologists, mainly taxonomists, as well as for biogeographers, local biol- ogists and naturalists. Äàííàÿ ðàáîòà ïðåäñòàâëÿåò ñîáîé ìèðîâóþ ðåâèçèþ ðîäà Yllenus, â êîòîðîé 65 âàëèäíûõ âèäîâ îïèñàíû èëè ïåðåîïèñàíû è èëëþñòðèðîâàíû. Ñîñòàâëåí äèàãíîñòè÷åñêèé êëþ÷ äëÿ òðåõ ãðóïï âèäîâ è äàíû êàðòû ðàñïðîñòðà- íåíèÿ äëÿ âñåõ âèäîâ. Âûäåëåí íåîòèï äëÿ Yllenus albifrons (Lucas, 1846). Îáîçíà÷åíû ëåêòîòèïû äëÿ ÷åòûðåõ âèäîâ: Attus albocinctus Kroneberg, 1885 ($); Attus vittatus Thorell, 1875 (#); Yllenus hamifer Simon, 1895 ($); Attulus validus Simon, 1889 (#). Òðè âèäà ñèíîíèìèçèðîâàíû: Yllenus somonensis Prószyñski, 1982 ñ Y. validus Simon, 1889; Yllenus israelensis Logunov, 1996 ñ Y. tschoni (Caporiacco, 1936); Yllenus probatus Weso³owska, 1996 ñ Y. mirandus Weso³owska, 1996. Ïðåäëîæåíî òðè íîâûå êîìáèíàöèè: Yllenus auriceps (Denis, 1966) comb.n. (ex Attulus), Yllenus maoniuensis (Liu, Wang et Peng, 1991) comb.n. (ex Philaeus) è Euophrys skalanicus (Dobroruka, 2003) comb.n. (ex Yllenus). Òðè âèäîâûõ íàçâàíèÿ òðàêòóþòñÿ êàê nomina dubia: Attus pusio Simon, 1871, Euophrys skalanicus (Dobroruka, 2003) è Yllenus kronebergi Roewer, 1951. Îïèñàíî 29 íîâûõ âèäîâ. Ðàáîòà ïðåäíàçíà÷åíà äëÿ àðàõíîëîãîâ, ãëàâíûì îáðàçîì òàêñîíîìèñòîâ, à òàêæå äëÿ áèîãåîãðàôîâ, êàðåâå- äîâ è íàòóðàëèñòîâ. All rights reserved. No part of this book may be reproduced, stored in a retrieval system or transmitted in any form by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior written permission of the copyrights owner. Cover illustration (front): landscape, sandy desert with Haloxylon sp. in Central Kyzylkumy (N. Uzbekistan), where a number of Yllenus species occur (a photo by DL); spider, the male of Yllenus arenarius from sand dunes in Poland (courtesy by Dr M. Bartos; £ód, Poland). © òåêñò, èëëþñòðàöèè, êàðòû, Ä.Â. Ëîãóíîâ, Þ.Ì. Ìàðóñèê, 2003 ISBN 5-87317-126-2 © èçäàíèå, èçäàòåëüñòâî ÊÌÊ, 2003 Contents Introduction .........................................................................................................4 Material and methods.....................................................................................4 Terminology ...................................................................................................4 Acknowledgements ........................................................................................5 Genus Yllenus Simon, 1868.................................................................................8 Definition .......................................................................................................8 Diagnosis and affinities................................................................................13 Composition .................................................................................................16 Distribution ..................................................................................................17 Natural History.............................................................................................20 Key to the species groups of Yllenus ...........................................................22 Survey of species ...............................................................................................23 The albocinctus species group .....................................................................23 The arenarius species group ........................................................................85 The hamifer species group ......................................................................... 118 Doubtful and invalid species names ..........................................................156 References .......................................................................................................160 Index to species ...............................................................................................166 4 D.V. Logunov & Yu.M. Marusik Introduction MMUM The Manchester Museum, The Uni- versity of Manchester, Manchester, UK (Dr D. V. Logunov) The genus Yllenus was erected by Simon MNHN Muséum national dHistoire naturel- [1868]. It was revised by Prószyñski [1968], le, Paris, France (Dr C. Rollard) who assigned 22 species to it. After Prószyñs- MRAC Musée Royal de lAfrique Centrale, kis revision a few species were added by Pun- Tervuren, Belgium (Dr R. Jocqué); da [1975; 2 species], Ponomarev [1978; 1 spe- MZSF Museo Zoologico de La Specola, Sez- cies], Liu et al. [1991; 1 species], Logunov ione del Museo di Storia Naturale, Firenze, Italy (Mrs S. Whitman) [1993b; 1 species], Logunov & Marusik NHMC Natural History Museum, University [2000a; 5 species] and Hu [2001; 1 species]. of Crete, Crete, Greece (Dr M. Chatzaki) Taking into account currently established syn- PSUN Department of Zoology of the Perm onymies, 32 valid species were known before State University, Perm, Russia (Dr S. L. Esyunin) this second revision, which covers 65 valid SMNH Swedish Museum of Natural History, species. Stockholm, Sweden (Dr T. Kronestedt) The main aim of the present work is to pro- SNMC Museum of Natural History, Slovak vide an up-dated taxonomic account of the ge- National Museum, Bratislava, Slovakia (Mr J. Sva- nus Yllenus which will (1) redefine the genus, toò) SZMN Siberian Zoological Museum of the In- (2) establish new synonymies (3) (re)describe stitute for Systematics and Ecology of Animals, all valid species, (4) refine their distributions, Novosibirsk, Russia (Ms G. N. Azarkina) and (5) comment on all invalid, doubtful or prob- YMPC The personal collection of Mr Yvan lematic species names. Montardi (Paris, France) ZISP Zoological Institute, Russian Academy of Science, St. Petersburg, Russia (Dr V. A. Krivo- Material and methods khatsky) ZMCD Zoological Museum, Copenhagen, Den- A total of 992 adult specimens (both museum and mark (Dr N. Scharff) newly collected ones) were used in this study; all of ZMHU Zoological Museum of the Helsinki them were borrowed from or distributed among the University, Helsinki, Finland (Dr J. Terhivuo) following depositaria and personal collections: ZMPA Institute of Zoology, Warsaw, Poland AMNH The American Museum of Natural (Dr T. Huflejt and Prof. J. Prószyñski) History,
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