Diptera: Empididae)

Total Page:16

File Type:pdf, Size:1020Kb

Diptera: Empididae) Eur. J. Entomol. 106: 441–450, 2009 http://www.eje.cz/scripts/viewabstract.php?abstract=1471 ISSN 1210-5759 (print), 1802-8829 (online) Revision of the Iteaphila setosa group (Diptera: Empididae) IGOR V. SHAMSHEV 1 and BRADLEY J. SINCLAIR2 1All-Russian Institute of Plant Protection, Podbelskogo 3, Pushkin, St. Petersburg 189620, Russia; e-mail: [email protected] 2Ottawa Plant & Seed Laboratories – Entomology, Canadian Food Inspection Agency, K.W. Neatby Bldg., C.E.F., 960 Carling Ave., Ottawa, ON, Canada K1A 0C6; e-mail: [email protected] Key words. Iteaphila, Empidoidea, Empididae, taxonomy, new species, distribution Abstract. Six species are identified in the Iteaphila setosa group [I. arundela sp. n., I. caucasica sp. n., I. italica Loew, 1873, I. kubaniensis sp. n., I. merzi sp. n., I. setosa (Bezzi, 1924)]. This species group is distributed from southern England in the west, through southern Europe and northern Africa, to the Caucasus and Middle Asia (Uzbekistan). INTRODUCTION London, England (BMNH); Canadian National Collection of Insects, Ottawa, Canada (CNC); Peter Hodge collection At higher northern latitudes and montane regions, the [private], Lewes, England (DODC); Museo Civico di Storia genus Iteaphila Zetterstedt, 1838 is one of the first polli- Naturale, Milan, Italy (MSNM); Muséum d’Histoire Naturelle, nators active in early spring, primarily visiting flowers of Geneva, Switzerland (MHNG); National Museum of Ireland, Prunus and Salix and other spring flowers. Apart from Dublin, Ireland (NMID); National Museum of Scotland, Edin- these collection records, little is known about the biology burgh, Scotland (NMSE); National Museum of Wales, Cardiff, of Iteaphila. There are currently some 19 recognized spe- UK (NMWC); Oxford University Museum of Natural History, cies of Iteaphila (Shamshev & Sinclair, unpubl. data), Oxford, England (OUMNH); World Museum of Liverpool, including at least two Holarctic species (I. nitidula Zet- Liverpool, England (WML); United States National Museum of terstedt, 1838, I. macquarti Zetterstedt, 1838). Iteaphila Natural History, Washington, DC, USA (USNM); Zoological Institute, St. Petersburg, Russia (ZIN); Museum für Naturkunde includes quite small (about 3.0 mm) blackish grey flies der Humboldt-Universität zu Berlin, Berlin, Germany (ZMHB); with long antennae, conspicuously lengthened proboscis Zoological Museum of Moscow State University, Moscow, and palpi (which are usually directed forward and sube- Russia (ZMUM). qual in length), slender legs and humped thorax clothed in All dissections were made in glycerine and tissues cleared numerous unmodified hair-like setae. Iteaphila is a primi- using hot 85% lactic acid. Terms used for adult structures pri- tive empidoid taxon currently unassigned to family in the marily follow those of McAlpine (1981), except for antennal Empidoidea (Sinclair & Cumming 2006) (see Discussion morphology which follows Stuckenberg (1999) and for the wing section below). venation where the terms of Saigusa (2006) are used. Conse- Ragas setosa Bezzi, 1924 was transferred to Iteaphila quently the following wing venation terms are used: M4 (= CuA ), cell cua (= anal cell or cell cup), and CuA+CuP (= anal by Sinclair & Saigusa (2001), highlighting an odd species 1 vein or A1). Homologies of the male terminalia follow those of with modified male hind femur, shortened, tapered anten- Sinclair & Cumming (2006). nae, and male terminalia with a distinct surstylus and Label data for primary types are cited in full, where labels are epandrial lobe. During a revision of the entire Iteaphila listed from the top downward, with data from each label group we discovered that I. setosa represented a distinct enclosed in quotation marks. Labels are cited with original species group and also included I. italica Loew, 1873. spelling, punctuation and date and lines are delimited by a slash Tuomikoski (1958) questioned whether I. italica Loew mark (/). Additional information is included in square brackets. was congeneric given the shape of the antenna and male The repository of each type is given in parentheses. The fol- terminalia as illustrated in Engel (1941) and the southern lowing abbreviations are used in the descriptions: dc – dorso- central bristle, pprn – postpronotal bristle, presut spal – presu- distribution of this otherwise northern or montane genus. tural supra-alar bristle, npl – notopleural bristle, pal – postalar We prefer to retain this group within Iteaphila until a bristle, psut spal – postsutural supra-alar bristle, and sctl – scu- thorough revision and phylogenetic analysis is completed tellar bristle. for the entire Iteaphila group. This is the first of several revisions on this genus group and describes five addi- ITEAPHILA SETOSA SPECIES GROUP tional species related to I. setosa. Recognition MATERIAL AND METHODS The species group is distinguished by the following combination of characters: postpedicel with broad base, This study is based on material borrowed from or deposited in the following collections: M. Barták collection [private], with apical half tapered; stylus nearly half length of pedi- Prague, Czech Republic (BARC); British Entomological and cel; apical mechanoreceptor (bristle) greatly reduced in Natural History Society (BENH); The Natural History Museum, length; scutellum with 2–3 pairs of apical bristles; male 441 hind femur usually modified; wing with basal costal anterior ocellars moderately long, proclinate and laterocli- bristle present; cell dm produced apically; male terminalia nate, posterior ocellars short. Postvertical and postocular with epandrial lobe and surstylus distinctly digitiform. bristles quite strong, longer than ocellars; additionally, Key to species of the Iteaphila setosa species group occiput covered with numerous similar setae of different lengths on lower part. Antenna black; scape short, sube- 1 Antennal stylus long, nearly half length of postpedicel, qual to globular pedicel, both with short setae; postpe- lacking apical mechanoreceptor (bristle). Scutellum with 2–3 pairs of bristles. Basal costal bristle present. Epandrial dicel rather broad at base, about 2.0 times longer than lamella with additional lobe [I. setosa group]........... 2 basal width, strongly tapered; stylus about 1/2 of postpe- – Antennal stylus very short relative to length of postpedicel, dicel length; segment 9 nearly 4 times as long as wide, with apical mechanoreceptor (bristle). Scutellum with at apical sensillum inconspicuous. Proboscis long, projected least 4 pairs of bristles. Basal costal bristle absent. Epandrial obliquely; labium about as long as head height; palpus lamella lacking additional lobe......................... projected parallel to labrum, about two-thirds length of ................. [I. macquarti group, not treated herein] labrum. 2 Male (holoptic)................................... 3 Thorax black in ground colour, with black setation; – Female (dichoptic) (unknown for I. setosa, I. caucasica, and scutum viewed dorsally matt black brown, with 2 hardly I. kubaniensis)....................................8 3 Hind femur unmodified. Epandrial lobe sinuous, slender, prominent paler vittae between acrostichal and dorsocen- nearly twice length of surstylus and strongly arched medi- tral bristles, viewed anteriorly pale vittae between acrosti- ally (Fig. 3)................................ I. italica chal and dorsocentral bristles; mesopleuron uniformly – Hind femur modified, with deep, broad, ventral constriction brownish grey pollinose. Proepisternum with 2–3 short basally (Fig. 7). Epandrial lobe similar in length to sursty- brownish setae on lower part and 1 long seta on upper lus............................................ 4 part. Postpronotal lobe with 1 long and 3–4 short pprn. 4 Hind femur with scattered setulae near base, lacking flat- Mesonotal bristles prominent; acrostichals biserial, some- tened bristles. Surstylus slender and finger-like (Figs 1, 2, 4, what irregular, slightly shorter than dc, lacking on prescu- 6)..............................................5 tellar depression; dc uniserial, offset from row anteriorly, – Hind femur with tuft of several flattened, curved long brist- les near base (Fig. 7). Surstylus broad, thumb-like and 1 pair anteriorly and 2 prescutellar pairs long; 1 posthu- highly setose (Fig. 5).................... I. merzi sp. n. meral, 1–2 presut spal, 3 strong and 3 thinner npl, 3–4 5 Halter knob pale. Gonocoxal apodeme very short, less than short psut spal, 1 pal, 2 pairs of sctl. one-third length of ejaculatory apodeme (Fig. 4); hypan- Legs brownish, knees of fore and mid legs brownish drium with deep and broad notch...... I. kubaniensis sp. n. yellow, finely greyish pollinose. Coxae and trochanters – Halter knob dark. Gonocoxal apodeme broad and greater with rather long, black, unmodified setae. Fore and mid than one-half length of ejaculatory apodeme; hypandrium femora with row of moderately long (about as long as with shallow or deep notch (Figs 1, 2, 6)............... 6 femur deep), uniform posteroventral setae. Fore and mid 6 Legs dark. Ventral margin of epandrial lobe uneven, sculp- tibiae with some prominent posterodorsal setae. Hind tured, apex narrower than base of lobe (Fig. 6).... I. setosa – Legs with “knees” pale. Ventral margin of epandrial lobe femur with deep broad excision near base, anterior smoothly curved, apex slightly expanded and broadly margin of excision not strongly produced, with row of rounded.........................................7 moderately
Recommended publications
  • ARTHROPOD COMMUNITIES and PASSERINE DIET: EFFECTS of SHRUB EXPANSION in WESTERN ALASKA by Molly Tankersley Mcdermott, B.A./B.S
    Arthropod communities and passerine diet: effects of shrub expansion in Western Alaska Item Type Thesis Authors McDermott, Molly Tankersley Download date 26/09/2021 06:13:39 Link to Item http://hdl.handle.net/11122/7893 ARTHROPOD COMMUNITIES AND PASSERINE DIET: EFFECTS OF SHRUB EXPANSION IN WESTERN ALASKA By Molly Tankersley McDermott, B.A./B.S. A Thesis Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Science in Biological Sciences University of Alaska Fairbanks August 2017 APPROVED: Pat Doak, Committee Chair Greg Breed, Committee Member Colleen Handel, Committee Member Christa Mulder, Committee Member Kris Hundertmark, Chair Department o f Biology and Wildlife Paul Layer, Dean College o f Natural Science and Mathematics Michael Castellini, Dean of the Graduate School ABSTRACT Across the Arctic, taller woody shrubs, particularly willow (Salix spp.), birch (Betula spp.), and alder (Alnus spp.), have been expanding rapidly onto tundra. Changes in vegetation structure can alter the physical habitat structure, thermal environment, and food available to arthropods, which play an important role in the structure and functioning of Arctic ecosystems. Not only do they provide key ecosystem services such as pollination and nutrient cycling, they are an essential food source for migratory birds. In this study I examined the relationships between the abundance, diversity, and community composition of arthropods and the height and cover of several shrub species across a tundra-shrub gradient in northwestern Alaska. To characterize nestling diet of common passerines that occupy this gradient, I used next-generation sequencing of fecal matter. Willow cover was strongly and consistently associated with abundance and biomass of arthropods and significant shifts in arthropod community composition and diversity.
    [Show full text]
  • Diptera, Empidoidea) 263 Doi: 10.3897/Zookeys.365.6070 Research Article Launched to Accelerate Biodiversity Research
    A peer-reviewed open-access journal ZooKeys 365: 263–278 (2013) DNA barcoding of Hybotidae (Diptera, Empidoidea) 263 doi: 10.3897/zookeys.365.6070 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research Using DNA barcodes for assessing diversity in the family Hybotidae (Diptera, Empidoidea) Zoltán T. Nagy1, Gontran Sonet1, Jonas Mortelmans2, Camille Vandewynkel3, Patrick Grootaert2 1 Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny (JEMU), Rue Vautierstraat 29, 1000 Brussels, Belgium 2 Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny (Ento- mology), Rue Vautierstraat 29, 1000 Brussels, Belgium 3 Laboratoire des Sciences de l’eau et environnement, Faculté des Sciences et Techniques, Avenue Albert Thomas, 23, 87060 Limoges, France Corresponding author: Zoltán T. Nagy ([email protected]) Academic editor: K. Jordaens | Received 7 August 2013 | Accepted 27 November 2013 | Published 30 December 2013 Citation: Nagy ZT, Sonet G, Mortelmans J, Vandewynkel C, Grootaert P (2013) Using DNA barcodes for assessing diversity in the family Hybotidae (Diptera, Empidoidea). In: Nagy ZT, Backeljau T, De Meyer M, Jordaens K (Eds) DNA barcoding: a practical tool for fundamental and applied biodiversity research. ZooKeys 365: 263–278. doi: 10.3897/zookeys.365.6070 Abstract Empidoidea is one of the largest extant lineages of flies, but phylogenetic relationships among species of this group are poorly investigated and global diversity remains scarcely assessed. In this context, one of the most enigmatic empidoid families is Hybotidae. Within the framework of a pilot study, we barcoded 339 specimens of Old World hybotids belonging to 164 species and 22 genera (plus two Empis as outgroups) and attempted to evaluate whether patterns of intra- and interspecific divergences match the current tax- onomy.
    [Show full text]
  • R. P. LANE (Department of Entomology), British Museum (Natural History), London SW7 the Diptera of Lundy Have Been Poorly Studied in the Past
    Swallow 3 Spotted Flytcatcher 28 *Jackdaw I Pied Flycatcher 5 Blue Tit I Dunnock 2 Wren 2 Meadow Pipit 10 Song Thrush 7 Pied Wagtail 4 Redwing 4 Woodchat Shrike 1 Blackbird 60 Red-backed Shrike 1 Stonechat 2 Starling 15 Redstart 7 Greenfinch 5 Black Redstart I Goldfinch 1 Robin I9 Linnet 8 Grasshopper Warbler 2 Chaffinch 47 Reed Warbler 1 House Sparrow 16 Sedge Warbler 14 *Jackdaw is new to the Lundy ringing list. RECOVERIES OF RINGED BIRDS Guillemot GM I9384 ringed 5.6.67 adult found dead Eastbourne 4.12.76. Guillemot GP 95566 ringed 29.6.73 pullus found dead Woolacombe, Devon 8.6.77 Starling XA 92903 ringed 20.8.76 found dead Werl, West Holtun, West Germany 7.10.77 Willow Warbler 836473 ringed 14.4.77 controlled Portland, Dorset 19.8.77 Linnet KC09559 ringed 20.9.76 controlled St Agnes, Scilly 20.4.77 RINGED STRANGERS ON LUNDY Manx Shearwater F.S 92490 ringed 4.9.74 pullus Skokholm, dead Lundy s. Light 13.5.77 Blackbird 3250.062 ringed 8.9.75 FG Eksel, Belgium, dead Lundy 16.1.77 Willow Warbler 993.086 ringed 19.4.76 adult Calf of Man controlled Lundy 6.4.77 THE DIPTERA (TWO-WINGED FLffiS) OF LUNDY ISLAND R. P. LANE (Department of Entomology), British Museum (Natural History), London SW7 The Diptera of Lundy have been poorly studied in the past. Therefore, it is hoped that the production of an annotated checklist, giving an indication of the habits and general distribution of the species recorded will encourage other entomologists to take an interest in the Diptera of Lundy.
    [Show full text]
  • A Revision of the New Genus Amiga Nakahara, Willmott & Espeland, Gen. N., Described for Papilio Arnaca Fabricius, 1776
    A peer-reviewed open-access journal ZooKeys 821: 85–152 (2019) A revision of the new genus Amiga 85 doi: 10.3897/zookeys.821.31782 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae) Shinichi Nakahara1,2, Gerardo Lamas2, Stephanie Tyler1,3, Mario Alejandro Marín4, Blanca Huertas5, Keith R. Willmott1, Olaf H. H. Mielke6, Marianne Espeland7 1 McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Flo- rida, Gainesville, Florida 32611 USA 2 Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru 3 School of Architecture, Rice University, 6100 Main Street, Houston, Texas 77005 USA 4 Departamento de Biologia Animal and Museu de Zoologia, Instituto de Biologia, Universidade Estadual de Campinas – UNICAMP. 13083-970 Campinas, São Paulo, Brazil 5 Life Sciences Department, Natural Hi- story Museum, Cromwell Road, London SW7 5BD, UK 6 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, Caixa postal 19020, 81.531 Curitiba, Paraná, Brazil 7 Arthropoda Department, Zoological Research Museum Alexander Koenig, Adenauer Allee 160, 53113 Bonn, Germany Corresponding author: Shinichi Nakahara ([email protected]) Academic editor: T. Simonsen | Received 20 November 2018 | Accepted 3 January 2019 | Published 31 January 2019 http://zoobank.org/ECFCCAF6-8D99-457B-B9F8-2443089D0182 Citation: Nakahara S, Lamas G, Tyler S, Marín MA, Huertas B, Willmott KR, Mielke OHH, Espeland M (2019) A revision of the new genus Amiga Nakahara, Willmott & Espeland gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae).
    [Show full text]
  • Vulnerability of At-Risk Species to Climate Change in New York (PDF
    Vulnerability of At-risk Species to Climate Change in New York Matthew D. Schlesinger, Jeffrey D. Corser, Kelly A. Perkins, and Erin L. White New York Natural Heritage Program A Partnership between The Nature Conservancy and the NYS Department of Environmental Conservation New York Natural Heritage Program i 625 Broadway, 5th Floor Albany, NY 12233-4757 (518) 402-8935 Fax (518) 402-8925 www.nynhp.org Vulnerability of At-risk Species to Climate Change in New York Matthew D. Schlesinger Jeffrey D. Corser Kelly A. Perkins Erin L. White New York Natural Heritage Program 625 Broadway, 5th Floor, Albany, NY 12233-4757 March 2011 Please cite this document as follows: Schlesinger, M.D., J.D. Corser, K.A. Perkins, and E.L. White. 2011. Vulnerability of at-risk species to climate change in New York. New York Natural Heritage Program, Albany, NY. Cover photos: Brook floater (Alismodonta varicosa) by E. Gordon, Spadefoot toad (Scaphiopus holbrookii) by Jesse Jaycox, and Black Skimmers (Rynchops niger) by Steve Young. Climate predictions are from www.climatewizard.org. New York Natural Heritage Program ii Executive summary Vulnerability assessments are rapidly becoming an essential tool in climate change adaptation planning. As states revise their Wildlife Action Plans, the need to integrate climate change considerations drives the adoption of vulnerability assessments as critical components. To help meet this need for New York, we calculated the relative vulnerability of 119 of New York’s Species of Greatest Conservation Need (SGCN) using NatureServe’s Climate Change Vulnerability Index (CCVI). Funding was provided to the New York Natural Heritage Program by New York State Wildlife Grants in cooperation with the U.S.
    [Show full text]
  • Nomenclatural Studies Toward a World List of Diptera Genus-Group Names
    Nomenclatural studies toward a world list of Diptera genus-group names. Part V Pierre-Justin-Marie Macquart Evenhuis, Neal L.; Pape, Thomas; Pont, Adrian C. DOI: 10.11646/zootaxa.4172.1.1 Publication date: 2016 Document version Publisher's PDF, also known as Version of record Document license: CC BY Citation for published version (APA): Evenhuis, N. L., Pape, T., & Pont, A. C. (2016). Nomenclatural studies toward a world list of Diptera genus- group names. Part V: Pierre-Justin-Marie Macquart. Magnolia Press. Zootaxa Vol. 4172 No. 1 https://doi.org/10.11646/zootaxa.4172.1.1 Download date: 02. Oct. 2021 Zootaxa 4172 (1): 001–211 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Monograph ZOOTAXA Copyright © 2016 Magnolia Press ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4172.1.1 http://zoobank.org/urn:lsid:zoobank.org:pub:22128906-32FA-4A80-85D6-10F114E81A7B ZOOTAXA 4172 Nomenclatural Studies Toward a World List of Diptera Genus-Group Names. Part V: Pierre-Justin-Marie Macquart NEAL L. EVENHUIS1, THOMAS PAPE2 & ADRIAN C. PONT3 1 J. Linsley Gressitt Center for Entomological Research, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817-2704, USA. E-mail: [email protected] 2 Natural History Museum of Denmark, Universitetsparken 15, 2100 Copenhagen, Denmark. E-mail: [email protected] 3Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW, UK. E-mail: [email protected] Magnolia Press Auckland, New Zealand Accepted by D. Whitmore: 15 Aug. 2016; published: 30 Sept. 2016 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 NEAL L.
    [Show full text]
  • Zootaxa, Empidoidea (Diptera)
    ZOOTAXA 1180 The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) BRADLEY J. SINCLAIR & JEFFREY M. CUMMING Magnolia Press Auckland, New Zealand BRADLEY J. SINCLAIR & JEFFREY M. CUMMING The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) (Zootaxa 1180) 172 pp.; 30 cm. 21 Apr. 2006 ISBN 1-877407-79-8 (paperback) ISBN 1-877407-80-1 (Online edition) FIRST PUBLISHED IN 2006 BY Magnolia Press P.O. Box 41383 Auckland 1030 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/ © 2006 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) Zootaxa 1180: 1–172 (2006) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1180 Copyright © 2006 Magnolia Press ISSN 1175-5334 (online edition) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera) BRADLEY J. SINCLAIR1 & JEFFREY M. CUMMING2 1 Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany. E-mail: [email protected] 2 Invertebrate Biodiversity, Agriculture and Agri-Food Canada, C.E.F., Ottawa, ON, Canada
    [Show full text]
  • Diptera, Empididae, Empidinae)
    Bulletin de la Société entomologique de France, 123 (1), 2018 : 119-123. A new species of Empis (Xanthempis) from Morocco (Diptera, Empididae, Empidinae) Fatima-Zohra BAHID1, Kawtar KETTANI2 & Christophe DAUGERON3 1,2 Équipe de recherche Écologie, Systématique et Conservation de la Biodiversité, Faculté des Sciences, Université Abdel Malek Essaadi, Tétouan, Maroc <[email protected]> <[email protected]> 3 Muséum national d’Histoire naturelle, Centre National de la Recherche Scientifique, Mécanismes adaptatifs et évolution, UMR 7179 MNHN-CNRS MECADEV, C. P. 50, 57 rue Cuvier, F – 75231 Paris cedex 05 <[email protected]> http://zoobank.org/C52AFF39-E81B-44A3-A86F-8C6988F39628 (Accepté le 6.II.2018) Abstract. – A new species of the subgenus Xanthempis Bezzi of the genus Empis Linnaeus is described from Morocco: E. (X.) widanensis n. sp. is the first Xanthempis species found in the Rif. It is the fourth species of this subgenus known for this country and North Africa as whole. A key including the four known Moroccan species, as well as two additional species distributed in Southern Spain, is given. Résumé. – Une nouvelle espèce d’Empis (Xanthempis) du Maroc (Diptera, Empididae, Empidinae). Une nouvelle espèce appartenant au sous-genre Xanthempis Bezzi du genre Empis Linné est décrite du Maroc: E. (X.) widanensis n. sp. est la première espèce de Xanthempis trouvée dans le Rif. C’est la quatrième espèce de ce sous-genre connue au Maroc et dans l’ensemble de l’Afrique du Nord. Une clé d’identification est proposée pour ces quatre espèces marocaines ainsi que deux autres espèces distribuées dans le sud de l’Espagne.
    [Show full text]
  • Hennig (1970) Insect Fossils from the Lower Cretaceous. II. Empididae
    ,- ~- . til. ."""r Secretary Secretariat of State d'Etat MULTILINGUAL SERVICES DIVISION - DIVISION DES SERVICES MUlTILINGUES TRANSLATION BUREAU BUREAU DES TRADLJCTIONS LIBRARY IDENTIFICATION - FICHE SIGNALETIQUE Translated from - Traduction de Into - En German English Author - ~uteur Hennig, Willi Title in Eraglish or French - Titre anglais ou fran~ais Insect Fossils From the Lower Cretaceous. II. Empididae (Diptera, Brachycera). Title in foreign language (Transliterate foreign characters) Titre en langue 'trang6re (Transcrire en caractAres romains) Insektenfossilien aus der unteren Kreide. II. Empididae (Diptera, Brachycera)~ Refere~ce in foreign language (Name of book or publication) in full, transliterate foreign characters. Rif4renc8 en langue ~trangtre (Nom du livre au publication), au complet, transcrire en caractAres romains. stuttgarter BeitrMge zur Naturkunde Reference in English or French - A6f6rence en anglais ou fran~ais stuttgart Contributions on Natural History Publisher .. Editeur Page Numbers in original DATE OF PUBLICATION Num'ros des pages dans DATE DE PUBLICAilON I'original not shown 1 12 Yeor Issue No. - Volume Place of Publication Annt1e Num6ro Number of typed pages Lieu de publieation Nombre de pages dactylographiees April Stuttgart 1970 214 25 Requesting Department Translation Bureau No. 1877017 Minist6re-CUent Agriculture Notre dossier nO _ Research. BranchOi~cdonoyor DivisionO~~lon ~_~t o_~~o_o~~~~_~~1 ~ Translator (Initials) AB __ Traducteur (lnitiales) __.,------------ Person requesting Jeff Curnmi.ng Demand6 par ~__--- ~_ NO\' 19 1986 Your Numbllr Vatr. dossier nO ~ _ Date of Request 13-08-86 Date de la derilande _ l •• Can. ada SEC 5·111 (84-10) Secretary Secretariat of State d'~tat MULT'L'NGUAL SERVICES DIVISION - DIVISION DES SERVICES MULTILINGUES TRANSLATION IlUREAU BUREAU DES TRADUCTIONS Client's No.-No du client Department Minist6re Division/Branch - Division/Direction City - Ville Agriculture Research ottawa, Entomology C.E.F.
    [Show full text]
  • Georg-August-Universität Göttingen
    GÖTTINGER ZENTRUM FÜR BIODIVERSITÄTSFORSCHUNG UND ÖKOLOGIE GÖTTINGEN CENTRE FOR BIODIVERSITY AND ECOLOGY Herb layer characteristics, fly communities and trophic interactions along a gradient of tree and herb diversity in a temperate deciduous forest Dissertation zur Erlangung des Doktorgrades der Mathematisch-Naturwissenschaftlichen Fakultäten der Georg-August-Universität Göttingen vorgelegt von Mag. rer. nat. Elke Andrea Vockenhuber aus Wien Göttingen, Juli, 2011 Referent: Prof. Dr. Teja Tscharntke Korreferent: Prof. Dr. Stefan Vidal Tag der mündlichen Prüfung: 16.08.2011 2 CONTENTS Chapter 1: General Introduction............................................................................................ 5 Effects of plant diversity on ecosystem functioning and higher trophic levels ....................................................... 6 Study objectives and chapter outline ...................................................................................................................... 8 Study site and study design ................................................................................................................................... 11 Major hypotheses.................................................................................................................................................. 12 References............................................................................................................................................................. 13 Chapter 2: Tree diversity and environmental context
    [Show full text]
  • Of the World
    OCCASIONAL PAPERS OF THE CALIFORNIA ACADEMY OF SCIENCES No. 147, 94 pages. December 2, 1991 GENUS-GROUP NAMES OF THE NEUROPTERA, MEGALOPTERA AND RAPHIDIOPTERA OF THE WORLD By John D. Oswald Department of Entomology, Cornell University, Ithaca, New York 14853-0999 and Norman D. Penny Department of Entomology, California Academy of Sciences, San Francisco, California 94118-4599 Abstract: Alphabetical listings of the genus-group names of extant Megaluptcra, Raphidioptera, and = Neuroptera (s. str. Planipennia) are presented. Taxonomic and nomenclatural data for each name are given. Summaries of new genus-group synonyms, unreplaced junior homonyms, names without valid type species fixations, and names based on misidentified type species are given. Complete bibliographic references are given for all names and nomenclatural acts. Contents Introduction Inlroduciion (1) The last worldwide species-level catalog of Scope (2) the order str. = Nomenclature (2) Neuroptera (s. Planipennia), and Format Arrangement of Entries (2) Hermann Hagen's 1866 Hemerobidarum Syn- General Arrangement (2) opsis Synonymica, has long been obsolete, as Subgenera (2) are the most recent revisions Synonymy (2) comprehensive Character Formals (3) of the orders Megaloptera (i.e.. Van dcr Publication Dates (3) Weele 1910) and Raphidioptera (i.e., Navas Type Species (3) [1919e] 1918). In the 120+ years since 1866, Unavailable Names (3) the number of available Homonymy (4) nomenclaturally Family-Group Taxa (4) genus-group names in the order Neuroptera Selected Taxonomic References
    [Show full text]
  • Download Download
    Tropical Natural History 20(1): 16–27, April 2020 2020 by Chulalongkorn University Endemism, Similarity and Difference in Montane Evergreen Forest Biodiversity Hotspots: Comparing Communities of Empidoidea (Insecta: Diptera) in the Summit Zones of Doi Inthanon and Doi Phahompok, Thailand ADRIAN R. PLANT1*, DANIEL J. BICKEL2, PAUL CHATELAIN3, CHRISTOPHE DAUGERON3 AND WICHAI SRISUKA4 1Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Mahasarakham 44150, THAILAND 2Australian Museum, Sydney, NSW 2010, AUSTRALIA 3Muséum national d’Histoire naturelle, 57 rue Cuvier, CP 50, F-75005 Paris, FRANCE 4Entomology Section, Queen Sirikit Botanic Garden, Mae Rim, Chiang Mai 50180, THAILAND * Corresponding author. Adrian R. Plant ([email protected]) Received: 7 August 2019; Accepted: 12 November 2019 ABSTRACT.– Composition and structure of communities of the Diptera superfamily Empidoidea (30,481 individuals of 511 species in 55 genera in the families Empididae, Dolichopodidae, Hybotidae and Brachystomatidae) were compared in Upper Montane Forests on Doi Inthanon and Doi Phahompok in northern Thailand. Based on taxon similarity (α-, β-diversity), structural diversity (the species abundance distribution and importance of dominant species), cluster analysis of community composition and the relative importance of inferred Oriental and Palaearctic influences, it was concluded that communities in Upper Montane Forest at 2,036 – 2,105 m near the summit of Doi Phahompok were most similar to those at 1,639 – 2,210 m on Doi Inthanon. Approximately 33% of species recorded at 2,036 – 2,105 m on Doi Phahompok were endemic to the mountain. Upper Montane Forest is a rare, dispersed and isolated habitat in southeastern Asia with scattered patches likely to experience comparable levels of β-diversity and endemism as found here.
    [Show full text]