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Home , Archaea, Cyanobacteria

DIVERSITY OF Current are classified into 3 Domains and/or 5-6 Kingdoms

Kingdoms Domains

1. 1a. Archaebacteria 1b. Eubacteria 2. Eukarya 3. 4. Fungi 5. Major groups are related

All alive today

All equally “evolved”

But also represent different depths of branching, since shared common ancestor Diversification also involved a series of innovations that added new forms of life

Older modes persisted, possibly largely unchanged

“advanced”vs. “primitive” Diverse species appeared then went extinct

Current diversity is a slice of a continuing process

3-1 1.Prokaryotes (bacteria) single celled DNA, lack nucleus no motility, incl. flagella closest to early life? Prokaryotes

No sexual () per se, but various modes of genetic exchange

The real “tree”of life? Prokaryotes

Much of evolved in this group

Both respiration and started here

______(“self feeding”) –use CO2 as source chemoautotroph – from chemicals (e.g. H2S) photoautotroph –energy from sun -photosynthesis

______(“other feeding”)–use organic molecules for carbon and energy

anaerobic –respiration without O2 aerobic –respiration with O2 Prokaryotes

Cyanobacteria (“blue-green ”) are free living and photosynthetic contain a, ancestor of ______Prokaryotes Some bacteria and cyanobacteriacan also fix ______

In aquatic and terrestrial systems

Even in some roots

Also involved in other transformations (discussed later) Important as ______

and

Often closely related species Prokaryotes

Archaea–extreme environments –salty, high temperature Prokaryotes “Black smoker”()

Archaea–“ancient”, http://www.ocean.udel.edu/kiosk/bsmoker.html origin of life? –Protists, Plants, Fungi, Animals

______and Endomembrane

Organelles –mitochondria and

Sexual reproduction (meiosis)

Multicellularity(tissues, organs)

Sociality 2. Protists –mostly small, single celled & multicellular -Eukaryotes –true nucleus (containing DNA) and endomembrane -Many are , aquatic –highly diverse -Various parasites (e.g., , malaria, helminths) Endosymbiotic origin of organelles

Mitochondria -______Chloroplast -______Key biochemistry done by bacteria or their descendants Protists

Includes true ALGAE –single and multicellularforms that are photosynthetic, and mainly ______(main “producers” in ______systems)

Unicellular algae -contain chloroplasts -, b, & c Multicellularalgae –“______” -various differentiated structures, conducting systems -not rooted -holdfast (nutrients from water) Complex life cycles &

Both diploid and haploid phases

Sexual(meiosis) and asexual () components

Brown Alga (Laminaria) Green Alga (Ulva) –haploid & diploid equivalent What is Diploid? Haploid?

Life cycles often switch between these two states.

______are haploid and fuse to form diploid zygotes.

The zygote is “diploid”because it contains two complete sets of , one “haploid”set from each parent. This means two copies of genes of each type (“locus”). The two copies need not be identical (alternate “alleles” are common at each locus) What is Meiosis? Mitosis?

When a phase of the life cycle duplicates itself asexually, it is a result of mitosis –the exact copying of the genes. Meiosis

Sexual reproduction results from a much stranger process, where the diploid produces haploid cells by meiosis. The genes replicate once, then divide twice, creating (four) cells with only one set of genes each (haploid). Both haploids and diploids can replicate asexually (mitotically) too

“Water Mold”(Oomycota) –mostly diploid Malaria (Plasmodium) mostly haploid Recombination –the real point?

At least as important as the production of haploid cells, is the recombination of alleles in meiosis. Before making haploid cells, the dividing cells undergoing meiosis essentially “shuffle”the two parental , creating new genotypes that are mixtures of the parents. Often, the resulting recombinant haploid gametes are fused with gametes from other individuals (______) to make the zygotes. This creates new genetic variation. The offspring are different from each other and from either parent. Independent assortment Shuffling whole Crossing over (chromosomes = blocks of genes) Shuffling genes within chromosomes

The paradox of sex -why meiosis? We have emphasized that favors (autocatalysts) that are good at transforming resources into more self –but here is a widespread mode of reproduction that seems designed to create offspring that are different from the parents, that are deliberately NOT self. Why? It is assumed there are good reasons, we just don’t know what they are for sure. But of course there have been many proposed explanations (hypotheses). This is another good example of the difficulty of verifying functional explanations. The Main Contenders 1. a.

b.

2. a.

b.

3.

4.

5. 3. Plants -primarily terrestrial, multicellular -evolved from ______-photosynthetic (chloroplasts), rooted in (nutrients) -extensive ______tissue -widespread, diverse and related species Ferns and others Includes tree ferns, once dominant Petro future here

Coal Age –Carboniferous 290-360 MYBP How long will current oil supplies last?

= ______years

http://www.eia.doe.gov/oiaf/ieo/oil.html Gymnosperms (conifers and others)

Ginko

Cycad Welwitschia Read Sacks Angiosperms –flowering plants Include most plants that dominate landscapes today 4. Fungi -eukaryotic, diverse ______-unicellular () and multicelled(mushrooms) -with bacteria, the most important ______(nutrient releasers) in aquatic and terrestrial systems

Read Sacks 4. Fungi

-important ______, plant root fungal symbiontsthat enhance nutrient and water uptake

-many fungi are plant & pathogens (mostly plants) 4. Fungi-include photosynthetic ______–a symbiotic association of a and a . Fungus gets for photosynthesis of algae. Live on bare substrate (rock, bark), tolerate drying. Largest ?

Blue Whale

Giant sequoia

Aspen clone (Populus tremuloides) –one

Honey mushroom (Armillariaostoyae) –one genotype 5. Animals -evolved from ______-multicellular, heterotrophic eukaryotes -35 “body plans”, most invertebrates, many marine -highly elaborated and organ systems -, , parasites Jelly

Anemone

Chiton(Mollusk) Scallop (Bivalve)

Sea Slug (Gastropod)

Sea Star

Cephalopods: Nautilus & Octopus Leech () -______-have a symbiotic association with “zooxanthellae”which are themselves a of a () and photosynthetic ().

Trilobite (extinct) Horseshoe Crab Barnacle Lobster

Shrimp Trilobite Optics Trilobites developed one of the first advanced visual systems in the animal . This may have promoted the spectacular diversification of animals.

The majority of trilobites bore a pair of compound eyes (made up of many lensedunits) Compound eyes in living arthropods such as insects are very sensitive to motion, and it is likely that they were similarly important in predator detection in trilobites. It has also been suggested that stereoscopic vision was provided by closely spaced, but separate eyes

http://www.trilobites.info/eyes.htm http://ebiomedia.com/gall/eyes/octopus-insect.html Some Trilobites “lost”vision, as have many existing species

crayfish

Astaynax

Yamamoto Y, Stock DW, Jeffery WR (2004) Hedgehog signalling controls eye degeneration in blind cavefish. 431:844-847. The eyes of the Octopus and Squid (Cephalopods) are very similar to ours in structure and function, with lens and image forming retina. Read Williams

Eyes of the vertebrates and cephalopod molluscs(squid and octopus) are remarkably similar, an example of ______evolution. One interesting difference –vertebrates have an inverted retina, the sensory cells lying beneath the nerve fibers. This results in the sensory cells being absentwhere the optic nerve is attached to the eye, thus creating a blind spot. The octopus eye has a non-inverted retina in which the sensory cells lie above the nerve fibres. There is therefore no blind spot in this kind of eye. http://en.wikipedia.org/wiki/Evidence_of_evolution The Blind Spot The blind spot is the area on the retina without receptors that respond to light. Therefore an image that falls on this region will NOT be seen. It is in this region that the optic nerve exits the eye on its way to the brain.

In the next two images, close your right eye. With your left eye, look at the numbers on the right side, starting with the number "1." You should be able to see the "sad face" (top image) or the gap in the blue line (bottom image) in your peripheral vision. Keep your head still, and withyour left eye, look at the other numbers. The sad face should disappear when you get to "4" and reappear at about "7." Similarly the blue line will appear complete between "4" and "7.“This is because your brain is "filling in" the missing information.

Here is another image to show your blind spot. Close your right eye. With your left eye, look at the +. You should see the red dot in your peripheral vision. Keep looking at the +with your left eye. The red dot will move from the left to the right and disappear and reappear as the dot moves into and out of your blind spot.

http://staff.washington.edu/chudler/chvision.html Arthropods –Spiders and Insects Vertebrates () Animals -Sociality End Part 3

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