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Transactions and Proceedings of the Palaeontological Society of

New Series No. 25

Palaeontological Society of Japan April 30, 1957 CONTENTS TRANSACTIONS Page 305. On the Miocene Lucinidae from the Mizunami Group, Japan ...... Junji hoJGAWA 1 306. Brachiopoda from the Daishaka and Tsurugasaka Fossil Zones. Minami Tsugaru-Gun, Aomori Prefecture ...... ]iro K

PROCEEDINGS ...... 40

President: Hisakatsu YABE Councillors: Kiyoshi AsANO, Riuji ENDo (Business and Accountant), Seido ENoo, Haruyoshi FuJJl\IOTo (Publication), Shoshiro HANZAWA, I\otora I·IATAI, Ichiro HAYASAKA, (General Affairs), Kazuo HuzJOI\A, Teiichi KonAYASIII (Editor), Jiro MAKIYAMA, Tatsuro MATSUMoTo, Masao MINATo, Tokio SIIIKAMA, Fuyuji T AKAI (General Affairs), Ryuzo ToRIYAMA.

All Communications relating to this Journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN Geological Institute, Faculty of Science, University of , Japan Trans. Proc. Palae:>nt. So:. Japan. N. S., Ko. 25, pp. 1-6, Pl. 1. April 30, 1957

305. ON THE MIOCENE LUCI:KlbAE FROM THE MlZUNAMI GROUP, JAPAN*

]UNJI ITOIGAWA

Kyoto Ur.iversity

fliJ~~l!fi!F.~~mttt l.ucinidae: i\\\1~ • !fft-J'j\ljjWlh">f/iJIJ'~fe'ili:J: I) il€ Lt.:. 6 flljQ) Lucinidae t:.llr, -t ~;,ft1.i J;l~t:. --:>~ • -c k'lt& r.HillH\1tfi L. ~t.:.-.k~Jilfn:. --:>i., -c ~~ L t~o '*. t.:... lVallucina .{!j(,Q).:....VJ(m · ~lii!.tH.t.:.o -!M~.Jll w=

Introduction and Acknowledgements Wallucina habti ITOIGAWA Wallucina okunptrai ·n. sp .. In the inside zone of southwestern Pillucina (Sydlorina) yokoyam~i (OUTKA) Japan, there are found scatterings of Saxoltlcina (Megaxinus) k-hataii (OTUKA) the Miocene strata that are assumptively Luci~ooma acutili11:ata (CoNRAD) representing the First Paleo-Setouchi Caviluchza (Monitilora)· kitamurai (HATAI supergroup (IKEBE, 1951) i. e. Mizunami, e~ NISIYAMA) Isshi, Ayukawa, Tsuzuki and Tsuyama The writer is indebted to Prof. J. groups etc. Yielding rich faunas and MAKIYAII.IA for his suggestions and revi­ floras, these strata are interesting sub· sion of English. Thanks are also due jects for the members of the Paleo­ to Dr. T. KuRODA and Dr. T. HABE for Setouchi research group. The Mizunami informations about the taxonomic mala· group. the type of the supergroup, was cology. developed in three mutually connected basins, Kani, Mizunami and Iwamura in Paleoecological t-.otes Gifu Prefecture. The present writer is studying the stratigraphy and paleon· Some paleoecologi~al aspects of these tology of this group and he has partly species were observed in field and labora­ reported the results. tory considering the data of stratigraphy This paper is the third result and and fossil fauna. Although these data prepared in view of taxonomic and paleo­ are not complete at present presumably ecological accounts of the Lucinid overlooking minute species, the conclu­ pelecypods from the Mizunami group. sion might not be very different from These are the interesting forms inasmuch that based upon the more accumulated as they are the indicators of various material. environments. Cavilucina ( Monitilora) kitamurai The following species are. collected

1 2 Junji lTOIGAWA m'sataictzsis 0TUKA and Turbo ozawai 0TUKA etc. Both valves of every indi­ Description of species vidual of these two species are attached Family Lucinidae snowing autochthonous origin in the Genus TVallucitza lREOAJ.E 1930 sandy environment. The fauna of Shuku­ nohora sandstone represents the warm Wallucina habei hoiGAWA 1955 sea waters and peculiar to this group. Pl. 1. figs. 3a -b. It seems that this two species inhabited 1955: Wallucin:J habei lTOJGA w A, Mem. Col!. in the warm and shallow waters with Sci., Kyoto U11iv. Ser. B. Vol. 22, No.2. sandy bottom. Art. 1, p. 139, pl. 6, figs. 1, 2. JVallucina hahei hoiGAWA is found in Geological occurrence:- the sandstones as a member of Dosinia­ 1. Kubohara sandstone, Iwamura basin. Nt'PP01zomarcia fauna. It is indicating 2. Togari formation, Mizunami basin. a sandy neritic environment, and the 3. Tsukiyoshi formation, Mizunami basin. attached valves show also the autoch­ Localities :- thonous origin. 1. J 3011I, Kamigiri, Iwamura-cho, Ena­ Pillucitw (Sydlorina) yokoyami (0Tu­ gun, Gifu Prefecture. KA) is found in the marginal facies 2. ] 40026, Nenga-hora, Togari, Mizunami developed near the basement with the City. following species: 3-a. ] 40041, Matsubara, Toki-cho, Mizu­ Nippmwmarcia nakamurai lKEBE, nami City. Vcnerupis siratm·iensis (OTUKA ). Nassa­ 3-b. J 40049, Kujiri, Toki City. rius simizrei 0Tuic-\. Turritella s·hataii Associated fonns :- NoMURA. 1. Dosiuia nomurai 0TUKA, NipP01zo­ Most probably this species sho\vs the marcia nakamurai IKEBE, Clitzocar­ similar environment to that of Wal­ dium slzinjiense (YoKOYAMA), lucitza lzabei hmGAWA. Glycymeris minoensis hoiGAw.~. Saxolucitza (Afega:r:itzus) ll-hataii Euspir:J meisensis MAKIYAMA, Tur­ (OTuKA) accompanies with Vicarya yoko­ rilella s-Jzataii NoMURA, Nassarius yamai TAJStero-dorsal side slightly convex, graduall}· descending to Pillucina (Sydlorina) yokoyamai (0TUKA) posterior margin : anterior margin round­ Pl. 1, figs. 4, 5. ed : ventral side arched : beak small, 1934: Lucina yokoyamai Ona;:A, Bull. Earthq. prominent; sculpture consisting of fine Res. ltzst. Tokyo Imp. Univ. Vol. 12. p. regular concentric lamellae and obscure 615. pl. 47. figs. 29-32. radiating striae ; dorsal area more or 1935: Luci11a (Lucinisca) yokoyamai OTUKA, less marked: lunule small, cordate, four. Fac. Sci. Imp Univ. Tokyo, Vol. dissymetric: hinge plate narrowly long, 5, p. 27. pl. •1. fig. 38. delicate: cardinal and lateral teeth frail, Shell small in size, suborbicular. as well developed: inner margin denticulate; long as high, moderately convex: beak inner side with fine radiating striae. smalL pointed, turned inwards, touching, Dimensions:- Height. 5 mm., length, placed slightly back of dorsal margin: 4.5mm. antero·dorsal margin short, concave, Holotype :- ]C 1500001, Paratype :- JC bluntly angled to subrounded anterior 1500002. margin: postero-dorsal margin also Remarlls :- This species is closely short. nearly straight descending to subtruncated J:OSterior margin ; ventral resembling TVallucina habei hoiGAWA, 1955, but it is distinguished in having margin regular:y rounded. connecting the thin and inflated shell with the fine both to posterior and anterior ends concentric lamellae, and the narrowly gradually ; sculpture consisting of many delicate hinge plate. Wallucina lamyi radiating striae · crossed by concentric CHAVAN, 1938 is another allied species incremental lines: radiating striae not but the present species has the thinner di\'aricate. weak at middle part; con­ and more orbicular shell, and its hinge centrics fine, ocassionally strong; lunule plate is more delicate v.ith the fine and ovate. deep, well marked; teeth of right frail teeth. valve consisting of a cardinal and 2 laterals : teeth of left valve consisting This species is named in honor of of 2 divaricating cardinals: inner side late Mr. Kikuo 0Kl'MURA who helped with fine radiating lin'3s; muscular the writer in the field. scar well-marked; inner margin fine:y Type Locality:- River-side cliff of crenulate. Shukubora valley about 100 m. SW of Dimensions:- Height, 6 mm., length, the bridge at S of Shukubora, Hiyoshi­ 6mm, cho, Mizunami City, Gifu Prefecture. (J 40020) (lwamura, 35° 24' 06" N, 137° Remarks:- This species was described 16' E) originally by 0TuKA from the lower Kadonosawa series (Miocene) as a speci­ Geological occurrence :- Shukunohora es of the Genus Lucina. 0TUKA states the sandstone, Mizunami basin. alliance with DALI.'s Lucinisca. Examin­ Associated forms :- Miogypsina kotoi ed in detail, it is clarified that it has HA:KZA w A, Operculina complattata japo­ the characters of the Genus Pillucina nica HANZAWA, Aloidis nisataiensis and the Subgenus Sydlorina i. e .. the 0TuKA, Cavt"lucina kitamurai (HATAI et teeth and its not divaricated sculpture. NISIYA,\IA), Turbo ozawai 0TUKA, Acteon The murricate sculpture of the Genus ozawai 0TUKA etc. Luc:'nisca is not at all seen in this Junji !TorGAwA: species. ric in form. deeply impressed, with fine, Geological occurretZce :- close. more or less irregularly crenated 1. Kubohara sandstone, Iwamua basin. striae parallel to the hinge margin ; 2. Togari formation, Mizunami basin. postero-dorsal area more or less distinct 3. Tsukiyoshi formation, Mizw1ami basin. extending to the postero-ventral margin: Localities :- Ligament impressed, broadly curved. 1-a. J 30246, Nakanislli, Yamaoka-cho, Remarks:- The remarkable features Ena.gun, Gifu Prefecture. of this species are as follows ; 1-b. J 3011L Kamigiri, Iwamura-cho, 1) very weak or obsolete teeth, 2) Ena·gun, Gifu Prefecture. well marked, dissymetric and deep 2. J 40026, Nenga-hora, Togari, Mizu· lunule, 3) well marked posterior area, nami City. 4) anteriorly 2 radiating lines, 5) smo­ 3-a. J 40041, Matsubora, Toki-cho, Mizu­ oth inner margin. nami City. This species belongs to the section 3-b. J 40049, Kujiri, Toki City. Megaxinus under the Genus Saxolucitza A~sociatcd forms:- 1-a. Barbatia ktt· on account of the above noted features. bam hme>AWA, NiPPorzomm·cia naka­ It is distinguished from the Gen).ls mzwai lKEBE, Venerupis siraton·cnsis Miltha by the weak or obsolete teeth. (0TVKA), Nassarius simizui 0TUKA, Geological occurrence:- Tsukiyoshi formation, Mizunami basin. Proto1·otella dep1·essa MAKIYAMA, 1-b. same to the associated fauna-(1) of Localities:- J-Vallucina habei hmGAWA, 2. same to 1. J 40008, Shobasama·hora, Tsukiyoshi, the associated fauna-(2) of Wallucina Mizunami City. habei lTmc;AwA, 3-a. same to .the asso­ 2. J 40041, Matsubora, Toki cho, Mizu­ ciate:! fauna-(3-a) of Wallucitta lzabei nami City. hoiGAWA, 3-b. same to the associated Associated fonns :- Vicarya yolwyamai fauna-{3-b) of Wallucitta habci hm- TAKEYAr-.tA, Vicaryella ishiiana (YoKo­ Ft:JITA GAWA. YAMA), Ceritltimn ka11eharai et Oc.osE, Cyclitta japonica KAMADA, San· guinaloria mitzoe1zsis (YOKOYAMA) etc. Genus Saxolucina STEWART 1930 Saxol:rcina (Megaximts) ll·hataii (OnJKA) Pl. 1. fig. 8. Genus Lucitwma DALL 1901 Lucinoma acutilineata (CoNRAD) 1934: Luci11a k-lzataii OTUK A, Bull. Earthq. Res. bzst. Tokyo Imp. U11iv .. Vol. 12, p. Pl. 1. figs. 9-12. 614. pl. 47, figs. 5, 6. 1849: Lttcina acutil;n:ala CoNRAD, U. S. Exp/. 1938: Luci11a (Mi/tha) k-hataii OTUKA, four. Exped. Vol. 10. Geol. p. 725. Fac. Sci. Imp. Un!v. Tokyo, Vol. 5. p. 1909: Phacoides acutililleata. DAJ.L, U. S. G. S. 27. pl. 4. fig. 38. Prof. P{lper. No. 59, p. 116. pl. 12. fig. Original description:- Shell r.ather 6. small, more or less thin, with the stir· 1931: Phacoides (Lucinom:~) awtilineata, Enm­ R!NGTON. U11.'v. Calif. Pub/. Geol. Vol. face more or less irregularly and con­ 20. p. 76. pl. 4. fig. 5. centrically lamellated ; anterior end longer, attenuated ; posterior end more Remarks :- This is one of the common plump, obscurely vertically truncated; species in the First Paleo-Setouchi super· beak low pointed : lunule small, crescent- group. The limits of this specie's have On the Miocene Lu;:inida~ from the Mizunami Group, Japan been discussed by American and Japan­ Genus Cavilucina P. FisHER 1887 ese authors. Cavilucina ( Monitilora) kitamurai The specimens at hand from the Mizu­

tion des Lucine3. jour. de Conchy. Vol. Paleont .. Vol. 23. No. 1, pp. 87-94. 81. pp. 133-153~ 198-216. 237-282. Vol. 82, HrRAYA,.,IA, K. (1954), On som~ Miocene pp. 59-97. 105-130, 215-243. Species of the Lucinoma from Japan, with DALL. W. H. (1901), Synopsis of the Lucinacea Description of Two New Species. ]apa11. and of the American Species. Proc. Cl. S. four. Geol. Geogr. Vol. 25, Nos. 1-2, pp. Nat. Mu$. Vol. 23. pp. 779-833. 101-115. -- (1909). Contributions to the Tertiary ITOIGAWA. J. (1955). The Cenozoic Strata in Paleontology of the Pacific Coast, 1. The the Iwamura Basin, Gifu Prefecture. Jap­ ~lio<:ene of Astoria and Coos Bay, Oregon. an. jour. Ceo/. Soc. japatl, Vol. 61, pp. U. S. Geol. Sun• .. Prof Paper, Ko. 59. 511-517. ETHERINGTON, T. ]. (1931). Stratigraphy and -- (1955). Molluscan Fauna of the Mizunami Fauna of the Astoria Miocene of Sou­ Group in the Iwamura Basin. Mcm. Coli. thwest Washington. U11iv. Calif. Pub/. Sci. Kyoto Univ. Ser. B. Vol. 22. No. 2. Geol.. Vol. 20, pp. 31-114. pp. 127-143. FUJITA, K. and OcosE, S. (1950, 1951), Libo· LAMY, E. (1920). Revision de:~ Lucinacea viv­ logic Classification of the Cenozoic Strata ants du Museum d'histoire naturelle de in the Northern Area of Mizunami-machi. Paris. ]our. de Conchy.. Vol. 65, pp. 71- Toki·gun, Gifu Prefecture. Japan. jour. 122, 169-222. 233-318. 335-388. Ceo/, Soc. japan, Vol. 56, pp. 481-492. 0TL'KA. Y. (1934). Tertiary Structure:; of the Vol. 57, pp. 99-l 10. Northwe;tern End of the Kitakami 1\loun­ GHANT. U.S. and GALE. H. R. (1931). Catalogue tainland. Iwate Prefecture. Japan. Bull. of the marine Pliocene and Pleistocene Earthq. Res. !nsf. Tokyo Imp. Uuiv.. Vol. l\·lollusca of California. Mcm. Sat! Diego 12. Pt. 3, pp. 566-638. Soc. JI.~·J/. Jlist. Vol. 1. -- (1938). Mollusca from the Miocene of HATAr. K. and NrsJ.YAMA, S. (1949), New Tyugoku. Japan. jour. Fac. Sci. Imp. U11iv. Tertiary Mollusca from Japan. jour. Tokyo. Sec. 2, Vol. 5. pp. 21-45. ------·------Explanation of Plate 1 Fi:;r. 1. Walluci11a okumurai !TorGAWA n. sp. Holotype. Reg. No. JC1500001. Left valve, x3. Loc. Shukubora, Hiyo:;hi-cho, l\olizunami City. Gifu Prefecture. Fig. 2. Walluci11a okumurai ITorGAWA n. sp. Paratype, Reg. No. JC1500002. Right valve, x 3. Loc. Same as above. Figs. 3a-b. Wtzll11cilla habei lTOJGAWA, a. Left valve. x 3. b. Internal view of 3a, x 3. Loc. Kamigiri, Iwamura-cho, Ena-gun. Gifu Prefecture. Fig. -1. Pilluci11a (Sydlorill!l) yokoyam:Ji (OTUKA). Right valve, x 3. Loc. Nakanishi. Yamaoka· cho, Ena·gun. Gifu Prefecture. Fig. 5. Pilluciwz (Sydlori~~rz) yokoyam1i (OTUK.-\.), Right valve. x 3. Loc. Same as a~?ove. Fig. 6. Caviluci11a (Mo11ililora) kitamr4rai (HATAI et N!SIYAMA). Right valve, x 1. Loc. Shukubora, Hiyoshi-cho, Mizunami City, Gifu Prefe..:ture. Fig. 7. Caviluciua (Mollifilor.J) kitamurai (HATAI et NISIYAMA), Left valve. x 1. Loc. Same as above. Fig. 8. Saxolucitw (Megaxinus) k·halaii (OTUKA). Right valve, x 1. Lo::. Matsubora, Toki-cho. Mizunami City. Gifu Prefecture. Fig. 9. Lucilwma aclliilimata (CoNRAD). Mold of Left valve. x 1. Loc. Hachiyato, Agi-mura. Ena-gun, Gifu Prefecture. Fig. 10. LuciiiOIIIO ac11filimata (CoNRAn). Right valve. x l. Loc. Same as above. Fig. 11. Luciuoma acutili11eata (CoNRAn). Left valve. x l. Lo~. Ebisuba, Haibara·cho. Yamabe­ gun. Nara Prefecture. Fig. 12. Luci11011W acutilineata (CONRAD), Mold of Right valve. x 1. Lo:. Hachiyato, Agi-mura, Ena·gun, Gifu Prefecture. J. hmGAWA: Miocene Lucinidae Plate 1

2 1 3a ,;···.,

5 4 3b

. .,·.,,.··.· ·..

6 7 8

. ' .. :"9o .."·~ . . ·. t.,~- . 10

12 Trans. Proc. Palaeont. So:. Japan, N. S .. 1\o. 25, pp. 7-10, April 3'). 1957

.306. BRACHIOPODA FROM THE DAISHAKA AND TSURUGASAKA FOSSIL ZONES, MINAMI-TSUGARU-GUN, AOMORI PREFECTURE.*

.TIRO KOTOH

Ishihara Sangyo Kaisha, Ltd., Tokyo Bran::!l.·Office

W~**~illlll!llEf:l. :k~iml:&V:~71!0Cft.:Ei#l::iiNt7.>~JJi~li'H::zt\,-c: l!im~ft.i;:.. 3(. fl'R.:Il~ :&:V:ili.X~~J: IJ, Pliocene c;i5;t t?:fL:<;,~i!JH:&U~71!t(ft1:imd~lil-ttJ.,}jjij)Em 6 fill Hemithyris psittacea (GIIIELI:o~). Coptothyris adamsi (DAvmso:-~), Terebratalia coreanica (ADAMS and REEVE), T. gou!di (DALL), Terebratulitta jopo11ica (SOWERBY), Laqueus rub?llus (SoWERBY) :.:gr\.,-c~~L. . .:.;tL~~~*.i-tt-?1Ui1\1.1J!ilifJ!~l3H~L.t.:.l{,v:>·CJ&7.>o ii ~ ~ rm

The Brachiopoda dealt with in this Here the coarse sediments occur in the article are from two localities, one (DS· lower part and the fine grained ones in 149) belonging to the Daishaka fossil the upper. zone, and the other CDS-140) to tbe The two fossil zones, from their stra­ Tsurugasaka fossil zone. The fossils tigraphic position, faunal content and were collected by the writer during the lithological characters, are regarded as summer of 1951 on the occasion of corresponding to the Shibikawa forma­ geological field work for the Nippon tion of Akita Prefecture, and to be of Mining Company. Pliocene age. The Daishaka and Tsurugasaka fossil Here I wish to r~cord my thanks to zones are exposed in the vicinities of Professors K. HuzwKA, T. INouE and T. the two mentioned lccalities as shown T.-\KAYAsu of the Akita University for in the text- figure. At Daishaka, the their encouragement, and to Mr. T. fossil zone consists of pebbly or cobbly. rvliYAJI:\1.\ of the Nippon Mining Com· coarse to medium grained sandstone pany for his interest taken in the work. with mudstone breccia and intercalates Thanks are also due to Dr. K. HATAI of thin mudstone layers. This zone is in the College of Education, Tohoku Uni­ fault contact with the adjacent forma­ versity, for kindly looking over the tions. At Tsurugasaka, the fossil zone manuscript. consists of granule conglomerate, coarse Leaving the micro-fauna for another to medium gra·ined sandstone and inter­ oppotunity, the brachiopods discriminat­ calates several thin mudstone layers. ed are: Daishaka Tsurugasaka Hemithyris psittacea (G:-.tELIN) ...... +- ... + Coptothyris adamsi (DA v 1 DSON) + + Terebratuli11a jopo11ica ( 3owERBY) · · · · · · · · ...... + Terebratalia corea11ica (ADAMS & REEVE) .. + ... + Terebratalia gouldi (DALL) ··· + Laqueus rub~llus (SOWERBY) ...... ······ ····· ...... ··· · ---- + * Read s~pt. 27, 1952-;- raceived May 7, 1956.

~ I 8 ]iro KOTOH

Text·fi~ure. Showing the precise localities: ...... Daishaka fossil zone. and 2 ...... Tsurugasaka fossil zone.

Of the above mentioned species. those though this species was found in both from Daishaka are of small size and fossil zones, each localities provided only few in individual number, while those a single specimen. The size of the spe­ frori1 Tsurugasaka are of lar~er size cimen measured about 7.2 mm in width, and common in individual number. 6.8 mm in length and 3.8 mm in thicl<­ From Daishaka, S. No1.tt:RA and K. ness; they have about 18 radial ribs in HATAI (1935) Iwve reported the occur­ the middle part ot the ventral valve of rence of llemitlzyris psittacea (GMELI1') the Tsurugasaka specimen and about a~d Terebratalia cct·emzica (AoA:o.ts and 14 of the same in the Daishaka specimen. REE\'E)- This species is known to inhabit a sea 1. Hemithyt·is psittacea (G:-.tELI:-> ). This bottom consisting of fine to coarse species is rare in the Daishaka fossil grained material providing it not be zonP., where it occurs only as broken or very muddy. The depth to which this isolated valves. In the Tsurugasaka species is known to extend at prerent, fossil zone it is found commonly either is from the tidal zone to about 37 fa­ as isolated or intact valve!:. In living thoms, and is one of the most common state, this species inhabits the seas of species of Brachiopoda known from the northern hemisphere, where it is northern Japan. This monotypic genus found from the tidal zone down to about is common in the tranquil waters 167 meters. It prefers a clean coarse of Mutsu Bay, Aomori Prefecture, but sea bottom. becomes less common in Matsushima 2. Coptotlty1·is adamsi ( DA v1oso;o.; ). AI- Bay, Miyagi Prefecture. and by going Brachiopoda from the Daishaka and Tsurugasaka Fos.>il Zones

still further south, the number of indi­ species is abundant, but generally occurs Viduals is much decreased, and at Ona­ as broken and isolated valves. Parta­ hama Bay in the ·southern part o~ king to a sea bottom consisting of fine 'Fukushima Prefecture, collecting is to coarse materials, this species has a difficult. Therefore, judging from the depth range of 40-499 meters in the seas frequency of occurrence, size of shell around Japan. and type' of radial ribs, it seems that · T~ese both fossil zones must be re~og. the types of these brachiopods resemble nized as the massing type of the so those in M~tsu ?ay, the thermal condi­ ·called ··Sweeping" accumulation in the tions during Daishaka and Tsurugasaka oceanic shallow water. While the zones was cooler than that 'of present ftowlngs of the wide 'spread may not be day Ma~s~shima Bay, the environmental 'considered because· of the species and conditions correlating with the. Mutsu the physical conditions that the fossils Bay, but deeper. are derived from them .. They may be 3. Tcrebratult'na. japonica (Sowr:RnY). seemed as the para-autochthonous fauna This species is commonly found in the .being accompanied with the allochtho­ Tsurugasaka fossil zone, and the ribs of nous one. and therefore, the sedimentary the younger specimens are rough and enviroments of these zones may be distinct. In living state it is known to considerable in general. be distributed entirely around Japan Namely, from the above mentioned and has been recorded from depths of data the Daishaka fossil zone can be 75-336 meters on the continental shelf. interpreted as containing a brachiopod' This species seems to prefer a sandy fauna which suggests, (a) accumulation sea bottom. in a sea \Vith a depth probably less than 4. Terebratalia coreanica (AoAMS and 80 meters, which is the most favorable REEvE). This species is represented by minimum depth for the brachiopoda it only fragments of the isolated and contains, (b) the thermal conditions broken valves of both valves, but of may be compared with present day different individuals. The specimens Aomori Bay but were deeper than th:c.t (restored) are smaller than those now and, (c) the paucity of species seems living in Mutsu Bay, Aomori Prefecture. to indicate that prevailing conditions This common shallow water brachiopod were not favorable for the flourishing is related to bottom control, but thermal of brachiopoda. The Tsurugasaka fossil barriers seem to have but little effect zone is characterized by, (a) an assem­ to its distribution. It ranges from 44- blage which accumulated in a sea with 296 meters on the continental shelf of a depth probably between 120-170 Japan where the predominating bottom meters, (b) the thermal conditions were material consists of fine sandy mud. similar to that of the Daishnlm fossil 5. Terebratalia gouldi (DALL). This zone but deeper, (c) the assemblage species is very abundant in the Tsuru­ consists of species which take to a mud­ gasaka fossil zone. In living state, this dy bottom and those which flourish on species inhabits on bottoms consisting a sandy sea bottom, (d) and finally the of mud within the range of 60 fathoms geological age may be Pliocene from to about 612 meters on the continental the absence of extinct forms, the evi­ shelf. dence afforded by the other fauna of 6. Laquezes rebellus (SowERBY ). This Foraminifera and Mollusca etc., and by 10 Jiro KoTCH the stratigraphic position of the fossil 2. Vol. 20, pp. 1-413. zones. -- (1950 ). Brachiopoda from Jizodo, Chiba Prefecture. Short Papers. IGPS., No. 1. NoMURA, S. and HATAI. K. (1935), On two Reference species of Brachiopoda from the Daishaka,

HATAI, K. (1937). The stratigraphic signifi­ shell beds of Daishaka. Aomori·ken. Saito cance of Tertiary Brahciopoda. japan. Ho-on Kai Mt1s., Res.Bull., 110. 5. ]our. Geol. Geogr., Vol. 14, 11os. 1-2. -- (1985), A Review of the Brachiopod Pale· -- (1988), The Tertiary and Recent Brachia· ontology of Northeast Honsyu, Japan. poda of Northeast Honsyu, Japan. Saito Saito Ho-011 Kai Mus., Res. Bull.. 110. 5. Ho-on Kai Mus., Res. Bull., No. 10. -- (1936), On Some Fossil Brachiopoda from -- (1939), A note on a Cenozoic Brachiopoda, Akita Prefecture, Northeast Honshu, Cop/athyris grayi (DAVIDSON). The Japan. Saito Ho-ott Kai Mus., Res. Bull. jubilee Publicatiatt it1 the commemoration 110. 1 o. pp. 183-194. of Prof. H. Yabe's 60th Birthday. rL\ Y ASAKA, I. (1933), Copt athyris grayi aomori· -- (1940), The Cenozoic Brachiopoda of ensis fossils. ]apmt. ]our. Geol. Geogr., Japan. Sci. Rep. Tohoku Imp. U11iv., ser. Vol. 10, 11os. 3-4, pp. 125-128. Trans. Proc. Palaeont. Soc. Japan, N. S., No. 25, pp. 11-16, Pl. 2, April 30, 1957

307. FOSSIL AND RECENT SPECIES OF THE GENUS PANOMYA FROM JAPAN*

SABURO KANNO

Tokyo University of Education

:<$:;f\lggf~:ri~l.:.l'.li!~ Pa11omya ~1.:.-0\.'"C: : Pm1omya ~O):fll!.1l{ll~HV.c l?tH::.:!tl!f.!l!i¥Jf.)-:ffi~~lMLt.:. 11' JE.f ~ f!ll

Introduction 2. P. turgida DALL, 1916. Proc. U. S. Nat. Mus., vol. 52, ·110. 2183. p. 416. In this paper is given a brief sum­ 3. P. beringia11a DALL, 1916. Ibid., p. 416. mary of the Japanese fossil and Recent 4. P. 11ipponica NOMURA and HATAI. 1933, species of the Genus Panomya and a Saito Ho-011 Kai Mus., Res. Bull., tiD. 5. p. description of a new species of fossil 20. pl. 1. figs. 7a-b. Panomya. 5. P. simotome11sis 0TUKA. 1934, Bull. Earlhq. Res. Inst .. 110. 12. pt. 3. p. 621, pl. 44·, fig~. The writer wishes to express his 66a-b. thanks to Profs. Haruyoshi FuJIMoTo 6. P. izumo NoMURA and HAT AI, 1938, fapmz. and Kotora HATAr of our Institute, for four. Geol. Geogr., vol. 16, 11os. 1-2, p. 6. their guidance and encouragement. pl. 1, figs. 2a-b. Thanks are also due to Messrs. Shigeru AoKI and Hiromi T AKAI of our Institute. Nos. 1-4 in the above list are living for kindly offering some specimens to species and the remaining ones are the writer. The writer is also indebted known only as fossil, but recently T. to Dr. Isao TAKI, Natural Science HABE (1955) lumped such Japanese Museum, and Mr. Sunao OcosE, Tokyo Recent species as" P. turgida DAr.L " and University, for their kind information P. tzipponica NoMURA and HATAI into P. on the bibliography. amp/a DAr.L.

Review of the previous works Distribution and Ecology

The following fossil and Recent spe­ The present genus is rather rare in cies of the geuns Panomya have been Japan both living and fossil. However, reported from Japan. the geological distribution of its living 1. P. ampla DALL, 1898, Trm1s. Wagner Inst. species seems to be restricted to the _Sci., vol. 3. pt. 4. p. 883. Pacific coast of northern Honshu (the main island of Japan). the Okhotsk and "' Read June 20, 1956; received June 21, 1956. the Bering Seas.

11 12 Saburo ·KANNo

·According toT. KuRoDA and T. HABE Family Hiatellidae (1952), the Recent species are distributed along the Pacific as follows; Genus Panomya GRAY, 1853 P. ampla DALL N. lat. 50°-68°, "P. turgida DALL" ~. lat. 43°-56°, P. bering­ Panomya GRAY, 1853, Figures oj Molluscus iaua DALL N. lat. 39°-64°, P. ttiP­ Animals vol. 5, p. 29. pottica NoMURA and HATAI N. lat. Pm10mya G1~AY, H. and A. ADAMS. 1858, The 39°. Ge1zera oj Recent Mollusca. vol. 2. pp. 351, Thus. from the known geographical 659. distribution it may be said that the Prmopaea MENARD, F. STOJ.JCZKA, 1871. Palellotzfologia bulica, ser. 4. vol. 3, p. 85. present genus is boreal in habitat. Panopaea. ME:SARD, L. REEVE. 1873. Mono­ The living species of Panomya are graph of the Genus Panopaea, pl. 5, fig. 7. usually buried in muddy bottoms com­ Pm10mya MENARD, s. P. WoonWARD, 1880, paratively offshore in cold water. and Manual of the Mollusca, pp. 492-493. are not known to bore into rocks. Panomya GRAY, W. H. DALL. 1898, Trans. The fossil species of Pattomya in Wagmr hzst. Sci. Phil.. vol. 3, pt. 4, p. Japan dale back to the Miocene of 832. Izumo, Shimane Prefecture, (N. lat. 35° Pmzomya G1~AY, T. HABE, 1952. Genera of 26'), which is the known southern limit Japanese Shells. 110. 3, p. 233. in Japan. the Miocene of Shiogarna, Genotype : Panopaea norvegica SPF.~­ Miyagi Prefecture (N. lat. 38° 14' 48"). GLER, Recent. the Miocene of Kurosawa, Akita Prefec­ According to W. H. DALL (1898), the ture (N. lat. 39° 14' 35'1) and the Plio­ genus Pmzomya has the following char­ cene of Suenornatsuyama, Jwate Prefec­ acters; •· Shell solid, large, irregular, ture (N. lat. 40° 171 30'') Accordingly, with a single cardinal tooth under the the distribut:on of the fossil species is beak in each valve: the pallial line of apparently more south than that of the unconnected, rounded impressions : the Recent one. The fossil species of animal larger than the shell, with large, Patwmya are usually entombed in fine­ united siphons, diverging slightly at the grained sandstone or mudstone. tips and covered with a wrinkled cori­ The present genus first appears in aceous epidermis." the Inferior Oolite in the United States Remm·ks : The present genus is of America, Europe and India. How­ somewhat related to the genus Mya ever, so far as Japan is concerned, the LINNE 1758, but it can be distinguished oldest record seems to be in the Miocene therefrom by having the pallial line of of Izumo and Shiogama, P. izumo and unconnected, rounded impressions, a P. simotomensis occur. The next single cardinal tooth in each valve. no younger occurence is in the Pliocene of chondrophre, and external ligament. the Suenomatsuyama and the Nakamura The present genus is also related to the mudstone (S. KANNo, 1955) where P. genus Panope MENARD, 1807, but it can simotomettsis and P. gigautca KANNo, be distinguished from the latter by hav­ n. sp. occur. No Pleistocene species of ing a medial depressed area, radial Patzomya have been reported from bounding folds, more shorter and non­ Japan. cylindt ical shell, and the pallial line of unconnected, rounded impressions. Systematic Description According to F. SToLJCZKA (1871), Fossil and Rec~nt Species of the Genus Panomya from Japan 13 the present group had been referred to 10. fig. 3. the genus Glycimeris of KLEI:-; (1753, Panomya amp/a, ABBOTT, 1951, Amer. Sea Tent. Meth., p. 130) based upon the shells, p. 454. Norwegian species. Subsequently, Pa11omya amp/a, HABE, 1955, Pub/. Akkeshi Mar. Bioi. Sta., 110. 4. p. 21, pl. 5, figs. LAMARCK struck with the pecular charac­ 3-4. ters of BRoN's Jl.fya glycimeris (=Chama Pauope (Panomyo) amp/a. GRA:ST and GALE, glycimeris ALDvANv), proposed a new 1931. Mem. San Diego Soc. Nat. Hist .. val. genus Glycimeris in 1799. However, a 1, p. 426, pl. 21, figs. lOa-b. few years back, when MENARD (1807) Panomya arctica turgida, KINOSHITA, 1937, Rep. proposed his name Panopaea, LAMARCK Fish. Surv. Hokkaido Fish. Exp. Sta .. no. dropped the name Glycimeris. The name 41, p. 30, pl. 9. fig. 53. Panopaea was proposed by MENARD in Pauomya turgida. HABE. 1952. Gen. Japan. Shell, 1807 for a Tertiary species, \vhich he 110. 3. p. 233, figs. 606. 607. calls P. Faujasi, but which has been Pa11omya 11ipponica NoMl.:RA and HAT AI. 1935. proved to be identical with the Recent Saito Ho-011 Kai Mus. Res. Bull., 110. 5, p. species Panopaea glycimeris of BRoN, or 20. pl. 1. figs. 7a-b. Panopaea Aldvatzdi of MENARD. Remarks : The so-called Japanese H. and A. AnAMS (1858) referred species Panomya ampla, P. turgida and these species to the genus Glycimeris, P. nipponica are merely transitional retaining the name Panopaea for P. forms as pointed out by T. ILo\nE (1955). norvegica of SPE:-

Earthq. Res. lnst., vol. 12. pt. 3, p. 621. over, this species closely resembles pl. 49, figs. 663.-b. DALL's P. ampla in the quadrate shell Panomya simolomensis, NoMURA. 1935, Saito outline, buf precise comparsion is dif­ Ho·Oil Kai. Mus., Res. Bull .. 110. 6, p. 223, ficult owing to the insufficient number pL 16, fig. 12. of specimens at hand. Panomya simotomensis, 0TUKA, 1941. jour. Distribution : Fujina, Tamayu-mura. japan. Assoc. Petrol. Tech .. vol. 9, 110. 2. p. 153, fig. 1, on p. 148. Yatuka-gun in Shimane Prefecture. Geologic 1·ange : Miocene. Remarks : The present species is a rather small type in the genus. This is characterized by the angle of the radial Panomya gigantea KANNo, n. sp. folds which are about 45°, and the beaks being situated at the anterior Pl. 2, Figs. 1, 2a -b. third of the shell. Distribution: Shimotomai, Nidatori, Shell large, rectangular in outline. all in the vicinity of Fukuoka·machi in inflated, gaping widely behind; anterior lwate Prefecture; Chiganoura, Shio· broadly rounded; posterior subvertically gama-shi in Miyagi Prefecture; Naka· truncated; ventral margin almost stra­ Nango, Sannai-gun, in Akita Prefecture. ight, being parallel with the postero­ Geologic range : Middle Miocene· dorsal margin. Beak situated at about Lower Pliocene. 4/10 from the anterior, rather small. somewhat swollen, turned-in and directed forwardly, not touching one another. Panomya izumo NoMURA and Surface ornamented with concentric, 1-IATAI, 1938 irregular undulations or rude concentric lines of growth, and the whole surface Pmwmya izumo No:.tURA and HATAI. 1938. with short, irregular, minor striations; japa11. jour. Ceo/. Geogr.. vol. 16. tzos. 1-2. p. 6. pl. 1, figs. 2a-b. medial portion of valves depressed and bounded by two radial folds, of which Remarks: This species is distin­ the anterior one is almost vertical to guished from P. simotomensis 0TUKA, the ventral border; angle of these two by its higher and shorter shell outline, folds is about 40°. Inner surface and and the beak which situated more cen· hinge not observed. tral in position than the latter. More- Dimensions (in mm) :

Reg. N'o. Length Height Thickness Valve Type 5600 130.5 85.5 25.0 Right Holotype 5601 ca. 137.0 98.0 Intact Paratype 5602 ca. 112.0 ca. 76.0 ca. 50.0 II ". deformed specimen

Remarks: Two specimens with both by its extremely larger and higher shell valves intact and three single valves (see text-figure 1). The present species, were examined. so far as the form ratio is concerned, This species is easily distinguished rather resembles P. beringiana and P. from such fossil and Recent species as turgida. The other species hitherto P. simotomemis, P. izmno, and P. ampla reported from Japan seem to be allied Fossil and Recent Species of the Genus Pm1omya from Japan 15

130 X

120

1!0

100

+

60

7 e 9 5') X

.\1 • 20 30 40 50 60 70 so 90 HEIGHT

Text-figure 1. Graph showing the form ratio of PatiOtiiJa giganlea KANNO, n. sp., and the known fossil and Recent species of genus Panomya.

1. P. giganlea KA:>:NO, n. sp. 7. P. amp/a of OLDROYD 2. P. beriugiana DAr.I. 8. P. 11ippouica NoMURA and HATA1 3. P. turgida DALL 9. P. izumo NoMURA and HATAT 4. P. turgida of KI:-;osniTA 10. P. simotomemis 0TUKA 5. P. amp/a of GRANT and GALE 11. P. simotometJSis of NoMURA 6. P. ampla of HABE 12. P. simotomensis of 0TUKA from Naka-Nango. to the Aleutian species, P. ampla. its site of burial. therefore, it may in­ The present species occurs rather dicate the thermal conditions of sea in rarely from the Pliocene Nakamura which it lived. The writer considers mudstone which is compcsed of grayish. that the molluscan fauna of the Naka­ pumiceous sandy mudstone. The new mura mudstone indicates cold water. species occurs in associated with These facts perfectly agree with the Patbwpecte1z sp., Fulgoraria sp., and thermal conditions indicated by the Echi11araclmius sp. The specimens with Tatunokuchi fauna (S. NoMURA, 1938). both valves intact are buried in the Reg. No. 5600 (Holotype), 5601 and strata in their natural position. From 5602 (Paratype). such evidence, it is inferred that the Locality : Railway-side, about 400 present species was not transported to meters south of Kornagamine station, 16 Saburo KANNO

Shinc11i·machi, Soma-gun, Fukushima In$[., T.okyo Univ. Educatio11, no. 4. pp. 11-23. Prefecture. KI:-

Explanation of Plate 2

(All figures in natural size) Fig. 1 Panomya gigontea KANNO, n. sp .. paratype. A left valve showing surface ornamentation. Figs. 2a-b. Panomya gigantea KANNO, n. sp., holotype. a. Right valve, surface more or les,; worn. b. Apical view of the same. S. K ANNo : Fossil and Recent Panomya Plate 2

S. A or

308. TWO CARBONIFEROUS CORALS FROM OKAYAMA PREFECTURE*

MASAO MINATO Geological and Mineralogical Institute. Hokkaido University and

KEIJI NAKAZA WA G.:ological and Mineralogical Institute. University of Kyoto

=of~ ':'~""" =.fi'"7 [U!]UJ* o Riiif :0:. c t 1.:.15~*C~I1<1) t <1) t ;7,..t..: i"'o ~~ lEI!Jii·~iR*=:

Amygdalophyllum giganteum ( YABE KA: On the identity of Echigophyl/um et HAYASAKA) and Clisaxophyllmn awa YABE et HAYASAKA (1924) and Amy­ MrNATo subsp. atetsuense nov. are gda/ophy/lu;n DUN and BENSON (1920) described. The former was newly found etc., four. Ceo/. Soc. ]apa11, Vol. 46, p. in a limestone developing at Niiyabara, !"39. 1955. Amygdalophyllum giganteum, Mit-:ATO: Tsuchihashi, Niimi City, Okayama pre­ Japanese Carboniferous and Permian fecture, while the latter was collected corals. four. Ji'ac. Sci., Hokkaido Univ., from the tufaceous limestone which ser. 4. Vol. !X. No. 2, p. 149, pl. 17, crops out near Morikuni, Niimi City, Fig. 9: text·tig. 16. Okayama prefecture. It is far from doubtful that the Description of Species present specimens now in concern are Amygda lopllyllum gigan teum wholly conspecific with Amygdaloplzyl­ lttm giganteum (YABE et HA v ASAKA) (YAnE et HAYASAKA) described and figured by HAYASAKA Pl. 3, Figs. 1, la. 2. from the Carboniferous limestone of 1924. Echigop/Jy/lum gigatlteum, HA YASAKA : Omi district. On the Anthracolithic limestone. etc., The writers have directly compared Sci. Rep. Tohoku Utriv .. ser. 2, Vol. 8. them with the holotype specimen now No.1, p. 20. pl. 4. Figs. 5. 6. 7. (On the deposited at the Inst. GeoL and Pala­ plate there were misnumbered as 9. 10 and 11. respectively.) eont., Tohoku Univ., Sendai. 1939. Amygdalophyl/um gigatrteum. HAY ASA- Echigophyllum giganteum was first established by YADE and HAYASAKA * Read June 20, 1956; received June 25, 1956 based on the coral specimens derived

17 18 Masao l\hNATO and Keiji NAKAZAWA from the limestone of Omi district, Nii­ Furthermore the dissepiments of the gata prefecture. This species. however, present specimens as well as those of was later transferred into the genus the holo':ype specimen of Amygdalo­ Amygdalopllyllum by HAYASAKA who phyllmn gigantcum are quite characte­ claimed the generic identify of Eclligo­ ristic, in the pattem of their arrange­ pllyllt!m with Amygdalophyllum. Thus ment. the generic name Echigophyllum has The dissepiments of this species. been abandoned. especially near the outer wall are The present senior author once fol· arranged in neither concentric nor lowed H,,YASAKA's view with slight angulo-concentric pattem, but they are doubt, and is now of the same opinion. ~arranged along the septa with convex The columella of Amygda/ophyllrmt sides faced inwardly. gigatzieum is observed to be composed Geological age: -Middle Carboni­ of numerous lamellar tissues. a very ferous? obscure median plate-like structure and This species has been regarded until septel lamellae-like tissues. The men· present day to indicate the Upper Lower tioned encircling lamellar tissues in ·the Carboniferous or Middle Carboniferous transverse section are never the usual in age, but the new specimens were axial tabellae. found in association with Waagerwphyl­ In the specimens presently at hand, lum, the latter of which is a quite defor· the columella is consisting of numerous med specimen and in ill-preservation, concentric lamellar tissues, and less although it is quite nearly related to numerous septal lamellae-like structures, lVaagenophyllum akasake1tsis (YABE), besides the obscure median plate. al· the Permian species. though the concentric lamellar tissues It is an open question, whether the are occas:onally discontinuous and inter· former species is a derived fossil brought rupted by a tmsting crack due to the in the Permina limestone as a pebble deformation of the specimens. or not, when the WaagNwpltyllum The concentric lamellar tissues in limestone was deposited. the transverse sections appear as a Reg. nos.:- 12460. 12461. Hokkaido tabellae-like strucutre in the longitu­ Univ., Department of Geology and dinal section; they are steeply ascen­ Mineralogy. ding towards the imaginary median plate. Clisaxoph~•llum awa MrNATO, However. in the usual species of subsp. atetsuense nov. genus Amygdalopltyllttm including the genotype, the concentric lamellar stru­ Pl. 3. Figs. 3. 4. cture is not so conspicuous in the Corallum simple, corallite ceratoid, transverse section of the columella as slightly CUT\'ed ? fairly large in size. in the present specimens and in the The septal number, calicular diamete1 holotype of Amygda/oplzyllum=Echigo­ and the diameter of the columella of Phyllum giganteum. Thus the structure each stage through the ontogeny of one of the columella in Amygdalophyllum specimen ate tabulated below : giganteum must be regarded to be much different from that of the usual species of Amygdalopltyllum. Two Carbonift!rous Corals from Okayama Prdecture I~

Sept:il number calicular diameter diameter or cotumella 31 31 15.0 mm 3.5mm 36 36 16.0 3.5 33 38 18.0 5.0 40 40 22.0 6. 5 43 43 26.0 9.5 45 45 28.0 10.0

Thus, the major septa increase in The septa, either major or minor number as the corallite grows. and ones, are very thick near the outer wall. become as many as 45, which are They gradually become thin distally in observed to be always alternating with early stage. However, the major septa the same number of minor septa. Dis­ show somewhat rhopaloid in the mature sepimentarium very narrow or almost stage, while such feature is never obser­ lacking in the early stage. while it vable in the minor septa even in the becomes much wider in the mature mature stage. stage. Cardinal septum is slightly shorter The columella is not solid in the than the other major septa; it is not neanic stage. In cross section. it is uniting with the median plate of the observed to be composed of the primor­ columella. Also the median plate is not dial columella or median plate-like connected with the counter septum at structure as well as septal lamellae- and any time in the ontogeny of the coral­ axial tabellae-like structures, all of lite, so far as observed. which show quite fibrous tissues. Dissepiments numerous, slightly an­ gular or sub-angulo-concentric in arran­ Also very fine vesiculate structure gement. Wall rather thick. are developed in this stage around the In the longitudinal section the diss­ primordial columella, and in the outer­ epimentarium is very wide in the most part of the columella. The feature mature stage where numerous dissepi­ is closely allied to the vesiculate colu­ mental vesicles are arranged in more mella observed in the corallites of than 10 rows., with their com·ex sides Clisaxop!tyllum azva MtNATo. faced inward near the tabularium, and Next. the sep';al lamellae-like stru­ upward as well as inward near the ctures become less distinct, and coarsely outer wall. The vesicles are variable in arranged vesicles develop well on both size, according to their situation; near sides of the mectian plate ; in the the outer wall they are very large in mature stage. the columella come to general, while they are smaller and sub­ consist of highly convex vesicles arran­ equal in size near the tabularium. ged concentrically, with their convex Septa with septal gratings are also sides faced outwards. clearly discernible in the longitudinal The feature is much like the colu­ section like the case of the cross section. mella of Clt'saxopltylltem awa MtNATo Columella in the mature stage con­ in the transverse section, as above sists of very sinuous median plate and stated, although there are observable quite numerous small vesiculate tabellae very fme and short trabeculae-like which are ascending towards the median ridges radially arranged in the columella plate, their convex sides being faced of the present species now in concern. outwards as well as upwards. Most of .20 Masao MINATO and Keiji NAKAZAWA

these structures are strengthened by awa MINATo, it is now ascertained by organic deposits. recent re-examination that such trabe­ Also there me observable numerous culae-like ridges in the columella are short ridges which are sporadically also developed, although they are never arranged in the columella, to diverge so prominent as in the present form. from the median plate. Their structures However there is no sign of the presence a1 e observed in cross section as if trabe­ of such definite :septal gratings in culae-like structures. Cli'saxoplzyllmn awa MINATo. Tabulae not complete, rather spora­ Geological rauge :-Middle Carboni­ dically arranged, counted 7 in a space ferous. of 5 mm, which are horizontal or gently Reg. 11os. :-12452-12459, Department inclined outwards. of Geology and Mineralogy, Hokkaido Remm·ks :-It is almost beyond doubt Univ. that the present form is most like Clisaxophyllum awa MINATO, but the Reference former is distinguishable from the latter in having trabeculae-like ridges in the M. MINATO: Japanese Carboniferous and columella. besides septal gratings. Permian corals. ]our. Fac. Sci .. Hokkaido In the holotype of Clisaxoplzyllum Univ., ser. 4, Vol. IX. No. 2,:1956.

------

Explanation of Plate :~

Figs. 1. la. 2. Amygdalophyllum giganteum (YABE et HAYASAKA) 1: (Cross section of corallite. x3.0) Reg. no. 12461. 2: (Cross section of columella, x5.0) Reg. no. 12461.3: (Longi· tudinal section of corallite. x 3.0) Reg. no. 12462, Loc.: Niiyabara, Tsuchihashi. Niimi City. Figs. 3. 4. Clisaxopllyllum awa l\hNATO var. atetsuense var. nov. 3: (Longitudinal section of corallite, x 3.0) Reg. no. 12453. ·~: (Cross section of corallite. x 5.0) Reg. no. 12454, Loc.: Morikuni, Niimi City. MINATo and NAKAzAwA: Two Carboniferous Corals Plate 3

M. MINATo photo. Trans. Pro::. Palaeont. Soc. Japan. N. S.. No. 25, pp. 21-25, Pl. 4, April 30. 1957

309. SOME MOLLUSCAN FOSSILS FROM THE EASTERN PART OF THE TANZAWA MOUNTAINLAND"

MATSUTARO SHIBATA

Kyo!Jashi Upper s~~ondary Chemical School

'1'1' .p 1'tiRtll~'!4!g).r.>1t:S: ftfRtli!JI!I::~N'5-t~~7' -ii-1!1!\"itftJ,i?~Hi-t ~1ti51:: --?1. '1:': it. -t-el:. : · ~~w.'iti:~J..o *~tt. $!1f!~lftJ,I?il¥.lli-t~1t;:et;:.-.,~, ,-c:I!JtiM t..t-:f!*· ~i?t-:1:: 3 rif~Z.~ c :-ht-:~~liC.lf~L-f-::o t.cio, ::$:ilfirmUfl~1t~·u. ll£i** · !ttf\l!!tt·~~.W:frtl?;tt."t:l.'7.>o ~t-:. $llli~"i 1lf~~.liil;il•l? Lepidocyclitw tZipponica HANZAWA tl;~.ei!.ttnt-:o t.tl.i?~~fl~i> c};: L."t:. ;$:!If! li,'l'/iif~~J'lo'lfft, ~ .t V:~Jt;:. ""'JI. '-!: ~ ~Zz L. t.:o ~ EE f.!: :t !'!&

(Crassostrea) gigas THUNBERG, and Ven­ Introduction ericardz'a panda (YoKOYA~IA): from the Reports in which the fossil mollusca ]ike formation were found ? Gibbula sp. from the so-called l'VIisaka series in the indet.. CoPiothyris grayi (DA\·msoN) and eastern part of the Tanzawa mountain· Echinoidea gen. sp. indet.: and from land have been listed but neither descri· the Fudojiri formation. only Lepidocy· bed nor illustrated are those by MrTsucHr clina nipponica HANZAWA and Gastro­ (1932), WATA::>~ABE, MrKA~II, Oso and poda gen. sp. indet. were discovered. SmNoKr (1952), and Sm:-;oKr and MrKA· :-.11 (1954). The discovery of Lepz'docy· Acknowll~dgements clina nippo11ica HANZAWA by MrKA:~u (1955) is noteworthy. The paleon­ !-or suggestions and help in many tological evidence of the mentioned ways the :writer wishes to thank the authors seem to be insufficient for deter­ following gentlemen. Profs. Haruyoshi mination of the geological age and cor­ FuJJ1\IOTO and Kotora HATAI of the In· relation, because their lists are not sup­ stitute of Geology and Mineralogy, plemented diagnostically. Tokyo University of EducaLion. Messrs. The writer was fortunate in collect· Masae O.MoRI, Sabur6 K.o\Nr-;o, Katsumi ing several characteristic fossils from HrRAYAMA, Shigeru AoKr and Hiroshi the Ochiai formation. The following UJIIE of the same Institute, 1\Iessrs. species were discriminated. namely: Kiyoshi KmKE, Yo NARUSE and Tokihiko Haliotis (Eulzaliotis) koikei SmaATA, n. MATst:oA of the Geological Institute, sp., Tegula (Chlorostoma) narusei Sm· University of Tokyo, Mr. Hisashi Ism­ BATA, n. sp., Tegula sp. indet., Turbo WARA, teacher of tl:e Second High School (Batillus) carnutus SoLANDER, Astraea of Hosei University. (Pachypoma) omorii SHIBATA, n. sp., !l!ytilus cfr. grayanus DuNKER, Osirea Stratiitraphic Sequence -----· ----- * Rea1 Jan 21, 1956; received June 4, 1956. The geological successions are as

21 22 Matsutaro St!IBAT.-\

Aikawa sub-group I fault Ochiai formation (350m) mainly composed of conglomerate, I fossiliferous . I .------mainly comJ)('sed of an alternation 0. ;::J ]ik~ formation (390m) of sandstone and mudstone, fossili- I 0... I ferous. I~ ------~ ~ ------«! -; ~ rJl I mainly composed of an alternation r.! Yataro formation (535 m) r.! of sandstone and fine tuff. :::N >. «! ~ ------E-< ;::J rJl mainly composed of an alternation ;::J C/) 63awa formation (800 m) of tuff (perhaps iron saponite tuff) and hard shale. I mainly composed of an alternation I Fudoiiri formation (938 m) ' of sandy shale and lappili tuff. fos- I siliferous. I 6yama sub-group follows: group. From thermal conditions and The stratigraphy of the northeastern the occurrence of Lepidocyclina, the and eastem part of the Tanzawa present group has characteristics com­ Mountainland will be reported in detail mon to those of Tertiary of Chichibu. by )~HIWARA in a later paper. particularly with the Chichibu-machi group. Therefore. the writer belleves Fossil localities that the Susugaya sub-group may safely be correlated to the Chichibu-machi The fos."3il localities of the Ochiai group. formation are along the Hayato-gawa It may be added that the discovery near Ocbiai, Miyagase-rnura, Aiko-gun, of Lepidocyclina from the base of the . Those of the Fudojiri formation is evidence sufficient }ike formation are around Hayato, Toya­ for stating that the Susugaya subgroup mura, Tsukui-gun, in the same Prefec­ is not older than the lower Miocene. ':ure, and the localities of the Fudojiri formation are near the dam of the Ochiai electric power station, Toya-mura, Systematic Description Tsukui-gun, also in Kanagawa Prefec­ ture. Genus IIaUotis LINNE:, 1758

Geological age and correlation Halioti1i (Euhaliotis) koikei The fossil fauna from the Susugaya SmBATA, n. sp. sub-group, as a whole, comprises warm Pl. 4. figs. 3a, 3b. \Vater inhabitants largely composed of those of rocky shores. Lepidocyclitza Sheil large, rounded oval in outline, occurs throughout the Susugaya sub- moderately convex. distance of maxi- Some 1\Iolluscan Fos3ils from the Eastern Part of the Tanzawa 1\Iountainland 23 mum height from margin about equal Shigarami PliOcene: MAKJYA:-.JA (1927) to one-third of greatest length, surface described Haliotis. kamtschatkana koya­ sculptured by subequal cords with mai n. subsp. from the Shigarami subequidistant interspaces and broad Pliocene: KutwnA (1931) listed Haliotis oblique wavy wrinkles: distance of apex kamtschatkana lwyamai MAKIY A:\JA from from margin equal to about one-seventh the Shigarami Pliocene; NoMURA and of greatest length: perforations on dor· NuNo (1932) described Haliotis lwm­ sal border, the last four holes abruptly tscluztlwna glabrosa n. subsp. from the but roundly elevated, the shape of the Yugashima Miocene: NO!t.tURA (1940) holes ellipsoid. other holes rounded in described Haliotis (Smzhaliati.s) japonica shape and nodulously elevated. REEVE from the Moniwa Miocene: \VA­ Dimensions:· Reg. No. 5280* (Bolo­ TANABE, ARAJ and HAYASHI (1950) re­ type) Maximum length 108 mm., mini­ ported Haliotis (Satthaliotis) n. sp. mum length 85 mm. (ca.), convexity 45 from the Haraya Miocene: " 7.-\TA:-.IABE, mm. (ca.). MmAllll and SuzuKI (1952) listed Haliotis Locality a11d formation :-Road-side kamtscltatkaua glabrosa No:-.wRA and along the Hayato-gawa near Ochiai, NIINO from the ltami Pliocene: and. Miyagase-mura, Aiko-gun. Kanagawa HANZAWA, HATAI, lwAt, KrrAML'RA and Prefecture. Ochiai formation. SHIBATA (1953) listed !ialiotis japo11ica Comparisons and affinities :-The pres­ REEVE from the Moniwa Pliocene. ent species closely resembles Haliotis naevosa iVI.-\RTY:-.1 and H aliotis gigantea CHEMNJTZ. But ·it differs from the for­ Genus Tegula Lr-:ssoN. 1832 mer by having less rolling of perfora­ tions, and from the latter by the follow­ Tegula ( Chlorostoma) narusei ing points, that is, 1) more rolled per­ SnrBATA, n. sp. forations. 2) lower elevation of every Pl. 4. figs. la, lb. lc. hole, last four holes ellipsoid in sha~e. 3) highest point leans towards apex. 4) Shell trochoid. more or less deformed, spiral cords subequal. interspace.<; sub­ whorls four in number, protoconch equidistant. missing: upper two whorls smooth, lon­ Remarlls :-The specific name is dedi­ gitudinal plicae developed on the surface cated to Mr. Kiyoshi KoiKE who gave of the penultimate whorl, but reduced me some instruction. Although the to mere oblique axial threads on the occurrences of fossil H aliotis are very last whorl: upper two whorls somewhat rare, the following reports are known:- compressed and the lower two whorls KocHJBE (1882) briefly described and steep: aperture depressed, rounded trig­ figured Haliotis gigantea CHE.l\JNITz from onal in outline, outer lip sharp. periph­ the Taga Miocene: YoKoY A'-IA (1925) eral. margin rounded; one strong tooth illustrated Haliotis gigantea var. llam­ on the inner lip: sub-umbilicated. tschatkana Jo::us (=Haliotis kamtschat­ Dimensions:-Reg. ~o. 5277 (Holotype) kana koyamai MAKJY.-\:>.IA) from the Height 35 mm., diam. 35 mm. Locality and for~nation :-Along the * All of the specimens are preserved in the Hayato-gawa near Ochiai, .Miyagase­ collection of the Institute of Geology and mura, Aik6-gun, Kanagawa Prefecture. Mineralogy, Tokyo University of Education. Ochiai formation. 24 Matsutaro SHIBATA

Comparisons mzd affinities :-The pres­ Ia som.ewhat concave, smooth, tooth­ ent species is allied to T e/[ttla n. sp. 2 like swelling on lower part. reported by WATANABE, A~

Explanation of Plate 4

Figs. la-c. Tegula (Chlorostoma) 11arusei SmBATA, n. sp. Holotype, Reg. No. 5278. a. Apical view x 1, b. Apertural view x 0. 85, c. Umbilical view x 1 Loc. Along the Hayato·gawa near Ochiai, Miyaga>e-mura. Aiko·gun, Kanagawa Prefecutre. Figs. 2a-c. Astraea (Pachypoma) omorii SHIBATA, n. sp. Holotype, Reg. No. 5276. a. Apical view x 0. 89. b. Apertural view x 0.89. c. Umbilical view x 0. 92. Loc. ditto. Fig3. 3a. b. Ha/iotis (Euhaliotis) koikei SHmATA, n. sp. Holotype, Reg. No. 5280. a. Apical view x 0. 81, b. Dorsal view x 0. 81. Loc. Road-side along the Hayato-gawa near Ochiai, Miyagasemura. Aiko-gun, Kanagawa Prefecture. M. SmsATA: Molluscan Fossils from Tanzawa Plate 4 Some Molluscan Fossils From the Eastern Part of the Tanzawa Mountainland 25

HATAI, K. and NrSIYAMA, S. (1952). Check REEVE, L.A. (1845). Conch. Icon., vol. III. List of Japanese TettiaT}" Marine Mol· Haliotis, 74 pis. lusca, Sci. Rep. Tohoku Imp. Univ., Smdai, SHINOKI. R. and !vhKAMI, K. (1954). Structure ]apa11. 2ttd ser., Special volume no. 3, p. of the Northeastern Border of the Tan­ 206. zawa Mountainland (Pt. 1), Studies from MAKIYAMA, J. (1927). Preliminary Report on the Geol. a11d Mitleral. Inst., Tokyo U11iv. the Tertiary Fossils from Kamiminochi­ of Educatio11, 110. 3. pp. 117-123. gun, Shinano, Chiky1t (Globe) vol. 8, no. 2, TRYON, G. W. (1890), Man. Conch., vol. XII, p. 188, Pl. III, fig. 4. p. 84, Pl. 7, fig. 42. MrKAMI, K. (1953a). Lepidocyc/i11a nipponica WATANABE. K., ARAI. J. and HAYASHI, T. from the Ochiai formation in the Eastern (1950), Tertiary Geology in Chichibu Border of the Tanza wa Mountainland, Basin. Bull. Chichibu Mus. Nat. Hist. 110. four. Geol. Soc. ]apm1, vol. 61, no. 717, 1. pp. 29-92. p. 274. --, MrKAMI, K. and SuzuKI, S. (1952a), - (1955b), Geology of the East Margin of sedimentation of the Shirahama Group· the Tanzawa Massif, Sci. Rep. Yokohama Geology of the Eastern Region of Shimoda Nat, Utliv. Sec. II, no. 4, pp. 41-64. Town, four. Geol. Soc. ]apa11, vol. 58, no. MrTSUCHI, T. (1932). Explanatory Text of the 678. p. 95. Geological Map of Japan, scale 1:75,000, -, --, 0No, K. and SHrNOKI, R. (1952b ), Hachioji. Imp. Geol. Surv .. japan, p. 34. On the so-called Misaka series of the NoMURA, S. and NnNo, H. (1932), Fossil Eastern Part of Tanzawa Mountainland. Mollusca from Izu and , Sci. Rep. Ibid. no. 681, p. 218. Tohoku Imp. Univ., Smdai, Japan, 2nd ser. WEINKAUFF, J, C. in MARTINI U. CHEMNITZ vol. 15, no. 3. p. 21, Pl. XII (II), fig. 12. (1883), Conchyl. Cab. Bd. VI, p. 26, Taf. OLDROYD, I. S. (1927), Marine Shells of the 4, Fig. 1, 2. West Coast of North America, Sta11j. Univ. WENZ, W. (1937), Handb. d. Palaeoz. Bd. 6, Pub/., vo/. 2, Pt. Ill, p. 165, Pl. 108, figs. Teil. I, p. 171. 5, 6. - (1938). Ibid. Teil Il, pp. 306, 361. OzAKI, H. (1954). On the Paleontology of the YOKOYAMA, M. (1925). Tertiary Mollusca Basal Conglomerate of Pliocene. Kanto from Shinano and Echigo, ]our. Foe. Sci .. Region, Bull. Nat. Sci. Mus. (Tokyo), N. S. Imp. U11iv. Tokyo. sec. II. vol. I, Pt. 1, p. vol. 1. (110 34), p. 12. 8, Pl. I. figs. 11, 12. Trans. Proc. Palaeont. Soc. Japan, N. S .. No. 25, pp. 26-30, Pl. 5, 1 text-fig., April 30, 1957

310. A MIOCENE FOSSIL CRAB, PARATYMOLUS YABEI N. SP. FROM NAGANO PREFECTURE*

RIKIZO IMAIZUMI

1st College of Arts and Sciences, Tohoku University

------~-

:lH!fJJil:rWmW"fft;J:;f:}!E:JII. "fft'1'~~:1ttl!dmi7J~.m11t:){(:JII.~J: IJi!il:l!l....f-: Paratymolus yabei n. sp. ~lm~l..... i-17J~llll~17Jilir~il:~:@B'~-e~Q.:. c~:mt.:t.:o ~17J P. pubescms lt. P. yabei l::l!!~L.. P. yabei :b>l?j!{tL.t..:~17Jc~k.l?;tL. W~O)JI.,~:OO~li P. sexspit~osus ltiii!Jt.fi:IIJll;:;tf!Y. L.f.:*~c~k.l?tLF.>o ~~$1::1t.n!!m17Ji'fiJmUf1:f!:~;tL"CI.'~ffil;til:~.;,o ~~.,. 311 ~o · ~:JR1Jit

ltllroduction :-Emeritus Professor reported as a fossil. Hisakatsu YABE kindly brought the fossil Acktzowledgements :-The writer is crabs frorn Chiyo-mura, Shimoina-gun, deeply indebted to Emeritus Professor Nagano Prefecture to the writer for his Hisakatsu YAsE who gave the specimen study. The fossil crabs were collected to the writer and has constantly guided by a school boy of the Chiyo Primary the writer's study. Thanks are due to School on April 12, 1949 and later they Dr. Tokio SmKAMA and Dr. Tune SAKAI were passed to the band of Mr. Yoshi­ who advised the writer for the further katsu SmMAOKA of the Yokohama study. The writer wishes to express National University, who was in the his thanks to Professors Sboshiro HA~­ course of the stratigraphic field work zA WA, Kiyoshi AsANo, Kotora H.nAI around Chiyo-mura. These fossil crabs and Kenzo YAGI Tohoku University, were studied by Dr. Tune SAKAI and for their continuous encouragements. Dr. Tokio SHIKAliiA of the Yokohama National University who determined them as a new genus and a new species, Family Majidae ALcocK "Ti'yocarcimts bifidus ", though their Subfamily Inachinae ALcocK detailed description was not given. The writer visited the locality of the Genus Paratymolus Mums, 1879 fossil crabs. in spring of 1956, observed Mnms, j. E:lward. 1879. On a Collection of the occurrence, and collected other Crustacea made by. Capt. H. C. St. ]OHN, species of fossil crabs. The fossil crabs R.N., in the Corean and Japanese Seas, are well preserved, showing their fine Proc. Zoo/. Soc. London, pp. 21, 45. and structures, and belong to the Paratymo­ 46, Pl. II. Fig. 6. Jus, which has hitherto never been SAKAI. T .. 1988. Studies on the Crabs of Japan, Ill, Brachygt~atha, Oxyrhyt~cha, pp. 207- * Read June 16, 1956; received July 9, 1956. 209, Pl. XXI. Figs. 1 and 2.

26 A Miocene Fossil Crab, Paratymolus yabei n. sp. from Ka5ano Prefecture. 27 compound eye is well preserved, and Paratymolus yab'ei, n. sp. square facets are about 20 p. in diameter. Pl. 5, Figs. 2-8, TeX:t-fig. 1 Dimemions (in mm) :-Length of Carapace elongate pentagonal and carapace, 8-9; with of carapace, 6-7; postero-lateral borders of the carapace length of cheliped, 4; length of appen­ longer than the antero-laterals. The dage, IG-13. ratio of length to width is 5 : 4. Surface Locality and geological horiznn :­ uneven and regions indistinct. Rostrum North cliff in the ground of the Chiyo large and acutely bilobated. The eye­ Primary School, Yonekawa. Chiyo-mura, stalk is exposed. Two strong spines at Shimoina-gun, Nagano Prefecture. IGPS the antero-lateral border, one on the loc. no. N-5, lat. 35° 25' 2" N., long. 13i0 anterior angle of the hepatic region is 51' 58". 6 E., middle part of Yonekawa larger and another smaller one behind formation, Tomikusa group l\liocene. of it is acute and directing forward. Repository :-IGPS coli. cat. no. 79485. The postero-lateral parts of the carapace Occurre1zce and preservation :-There are not completely preserved. Proto­ are about twenty specimens in the slab gastric region and the anterior part of which is 65 mm x 195 mm in size. The the meso.gastric region are covered with lithology of the slab beating_ the fossil granules. The granules on the former crab, is dark brownish gray fine siltstone. region are rotmd and larger than those Laminations are indistinct and perhaps on the latter. Epi-branchial and meso­ very thin, about 2 mm. thick. Between branchial region are also covered with the natural parting, the rock is massive small granules. Granules on the postero­ q.nd has concoidal fracture. Most of the branchial region are obliquely oblong, specimens are the impressions of the and under magnification, these oblong carapaces and some of them have been granules have two or three small granu­ doubly pressed by its theracic sternums les on the crests. with the abdominal somites on the Antero-lateral, lateral, posterior part impression of the carapaces. The fossils of the postero·lateral and posterior bor­ occur within so uniform and homogene­ der are rimmed with granules. Antenna ous matrix that the minor structure seems to be long. Thoracic sternum is such as the compound eye is well relatively narrow. Chelipeds are mode­ preserved, and the carapaces have rate in size, fingers are longer than retained some of its original relief. manus. movable finger with three or four Some specimens show the sternums with blunt teeth. narrower than immovable the abdomens of male, which are repla­ finger, and has a longitudinal row of ced by the carbonate material. Depres­ granules on its outer-upper part. Carpus sion of the sediment has produced of the cheliped has a spine on its inner considerable spreading and distortion. distal angle. Pereiopods long and Some of the specimens compressed slender, merus long and broad, with a obliquely and the sternums with the longitudinal groove on its postero-outer abdominal somites are displaced from part : carpus short, propodus broad at the impressions of the carapaces, but its distal half. shorter than dactylus ; the directions of the displacements are dactylus narrow. The· first and fourth not constant. pereiopods are longer than the second The impressions of carapaces are and the third. Minor structure of the separated from their sternums and Rikizo l~tAIZUMI

Text-figure 1-Paratymolus yabei. n. sp., showing dorsal view. the same specimen as figured in pl 5, Fig. 4. abdominal parts, and therefore the the thin laminae. These specimens of ~pecimens bearing ventral parts must fossil crabs are the same in size or have have their impressions of carapaces attained the same stage of growth. under the venbal surfaces. Regardless These grouped assemblage of the fossil of whether the specimens show their crabs is thought to be resulted from dorsal views or their ventral views, the their ecological condition. fossil crabs on the handspecimen always At the north cliff in the ground of lay their outer surfaces of carapaces the Chiyo Primary School (Fig. 1), a down on the surface of the slab. It is white siltstone is exposed lying confor­ concluded from this fact that these mably on this dark brownish gray fossil crabs had been living on some siltstone bearing Paratymolus yabei. plane of sediments. and were embedded Another species of the fossil crabs rarely later without an upside down arrange­ occur here with accompanied with the ment. Almost all the specimens have abundant mollusc remains, such as their appendages. These facts suggest So!emya tokunagai Y., and Yoldia that they were rapidly embedded in the ( Yoldia) laudabilis Yot<., Lt"nthia 1tippo­ sediments after death without agitation. nica YosmwARA, and plant impressions The field observations show that the such as Accr, Querc:ts, and algae. horizon of the fossil crabs, Paratymolus Callitmassa sp. aff. inornata NAGAo yabei is restricted to two or three met­ and H uzJOKA was reported from the res in thickness, probably, in the middle At·akida formation, which is correlated part of the Yonekawa formation. The to the Yonekawa formation, at Idozawa, fragments of the appendages of the Furujyo, Tomikusa-mura, Shimoina­ fossil crabs, Paratymolus yabei occur gun,*l (Smt

but such well preserved specimens as in ------~- this slab are limited to some parts of A Miocene Fossil Crab, Paratymolus yabei n. sp. from Nagano Prefecture. 2Q specimen is shorter than the manus, but ted, but is smaller than one of Paratymo­ Callianassa inonzata has the carpus of lus yabet'. The eyestalks of Paratymolus the cheliped that is longer than the pubescens is longer and slender than manus. This specimen is nearer to those of Paratymolus sexspitzosus, and Callianassa titaensis NAGAO than to those of Paratymolus yabei are long and Calliatzassa i11ornata, and the specimen stout. perhaps may belong to a new species The anterior hepatic spine of Pm·aty­ different from these species. Tymolus molus pubescens is smaller than the sp. also was reported from the Awano posterior one, which is large and acute, formation, which conformably overlies but in Paraty11zolus yabei, the anterior the Arakida formation, at Shonosawa. one is larger and broad at its base and Mombara, Tomikusa-mura, (SHIK.-niA, the posterior one acute and directing 1954, Pl. VIII. Fig. 7, List 16).*2 This more forward than the one of Paraty­ specimen seems to belong to Tymolus ·molus pubescens. kamadai IMAIWMI, collected from the Miocene Numanouchi formation, at east In Paratymolus sexspitzosus, there are two projections on the hepatic region cliff of Kosuganotsutsumi, Takalru-mura, Iwaki-gun, Fukushima Prefecture and and the anterior projection is larger Misaki, Ena-machi, Iwaki-gun.*3 than the posterior one. In the general Remarks :-From Japan. two Recent outline of the carapace, which is more species of the Paratymolus have been important feature than the eyestalk or the sizes of lateral spines, Paratymolus known, Paratymolus pubescens MrERs, yabei is nearer to Paratymolus prebescetzs 1879 and Paratymolus sexspinosus MrERs, 1884. The former is known from various which is far longer than width and of parts of the sea bottom, 10 to 100 metres which one specimen is 7.5 rnm in length deep, south of the Tokyo Bay. Sagami and 6 mm in width. Paratymolus sex­ Bay, Izu Peninsula of Shizuoka Prefec­ spitzosus is slightly longer than wide, ture, Mie Prefecture; Kagoshima, Naga­ of which one specimen is 10 mm in saki, coast of Kyushu and to Thursday length and 9.5 mm in width. The Island, northern extremity of Queensland. anterior width of the carapace of and the Gulf of Siam, and the latter Paratymolus pubesce1tS is almost the from rocky or weedy ground, not far same as the width of the postero. from the shore line, Shimoda of Shizuoka lateral part but the anterior one of Prefecture, India and Australia. Paratymoltts sexspitzosus is wider than The rostrum of Paratymolus pubes­ the postero-lateral part of the carapace. cens is larger than that of Paratymolus In these features, Paratymolus yabei is sexspitzosus, which is indistinctly biloba- more nearer to Paratymolrts pubescem; than to Paratymoltts sexspinosus. These *2 ~F fJl»!!llllt.'f:f;i:ftt!i~r~J.Il{EE / iTUi~~ w relations of the features suggest, *3 tiJiiJJII!l.;sJJ,tmi9i~:M,Hf~M~]!(u"ii~ i!IT hypothetically the following phylogeny =-mzrJ:Jri!P!filt7 about these three species.

Paratymolus yabei;; ...... ··· ...... Paratymolus pubescem

··· ...... ··· ... Paratymolus sexspinosus so Rikizo biAIZUMI Paratymolus pupescens has more dose relations of the phylogeny~to Paratymo­ References lus yabei than to Paraty'inolus sexspino· MIERS. J. Edward (1879). On a Collection of sus, and the latter is the one that has Crustacea made by Capt. N.C. St. ]OHN. not evolved from Paratymolus pubescens R.N .. in the Corean and Japanese Seas, but branched from Paratymoltts yabei, Proc. Zoo/. Soc. Londotr. pp. 21. 45 and 46, as the time of tremble, as Paratymoltts Pl. II. Fig. 6. pubescetts evoluted from Paratymolus IMAIZU:-.n. Rikizo (1938). Fossil Crayfishes yabei. from Jehol, Sci. Rep., Tohoku U11iv .• Seco11d The eyestalk of crab is related to the Ser. (Geol.). Vol. XIX. No.2, pp. 173-178, hormone gland controlling the ecdysis Pis. XXII and XXIII. and comparatively important organ. -- (1952). A Miocene Crab, Tymolus kama­ One can see the orthogenetic change of dai n. sp. from the Numanouchi Formation the eyestalk among these three species, of the Joban Coal·field, Tram. Proc. Palet:t. Soc. ]apatl, N. S .. No. 7, pp. 201- that is, in Paratymolus yabei the eyes­ 204, 5 Text-figs .. talk is large, in Paratymolus pubescens NAGAO, Takumi and HUZIOKA, Kazuo (1938), eyestalk is sle11der and in Paratymolus A ~ew Species of Cal/iat1assa from the sexsPi1_zosus it is small. There is no Neogene Tertiary of Hokkaido, ]our. such relation as above described in the Facult. Scie. Hokkaido U11iv .. Ser. IV, Vol. subordinate feature as the lateral spines. IV. Nos. 1-9, pp. 63-67, Pl. VI. Square facets of the compound eye of NAGAO, Takumi (1941). On Some Fossil Paratymolus yabei are about 20 p. in Crustacea from Japan. jour. Facult. Scie. diameter and smaller than the facets of Hokkaido Univ .. Ser. IV, Vol. VI, No. 2, the compound eye of Astacus spiniro­ pp. 85-lO:J, Pl. XXVI. stritts, which the writer has reported SAKAI, Tune (1938). Studies on the Crabs of Japan, III, Brachygtzalha. O:cyrhyt:cha. pp. 1938 from the Lycoptera davidi forma­ 19-l-364. Pis. XX-XLI. 56-Text-figs .. tion, Upper Jurassic or Lower Cretace­ SHIKAli!A, Tokio (1953). On the Tertiary ous, at Nieshutsekow, near Lingyuan, Formations of Tomikusa in South Nagano Jehol, (IMAIZUMI, 1938, Pl. XXIII, Fig. Prefecture, Sci. Rep. Yokohama Nat. U11iv .. 10); the diameters of the square facets pp. 71-108, Pis. IV-VIII. 16-lists. 2-Text­ are about 50 p.. figs.

------

Explanation of Plate 5 Figure 1. North cliff of the ground of Chiyo Primary School, Yonekawa, Chiyo-mura, Nagano Prefecture. Figures 2-8. Paratymol11s yabei, n. sp. 2. Slab bearing specimens, nat. size. 3. Well preserved specimen showing dorsal view. in the left of the slab, x 3. 4. Impressions of carapaces, showing dorsal views. specimens in a part, a little to the center of the slab, x8/3. 5. A specimen showing ventral view in the right of the slab, x 3. 6. A specimen showing ventral view in the right of the slab. x 8/3. 7. Fragments of a cheliped and an anterior part of carapace with eyestalks, in the right lower border of the slab, x 2. 8. Square facets of compound eye of the same specimen as figured in fig. 7, x 18. R. fMA I ZUMI : Paratymolus yabei, n. sp. Plate 5

1 Trans. Proc. Palaeont. Soc. Japan, N. S., No. 25, pp. 31-39. Pis. 6-7. 2 text·figs., Apr. 30. 1957

311. AMUSSIOPECTAN IITOMIENSIS (OTUKA) AND ITS ALLIES FROM JAPAN*

MASAHIKO AKIYAMA

Geological and 1\lineralogical Institute, Tokyo University of Education

Amussiopectetl iitomiemis (OTUK A) (: -=t o:>ili~fil!l:::. "?1. '"C : *~5'!J'Z.llt.liJ ;M/IIlfJ;~tJ' I? Pecten iitomiemis OTUK A (: t.. "C%~ t..tdt."El'i!Ht Amussiopecletl ~~:::.~ t... Amussiopecten hyugaensis SHU TO (t~c l"i.J-fl!l~~i .:::_ c :zomJi!i L, -=t"o:>:j!j:j;ClfU~fT"?t.:o "f. t.:, ;;f;:RJto:>f!go:> 3 .N!~1JJ.ll1t!ft L, -=ttL I? o:>;fA~B :Z.~ l?tJ,I:::. Lt.:o tl\ ill mE ~

The genus Amussiopecten has been locality should be referred to another reported from many localities of Japan species. and Formosa (Text·fig. 1). Amussiopecten planicostulatum (No­ Amussiopecten praesi'g1tis (YoKoYA- MURA and Nmw) was first described by 1\IA) was first described by M. YoKOYAMA S. NoMURA and H. NnN<> in 1932 from in 1922 from an unknown locality. In the Shirahama group at Ichiya near 1926 he pointed out ··that its locality Yugashima, Izu Peninsula, but no subse­ lies somewhere in Southern Totomi ", quent record of occurrence has been . This species is published. also recorded by the same author from Amussiopecten yabei (NoMURA) was the Lower Byoritsu beds in Taiwan first described by S. NoMURA in 1933 (1928) and Tonohama in Tosa (1929). from Taiwan. As was pointed out by T. KuRODA (1931), Recently, T. SnuTo (1955) described the specimen recorded from the former Amussiopf!cten lzyugaensis SnuTo and locality should be referred to another Amussiopecten praesignis (YoKOYAMA) species. In 1932 S. NoMURA and H. Nn­ from Hyuga, Miyazaki Prefecture, and No reported the present species from discusseq their stratigraphic significance. the Nawachi gold mine, Izu Peninsula, The writer studies some specimens but this specimen also represents ano­ of Amtt!isiopecten praesignis which are ther species. S. NoMURA and N. ZINBO probably topotypes, many individuals of recorded this species from the Shimaziri "A. pra_esignis" and some of "Pecten beds, Okinawa in 1936. Y. 0TUKA also iitomiens,is 0TuKA " from the southern reported the present species from the foot of Mt. Minobu, Yamanashi Prefec­ Tanna tunnel, Shizuoka Prefecture ture, and some of A. planicostulatum (1933), northern foot of Mt. Minobu. fro:n Abuzuri, Zushi City, Kanagawa Yamanashi Prefecture (1934), and south Prefecture. of Murono, Shizuoka Prefecture (1938), As -the result of his study, it becomes but the specimens from the second clear that the topotype specimens of .. Pecten iitomiensis 0TUKA " represent * Read June 20, 1956; received Sept. 8, 1956. Amussiopecten iitomie1lSis (OTuKA),

31 32 Masahiko AKIYAMA~

..

130 140

Text-figure 1. Map showing the distribution of the genus Amusliopectetr in Japan and Formosa. 0: A. praesignis (YoKOYAMA). x : A. iitomiensis (OTUKA). e: A. planicostulatum (NOMURA and Nmm). A: A. yabei (NoMURA). 1. Shizukawa sandstone. 2. Zushi formation. 3. Yugashima group. 4. Shirahama group. 5. Hamaishidake conglomerate. 6. Dainichi sand. 7. Tonohama group. 8. Takanabe formation. 9. Miyazaki group. 10. Shimaziri beds.* 11. Byoritsu beds.

* It is questionable whether these specim!!ns can be assigned to Amussropecten praesignis (YoKOYAMA).

including the individuals from the Acknowledgements are due to Dr. northern foot of Mt. Minobu which Y. Haruyoshi FuJIMOTo, Dr. Kotora HATAr 0TUKA (1934, 1936) once assinged to A. and Mr. Masae OMoRI of the Geological praesignis, and that Amussiopecten and Mineralogical Institute, Tokyo hyugaetzsis SnuTo is a synonym of this University of Education, for their species. valuable su'!gestions and reading of this In this paper the writer ~ discusses manuscript. Thanks are due to Mr. S. the above mentioned four species and AKAGI, Mr. T. MATSUZAKI and Mr. T. clarifies their variations and interspecific OMORI, for co-operating with him in col­ relationships. lecting some of the present specimens. Amussiopectan iitomiensis (OTUKA) and Its Allies from Japan 33 Shell large, rather thin, longer than Family Pectinidae high, orbicular in outline; right valve Subfamily Anrlisiinae more convex than the left; both valves Genus Amussiopectett SAcco, 1897 radiately ribbed. Right valve equilateral. Amttssiopectetz praesigttis (YoKOYAMA) with 16-17 broad, elevated, flat-topped, straight ribs, which become gradually Pl. 7, Figs. 5. 6. lower towards the ventral and the lateral 1922 Pecten praesig11is, YoKOYAMA. four. Geol. margins, and sometimes bifurcate at the Soc. Tokyo, vol. 29, 110. 530. pp. 1-2, pl. lateral borders and sometimes are 5, figs. I-8. single ; interstices narrower than ribs 1926 Pecten praesig~zis, YOKOYAMA, jour. Fac. themselves; internal ribs 13 pairs in Sci., Imp. Univ. Tokyo, Sec. 2, vol. 1, pt. number, wide and flat at the beginning 9. pp. 357-358, pl. 40. figs. I. 2; pl. 4I, but later becoming prominent and con­ fig. 1. cave to have prominent ridges on both 1931 Amussiopecten praesignis. KuRODA, Ve1ms vol. 3, appe11dix. pp. 76-77, fig. 80. sides: interstices becoming as wide as 1938 Amussiopecten praesignis, 0TUKA, jour. the ribs at the ventral border. Growth Fac. Sci., Imp. Univ. Tokyo, Sec. 2, vol. line weak but distinct. Ears subequal; 5, pt. 1' pp. 6-7, pl. 1. fig. 2. anterior ear being somewhat wavy and 1955 Am11ssiopecten praesignis, SHUTO. Trm1s. with weak but distinct byssal notch. Proc. Pa/aeo11t. Soc. ]apa11. N. S .. no. 20, Hinge line wing-like. Left valve almost pp. 103-104, pl. I7, figs. 1, 3. flat, ornamented with external and The six specimens collected by the internal ribs and growth liJ!es; external writer and his friends from the Dainichi ribs rounded and as wide as interstices. sand, · Shizuoka Prefecture take the The measurements are shown in following description. Table 1 (in mm.).

Table I

1 Reg. No. _L_.-:~ ~ J\~~:_~~~~~-1-l_in_g_ei_L__ N_;o_ibs_·_of_, __i_~~i~-~- '_V_a_l_ve_

5391--- 104 I 102 14 : 46 I .98 I .13 .4-1 17 127 : R. 5392 93 I 80 8 48 . 86 . 09 . 52 16 127 I R.

5393 114 : 108 7 't . 95 . 07 I R. 5394 66 64 10 I .97 .15 R. I 5395 84 73 8 36 .87 .10 .43 16 120 ' R

Remarks:-This species is character­ and the ratio of height to length of the ized by the rather thin, large shell which shell varies between .86 and .98. is provided with 16-17 elevated, flat­ The specimen figured by M. YoKo­ topped, bifurcating. radial ribs which YAMA (1928) from Lower Byoritsu beds have prominent ridges on both sides. shquld not be included in this species, The ribs become obsolete towards the as was suggested by T. KuRoDA (1931). ventral and the lateral margins and are The specimens reported by Y. 0TUKA broader than the interstices. The (1934) from the Shizukawa formation umbonal angle exceeds 120 degrees. It should be assigned to A. iitomie11Sis is noteworthy that the bifurcating ribs (0TUKA), and the one recorded by S. are hardly recognized in one specimen No:.IURA and H. Nm;o from the Nawachi 34 Masahiko AKIYAMA

gold mine may be also included in A. are sometimes dichotomous or bran­ iitomiensis. ching : in some specimens the ribs are Locality and horizon :-In the tunnel almost obsolete near the ventral mar. near Nishiyama. Haranoya-mura, Ogasa­ gin; interspaces subequal, very shallow, gun, Shizuoka Prefecture : Dainichi much narrower than the ribs, rarely sand.· provided with interstitial riblets; whole Repository :-The Geological and surface covered by numerous fine growth Mineralogical Institute, Tokyo Univer­ lines and a few stronger periodic ones: sity of Education. Reg. No. 5390-5395. hinge line less than one-half of disk­ Distribt~tiotz*: - 1) Hamaishidake length. Ears subequal in length ; ante­ conglomerate, Shizuoka Prefecture: 2) rior ear rounded in front and ornamen· Tonohama group, Kochi Prefecture; ted by almost obsolete radials as well 3) Takanabe member, Miyazaki Prefec­ as concentric lines of growth ; byssal ture. notch rather shallow and broad : post­ erior ear is obliquely truncated, orna­ mented by only concentric lines. Left valve quite similar to the right except Amussiopecten p!atzicostulatum for almost unequal ears." The four (Nol\WilA and NnNo) specimens of the right valve and a single one of the left which were col­ Pl. 7, Figs. 2. 3, 4. lected by the writer ·from the Zushi

1932 Pecte11 pla1ricostulatus NoMURA and Nn­ formation at Abuzuri, Kanagawa Prefec­ NO. Sci. Rep., Tolzoku Imp. U11iv., 211d. ture take the following description. Ser., vol. 15, 110. 3, p. 117, pl. 12, figs. Shell large, rather thick, longer than 2. 3. •l. 5. high, orbicular in outline; almost equi­ 1952 Anmssiopecte11 p/atricostulatrmr, HATAJ lateral and inequivalved: umbonal angle and NJSHIYAI\IA, Sci. Rep., Tolzoku Imp. llQ-120 degrees. Right valve with 15- U11iv., Spec. Vol., No. 3, p. 116. 18 broad, rounded, straight ribs, which The original description of Amus· become gradually obsolete towards the ventral and the lateral margins and are siopecten platzicosttelatum (No:o.mRA and rarely provided with interstitial riblets; NnNo) is as follows. interstices much narrower than ribs "Shell large, about 110 mm. in height, themselves. Internal ribs with promi­ compressed, length nearly equal to nent ridges on both sides at the ventral height, subequivalve, subequilatcral ; side. The shell surface has fine distinct sides straight, with margins smooth : concentric lines near the ventral margin. umbonal angle about 110 degrees. Test Ears subequal and wing-like; anterior rather thin. Right valve ornamented ear larger than posterior one.· Byssal by 18 low. rounded, subequal ribs which notch shallow but distinct. Imperfectly preserved left valve with low, rounded, • This species was reported from the radial ribs which are as wide as inter­ Neogene of Tanna tunnel (OTUKA 1933) and from the Shimaziri beds (NoMURA stices and provided with interstitial and ZDmo 1936), but thes: specimens riblets. Concentric Jines are weak but were not figured, so it is questionable distinct. whether they can be assigned to this The measurements are shown in species. Table 2 (in mm.). .4mussiopectetl iitomiensis (OniKA) and Its Allies from Japan 35

Table 2. No. of Apical 0 D. [Hinge H/L D/L Hinge/L Valve line Ribs -~~~ --- R::9~ -.l-i~i-\ : I 7 .87 .07 18 110 R. 5398 94 92 11 46 .98 .11 . 49 15 110 R . 5399* 91 87 9 48 .96 . 10 .53 18 120 R. 5400 ! 106 95 9 55 .90 .09 . 52 17 120 R. I -----~- - --' Remarks :-This species is characteri­ The type specimens figured by S. zed by the rather thick. large shell which NoM(;RA are ill preserved. S. NoMURA is provided with 15-18 low, rounded, pointed out that it is distinguishable radial ribs and with interstitial riblets from A. praesignis by the numerous on the left valve. The ribs become internal ribs, however it may be sug­ obsolete towards the Yentral and the gested from Text-figure 2 that his lateral margins, and are much broader specimens should be included in A. than the interstices. The umbonal planicostulatum. angle is less than 120 degrees. Th~ type specimens have sometimes dichotomous ribs which are never recog­ Api l Ar~ nized in the present specimens from the 0 Zushi formation. The left valve (NoMuRA 140 and NnNo 1932, pl. 12, fig. 5) to which S. NoMURA and H. N11No assigned their 1ao specimen may belong to the right valve. Amussiopecten praesig1tis ( Y oKOY A­ 1.20 MA) is closely allied to this species, but the former has more elevated, fiat­ topped, radial ribs, larger umbonal angle 110 and broader interstices which are never provided with interstitial riblets. IOC lAJcality mzd lzorizmt :-Abuzuri, Zu­ shi City, Kanagawa Prefecture; Zushi formation. 10 I~ Repository :-The Geological and Mineralogical Institute, Tokyo Univer­ Text-figure 2. Variation diagram of the apical sity of Education : Reg. No. 5396-5400. angle to the number of ribs with regard to Distributi(m :-Shirahama group near right valves of four species of Amussiopectm. Yugashima, Izu Peninsula. 0: Amussiopectetl praesigtlis (YoKOYAMA). x : A. irtomienSIS (OTUKA). e: A. plani­ costu/atum (NoMURA and NuNo). D.: A. Amussi"opecten yabei (No:-..IURA) yabei (NoMURA). 1928 Pecletl (Amusimn) praesignis, YoKOYAMA, Imp. Ceo/. Surv. ]apa11. Rep .. No. 101, p. * This specimen was collected by teachers 96. pl. l!'i, fig. 1. of natural science al the Meguro Fourth 1933 Pecten (? Amussiopecte11) yabei NoMURA. .Middle School in Tokyo, to whom the Sci. Rep., Tohoku Imp. Univ., 2nd Ser.. writer expresses his sincere thanks for Vol. 16. No. I, p. 59, pl. 2. figs. 3. 4. their gift of it. .36 Masahiko AKIYAMA

Y. 0TUKA 0934) assigned some of Amusswpccten iitomiensis (0TUKA) the scallops from the Shizukawa for­ Pl. 6, Figs. 1. 2, 3, 4. 5, 6; Pl. 7. Fig. 1. mation to Amussiopecten praesignis (YoKOYAMA) and Pecten iitomietzsis 1934 Pecten iitomiensis 0TTJKA, four. Geol. Soc. 0TUKA, but as the result of the writer's Tokyo. Vol. 41, No. 492, pp. 566-567, study, it becomes clear that they repre­ text-fig. 1. sent the same species, Amussiopecten 1934 Amussiopecten praesigm. 0TUKA. ibid., p. iitomiensis (0TuKA). 567. The original description of "Pecten 1936 Amussiopecten praesignis, 0TUKA, Geogr. iitomiensis 0TuKA" (in Japanese) is as Rev.. Vol. 14. No. 12, p. 978. follows. " This new species resembles 1952 Patinopecten iitomiensis, HATAJ and NISI· Pecten nagattumana YoKOYAMA, but the YAMA, Sci. Rep., Tohoku Imp. U11iv .. 2nd former has a flatter and higher shell Ser.. Spec. Vol .. No. 3, p. 110. 1955 Amussiopecten hyugaetiSis, SnuTo. Trans. than the latter. Patitwpectetz yessoensis Proc. Palaeont. Soc. Japan, N. S., No. 20. ]AY is allied to the present species, which pp. 105-108. pl. 16. figs. l, 2. 3, 4, 5; pl. has narrower and shallower interstices. 17, figs. 2. 4, 5. The measurements are as follows; height length thickness Specimen 1; (R. G. No. 1879) 50 47 6.5-5mm. (one valve) Specimen 2; (R. G. No. 1880) 57 t9 6.5mm. (one valve)

Ribs 14-15 in number. The specimens collected by the writer The apical portion of this species is from the Shizukawa sandstone take the flat and resembles Pecten laqueatus following description. SowERBY. Ears are simple and trian­ Shell large, rather thin, varying in gular; byssal notch indistinct. Radial outline from opithocline to prothocline ribs rounded and become gradually through orthocline. Right valve a little flatter and obsolete towards the ventral convex, slightly curved near the beak, and the lateral margins. The interstices provided with 12-15 low, rounded. narrower than the ribs themselves." straight ribs, which become obsolete

Explanation of Plate 6

Fig. 1. Amussiopecten ii!omie11Sis (OTUKA), Topotype, Reg. No. 5312, Right valve, x 1. Loc. near Kannon·bashi, west of Hayakawa-bashi, Nakatom:-machi. i\Iinamikoma-gun. Yamanashi Prefecture. Fig. 2. AmilSSiopectell iitomieusis (OTUKA), Reg. No. 5315, Right valve, x 2/3, Loc. Osozawa, Nakatomi-machi. Minamikoma-gun, Yamanashi Prefecture. Fig. 3. Amussiopecten iitomiensis (OTUKA), Reg. No. 5316, Right valve, x 1/2. Loc. Same as above. Fig. 4. Amussiopecteu iitomiensis (OTUKA), Reg. No. 5317, Right valve, x 2/3, Loc. West of Oharazima. Nakatomi-machi. Minamikoma-gun, Yamanashi Prefecture. Fig. 5. Amttssiopectell iitomiemis (OTUK A), Reg. No. 5324. Inner surface of right valve. x 2/3. Loc. Osozawa. Nakatomi·machi, lVIinamikoma-gun, Yamanashi Prefecture. Fig. 6. Amussiopecten iitomiensis (OTt.iKA). Topotype, Reg. No. 5311, Left valve. x 1, Loc. Near Kannon·bashi, west of Hayakawa-bashi Nakatomi-machi, rviinamikoma-gun, Yamanashi Prefecture. M. A K I YAMA: Amussiopecten Plate 6 Amussiopecten iitomiensis (OTUKA) and Its Allies from Japan 37

Table 3.

H ~~Hinge No. of Apical i I 11equi- Reg. No. L. ! H/L I Hinge/L Valve __·_ • line Ribs Angle • larity i 1 * 1- 5311 48 50 10 28 1.04 .58 13 105 .44 R. 5312 42 48 22 1.10 I .52 12 112 .52 L. I 5315 107 99 9 I . 93 13 102 I .62 R. 5316 134 119 7 60 .87 . 45 14 115 .43 R. 5317 118 98 7 48 . 83 .41 15 120 .46 R. 5318 55 11 120 L. 5319 89 76 8 .84 H 115 .45 R. I i 5320 52 41 5 26 .79 I .50 15 120 .40 R. 5321 71 59 7 .83 I 14 120 .54 ! R. I 5322 fi2 49 5 26 .79 .42 13 120 . ·12 R. 5323 50 42 3 30 .84 .60 15 120 . 49 I R. I 5324 Bl 98 12 .70 13 137 .33 I R. 5326 69 59 7 .85 15 117 .4.0 I R. 5327 13 100 I R. 5328 45 45 8 1.00 12 110 .38 R. I I ' 5329 94 71 7 37 . 76 I .39 15 130 .59 R. 5330 56 57 28 1.09 I . 52 11 110 : .47 L. I 5332 35 35 4 20 1.00 14 100 ' .50 R. I 5333 81 I 81 1. 00 14 110 .4fi R. I I gradually towards the ventral and the straight. radial ribs. The ribs become lateral margins: interstices much nar­ obsolete at the ventral and the lateral rower than ribs themselves. Internal margins, and are much broader than ribs have inconstant width and promi­ the interstices. The present species, as nent ridges on both sides near the was pointed out by T. SnuTo (1955), ventral bol'der. Concentric striations has variable characters. Above all, the weak but distinct. Ears subequal. ante­ shell outline varies from opithocline to rior one being somewhat wavy and with prothocline through orthocline and from weak but distinct byssal notch. Hinge .33 to .62 in lnequilarity. line wing-like. Left valve flat, orna­ Amussiopecten hyugaensis SuuTo is mented with 11-12 round-topped. radial synonymous with this species. ribs and a few interstitial riblets: A. praesignis (YoKOYAMA) is closely interstices broader than ribs at the upper allied to the present species, but the half of the disc ; on the other hand, latter has less numerous, rounded, radial near the ventral side interstices as wide as ribs. The measurements of the selected * Inequilarity :-This term is propos:'!d by the writer to express a degree of an specimens are shown in Table 3 (in inequilateral shell. 'Ve draw a perpen· mm.). dicular from th:'! beak to length-line ab, Remarks :-This species is charac­ where the point a is at the anterior, and terized by the rather thin, large shell call a point of intersection c. Here, ac/ which is provided with 11-15 rounded, L is defined as " I nequilarity ". 38 Masahiko AKIYAMA ribs which never split into riblets and (YoKoYAMA) is characteristic in the the interstices are nmch narrower. Lower Pliocene deposits along the Pacific A. pla1licostulatum (NoMURA and coast of Japan. The genus Amussio­ NuNo) is also allied to the present pectm. therefore. is to be considered to species, but the latter has less numerous have valuable stratigraphic significance. ribs which never split into riblets. Localities and horizon :-Osozawa and Geological significance :-T. SnuTo near Kannon-bashi in Nakatomi-machi has stated that " A. lzyugaetzsis " occurs and Oharazima in Minobu-machi, Mina­ from the lower part of the Miyazaki mikoma·gun, Yamanashi Prefecture: group in south-east Kyushu, while A. Shizukawa sandstone. praesigtzis is found from the upper hori­ Repository :-The Geological . and zon of the Takanabe member. Consider­ Mineralogical Institute, Tokyo Univer­ ing the above mentioned facts and the sity of Education. Reg. No. 5311-5313, morphological relationship of these two 5315-5324, 5326-5334. species, it may be suggested that A. Distribution*:- Lower part of the praesigtzis is derived from A. iitomiensis Miyazaki group. (A. hyugaettsis). If this is true, the Shizukawa sandstone (OTuKA 1955) in Literature Cited Yamanashi Prefecture which yields A. KuRODA, T. (1931), Catalogue of the Japanes~ iitomiemis may be Upper Miocene in Mollusca (in Japanese). Venus, Vol. 3, age. Unfortunately, the writer has had (appendix). no chance to examine A. praesignis (?) NoMURA, S. (1~33), Catalogue of the Tertiary reported from the Sagara formation, but and Quarternary Mollusca of Taiwan. Sci. in the future, he wishes to collect and Rep., Tolzoku Imp. U11iv., 211d Ser.. Vol. 16. study those specimens and to clarify the No. 1, pp. 1-108, pis. 1-4. phylogenetic relationship between A. * The specimen recorded by S. NoMURA Pmesignis and A. iitomiensis. and H. NnNo from the Nawachi gold Y. 0-ruKA (1938) stated that the mine, Izu Peninsula may be assigned to fauna occurring in association with A. this species. ronsidering that it has less praesigtzis and Veneri'cardia panda numerous ribs.

----.- -- ... ------

Explanation of Plate 7

Fig. 1. Atmlssiopecteu iitomiemis (OTUK.-\). Reg. No. 5318, Left valve. x 2/3. Loc. Osozawa, Nakatomi-machi, Minamikoma-gun. Yamanashi Prefecture. Fig. 2. Amussiopecten plauicostulatum (NoMURA and NriNo), Reg. No. 5400, Right valve, x 1/2, Loc. Abuzuri, Zushi City, Kanagawa Prefecture. Fig. 3. Amussiopecten p!atricostu/atum (NoMURA and NuNo), Reg. No. 5399, Right valve, x 1/2. Loc. Same as above. Fig. 'l. Amussiopecten plauicostrllatum (NOMURA and 1\uNo), Reg. No. 5396, Left valve, x 1, toe. S:~me as above. Fig. 5. Amussiopecteu praesiguis (YoKOYA:O.tA), Reg. No. 5391, Right valve, x 1/2, Lo;:. In the tunnel near Nishiyama, Haranoya·mura, Ogasa-gun, Shizuoka Prefecture. Fig. 6. Amussiopecte11 praesig11is (YOKOYAMA), Reg. No. 5392, Right valve, x 1/2, Loc. Same as above. Plate 7 M . AKIY AMA: Amussiopecten Amussiopecten iitomicnsis (OTUKA) and Its Allies from Japan 39

NOMURA, S. and NnNo, H. (1932), Fossil Crustal Movements in Japan (in Japanese). Mollusca from Izu and Hakone. Ibid .. Bull. Eartlzq. Res. hrst .. No. 33, pt. 3. pp. Vol. 15, No. 3, pp. 169-192. pis. 11 ·12. 4•19-169. -- and Zt:mo. N. (1936), Molluscan Fossils SnuTo, T. (1935). Amussiopecten from the from the Shimaziri Beds of Okinawazima. ~[iyazaki group, l\liyazaki Prefecture, ibid.. Vol. 18. No. 3, pp. 229-265. pl. 11. Japan. Trmrs. Proc. Palaeo11t. Soc. Japml, OTUKA, Y. (193•1). Some Fossils from the N.S., No.20, pp.l01-110, pis. 16-17. Northern Foot of Mt. Minobu and the YoKoYAMA, M. (1922), On a new species of Hayakawa Tuffite of Hakone(in Japanes.:). Pecten from the Neogene of Japan. Jour. Jour. Geol. Soc. Tokyo. Vol. 41, No. 492, Geol. Soc. Tokyo, Vol. 29, No. 530. pp. pp. 562-568. 1-2. pl. 5. -- (1936}, Geological Investigation of the -- (1926). Tertiary Mollusca from Southern Fuzimiyama Fault Scarp in the South· Totomi. four. Fac. Sci .. Imp. llniv. Tokyo, western Part of Yamanashi Prefecture (in Ser. 2. Vol. 1, pt. 9. pp. 313-364, pis. 38- Japanese). Geogr. Rev., Vol. 14, No. 12, 41. pp. 969-984. -- (1928). Mollusca from the Oil-Field of -- (1938), Neogene Fossils of the Ihara the Island of Taiwan. Imp. Geol. Surv. District, Shizuoka Prefecture. Japan. Jour. Japa11, Rep. No. 101. pp. 1-112. pis. 1-18. Fac. Sci .. Imp. Univ. Tollyo. Sec. 2. Vol. 5, -- (1929). Pliocene Shells from Tonohama, pt. 1. pp. 1-19, pis. 1-2. Tosa. ibid. 1\"o. 104. pp. 9-17, pis. 7-8. -- (1955). Shizukawa Group and Tertiary 40

PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY OF JAPAN

rs*tl.?!:.!/fJJ!!f-~ 1956 ~A~J 1957 <1f-2 JJ io. Radulotzeclites. a New Pectinid Genll!Y 9 s JfCJ.ltx~l!l!~tffl:r&j(~~~~'L~""'-c~tm from the Liassic Kuruma Group in Central Japan ...... ltaru HAY AMI L.fc: (~~:f'f 38 tJ)o <1f.~tcb"NT .Oh~:Mz 11. Liassic Chlamys, Camptonecles and other t;tc•Jmml'l~ 15. Trigonian Faunule from Mindro in the 4. :ltJ::Ill~im Sciophyllum (I'YMJilllfMl) (ft!'l) Philippines ...... Teiichi KOBAYASHI ...... ~ IE~ 16. Bemerkungen zur systematischen Stell­ 5. Occurrence of Jurassic Burmirhynchia ung von Motzotis. Etztomotzotis, Claraia. Emlzor­ BUCKAMAN from Japan ... Akira ToKUYAMA photis und einigen anderen Triasgattungen 6. On some Triassic Spiriferinids from the der "Pteriidae" (~!!) ... Koichiro IcHIKAWA Sakawa Basin, Prov. Tosa ... Akira ToKUY AMA 17. Cambrian Fossils from Peninsular 7. On some New Species of Palinopecletz Thailand ...... Teiichi KOBAYASHI from the Tertiary formations of Chichibu 18. tt~U'Iillilli o:> itEit.!R1ftO)--t/ifll::. -?1., -c Basin, Saitama Prefecture. Japan ...... ,i;Jill.j~;flr ...... Saburo KANNO 19. Some Notes on a rare Species Tricera­ 8. The Molluscan Fauna from the Environs tiwn simplex J. Brun ( fta) ... Wataru IcmKA w A of the Zenkoji Hotspring, Nagano Prefecture, 20. J;l!~~gg*C~im7t'I3HtE ...... 1Jf~B=fl~ Japan ... Saburo KANNO and Tsuneo TOMIZAWA 9. Cretaceous Pennatae Trigonians in Japan t!YBOi/4~ m 4 @l§!i!~fi!r~~~.itt!-&l-l.:.llilm L -c ...... Mitsuo NAKANO (i]j:'ffj!ffl) ...... ~~~

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On the occasion of the Annual Meeting of the Palaeontological Society of Japan, held at Tokyo, February 9, 1957, it was decided upon to revise Article 1 and Regulation for Publication 3 as indicated (in italic) below. Article 1. Name. The Society shall be known as the Palaeontological Sot'iety of Japan. ("The Society is a section of the Geological Society of Japan" will be omitted.) Regulations for Publication. 3. Manuscripts (includiue of text-figures. maps and tables) will be limited to 8 printed pages (less tha11 18 type-written pages).