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ROHACEK8 MARSHALL: 31 3-372 Studia dipterologica 7 (2000)Heft 2 0 ISSN 0945-3954

A world revision of the seaweed genus DUDA (Diptera: : ) Part 2: Palaearctic species

[Revision der tangbewohnenden Dungfliegen der Gattung Thorucochuetu DUDA der Weltfauna (Diptera: Sphaeroceridae: Limosininae) Teil 2: palaarktische Arten]

by Jindiich ROHACEK and Stephen A. MARSHALL Opava (Czech Republic) Guclph (Canada)

Abstract Thc Palaearctic species of the seawecd fly gcnus Thomcochaera DUDA. 1918 are revised; the 13 species recognized in the rcgion are keyed, (re)described and thcir male and female genita- lia illustrated. Thomcochaera nigripennis (DUDA.1918), L rudis VANSCHUMBROECK, 1951, firrknui (REMMERT,1935) and Z tunísica PAPP,1978 are synonymised witb Z bmchysroma (STMHAMMAR,1855), and ZpoipebrLr CARLES-ToLRA,1994 is synonymised with Z medirenunea MUNARI,1989. The following seven new species are described Thoracochoeta accola [Gnece, Ukraine, Georgia]. Z acinoces [China, Japan]. Z cercuILr and Zgemina [Rusia: Kuril Is.]. T. miranda [Japan], Z Qnx [England], and X valentinei [Great Britain, Belgium]. Variability of sorne spccies is discussed. Notes on the relationships, biology and distribution (including new - records) of al1 species are given. T. bmchysroma is newly recorded ñom indonesia (Sumatra). Key words Diptera, Sphaeroceridae, Thomcochoeta, , nnv species, new synonyms, key, distri- bution, Palaearctic, marine shorelines, supralittoral

Zusarnmenfassung Die palaarktischen Arten der tangbewohnendcnDungfliegen der Gattung Thomcochaera DUDA, 1918 wcrden revidiert. Es handelt sich um insgcsamt 13 valide Spezies. Ein Bcstimmungs- schlüssel diescr Arten wird vorgestellt; die Artcn werdcn detailliert beschrieben und dic mhlichen und weiblichen Genitalien abgcbildct. Thomcochaera nigripennis (DUDA,1918), T rudis VANSCHUYTBROECK,1951, tfiftkaui(RE~~ER~, 1955) und L tunisica PAPP, 1978 wrden mit L brachysroma (STENHAMMAR,1855) und Z paipebris CARLES-TOLLA,1994 mit L medi- rermnea MUNARI,1989 synonymisiert. Es erfolgi die Beschreibung von sieben neuen Arten: Thoracochaeru uccola [Gricchenland, Ukraine, Georgien], Z acinaces [China. Japan], Z cercalis und T. gemina [Russland: Kurilen], Z miranda [Japan], Z knr [England] und Z vafenrinei (GroBbritannien, Belgien]. Die Variabilitgt ciniger Arten ist Gegenstand der Diskussion. Anmerkungen N denVewndtschaftsbczichungen, der Biologie und Verbreitung (nebst neuer Fundmeldungen) von allen Artcn erganzen dic Ausführungen ZUT Taxonomie. Thomcochaeta bMChyStoma wird erstmals aus Indonesien (Sumatra) gemeldet. Stichwarter Diptera, Sphaeroceridae, Thomcochaeta, Taxonomie, neue Arten, ncue Synonyme, Bestim- mungsschliissel, Verbreitung, Paliiarktis, Meercsufer, Supralitoral

Introduction h4hs1.1~~(1 982) revised the Nearctic species of the genus Thomcochuetu DUDA, 19 18 and MAR- SHALL & RoHÁ~(2000) studied the world species with the exception of species found in the Holarctic Region This paper revises the Palaearctic Thorucochaetu species, thus completing a world revision of the genus. Many of the Palaearctic species now placed in Thomcochuetu(Limosinu zosteme HALJDAY,1833; L.fulvipes MEIGEN,1838; L. setulosu ZETERSTEDT,1847; L. bmchystomu S~~AMMAX,1855; L. piligera STEMM&IAR,1855; L. ciliosa RONDA,,,1880 and L. undalusiuca STROBL, 1900) were descnbed in the 19th century, and problems in establishing the identity of

7 313 scrratc apex. Fciiiaie tcrgite 7 enlargcd and bulgiiig dorsolaterally (Fig. 75), steriiite spine-like sctac iii front of a largc, postcriorly angular, posteroiiicdial lobc; iiialc 7 posteroiiicdially coiicavc aiid stcriiitc 8 (Fig. 76) spatulate (Great Britaiii: Eng- ccrci large, proiniiieiit aiid richly sctosc (Figs 80, 81); surstylus (FIg. 87) unilobatc, laiid) ...... I: farix spec. nov. with 2 strong ventral spines. Feniale tergitc 7 (Fig. 84) with incdial unpigmcntcd Arista loiig-ciliate; male f, and t, ventrally simply setulose, t, with only 1 vu seta. stripe; sternite 7 (Fig. 85) with inembranous, traiisvcrscly roundcd postcromcdial Male sternitc 5 with posteromedial bifid lobe and stout spines in front of it; surstylus area (Russia: Kuril 1s.) ...... T. cercafis spec. nov. witli botli anterior and posterior lobes slendcr. Feiiiaic tergite 7, sternite 7 and sternite 4 (2+2) long dc; sc setae longer (laterobasal sc 1.1 X as long as scutellum). Male 8 simple (Figs 43, 50) ...... 8 sternite 5 (Figs 105,110) sparseiy and shortiy setose, with a row oferect or proclinate Larger (wiiig ieiigth 1.5-2.0 inni); veiiis on wing brownish; dc macrosetae (1+2) long. setae in front of posteriorly rounded posteromedial lobe; male cerci rcduced; surstylus Malc sternite 5 (Figs 34,36) with a coinb of dense stout spines in front of posterome- trilobatc (Figs 108, 117), with only 1 posteroventrai stout spine. Fcmale tergite 7 dial lobe; surstylus (Fig. 33) witli anterior lobe slightiy bent and apically tapcred; and sternite 7 simply pigmented ...... 12 postgonite (Fig. 3 1) niore slender, siiglilly sinuously margined. Femaie postabdominal 12(11) g seta about 1.5 X as long as subvibrissa. Posteromediai iobe of inale sternite 5 (Fig. sclcrites (Figs 40,43) narrower and ccrci (Fig. 40) longer and more slender (Balcaric 105) with bare and slightiy emarginate posterior margin; inale cerci ventrally rouiidcd Is., Pantelleria I., Malta, ) ...... Z rrreúiterrariea MUNARI,1989 (Fig. 104); surstylus (Fig. 108) long, witli small antcrior lobc; postgonite (Fig. 102) Sinaller (wing lengtli 1.2-1.6 min); veins on wing (cxcept for C) strikingly pale, dirty with naiyow apcx. Female sternite 8 (Fig. 109) rounded-hexagonal with deep postero- wliitisli; dc macrosetae (2+2) sliorter. Male sternite 5 (Figs 47-48) with only scveral inedia1 incision; spermathecae (Fig. 107) traiisvcrsely oval, witli groovcd surface and scattcred spines in front of posteromedial lobe; surstylus (Fig. 52) witli anterior lobe distinct cup-shaped basa1 part (Holarctic) ...... 3: seticosta (SruLm, 1925) sinuous and blunt on apex; postgonite (Fig. 46) more robust, with broadened middle - g seta about 2 x as long as unusually sliort subvibrissa. Posteroinedial lobe of male stemite and apical parts. Feinaie postabdominal scierites (Figs 49,50) broader and cerci (Fig. 5 (Fig. 110) witli spinuiose and medially deepiy emarginate posterior iiiargh; surstylus 49) sliort and stout (Greece, Ukraine, Georgia) ...... T. accofa spec. nov. (Fig. 117) short, with largc flat antcrior sabre-sliaped lobe; inalc cerci ventrally angular Eyc uiiusually iarge and convcx (its Iieight about 3.6 x as long as genal height); poste- (Fig. 113); postgonite (Fig. 112) with broad rounded apex. Fcmale sternite 8 (Fig. 1 15) rior notopleurai seta conspicuously reclinate. Male stcrnitc 5 (Fig. 93) with 4 robust broad antcriorly and tapered posteriorly; spermathccae (Fig. 118) almost spliaerical, with spines in front of a narrow rounded posteromedial lobe; surstylus (Fig. 91) with pointed sparsely and fiely wrinkled surface (China, Japan) ...... Z aciriaccs spec. nov. antcrior lobc and 2 robust ventral spines; dista1 portion of basipiiallus with lateral wing-like, toothy projections. Female postabdomen (Figs 94, 95) very slender, with cerci tapcrcd in the niiddle (Russia: Kuril 1s.) ...... Z gcrniria spec. nov. Descriptions and notes o11 tlie species Eye sinallcr and flatter (its height at most 2.8 X as long as genai height); posterior notopieurai seta erect (as usual in Tltorucoc.haefa).Male sternite 5 with different Tlioracocliaeta bracliystorria (STENHAMMAII,1855) postsromedial lobe and more nuinerous spines in froiit of it; surstylus either with (Figs 4.9-22) only 1 posteroventral spinc or no anterior lobe developed. Female postabdoiiien Liniosina bmchystorrra STENIIAMMAR,1855: 393. broader and cerci different ...... 10 Lirtiosina (Thomcochaeta)bruchystoma. -DUDA,1918: 100; DUDA, 1938: 98; REMMERT,1955: 471. Leptocem (Thoracochaeia)bmchystorna. -DUDA, 1Y25: 198, SruLER, 1925: 120; TENORIO, 1968: 183,190;RICHAKOS, Both presutural dc sctae slroiigiy inclinate (never reclinate); 1 ac microseta present 1973: 329; MARSHALL, 1982: 69-71. between prescutellar ac and dc setae (Fig. 2); male f, and t2 ventrally witliout special- Leptocern bmchystoma. - Sttiuv, 1934: 464. ized sctae or spines. Male sternite 5 (Fig. 63) with 2 separate combs of spines in front Thomcochaeta bmclrysioma. - FREY,1941: 26; Pm,1973: 34; Pm,1984: 85;MARSIIALL & ROIIACEK,2000: 268-273. Limosino anúalusiaca STROBL,1900: 69; DUDA, 1918: 100 [synonymy]. of sinall posteromedial lobe; surstylus (Fig. 69) posteroventrally witli only 1 long seta (Thomcochoefu)bmcliysfon~a var. nigripennis DUDA, 19 18: 102 syn. nov. in addition to usual stout spine; postgonite (Fig. 65) robust and apically bilobed. Fe- Thomcocliaeta nigriyennis. - Pm, 1984: 86. maie tergitc 8 projecting on ventral side of abdomen (Fig. 70), ccrci (Fig. 67) with Leptocem ruJa SruLER. 1925: 122; MARSHALL.1982: 69 [syiionymy]. Lepfocem (Thomcochaeta)m/o. - RICHARDS,1973: 329. sliort thick setae at apex (subcosmopoiitan) ...... Z zosterae (HALIDAY, 1833) Thomcochaeta rudis VANSCIIUYTBROECK,1951: 186 syn. nov.; PArr, 1984: 85. At lcast 1 (posterior) presutural dc more reclinate than inclinate; usually no ac Limosino (Thomcochaeta)jittkauiREMMERT. 1955: 471 syn. nov. inicroseta bctween prcscutellar uc and dc setae (it rarely occurs in Z ucinaces spec. Lepfocera (Thomcochaeta)jitt~ui.- RICHARDS, 1963: 120; RICHARDS,1973: 329. nov.); inaie fi and tz ventrally witli rows of sliort tliickened setae or spines. Male Thomcochaetajitlui. - PArr. 1984: 85. Eíachistonta spinicosla COLLIN.1966: 36; Pm,1984: 83; MUNARI,1994: 195 [synonyiny]. sternite 5 with 1 comb or row of spines; surstylus posteroventrally with numcrous Leplocera (T.)fircico/aRlctinRus, 1973: 32ú; MAKSMALL& I~oHACEK,2000: 268 [synonyiny]. iong setae; postgonite with apex simple. Female tergite 8 without ventral projec- Thomcocliaeta tunisica PAw, 1978: 198 syn. nov. tioiis and cerci wilh long setae apically ...... 11 ’Qpe material L(10) Qnly 3 (1+2) long dc; sc setae sliorter (laterobasal sc 0.8 x as iong as scutellum). Limosina brochysrorna STENIIAMMAR:Neotype niale (MZLU, hereby dcsignaicd, gcnit. prcp.), labeiled: “S: Sk. Lomiiia. 6.9.1975 Stranden. leg. H. Andersson”, ‘Thoracochaeta brachystoma (Sienhanimr, I854), 6 dci. F. Florén 1983” Male sternite 5 (Fig. 82) densely and long setose, with a row of reclinate to exclinate and “Neotypus 8,Liniosina brachvstoma Stenli.. J. RoháEek des. 2Wflahel wiih red niargin niid “Neotvaus”). ., .----.--* .-”.,. ...., _..... zII

;22 323 ROHACEK& MARSHALL:Aworld revision of ... Thoracochaefa DUDA (... Sphaerocendae ...) ... Part 2 _._ RO~CEKB MARCHALL: Aworld revision of ... Thoracochaeta DUDA(... Sphaerocendae ...) ... Part2 ... setae; cerci reduced and fused to form subanal platc, eacli with 1 long and 2-3 short weak setae. PArp, 1978 is also a new synonym of T. bmchysforira. It was describcd from 17 specimens Hypandriuni short; its anteromedial apodeme flat, triangular in dorsal view. Surstylus (Fig. 12) with Iiolotypc and scveral paratypes werc lost during the post rcturn froin BMNH (B. R. Pitkin slender, apicaliy pointed anterior lobe and with lcss tapered posterior lobe beariiig distinct lateral Papp, personal commiication, 2000) and, thcrefore, a ncotypc was selectcd and designated kcel and 1 apical stout but short spinc. Sctae on external and ventral sides of surstylus long and paratypcs loaiicd froni HNHM. Tlie iiialc neotype has stcrnitc 5 arnied by 5 spiiic-likc se posteroextcrnal area of posterior lobe (behiiid lateral keel) finely pubcsccnt. Acdcagal coinplex (Fig. relatively short setac on surstylus and scutelluin but al1 thesc fcatures fall witliiii tlic variabilit i 1). Phallapodeme simple; basiphallus with fuscd posterior part aiid aiilcrior projcction (ovcrlapping brachysforna.AII othcr paratypcs exaniiiicd (2 males, 3 fcinalcs) bclong to tlic saiiic spccics. but not fused with proximal part of distipliallus), ventrally broadly, rouiidedly rectangular. Distiphdi- Hiology: One of tlie coininoncst specics of Thorucochaefa worldwidc. Altliougli it lus shorter (han basiphallus, relatively simply sclcrotized, funncl-shapcd. Postgoiiitc wcakly sinuatc, niasse oii coastal wrack and its clouds can bc blowii far inlaiid by wiiid, it is only rarcly scci with distal third iiarrowed, quadrate (witli iiiinutc subapical point) apically; antcrior margin of the from sca shores becausc it probably is unable to pcrsist iii inland Iiabitats. Thc fc middlc broad part distinctly finely Iiaircd (see Fig. 11). Ejaculatory apodcme short, witli broad prox- (TESCIINEK1958; FLORBN1989, four spcciinens exaiiiiiicd) dcal witli single spcciiiicns col imal part and narrow rod-likc distal part. dung-baited traps, a cowslied, a sawdust pilc, a potato ficld, aii urinal and on dccaycd fr Feniale abdoiiieii: Posiabdomcn relatively broad; its tergites beconiing only sliglitly iiarrowcr poste- plants. Tlierc is also onc qucstioiiablc rccord froni a subiiioiitaiic cave (PAX & MAsctiKa 1935) riorly. Tergitc 6 simple, setae few and sliort; tcrgite 7 usually with only 6 setae at posterior margin. of this species are parasitised by cucoiiid wasps in Lhc gciius Kleiúoforna. Tergilc 8 complete bui dark pignientcd and sctose only laterally (Fig. 17). Tcrgitc 10 broad but short, Distribulion: Cosmopoiitan in coastal arcas, bu1 rarcr iii tropical arcas (see MARsiiALL & R with cerci inscrted in cmarginaled posterior niargin and with a pair of sliort setac and some niicropu- 2000). Afrotropical: Cameroon, Ghana, Scyclielles, South Yciiicii, Soutli Africa, Tristan da bescence oii disc. Sternitc 6 broadcr and with morc setae than tergite 6; sternitc 7 longer than tergite AustralasiadOccanian: Australia, Cook Islaiids, Hawaii (USA), New Caledonia; 7, simple, witli a iiuinber of short setae. Sternitc 8 (Fig. 18) largely membranous witli niicropubcs- (NB, NS, PE, QB,), USA (CT, FL, MA, NH, NY, RI, TX); Ncotropical: Argcntina, Antigua cence in distinct pattern, a pair of long lateral setac and 4-6 sliort posterior sctulae. Sternitc 10 Beiize, Bcriiiuda, Chile, Doiniiiica, Eastcr lsland, Jamaica, Juan Fernandcz 1s. (Cliilc), relatively largc, rounded pentagonal, pale-pigmented, with micropubescence rcstrictcd to postcrior QR), Peru, Venezuela; Oriental: Pliillipincs, liidonesia (ncw); Palaearctic: Adcn, Algcria, Azore half and several setulae at posterior margin. Interna1 “spcctacles-sliapcd” sclerite (Fig. 19) largely ‘ ‘ tugal), Bclgiuin, Bulgaria, Caiiary 1s. (Spain), Croatia (iicw), Czcch Rcpublic, Dcniiiark meinbranous, with narrow rings and small paie scieritc at inscrtion of spermatliccal ducts. Spcrina- land, , Geriiiany, Gcorgia (new), Great Britaiii, Greccc (ncw), Italy, Lcbaiion ( thecac (Fig. 22) dark, almost sphaerical, flat-toppcd, with deeply grooved surfacc, shallow terminal Madcira (Portugal), Malta (iiew), Polaiid, Rouniaiiia, Spaiii, Swcdcii, Tunisia, Ukraiiic (iicw), invagiiiation aiid short but relatively narrow sclcrotised parts of ducts. Cercus robust, soinewhat flattencd dorsovcntrally, with wcak and sliort setae, only apical seta soinewhat loiiger. Tiioracociiacta valciitiiiei spcc. nov. Cominenls: Iii spite of Iiigli variability in external features (see below) Thoracocliaefabrachysfonra (Figs 23-30) can bc easily recogiiised bccause 1: valenfinei spcc. nov. is the only specics in Uie area witli similar mid tibia1 chaetotaxy. Tliese species are distinguished by characters given in tlie key and by tlic Type material: Holotype male (BMNH) labelled: “U.K.. Sornersct, Berrow Saiids, under seawced, 2S.xi. 199 ignated ‘T2’, Jolin VALENTINE”.Paratypes: 7 malcs 8 fcmales (UMNH, DEBU, SMOC), sanie data as for Iiolo different iiiale and feinale gcnitalia (see under L valenfinei spec. nov.). males 1 female (BMNH, DEBU, SMOC). GREAT BRITAIN: Esscx, Walton, under seawced, 8.iv.1992. dcsi Variability: 1: bmchysbma has unusual variation in body lenglh, lengtli of sctac, wing characteristics ‘T3’,John VALENTINEleg.; 1 íeinalc (BMNH), GREAT BRITAIN: Scoilaiid, Barra, Ardvyran, o11 Ln‘nlcm~ti and cliactoiaxy of tlie male fiNi stcrnite. A relatively higii portion of southcrn hcniisphcrc specimens of S.viii.1976,A. R. WATERSTONkg.; 1 icmalc (ISNB), BELGIUM: Kiiockc dm,1 i .xi.lY49, M. BEQUA~RTIcg. L bruchysloina liad oiily 2-3 stout sliort spiiic-like setae on sternitc 5 while Holarclic specinicns nor- Budy lenglli: 1.3-2.0 inin, wing Iciigtli 1.1-1.6 nim; gciicral colour dark brown, iiiic iiially Iiavc 4 lo 6 spiiics and tlie largcst exaiiiined speciincns (1 male from Soutli Africa, 2 froin Bcl- dull; lcgs and thoracic pleura (particularly veiitrally and plcural sutures) paler brown; giuiii, 2 froiii Bulgaria) liad 7-8 spiiies. We havc, Iiowcver, also secii 2 very small iiialcs from Fraiice and ish-opaque. Head: Frons broad but frontal triaiigle vcry narrow, greyish microtoinciitose; Canary 1s.. with only 2 of Uiese spiiics. Tlie variability of this feature scenis to bc loosely corrclaíed taiia separated froiii orbits and frontal trianglc by narrow blackisli browii stripcs foriniiig wiUi body size (smailer spccinicns have fewcr spines on sternitc 5). both in brucltystorrta and in otlier L pattern. Interfroiital area aboul as broad as high, margiiicd by 4 pairs of i/; setac, tli of Uie gciius (sce accola spec. nov. or jolansorii for example). Tlie rclativc lengths of Cs, and species L L soinchat shorter (not iii holotype) and flaiiked by lateral setula. Distancc bctween posterior 0.7 0.75-1.4 X) Cs, (ratio rangiiig bctween and 1.6), and of apical scutcllar setac versus scutelluiii (ratio only 0.6-0.7 X as iong as distancc bctwccn anterior ocellus aiid frontal suture, thus ocellar tri exhibit coiispicuous variability that has led Lo maiiy syiionyms of E bmcliystoma (sce bclow). distinctly iiarrower thaii tliat of T bruchysforna;oc sctac insertcd outsidc tlic border of occllar ir¡ Notes on synonyms: The identity of Thomcocliaeiu bmchysionia is fixed herc by neotype dcsigna- but not in iine with fr setae. Macrosctac oii froiis rciaíively sinail while inicrosctac weil dcvcl UDA C UDA to tion. D (1918, 1938) considered nigripennis (D , 1918) be a dark-coloured variety of L iiiciinate ori (5-6) in a distinct row, wliilc excliiiate uds iii two groups: posterior iicar ors bruchystoma, but PArr (1984) resurrected it as a valid species. Examination of tlic female holotype longer setulac and anterior in front of eye with shorter 3-4 setulae. oc (strongly cxciinat (including its postabdoininal structures) revcalcd that DUDA (I.c.) was right; thc specimcn belongs to both ors similar in length; occi almost of tlie samc Iciigtli but wcakcr, occe distiiictly sliortc 1: brachysfonia altliough it has darker wings than typicai specimens of the latter species. However, weakcr; ors cqual in sizc and equidistant froiii eyc margin; true yvi absciit. Oceilar trianglc flai darkcr specinicns are not rarc in (particularly northern) populations of L 6racliystoriia. Thc Iioloiypc a few (3-4) proclinatc and cxclinate microsctae; about 5-6 siinilar microsctae also bc of Tlioracochaefaruúis VANSCHUYTBROKK,195 1 was found to be only a large (body length 1.75 mm, ocelli. Facc vcry broad, rcddish or oraiigc browii like frontal lunulc cxcept for incdial d wing Iciigth 1.8 mm) female of L bracltysfonia.Ljitfhui (REMMERT,1955) is syiionyinized hcre with contrasting with dark brown froiis; carina distinctly buigiiig bctween aiitcnnac. vi stout, 1.8- L brachystorna oii tlie basis of 6 paratypcs found iii BMNH. Gcnitalia of thc disscctcd male paratype gcnal hcight; g scta sliglitly longer than gciial Iiciglit aiid as long as siibvtbrissa. Eyc siiboval aaree perfectly with typicñl specimeiis of T brncliysforiia iii having 6 spine-iikc setac on stcrnite 5 *. -

lkliilc iil~~ll~iiicii:slcriiilc 5 (l:ig. 54) rcl;itivcly Iliiig. Willi lqc. pdc pigiilciiicd ;iiitI I'ii bcscciii posicroiiictliiil iircii l~iiicoiiiplcicly liichiiig iiiiy iiriiiiiiiiic (11' rohiisi sct;ic OIspiiics iiiii (I'igs 55-56) sliori aiid hroad. witli dciisc iiiid loiig sciiie; aiiol fissurc largc; ccrci rctli projcciiiig vciiiraily, cacli ariiicd by I lotig aiid scvcral siiiali sctac (Fig. 56). I lypaiidriuiii apodciiic llai. triaiigular iii dorsal vicw. Surstylus (Fig. 61 ) slciidcr, dccply bilobcd, iis cxt witli loiig sciac bui wiilioui iiiicropubcscciicc; boili aiilcrior aiid posicrior lobcs siiiiiiar. ve ilic postcrior without robusi spinc aiid wiili oiily a fcw setulac. Acdcagal coinplcx (Fig. 57). dciiic siiiall; basipliallus hipartitc: iis posicrior pnri cxpiiiidcd vciitrally aiid aiiicrior piii ovcrlappiiig basai par1 (ir disiiplialliis. üisiipliiiilus siiiipiy sclcroiizcd. apicaiiy imaticiic iikc. I1osigoiiiic broadcsi iii iiic iiiiddlc, siiiiiously iiiiirgiiicd, iiipcrcd disially biii apcx bliiiii tory apodciiic siiiiplc, wiilioiii disiiiict capiicliiiiii. Fciiialc abduiiicii: I>osiabdoiiicii (ligs 58-59) distiiiciivc. 'Icrgiic 0 cssciiiially Iiirgcr iliiiii aiiil ariiicd by loiig sioui sciac. Tcrgiic 7 strikitigly proloiigcd aiid coiivcx. wiili posicrior 11 tiiriicd up. bui iiarrowcr iliaii botli icrgitc 6 aiid 8. 'Icrgiic 8 broiitl, coiisisiiiig ol' 3 tlarkly pl;iics: IWO coiivcx Iiiicral sclcriics iiiid a siiiail. ii-aiisvcrsc iiicdi;il oiic. -Icigiic 10 roiiglily wiili ii pair ol' siiiiill dorsal sciiilac. iiiicropubcscciicc rcsiricictl (o posicrior Iiiill: Sicriiiic iis sicriiiic 7. wiili loiig sciuc iiiid ixilc posterior iiiiirgiii. Sicriiiic 7 cliiir;icicrizcd by rcdii o11 disc. Slcriiiic 8 iraiisvcrsc. liglitly pigiiiciiicil. largcly iiiciiil~rmoiis,witli cciiiral piiri iiii cciit. wiili loiig Iiiicral scia mid scvcriil sliori sciiic. Stcriiiic I O siiiiplc, -sciiiicirciiliir, wiili loiig sciac. Spcciaclcs-shapcd sclcriic largcly iiiciiibraiioiis. also iis cciitral sclcritc w iizcil üiid poorly visiblc. Spcriiiailiccx ( Figs 60, 62) siiiiiiar io iliosc of 7: lllCd¡/CJ'J'ílJJ~ll. Ixisril part, sparsc wriiiklcs o11 surlncc aiid distiiict tcriiiiiiiil iiivagiiiaiioii; sclcroiizcd par vcry sliori. Ccrcus robusi, aiso its sciac rclaiivcly sioui, apical scie hciiig loiigcr iliaii ccrc

Cuiiiiiiciiis: A disliiictivc spccics rcliiicd lo 7: iii~~~li/~~/~J'u/i~,íl- 7: lcbcciiiise of iis crcci loicroilasal su scia aiid oilicr cxicriiai kcy) aiid by uiiariiicd iiialc sicriiiic 5. surstylus lackiiig posierovciiiial spiiic. fciii;ilc i siroiig sciac, vcry coiivcx icrgiic 7 aiid coiispiciious stcrnitc 8. Disti'ibuiioii: 7: cprw~liseluscciiis lo bc witlcsprcad iii Atlaiiioiiicdiicrraiicaii arca; ii w;is (irciii hiiaiii: Eiigkiiid (RoiiAiii~i~8:i); Spiiiii: ('iiiiiiry is.-'ikiicril'c (iicw); i~iilc;iricis.-ii)izi l'oi.itA 1994); Iialy: Sicily (iicw).

Ilrorawcltaclu zostcr(ic (1 I ALIIMY, 1833) (I:igS 1-2. S-h. 63-70) hJ'/iilJ'JJS %J.WJ'lJC 1 [Al IIIAY, 1x33: 178. /~~JJI~~.~¡JiAY, 1936: 333. /~llJJfJS¡JiA, 1925: 1 YX. /X/ll>lO<.~~J'li(7~l~JJ'U~fJ<'~Jr, 1x47: 2502; zl:l IliWSII:I)I. 1x60: 64 14 (Synoiiyiiiyl. /~~JililS~JJfJ/I;/;X(WJ Sil~NllAhlMAR. 1855: 386: ~l~.l'll.~Sll~l~l,1x60: 64 14 [piOlId>k! S~iitiii~ili~];l)ili)A, 1Y3 »iiylilyJ. /.~JJJOS~~J~J<.f/ioJs. IiCi-Cl?y dcS¡giiiilC