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Pollinator Diversity and Reproductive Success of [I]Epipactis Helleborine[I]
Manuscript to be reviewed Pollinator diversity and reproductive success of Epipactis helleborine (L.) Crantz (Orchidaceae) in anthropogenic and natural habitats Agnieszka Rewicz Corresp., 1 , Radomir Jaskuła 2 , Tomasz Rewicz 3 , Grzegorz Tończyk 2 1 Department of Geobotany and Plant Ecology, University of Lodz, Łódź, Poland 2 Department of Invertebrate Zoology and Hydrobiology, University of Lodz, Łódź, Poland 3 Laboratory of Microscopic Imaging and Specialized Biological Techniques, University of Lodz, Łódź, Poland Corresponding Author: Agnieszka Rewicz Email address: [email protected] Background. Epipactis helleborine is an Eurasian orchid species which prefers woodland environments but it may also spontaneously and successfully colonise human-made artificial and disturbed habitats such as roadsides, town parks and gardens. It is suggested that orchids colonising anthropogenic habitats are characterised by a specific set of features (e.g.: large plant size, fast flower production). However, as it is not well known how pollinator diversity and reproductive success of E. helleborine differs in populations in anthropogenic habitats compared to populations from natural habitats, we wanted to compare pollinator diversity and reproductive success of this orchid species between natural and anthropogenic habitat types. Methods. Pollination biology, reproductive success and autogamy in populations of E. helleborine from anthropogenic (roadside) and natural (forest) habitats were compared. Eight populations (four natural and four human-disturbed ones) in two seasons were studied according to height of plants, length of inflorescences, as well as numbers of juvenile shoots, flowering shoots, flowers, and fruits. The number and diversity of insect pollinators were studied in one natural and two human-disturbed populations. Results. -
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
Pdf of JHOS January 2012
JJoouurrnnaall of the HHAARRDDYY OORRCCHHIIDD SSOOCCIIEETTYY Vol. 9 No. 1 (63) January 2012 JOURNAL of the HARDY ORCHID SOCIETY Vol. 9 No. 1 (63) January 2012 The Hardy Orchid Society Our aim is to promote interest in the study of Native European Orchids and those from similar temperate climates throughout the world. We cover such varied aspects as field study, cultivation and propagation, photography, taxonomy and systematics, and practical conservation. We welcome articles relating to any of these subjects, which will be considered for publication by the editorial committee. Please send your submissions to the Editor, and please structure your text according to the “Advice to Authors” (see website www.hardyorchidsociety.org.uk , January 2004 Journal, Members’ Handbook or contact the Editor). Views expressed in journal arti - cles are those of their author(s) and may not reflect those of HOS. The Hardy Orchid Society Committee President: Prof. Richard Bateman, Jodrell Laboratory, Royal Botanic Gardens Kew, Richmond, Surrey, TW9 3DS Chairman: Celia Wright, The Windmill, Vennington, Westbury, Shrewsbury, Shropshire, SY5 9RG [email protected] Vice-Chairman: David Hughes, Linmoor Cottage, Highwood, Ringwood, Hants., BH24 3LE [email protected] Secretary: Alan Leck, 61 Fraser Close, Deeping St. James, Peterborough, PE6 8QL [email protected] Treasurer: John Wallington, 17, Springbank, Eversley Park Road, London, N21 1JH [email protected] Membership Secretary: Moira Tarrant, Bumbys, Fox Road, Mashbury, -
Orchids of Lakeland
a field guide to OrchidsLakeland Provincial Park, Provincial Recreation Area and surrounding region Written by: Patsy Cotterill Illustrated by: John Maywood Pub No: I/681 ISBN: 0-7785-0020-9 Printed 1998 Sepal Sepal Petal Petal Seed Sepal Lip (labellum) Ovary Finding an elusive (capsule) orchid nestled away in Sepal the boreal forest is a Scape thrilling experience for amateur and Petal experienced naturalists alike. This type Petal of recreation has become a passion for Column some and, for many others, a wonder- ful way to explore nature and learn Ovary Sepal (capsule) more about the living world around us. Sepal But, we also need to be careful when Lip (labellum) looking at these delicate plants. Be mindful of their fragile nature... keep Spur only memories and take only photo- graphs so that others, too, can enjoy. Corm OrchidsMany people are surprisedin Alberta to learn that orchids grow naturally in Alberta. Orchids are typically associated with tropical rain forests (where indeed, many do grow) or flower-shop purchases for special occasions, exotic, expensive and highly ornamental. Wild orchids in fact are found world-wide (except in Antarctica). They constitute Roots Tuber the second largest family of flowering plants (the Orchidaceae) after the Aster family, with upwards of 15,000 species. In Alberta, there are 26 native species. Like all Roots orchids from north temperate regions, these are terrestrial. They grow in the ground, in contrast to the majority of rain forest species which are epiphytes, that is, they grow attached to trees for support but do not derive nourishment from them. -
Rare Plant Monitoring 2017
RARE PLANT MONITORING 2017 Ajuga pyramidalis Ophrys insectifera © Zoe Devlin What is it? In 2017, we decided to carry out a small pilot scheme on rare plant monitoring. Where experienced plant recorders had submitted recent casual records of rare plants to the Centre, they were asked if they would be willing to visit their rare plant population once a year during its flowering period and to count the total number of individuals present. The response to the scheme from the small number of recorders contacted has been overwhelming positive and it has resulted in very valuable data being collected in 2017. Data on the rare plant location, the count and additional information about the site is submitted online through a dedicated web portal set up by the Data Centre. The project was discussed and agreed with the NPWS. It is framed around the 2016 Vascular Plant Red List and is mainly focused on monitoring vulnerable, near threatened and rare least concern species. Why is it important? When assessing the national FAST FACTS 2017 conservation status of very rare species according to IUCN Red List methodology, it is recommended that 37 you use annual population count data. That’s the total number of rare plant Given the numbers of rare plant populations that were monitored in the species a country might have, this 2017 pilot information can be difficult to collect in any volume. This citizen science project relies on the generosity of 22 expert volunteers to ‘keep an eye’ on That’s the number of rare plant species rare populations near them and to that were monitored in 2017 submit standardised count data once a year. -
“Dynamic Speciation Processes in the Mediterranean Orchid Genus Ophrys L
“DYNAMIC SPECIATION PROCESSES IN THE MEDITERRANEAN ORCHID GENUS OPHRYS L. (ORCHIDACEAE)” Tesi di Dottorato in Biologia Avanzata, XXIV ciclo (Indirizzo Sistematica Molecolare) Universitá degli Studi di Napoli Federico II Facoltá di Scienze Matematiche, Fisiche e Naturali Dipartimento di Biologia Strutturale e Funzionale 1st supervisor: Prof. Salvatore Cozzolino 2nd supervisor: Prof. Serena Aceto PhD student: Dott. Hendrik Breitkopf 1 Cover picture: Pseudo-copulation of a Colletes cunicularius male on a flower of Ophrys exaltata ssp. archipelagi (Marina di Lesina, Italy. H. Breitkopf, 2011). 2 TABLE OF CONTENTS GENERAL INTRODUCTION CHAPTER 1: MULTI-LOCUS NUCLEAR GENE PHYLOGENY OF THE SEXUALLY DECEPTIVE ORCHID GENUS OPHRYS L. (ORCHIDACEAE) CHAPTER 2: ANALYSIS OF VARIATION AND SPECIATION IN THE OPHRYS SPHEGODES SPECIES COMPLEX CHAPTER 3: FLORAL ISOLATION IS THE MAIN REPRODUCTIVE BARRIER AMONG CLOSELY RELATED SEXUALLY DECEPTIVE ORCHIDS CHAPTER 4: SPECIATION BY DISTURBANCE: A POPULATION STUDY OF CENTRAL ITALIAN OPHRYS SPHEGODES LINEAGES CONTRIBUTION OF CO-AUTHORS ACKNOWLEDGEMENTS 3 GENERAL INTRODUCTION ORCHIDS With more than 22.000 accepted species in 880 genera (Pridgeon et al. 1999), the family of the Orchidaceae is the largest family of angiosperm plants. Recently discovered fossils document their existence for at least 15 Ma. The last common ancestor of all orchids has been estimated to exist about 80 Ma ago (Ramirez et al. 2007, Gustafsson et al. 2010). Orchids are cosmopolitan, distributed on all continents and a great variety of habitats, ranging from deserts and swamps to arctic regions. Two large groups can be distinguished: Epiphytic and epilithic orchids attach themselves with aerial roots to trees or stones, mostly halfway between the ground and the upper canopy where they absorb water through the velamen of their roots. -
Phytogeographical Analysis and Ecological Factors of the Distribution of Orchidaceae Taxa in the Western Carpathians (Local Study)
plants Article Phytogeographical Analysis and Ecological Factors of the Distribution of Orchidaceae Taxa in the Western Carpathians (Local study) Lukáš Wittlinger and Lucia Petrikoviˇcová * Department of Geography and Regional Development, Faculty of Natural Sciences, Constantine the Philosopher University in Nitra, 94974 Nitra, Slovakia; [email protected] * Correspondence: [email protected]; Tel.: +421-907-3441-04 Abstract: In the years 2018–2020, we carried out large-scale mapping in the Western Carpathians with a focus on determining the biodiversity of taxa of the family Orchidaceae using field biogeographical research. We evaluated the research using phytogeographic analysis with an emphasis on selected ecological environmental factors (substrate: ecological land unit value, soil reaction (pH), terrain: slope (◦), flow and hydrogeological productivity (m2.s−1) and average annual amounts of global radiation (kWh.m–2). A total of 19 species were found in the area, of which the majority were Cephalenthera longifolia, Cephalenthera damasonium and Anacamptis morio. Rare findings included Epipactis muelleri, Epipactis leptochila and Limodorum abortivum. We determined the ecological demands of the abiotic environment of individual species by means of a functional analysis of communities. The research confirmed that most of the orchids that were studied occurred in acidified, calcified and basophil locations. From the location of the distribution of individual populations, it is clear that they are generally arranged compactly and occasionally scattered, which results in ecological and environmental diversity. During the research, we identified 129 localities with the occurrence of Citation: Wittlinger, L.; Petrikoviˇcová, L. Phytogeographical Analysis and 19 species and subspecies of orchids. We identify the main factors that threaten them and propose Ecological Factors of the Distribution specific measures to protect vulnerable populations. -
HAUSTORIUM 76 1 HAUSTORIUM Parasitic Plants Newsletter ISSN 1944-6969 Official Organ of the International Parasitic Plant Society (
HAUSTORIUM 76 1 HAUSTORIUM Parasitic Plants Newsletter ISSN 1944-6969 Official Organ of the International Parasitic Plant Society (http://www.parasiticplants.org/) July 2019 Number 76 CONTENTS MESSAGE FROM THE IPPS PRESIDENT (Julie Scholes)………………………………………………..………2 MEETING REPORTS 15th World Congress on Parasitic Plants, 30 June – 5 July 2019, Amsterdam, the Netherlands.………….……..2 MISTLETOE (VISCUM ALBUM) AND ITS HOSTS IN BRITAIN (Brian Spooner)……………………………10 PHELIPANCHE AEGYPTIACA IN WESTERN IRAN (Alireza Taab)……………………………………………12 NEW AND CURRENT PROJECTS Delivering high-yielding, disease-resistant finger millet to farmers…………………………………………….…..13 N2AFRICA – new Striga project – update……………………………………………………………………….…...14 Striga asiatica Madagascar fieldwork summary 2019……………………………………………………………......14 Pea (Pisum sativum) breeding for disease and pest resistance ………………………………………………….......15 REQUEST FOR SEEDS OF OROBANCHE CRENATA (Gianniantonio Domina)…………………………...…..15 PRESS REPORTS Metabolite stimulates a crop while suppressing a weed………………………………………………………….…..16 Dodder plant poses threat to trees and crops (in Kenya)………………………………………………………...….17 PhD OPPORTUNITY AT NRI (Jonne Rodenburg)…………………………………………………………………18 THESIS Sarah Huet. An overview of Phelipanche ramosa seeds: sensitivity to germination stimulants and microbiome profile. …………………………………………………………………………………………………………………..18 BOOK REVIEW Strigolactones – Biology and Applications. Ed. by Hinanit Koltai and Cristina Prandi. (Koichi Yoneyama) …………………………………………………………………………………………………………………………....19 -
Cypripedium Parviflorum Salisb
Cypripedium parviflorum Salisb. synonym: Cypripedium calceolus L. var. parviflorum (Salisb.) Fernald yellow lady's-slipper Orchidaceae - orchid family status: State Threatened, USFS sensitive rank: G5 / S2 General Description: Perennial with showy flowers; stems 7-70 cm tall, sparsely pubescent, somewhat glandular. Leaves several, alternate, bases slightly sheathing the stem, broadly elliptic to elliptic-lanceolate, 6-17 x up to 7 cm, lightly pubescent, usually glandular. Floral Characteristics: Flower 1 (rarely 2), terminal, subtended by an erect, leaflike bract. Sepals and petals greenish or yellowish, often marked with dark reddish brown or purplish spots, blotches, or streaks. Upper sepal broadest, 19-80 x 7-40 mm; the lateral pair of sepals completely fused or with only a notch at their tip. Petals somewhat narrower and longer than the sepals, 24-97 x 3-12 mm, often wavy-margined and spirally twisted. Lip strongly pouched, 15-54 mm long, pale to deep yellow (rarely white), sometimes with reddish or Illustration by Jeanne R. Janish, purplish spots around the orifice. Flowers May to June. ©1969 University of Washington Press Fruits: Ellipsoid to oblong-ellipsoid capsules. Identif ication Tips: Cypripedium montanum has a white lip, rarely suffused with magenta. It may hybridize with C. parviflorum, resulting in individuals with very pale yellow lips. The habitat of C. montanum is typically well-drained upland, while that of C. parviflorum is wetland/riparian or the ecotone between wetland and upland. Two varieties, var. makas in and var. pubes cens , are both reported from WA ; their relative abundance and distribution is under review. Range: East of the C ascade crest in B.C ., WA , and O R, to the eastern U.S. -
The Orchids of the Isle of Formentera
Jour. Eur. Orch. 37 (4): 501 – 528. 2005. Barbara Klahr Translated by Dr.Eckhard Kiehl The Orchids of the Isle of Formentera Keywords Orchidaceae; Aceras anthropophorum, Barlia robertiana, Gennaria diphylla, Neotinea maculata, Ophrys bombyliflora, O. dyris, O. cf. fabrella, O. fusca, O. speculum, O. tenthredinifera, Orchis collina, O. fragrans, Serapias parviflora; Formentera, Pityuses, Balearic Islands; distribution; ecology; endangerment; protection. Summary Klahr, B. (2005): The orchids of Formentera.- Jour. Eur. Orch. 37 (4): 501 XXX. A short review of history and geography of Formentera is given; aspects of flora and fauna of a south mediterranean Island, survival of plants in a half-arid zone without any wells or fresh ground water. As the most southern Island of the Pityuses, Formentera belongs climatically more to Africa than to Europe. It is a very rocky eroded Island after having had a long period without agriculture. Endemic species in flora and fauna are mentioned. Wild Orchids are growing on this small Island mainly in the higher regions of the plateau La Mola and Cap de Barbaria. There is not as much pastureland as in other mediterranean countries. Ophrys and Orchis predominate.. Most areas of Formentera were explored during the whole year, also difficult to reach regions like “Torrentes” (Canyons). The period of my observation is 2002 - 2005. The Orchids are endangered as a result of the reinforced agriculture and chemical fertilizing, subsidized by the EU, and by increasing building activities. Centuries fallow agricultural land that makes very interesting biotopes will be destroyed. The orchids of Formentera are bibliographically quoted; the results are to be supplemented with four new found species and new sites. -
Appendix LEA10 Local Wildlife Site Maps and Citations KENT LOCAL WILDLIFE SITE Site Ref: AS01 Site: Hemsted Forest Grid Ref: TQ819360
Appendix LEA10 Local Wildlife Site maps and citations KENT LOCAL WILDLIFE SITE Site Ref: AS01 Site: Hemsted Forest Grid Ref: TQ819360 580000 580500 581000 581500 582000 582500 583000 583500 138500 138500 138000 138000 137500 137500 137000 137000 136500 136500 136000 136000 135500 135500 135000 135000 134500 134500 134000 134000 580000 580500 581000 581500 582000 582500 583000 583500 0 250 500 1,000 Kent Wildlife Trust 2017 Meters This map is based upon Ordnance Survey material with the permission of Ordnance Survey on behalf of the Controller of Her Majesty's Stationery Office ± © Crown Copyright. Unauthorised reproduction infringes Crown Copyright and may lead to prosecution or civil proceedings. 100019238. 2017. Approved: 2017 Jun AS01 Hemsted Forest Page 1 of 3 KENT LOCAL WILDLIFE SITE KWT File Ref: 820362 Site Name: Hemsted Forest Site Ref. No: AS01 LPA: Ashford / Tunbridge Wells Central Grid Ref: TQ 819360 Parish: Cranbrook / Benenden / Category: Woodland, grassland Biddenden Owner: Forestry Commission Natural Area: High Weald Area: 509.25 ha AONB: No Date first notified: 1986 TPO: Yes Dates revised: 1993, Feb 2000, June 2016 Date last approved: June 2017 REASON FOR DESIGNATION Much of the woodland within this site is included on Natural England’s Provisional Ancient Woodland Inventory and is known to support at least 42 ancient woodland vascular plant indicator species. The site also includes areas of secondary woodland which are contiguous with the ancient woodland and which are considered to have the potential for colonisation by species associated with the ancient woodland. RATIONALE FOR SITE BOUNDARY The site boundary stretches from Tottenden Wood in the far south to Hammer Wood at the north. -
Number English Name Welsh Name Latin Name Availability Llysiau'r Dryw Agrimonia Eupatoria 32 Alder Gwernen Alnus Glutinosa 409 A
Number English name Welsh name Latin name Availability Sponsor 9 Agrimony Llysiau'r Dryw Agrimonia eupatoria 32 Alder Gwernen Alnus glutinosa 409 Alder Buckthorn Breuwydd Frangula alnus 967 Alexanders Dulys Smyrnium olusatrum Kindly sponsored by Alexandra Rees 808 Allseed Gorhilig Radiola linoides 898 Almond Willow Helygen Drigwryw Salix triandra 718 Alpine Bistort Persicaria vivipara 782 Alpine Cinquefoil Potentilla crantzii 248 Alpine Enchanter's-nightshade Llysiau-Steffan y Mynydd Circaea alpina 742 Alpine Meadow-grass Poa alpina 1032 Alpine Meadow-rue Thalictrum alpinum 217 Alpine Mouse-ear Clust-y-llygoden Alpaidd Cerastium alpinum 1037 Alpine Penny-cress Codywasg y Mwynfeydd Thlaspi caerulescens 911 Alpine Saw-wort Saussurea alpina Not Yet Available 915 Alpine Saxifrage Saxifraga nivalis 660 Alternate Water-milfoil Myrdd-ddail Cylchynol Myriophyllum alterniflorum 243 Alternate-leaved Golden-saxifrageEglyn Cylchddail Chrysosplenium alternifolium 711 Amphibious Bistort Canwraidd y Dŵr Persicaria amphibia 755 Angular Solomon's-seal Polygonatum odoratum 928 Annual Knawel Dinodd Flynyddol Scleranthus annuus 744 Annual Meadow-grass Gweunwellt Unflwydd Poa annua 635 Annual Mercury Bresychen-y-cŵn Flynyddol Mercurialis annua 877 Annual Pearlwort Cornwlyddyn Anaf-flodeuog Sagina apetala 1018 Annual Sea-blite Helys Unflwydd Suaeda maritima 379 Arctic Eyebright Effros yr Arctig Euphrasia arctica 218 Arctic Mouse-ear Cerastium arcticum 882 Arrowhead Saethlys Sagittaria sagittifolia 411 Ash Onnen Fraxinus excelsior 761 Aspen Aethnen Populus tremula