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080057-10.016.Pdf 'Irclep 'sBuIpuIJ ur pauttrr€xeeJe,r ecuullls aqt lo srequreur1p;o fiuouoxsl puu ernleltueruoueql eseql 8ul$olloJ pue (JVIA1)sn Jo euo Fq pa{Jeqcflluenbesqns uIBBu erer'r suaultcads ed,fi eq1 reltu.tts,fte,t 'lpu'IalDor8 ara^\(wntqcotsotJDui 4,goru,{uou,{s u spsJoqlnu tueceJ dq peteeJ4) d PuesuaSuu d lo 'fJe^ocslp sedfl eqtrtruqtr trueruddB etuef,oq lI ,,'\eueqt etEcqueqlruol dnoJ8eql ul ExEtIIB Jo sod,{leqlJo sqdurSoloqd?urlceqcer uod11 elJeqeHeullned put uo1 fq sJoqlnuerlt Jo uonue " aql o1lq8norq sB,l\ecusrllB srql ur serceds:eqloue6361 uI {Jo,,$plelJ Suunc seleg'I'd t4ulJlt2J dpue sel€g fd ( qqJU) ,qa8r! 'd 'utn(qcolsoJJolu'd '2f^'GB6l sel"g) ecuerlleslql ut serradsaerqt eq o1peleplsuoc aru ereqt fpuarrn3 sre/!\og ,,rroJr€u'lplus ,{lo^It€leJpu" llq€I{ repuels d:e,l e ,{q elquzruSoce: orBecu"rllB srql Jo sJequ?],{ ( V S) erFl]SnV qlnos puE( V i(\) PIIEI]SnVuJelsa \ uI sJnccoqcrq,|d 'd sercads;o ecuElllB Jg uttl|qcotsoroDutwn tqdosoJJ erF ueeq sEqeruoselqnoJl flrBln3lu€d '(ss?rduI seuof) peSeydruoc,trou xeldutoc '4 'roqtnE wnuldlo er11uoselpnts esaql Jo lsrg eql qlta Jolueseql ,(q peult[exe ?uteq ere sexelduoc 'snue8 snouel eqt Jo ,{pnls1p;e,ro ue ;o ped sy suorurcadsurnueqreq paup 3ut,(gnuapt;o ,(tlnctg;tp eql puB uxsl uee,rteq FSoloqdrou prol+ Jo fluElruJrsIIeJeAo lueJBdde eql Jo esneceqsJeIJo,{\ plag pur slsruouoxel fq pecuerredxeueeq e,lEq sxBt Jo uonscgrlueptqlt,,rr surelqord elqereplsuoJ €leue8 prqcJouEr[pISnV Jo llncuJrp lsoru erl] Jo euo ,{llecluouoxet s|tg'A unl\Kqdosrzla,snue8 eql uogJnpoJlul 'erleJlsnv uJelsad\Jo lse^\ ' 'd 'aDurlnod -qtnoseqt ur seJ" palculser€ tuo{ pequcsepsI Jg d art?,{4rpLsoirDut ol pelel€.J(lesolc 'sercads un11Kqdosot4 ,tou eru: y eddloloq €ql Jo uoqeultuuxeI"cItIJc " regu peIJLIs[csI IpuI-I alnot8 wnl1|qdosotd prqcrou€rle.qsnv erpJo sn1€tscnuouowt eqJ (966I) 8It-60t:(t) 0I sIS nN 'Erl€Isnv 'aoutlmtd '(eEscBp[lcJo) uJetselAlse,\\-qlnos uo.rJ serceds 1t1Leu e 7;o uondrrcsepeql r0L,[ 'I wn(qtolsotcow 4 puu a1aot8 wn11tt1dosudJo tueussess€e.IV V tr{'slueurolJ pull C'seuof psrlsqY '009 t09z Ertxllsnv'ereqwJ t xog v J N v urnrreqreHlEuortBN rrBrls4snv q.rEesad ^llsre^lpolg lrr?ld roJ e4ue3 'I slueuelJ v {.r"JI pu€seuol pr^€c sllarlsnY uralsal\ lsa^r-qlnos I '(eBeJEplqJJO) ruo{ salJedsL\eu e aounnDdA Jo uolldlJJsepaq} ql.r,r{ 'd unttlcDtsolcma pue aflcDJBMryItqdosDld lo tuaussasseeJV (966I) 8lt'60i:(€) 0l EIs nN NuytsiaVol. 10,No. 3 (1996) Methods This study is based on field observationsand herbarium collections (spirit and dried) from the following Australian herbaria (AD, CANB, MEL and PERTH). Type specimensof all previously described taxa have been examined including those in overseas herbaria (BM, K, K-L, W). Measurementsgiven in descriptionsare fron living plants or spirit-preservedspecimens. Notes on distribution, habitat(particularly soil andplant association)and conservation status were derived fron our own field studiesand consultationswith A.P. Brown and R. Heberle. Taxonomic history Prasophyllum macrostachyumwas describedby Brown in 1810based on collections he made in swampsnear Albany, W.A. In 1840Lindley describedP. gracilebasedon matedalcollected by James Drummond, probably from near Perth. Six years later Endlicher (1846), determinedtwo Preiss collectionsfrom Westen Australia asrepresentative ofP. algrlcansR. Br andP. graclle, eventhough both collections are in faot the same species. Reichenbach(1871), after studying the same two collections,determined that the specimenattributed by Endl cherto P. nigricans was rnfact P. gracile Lindley and the secondspecinen, determinedby Endlicher as P. gracile, was a new taxon which he described as P. gracile var. ringens. George (1971), after a study of the types, made the new combination of P. macrostachyumvar. ringens and reduced P. gracile to a synonym of P. macrostqchyumvar. macrostqchyum.In a revision of the alliance,Bates (1989) describeda third species,P. calcicola, from matedalcollected on YorkePeninsula, S.A but which also occursin W.A. In the samepaper he elevatedP. macrostachlum var. ritge,?sto specific status. Clementsand Jones (in Clements 1989) independentlyreached a similar conclusionabout the statusof this taxon and renamedit as P. antennqtum. A subsequentrc-assessment of the type collection of Pra.sophyllum gracile by one of us (MAC), housed in the Lindley Herbarium, Kew, showed that the specimenshave lateral sepalswith long, acicular terete apicesand this speciesis conspecific wlth P. ringens, and not P macrostachJumas previouslyinterpreted by George(1971) and Clements (1989). Accordingto article11of ths 1994 International Code of Botanical Nomenclature,Prasophyllum gracile Lrndley has priority over P. ringens (H.G. Reichb.) R.J. Bates. Studies of the types of other speciesin the complex have confirmed that their taxonomicstatus agrees with that accordedto them by Clements( 1989),and show that the taxon rccently discoveredby Ron and PaulineHeberle is new. Subsequentfield studiesby the seniorauthor have allowed the comparisonoffresh materialof P. mqcrostachyum,P. gracile and the new taxon. These studieshave supportedthe recognitionof the three taxa detailed in this paper and have revealed further distinguishing charactersthat are not readily evident in herbarium specimens.The taxonomy and morphology of the three taxa is presentedhere. Prasophyllum calcicolo was the subjectof a recentstudy (Bates 1989) and is not describedhere but is included in the key. Taxonomic treatment Key to species L Flowers6-10 rnm long, column wings well developed ...............................2 Flowersc. 5 mm long,colunn wingsvestigial ................. ........P. calcicola '886I-0t-l 's{cod '2961 's{cou EuolcIA T LL6Z sauot I s€pun(I:(HJdAd) E6IV aSroaD 6-ZZ '0g6I-6- 'ur8"td 'tuEql€'l 'CV) sawg :(CgJ) EL\Z stuawalJ LZ Jo tsa.^\UDI a lhtr :(SNVJ t6ll 't86I patalag 6 Z leuBEINlhlJo id\lN slcol :V'I-IVUISnV NUAJS8II (peulu€xr 6t) suawtcads 'epr,^d 'plo^oqo (I ernBIC) uae:8 ,{urqs lrllu 9 €-€'Buol ttttrt ,-9 alnsdDS 'se?upueddEeql uBql req8tq q3nul ulnllelsoJeql'epl^\ utrl 9 g'3uo1 uur Z l ,'lrlJoJiuar ',$olle,{'8uol 'plo^€lc-l€eu11erur11od:e1u1n3r1'3uo1 ,(lasrelsue.q12 8?JS eluces tutx I J ruu 91 9 r '3uo1 snlnupq :ellq,\^ urut 16 c'oqdqle uInIPrJsI^:3uol tutu Zl J wnltDulllod u'{\orq{I?p 'epr,r '8uol 'e.tu^o 'ueel8 'epl,l^ '8uol '3]BloeJuul rrtu Z l rlrtu I 2 taqwv esnlqo elEd uul ? 0 rrrtll 1 J 'epr '3 -8uo1qo,{1,{ouuu se8epueddp : slEted puu pdes psrop eqt ,{q uepplqlou ^\ urur8 1 uoI unuI I J ',{l1?^o Burpualxr eq] Jo pua eqt rxo{ tco[od runlo3 xederunllaqel eql ol ecuEtslpJqlJo spltql-o^u 'flpluec 'ueeJ3 '^qseu 'pesl€r pur?pueq eql puo.(eq pe,{orreu'eJque sulSlEul pelleuuuqc lqSII 'epr,{\ 'uoqces 'esBq uur 7 y r'3uo1 tulll 8 l-9 1 Suolqo-]€eullP ol Pe^tro]JPuuJql aql lEJu Jrldllla 'spdes -el€^o snlpa :esnlqo ,{l,^\otrEu 1u:e1e1eq} Sulqcnot1sn[ pue xedt 3ql J"eu p3^lnce] ,([lBnsn 'sel8w 'sul8.lEru 'epIS 'sur8reruelElnuar. ,{l,^dolleqsqll{\ lq8u 1€ pa^JncajJI€q lPlslp arnua r0l{\ eql '1ce::od;pq 'uoqs 'epl^\ uorJ pe,^\er^uerJ,r snoqqtS 1ou purrxord "'ne1cleseq e1e1n3ll 3 01pe,toJJ€u '3uol 'peuellEu glBloecuelew^o (pe^lnJul rurrrZ 8 I tutu g g-g ueq,{ eulllno uI wn aE/7 elnJPqns 'tcoJJod'epl,!\ '8uol 'e]€loecuEt uIuI I ruul t , ,{l,t\o!€u sp1a4 ?uo1urur 9Z-S I secrdl?eleJel 'lelncrJE 'etnlo^ul 'eeq 'epl^\ '3uo1 qtr^ suISr€uJl€lslp ueql;pq purtxotd roJ el€uuor uru 7 1 9 1 'etBloecu€l-Suolqo 'pexegep '{1enb11qo ruu E 9-g g ,{I,{oJ.I"ur/rdas PJaJtl elgln.i.duol elnJEqns 'e^pcuoJ'epl^d '3uo1ruru 'stBloacu€l-els^o rIJuI/ 1-2 1 9 E 9 t ,{l,i!\oJ.rEulDdas l,sroQ|ellsses "{lepl^\ 'ssorrc Suruedo'por Jo u,toJq Flloq,|d,(lerur'ueerB qsl{olle,{ ol ueer8epd,{1lunsn uru g ;'3uo1 'srqceq: ',uee;3 'epl4 '8uol ruru ' uu orL srat$olg eql o1searSep 09 lnoq€ ]E les ,{utqs lfltrr 9 I 9 € 'eplla '8uol 'elB^o 'pro^oqo trr|2,1oet€lnclds urul € l ulru € l , flesJe,\suu4.ryrrlq /otoll sJa^\ol+ '3uo1 'esuep aJuaJS oL{uI 0g J - SJo Suqsrsuoc urc E7-ge>1lds .lEaull ,{lelerepouro1esreds'mc].)uuN 'srseqtu€ 'peJ 'qslllq'{\ lP pa:eqtr,tr,{prud ueuo'JePuelsfle^'lceJe'3uo[ uIJ 9I-9 €ullxel ee4 ,{lell]l 'ssoJJE 'ueal8 '8uol 'ele]el-JP}u\ '"pt,t. 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